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<document id="903313A148CF30BE3FE59395CD39CF81" ID-CLB-Dataset="22518" ID-DOI="10.11646/zootaxa.4683.2.8" ID-GBIF-Dataset="ee200a72-86bd-47cf-ac62-a234d97c394a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3772813" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588058770637" checkinUser="plazi" docAuthor="Polotow, Daniele &amp; Brescovit, Antonio D." docDate="2019" docId="03E087ED3C392864FF091AB3FD9665E0" docLanguage="en" docName="zootaxa.4683.2.8.pdf" docOrigin="Zootaxa 4683 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Isoctenus guadalupei Polotow &amp; Brescovit 2019, comb. nov." docType="treatment" docVersion="7" lastPageNumber="293" masterDocId="FFD9FF953C382867FF9E1A51FF876535" masterDocTitle="Almost eight decades later: description of the male of Isoctenus guadalupei (Mello- Leitão, 1941) comb. nov. and redescription of the female" masterLastPageNumber="294" masterPageNumber="291" pageNumber="292" updateTime="1698837755131" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="44C903DBC3BD57DAC30CA828D7D76579" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E41163C2BD74D3B9941136E0BFC3ED15">
<mods:title id="7DDB839512F73BEB9D30B7E8744DEC56">Almost eight decades later: description of the male of Isoctenus guadalupei (Mello- Leitão, 1941) comb. nov. and redescription of the female</mods:title>
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<mods:name id="06B0EB15A7631B6B565AED2005E20947" type="personal">
<mods:role id="FCE38AD9494154141DAE7115160AF7CF">
<mods:roleTerm id="2C6040736F023C5A1EEB641DDCD35681">Author</mods:roleTerm>
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<mods:namePart id="1108F924BF00C46F2D652E9054BB37C3">Polotow, Daniele</mods:namePart>
<mods:affiliation id="B8C5A741BAB6B22B29217B4D754EBACB">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), campus São Carlos, Rodovia Washington Luís, Km 235, CEP 13565 - 905, São Carlos, SP, Brazil.</mods:affiliation>
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<mods:role id="B14AE455C74AA8A38A3D74B42AE8ABF4">
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</mods:role>
<mods:namePart id="ED675E26E447EB8FAF236149AABB9CD4">Brescovit, Antonio D.</mods:namePart>
<mods:affiliation id="6E7857001EA1B9E65938ABD35CB061FE">Laboratório Especial de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil 1500, CEP 05503 - 900, São Paulo, SP, Brazil. E-mail: danielepolotow @ gmail. com.</mods:affiliation>
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<mods:typeOfResource id="21B6E13CF940DE910E31B1E03E348CBD">text</mods:typeOfResource>
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<mods:title id="B49DE61D6E2318D6F9F6243BF91140AE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FD7054206832A87AC32D0ECB79BD38C4">
<mods:date id="74EAF61CD58AD3BEAB77C82DAA80C3F2">2019</mods:date>
<mods:detail id="BF3F72FC5A9D366092D4E7864283AEA1" type="pubDate">
<mods:number id="4825E29EF72E1C95E921871A364A619A">2019-10-08</mods:number>
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<mods:detail id="10060FF38F33992CB04CEE81C36634D4" type="volume">
<mods:number id="FF63B8DA7FE3E73B3292830D4D2CD4B6">4683</mods:number>
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<mods:detail id="F587C3C20CA48AB29998259EA6E2C147" type="issue">
<mods:number id="90E293707AA8ED7CB04E1959E6CB9D15">2</mods:number>
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<mods:classification id="03E74C5067FD85C02C284C9331399C71">journal article</mods:classification>
<mods:identifier id="16DC73E440004C4832CC3CA7FA23E117" type="CLB-Dataset">22518</mods:identifier>
<mods:identifier id="B9948CA569144207BBAE846BBF8C5959" type="DOI">10.11646/zootaxa.4683.2.8</mods:identifier>
<mods:identifier id="D07D71248BA6968413386A4F0D09D2A4" type="GBIF-Dataset">ee200a72-86bd-47cf-ac62-a234d97c394a</mods:identifier>
<mods:identifier id="C4292B59C04A26535CA483C93C367AF9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D4A33F182F8421DBC83BE424CBA1B788" type="Zenodo-Dep">3772813</mods:identifier>
<mods:identifier id="B29CE391902E6CB680EB405608BC3450" type="ZooBank">5C967EE8-61AB-4ED1-BD56-B0B60074CBA7</mods:identifier>
</mods:mods>
<treatment id="03E087ED3C392864FF091AB3FD9665E0" ID-DOI="http://doi.org/10.5281/zenodo.3797573" ID-GBIF-Taxon="163830165" ID-Zenodo-Dep="3797573" LSID="urn:lsid:plazi:treatment:03E087ED3C392864FF091AB3FD9665E0" httpUri="http://treatment.plazi.org/id/03E087ED3C392864FF091AB3FD9665E0" lastPageId="3" lastPageNumber="293" pageId="1" pageNumber="292">
<subSubSection id="C35365703C392866FF091AB3FCAA65C9" box="[151,813,225,252]" pageId="1" pageNumber="292" type="nomenclature">
<paragraph id="8BF636FB3C392866FF091AB3FCAA65C9" blockId="1.[151,813,225,287]" box="[151,813,225,252]" pageId="1" pageNumber="292">
<heading id="D0BE81973C392866FF091AB3FCAA65C9" bold="true" box="[151,813,225,252]" fontSize="11" level="1" pageId="1" pageNumber="292" reason="1">
<emphasis id="B93DEAE93C392866FF091AB3FCAA65C9" bold="true" box="[151,813,225,252]" pageId="1" pageNumber="292">
<taxonomicName id="4C494D783C392866FF091AB3FD1865CE" ID-CoL="98ZQM" authority="(Mello-Leitao, 1941)" authorityName="Polotow &amp; Brescovit" authorityYear="2019" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1941" box="[151,671,225,252]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="guadalupei" status="comb. nov.">
<emphasis id="B93DEAE93C392866FF091AB3FE1E65C9" bold="true" box="[151,409,225,252]" italics="true" pageId="1" pageNumber="292">Isoctenus guadalupei</emphasis>
(
<bibRefCitation id="EFD84B0A3C392866FE371AB0FD1065CE" author="Mello-Leitao, C. F. de" box="[425,663,225,251]" pageId="1" pageNumber="292" pagination="199 - 225" refId="ref2105" refString="Mello-Leitao, C. F. de (1941) Las aranas de la provincia de Santa Fe colectadas por el Profesor Biraben. Revista del Museo de La Plata, New Series (Zoologia), 2, 199 - 225." type="journal article" year="1941">Mello-Leitão, 1941</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20E57923C392866FD381AB3FCAA65C9" box="[678,813,226,252]" pageId="1" pageNumber="292" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35365703C392866FF091B56FE9F642A" box="[151,280,263,287]" pageId="1" pageNumber="292" type="description">
<paragraph id="8BF636FB3C392866FF091B56FE9F642A" blockId="1.[151,813,225,287]" box="[151,280,263,287]" pageId="1" pageNumber="292">
<figureCitation id="13722A7E3C392866FF091B56FE9F642A" box="[151,280,263,287]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1300,1325]" captionStartId-1="2.[151,250,1637,1662]" captionStartId-2="3.[151,250,860,885]" captionTargetBox-0="[151,1436,308,1273]" captionTargetBox-1="[161,1422,393,1596]" captionTargetBox-2="[289,1289,234,833]" captionTargetId-0="figure@1.[151,1436,308,1273]" captionTargetId-1="figure@2.[151,1436,379,1611]" captionTargetId-2="figure@3.[289,1299,234,834]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov., male:A, dorsal habitus; B, ventral habitus; C, lateral view of venter, with arrows pointing to the projection on coxae IV; D, male palp, ventral view." captionText-1="FIGURE 2. Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov.: A, male palp, ventral view; B, epigynum, ventral view; C, female internal genitalia, dorsal view." captionText-2="FIGURE 3. Collection localities of Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov. and I. ordinario Polotow &amp; Brescovit, 2009: A, map representing South America (partial), with Chacoan and Parana dominions colored; B, detail of Chacoan and Parana dominions with distributional records of Isoctenus guadalupei and I. ordinario." figureDoi-0="http://doi.org/10.5281/zenodo.3772815" figureDoi-1="http://doi.org/10.5281/zenodo.3772817" figureDoi-2="http://doi.org/10.5281/zenodo.3772819" httpUri-0="https://zenodo.org/record/3772815/files/figure.png" httpUri-1="https://zenodo.org/record/3772817/files/figure.png" httpUri-2="https://zenodo.org/record/3772819/files/figure.png" pageId="1" pageNumber="292">Figures 1−3</figureCitation>
</paragraph>
</subSubSection>
<caption id="DF3666733C392866FF091F45FB9A6064" ID-DOI="http://doi.org/10.5281/zenodo.3772815" ID-Zenodo-Dep="3772815" httpUri="https://zenodo.org/record/3772815/files/figure.png" pageId="1" pageNumber="292" startId="1.[151,250,1300,1325]" targetBox="[151,1436,308,1273]" targetPageId="1">
<paragraph id="8BF636FB3C392866FF091F45FB9A6064" blockId="1.[151,1436,1300,1361]" pageId="1" pageNumber="292">
<emphasis id="B93DEAE93C392866FF091F45FE946018" bold="true" box="[151,275,1300,1325]" pageId="1" pageNumber="292">FIGURE 1.</emphasis>
<taxonomicName id="4C494D783C392866FE871F44FD4B6019" authority="(Mello-Leitao, 1941)" authorityName="Polotow &amp; Brescovit" authorityYear="2019" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1941" box="[281,716,1300,1325]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="guadalupei" status="comb. nov.">
<emphasis id="B93DEAE93C392866FE871F44FE766019" box="[281,497,1300,1324]" italics="true" pageId="1" pageNumber="292">Isoctenus guadalupei</emphasis>
(
<bibRefCitation id="EFD84B0A3C392866FE631F45FD426019" author="Mello-Leitao, C. F. de" box="[509,709,1300,1325]" pageId="1" pageNumber="292" pagination="199 - 225" refId="ref2105" refString="Mello-Leitao, C. F. de (1941) Las aranas de la provincia de Santa Fe colectadas por el Profesor Biraben. Revista del Museo de La Plata, New Series (Zoologia), 2, 199 - 225." type="journal article" year="1941">Mello-Leitão, 1941</bibRefCitation>
)
</taxonomicName>
<emphasis id="B93DEAE93C392866FD4C1F44FCC16018" bold="true" box="[722,838,1301,1325]" pageId="1" pageNumber="292">
<taxonomicNameLabel id="A20E57923C392866FD4C1F44FCC16018" box="[722,838,1301,1325]" pageId="1" pageNumber="292" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, male: A, dorsal habitus; B, ventral habitus; C, lateral view of venter, with arrows pointing to the projection on coxae IV; D, male palp, ventral view.
</paragraph>
</caption>
<subSubSection id="C35365703C392866FF091FD0FB1E6331" pageId="1" pageNumber="292" type="reference_group">
<paragraph id="8BF636FB3C392866FF091FD0FAF96088" blockId="1.[151,1437,1408,2045]" pageId="1" pageNumber="292">
<treatmentCitationGroup id="AB5911D53C392866FF091FD0FAF96088" pageId="1" pageNumber="292">
<treatmentCitation id="0AE810EA3C392866FF091FD0FDF460AD" author="Mello-Leitao, C. F. de" box="[151,627,1408,1433]" page="215" pageId="1" pageNumber="292" year="1941">
<taxonomicName id="4C494D783C392866FF091FD0FDF460AD" ID-CoL="327TV" authority="Mello-Leitao, 1941: 215" authorityName="Mello-Leitao" authorityPageNumber="215" authorityYear="1941" box="[151,627,1408,1433]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="guadalupei">
<emphasis id="B93DEAE93C392866FF091FD0FEE660AC" box="[151,353,1409,1433]" italics="true" pageId="1" pageNumber="292">Ctenus guadalupei</emphasis>
<bibRefCitation id="EFD84B0A3C392866FEF71FD1FDF460AD" author="Mello-Leitao, C. F. de" box="[361,627,1408,1433]" pageId="1" pageNumber="292" pagination="199 - 225" refId="ref2105" refString="Mello-Leitao, C. F. de (1941) Las aranas de la provincia de Santa Fe colectadas por el Profesor Biraben. Revista del Museo de La Plata, New Series (Zoologia), 2, 199 - 225." type="journal article" year="1941">Mello-Leitão, 1941: 215</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 23–24 (female
<typeStatus id="54F288593C392866FCCA1FD1FC3660AD" box="[852,945,1408,1432]" pageId="1" pageNumber="292" type="holotype">holotype</typeStatus>
from
<collectingCountry id="F35E766B3C392866FC681FD1FBED60AD" box="[1014,1130,1408,1432]" name="Guadeloupe" pageId="1" pageNumber="292">Guadalupe</collectingCountry>
Norte [
<geoCoordinate id="EE7D503C3C392866FB221FD0FAA660AC" box="[1212,1313,1409,1433]" degrees="28.9408" direction="south" orientation="latitude" pageId="1" pageNumber="292" precision="5" value="-28.9408">-28.9408</geoCoordinate>
,
<geoCoordinate id="EE7D503C3C392866FAB31FD1FA1560AD" box="[1325,1426,1408,1432]" degrees="59.5624" direction="west" orientation="longitude" pageId="1" pageNumber="292" precision="5" value="-59.5624">-59.5624</geoCoordinate>
],
<collectingRegion id="498DF8193C392866FF091FF5FE046088" box="[151,387,1444,1469]" country="Argentina" name="Santa Fe" pageId="1" pageNumber="292">Província de Santa Fé</collectingRegion>
,
<collectingCountry id="F35E766B3C392866FE121FF5FE7E6089" box="[396,505,1444,1468]" name="Argentina" pageId="1" pageNumber="292">Argentina</collectingCountry>
, M. Birabén coll., deposited in MLP 15180, examined);
<bibRefCitation id="EFD84B0A3C392866FBCD1FF5FAFD6088" author="World Spider Catalog" box="[1107,1402,1444,1469]" pageId="1" pageNumber="292" refId="ref2304" refString="World Spider Catalog (2019) World Spider Catalog. Version 19.5. Natural History Museum Bern, Bern. Available from: http: // wsc. nmbe. ch (accessed 1 January 2019)" type="url" year="2019">World Spider Catalog 2019</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF636FB3C392866FF591F98FB1E6331" blockId="1.[151,1437,1408,2045]" pageId="1" pageNumber="292">
<emphasis id="B93DEAE93C392866FF591F98FE8460D4" bold="true" box="[199,259,1481,1505]" pageId="1" pageNumber="292">Note.</emphasis>
<typeStatus id="54F288593C392866FE971F98FEB960D4" box="[265,318,1481,1505]" pageId="1" pageNumber="292">Type</typeStatus>
specimen with faded coloration and missing leg IV segments, except the femur and coxae. The male was matched by similarities of size and coloration, and by the close proximity from the
<typeStatus id="54F288593C392866FB911FBCFBBA6330" box="[1039,1085,1517,1541]" pageId="1" pageNumber="292">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C35365703C392866FF591C41FE236345" pageId="1" pageNumber="292" type="materials_examined">
<paragraph id="8BF636FB3C392866FF591C41FE236345" blockId="1.[151,1437,1408,2045]" pageId="1" pageNumber="292">
<emphasis id="B93DEAE93C392866FF591C41FDA6631D" bold="true" box="[199,545,1552,1576]" pageId="1" pageNumber="292">Additional material examined.</emphasis>
<materialsCitation id="3B213CA63C392866FDBB1C40FE276345" ID-GBIF-Occurrence="2609012201" collectingDate="2011-05-27" collectingDateMax="2011-05-30" collectingDateMin="2011-05-27" collectorName="G. Rubio &amp; M. Izquierdo &amp; L. Piacentinni" country="Argentina" elevation="85" latitude="-28.0049" location="Parque Nacional Mburucuya" longLatPrecision="6" longitude="-58.0365" municipality="Mburucuya" pageId="1" pageNumber="292" specimenCount="86" specimenCount-male="1" stateProvince="Corrientes">
<collectingCountry id="F35E766B3C392866FDBB1C40FD43631C" box="[549,708,1553,1577]" name="Argentina" pageId="1" pageNumber="292">ARGENTINA</collectingCountry>
.
<emphasis id="B93DEAE93C392866FD531C40FCB8631D" box="[717,831,1553,1576]" italics="true" pageId="1" pageNumber="292">
<collectingRegion id="498DF8193C392866FD531C40FCB8631D" box="[717,831,1553,1576]" country="Argentina" name="Corrientes" pageId="1" pageNumber="292">Corrientes</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6B92AC813C392866FCD31C41FC4C631D" box="[845,971,1552,1576]" pageId="1" pageNumber="292">Mburucuyá</collectingMunicipality>
,
<location id="8E9660203C392866FC4A1C40FA82631D" LSID="urn:lsid:plazi:treatment:03E087ED3C392864FF091AB3FD9665E0:8E9660203C392866FC4A1C40FA82631D" box="[980,1285,1552,1576]" country="Argentina" latitude="-28.0049" longLatPrecision="6" longitude="-58.0365" municipality="Mburucuya" name="Parque Nacional Mburucuya" pageId="1" pageNumber="292" stateProvince="Corrientes">Parque Nacional Mburucuyá</location>
(Sendero Che Roga,
<geoCoordinate id="EE7D503C3C392866FF421C64FEB96378" box="[220,318,1589,1613]" degrees="28.0049" direction="south" orientation="latitude" pageId="1" pageNumber="292" precision="5" value="-28.0049">-28.0049</geoCoordinate>
,
<geoCoordinate id="EE7D503C3C392866FED91C65FE2E6379" box="[327,425,1588,1612]" degrees="58.0365" direction="west" orientation="longitude" pageId="1" pageNumber="292" precision="5" value="-58.0365">-58.0365</geoCoordinate>
,
<specimenCount id="9D4FFD723C392866FE2D1C65FE746379" box="[435,499,1588,1612]" pageId="1" pageNumber="292" type="generic">
<quantity id="4CB19B1E3C392866FE2D1C65FE686379" box="[435,495,1588,1612]" metricMagnitude="1" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="292" unit="m" value="85.0">
<elevation id="0064D1C83C392866FE2D1C65FE686379" box="[435,495,1588,1612]" metricMagnitude="1" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="292" unit="m" value="85.0">85 m</elevation>
</quantity>
.
</specimenCount>
a.s.l., manual collecting),
<specimenCount id="9D4FFD723C392866FD601C64FCC06379" box="[766,839,1588,1613]" pageId="1" pageNumber="292" type="male">1 male</specimenCount>
,
<date id="FFF7103B3C392866FCCF1C64FC6F6378" box="[849,1000,1589,1613]" pageId="1" pageNumber="292" value="2011-05-27" valueMax="2011-05-30" valueMin="2011-05-27">
<collectingDate id="EFB3E9D33C392866FCCF1C64FC6F6378" box="[849,1000,1589,1613]" pageId="1" pageNumber="292" value="2011-05-27" valueMax="2011-05-30" valueMin="2011-05-27">27–30.V.2011</collectingDate>
</date>
,
<collectorName id="26BC532D3C392866FC6F1C64FBD26379" box="[1009,1109,1588,1613]" pageId="1" pageNumber="292">G. Rubio</collectorName>
,
<collectorName id="26BC532D3C392866FBC01C64FB6F6379" box="[1118,1256,1588,1612]" pageId="1" pageNumber="292">M. Izquierdo</collectorName>
&amp;
<collectorName id="26BC532D3C392866FA961C64FA1B6379" box="[1288,1436,1588,1612]" pageId="1" pageNumber="292">L. Piacentinni</collectorName>
coll. (MACN-Ar 30496)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35365703C392866FF591C2DFBFE6258" pageId="1" pageNumber="292" type="diagnosis">
<paragraph id="8BF636FB3C392866FF591C2DFBFE6258" blockId="1.[151,1437,1408,2045]" pageId="1" pageNumber="292">
<emphasis id="B93DEAE93C392866FF591C2DFEBB63A1" bold="true" box="[199,316,1660,1684]" pageId="1" pageNumber="292">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C494D783C392866FE3D1C2CFD0263A0" authorityName="Polotow &amp; Brescovit" authorityYear="2019" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1941" box="[419,645,1661,1685]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="guadalupei" status="comb. nov.">
<emphasis id="B93DEAE93C392866FE3D1C2CFD0263A0" box="[419,645,1661,1685]" italics="true" pageId="1" pageNumber="292">Isoctenus guadalupei</emphasis>
</taxonomicName>
<emphasis id="B93DEAE93C392866FD141C2CFC8363A0" bold="true" box="[650,772,1661,1685]" pageId="1" pageNumber="292">
<taxonomicNameLabel id="A20E57923C392866FD141C2CFC8363A0" box="[650,772,1661,1685]" pageId="1" pageNumber="292" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13722A7E3C392866FC8F1C2DFCE463A0" box="[785,867,1660,1685]" captionStart="FIGURE 2" captionStartId="2.[151,250,1637,1662]" captionTargetBox="[161,1422,393,1596]" captionTargetId="figure@2.[151,1436,379,1611]" captionTargetPageId="2" captionText="FIGURE 2. Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov.: A, male palp, ventral view; B, epigynum, ventral view; C, female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772817" httpUri="https://zenodo.org/record/3772817/files/figure.png" pageId="1" pageNumber="292">Fig. 2A</figureCitation>
) resemble those of
<taxonomicName id="4C494D783C392866FBB11C2CFB2E63A0" authorityName="Polotow &amp; Brescovit" authorityYear="2009" box="[1071,1193,1661,1685]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="ordinario">
<emphasis id="B93DEAE93C392866FBB11C2CFB2E63A0" box="[1071,1193,1661,1685]" italics="true" pageId="1" pageNumber="292">I. ordinario</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD84B0A3C392866FB2B1C2DFF57638C" author="Polotow, D. &amp; Brescovit, A. D." pageId="1" pageNumber="292" pagination="583 - 614" refId="ref2184" refString="Polotow, D. &amp; Brescovit, A. D. (2009) Revision and cladistic analysis of Isoctenus and description of a new neotropical genus (Araneae, Ctenidae, Cteninae). Zoological Journal of the Linnean Society, 155, 583 - 614. https: // doi. org / 10.1111 / j. 1096 - 3642.2008.00452. x" type="journal article" year="2009">Polotow &amp; Brescovit, 2009</bibRefCitation>
: fig. 9A–B) by the elongated dorsal branch of RTA, bifid tip of embolus and shape of median apophysis, but it can be distinguished by the presence of a basal retrolateral projection of embolus (RPE), shorter tibia of palp and cymbium, and embolus with larger base. Males of
<taxonomicName id="4C494D783C392866FDA01CB8FCA76234" authorityName="Polotow &amp; Brescovit" authorityYear="2019" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1941" box="[574,800,1769,1793]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="guadalupei">
<emphasis id="B93DEAE93C392866FDA01CB8FCA76234" box="[574,800,1769,1793]" italics="true" pageId="1" pageNumber="292">Isoctenus guadalupei</emphasis>
</taxonomicName>
also present a projection on each coxae IV (
<figureCitation id="13722A7E3C392866FB6B1CB9FAC16234" box="[1269,1350,1768,1793]" captionStart="FIGURE 1" captionStartId="1.[151,250,1300,1325]" captionTargetBox="[151,1436,308,1273]" captionTargetId="figure@1.[151,1436,308,1273]" captionTargetPageId="1" captionText="FIGURE 1. Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov., male:A, dorsal habitus; B, ventral habitus; C, lateral view of venter, with arrows pointing to the projection on coxae IV; D, male palp, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772815" httpUri="https://zenodo.org/record/3772815/files/figure.png" pageId="1" pageNumber="292">Fig. 1C</figureCitation>
), which is absent in
<taxonomicName id="4C494D783C392866FE841D5CFE116210" authorityName="Polotow &amp; Brescovit" authorityYear="2009" box="[282,406,1805,1829]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="ordinario">
<emphasis id="B93DEAE93C392866FE841D5CFE116210" box="[282,406,1805,1829]" italics="true" pageId="1" pageNumber="292">I. ordinario</emphasis>
</taxonomicName>
. Females of
<taxonomicName id="4C494D783C392866FDBA1D5CFC8F6210" authorityName="Polotow &amp; Brescovit" authorityYear="2019" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1941" box="[548,776,1805,1829]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="guadalupei">
<emphasis id="B93DEAE93C392866FDBA1D5CFC8F6210" box="[548,776,1805,1829]" italics="true" pageId="1" pageNumber="292">Isoctenus guadalupei</emphasis>
</taxonomicName>
resemble those of
<taxonomicName id="4C494D783C392866FC491D5CFBD46210" authorityName="Polotow &amp; Brescovit" authorityYear="2009" box="[983,1107,1805,1829]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="292" phylum="Arthropoda" rank="species" species="ordinario">
<emphasis id="B93DEAE93C392866FC491D5CFBD46210" box="[983,1107,1805,1829]" italics="true" pageId="1" pageNumber="292">I. ordinario</emphasis>
</taxonomicName>
by the cordiform shape of the median field of epigynum and anterior position of lateral fields projections, but can be distinguished by the median field less excavated apically (
<figureCitation id="13722A7E3C392866FE041D05FE6B6258" box="[410,492,1876,1901]" captionStart="FIGURE 2" captionStartId="2.[151,250,1637,1662]" captionTargetBox="[161,1422,393,1596]" captionTargetId="figure@2.[151,1436,379,1611]" captionTargetPageId="2" captionText="FIGURE 2. Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov.: A, male palp, ventral view; B, epigynum, ventral view; C, female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772817" httpUri="https://zenodo.org/record/3772817/files/figure.png" pageId="1" pageNumber="292">Fig. 2B</figureCitation>
) and the larger spermathecae and copulatory ducts (
<figureCitation id="13722A7E3C392866FB861D05FBEB6258" box="[1048,1132,1876,1901]" captionStart="FIGURE 2" captionStartId="2.[151,250,1637,1662]" captionTargetBox="[161,1422,393,1596]" captionTargetId="figure@2.[151,1436,379,1611]" captionTargetPageId="2" captionText="FIGURE 2. Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov.: A, male palp, ventral view; B, epigynum, ventral view; C, female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772817" httpUri="https://zenodo.org/record/3772817/files/figure.png" pageId="1" pageNumber="292">Fig. 2C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35365703C392865FF591D29FEB062F7" lastPageId="2" lastPageNumber="293" pageId="1" pageNumber="292" type="description">
<paragraph id="8BF636FB3C392865FF591D29FB626453" blockId="1.[151,1437,1408,2045]" lastBlockId="2.[151,1436,153,358]" lastPageId="2" lastPageNumber="293" pageId="1" pageNumber="292">
<emphasis id="B93DEAE93C392866FF591D29FED562A5" bold="true" box="[199,338,1912,1936]" pageId="1" pageNumber="292">Description.</emphasis>
<emphasis id="B93DEAE93C392866FEC21D28FE1462A4" box="[348,403,1913,1937]" italics="true" pageId="1" pageNumber="292">Male</emphasis>
(MACN-Ar 30496). Prossoma uniformly orange, darker in the dorsum; with brown thoracic groove and black rings around eyes; greyish dorsal opistossoma with light brown dorsal, folium-like marking (
<figureCitation id="13722A7E3C392866FAEF1DCDFF2762EC" captionStart="FIGURE 1" captionStartId="1.[151,250,1300,1325]" captionTargetBox="[151,1436,308,1273]" captionTargetId="figure@1.[151,1436,308,1273]" captionTargetPageId="1" captionText="FIGURE 1. Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov., male:A, dorsal habitus; B, ventral habitus; C, lateral view of venter, with arrows pointing to the projection on coxae IV; D, male palp, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772815" httpUri="https://zenodo.org/record/3772815/files/figure.png" pageId="1" pageNumber="292">Fig. 1</figureCitation>
A−B). Total length 10.20. Carapace 5.60 long and 4.35 wide. Eye diameters: AME 0.20, ALE 0.20, PME 0.25, PLE 0.24. Leg measurements: I: femur 6.90/ patella 2.80/ tibia 7.10/ metatarsus 6.87/ tarsus 3.00/ total 26.67; II: 5.93/ 2.55/ 5.80/ 5.90/ 2.55/ 22.73; III: 5.16/ 2.00/ 5.00/ 5.65/ 2.15/ 19.96; IV: 7.00/ 2.25/ 6.70/ 8.90/ 2.90/ 27.75. Leg spination: tibia I-II v2-2-2-2-2, r0, p0, III-IV v2-2-2, r1-1, p1-1; metatarsus I-II v2-2-2, r0, p0, III v2-2-2, r1-1-1, p-1-1-1, IV v1-1-1-1- 1-2, r1-1-1, p1-1-1. Coxa IV with a ventral projection (
<figureCitation id="13722A7E3C3A2865FD711AB3FCC465CF" box="[751,835,226,250]" captionStart="FIGURE 1" captionStartId="1.[151,250,1300,1325]" captionTargetBox="[151,1436,308,1273]" captionTargetId="figure@1.[151,1436,308,1273]" captionTargetPageId="1" captionText="FIGURE 1. Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov., male:A, dorsal habitus; B, ventral habitus; C, lateral view of venter, with arrows pointing to the projection on coxae IV; D, male palp, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772815" httpUri="https://zenodo.org/record/3772815/files/figure.png" pageId="2" pageNumber="293">Fig. 1C</figureCitation>
). Palp (
<figureCitation id="13722A7E3C3A2865FC061AB3FC7665CF" box="[920,1009,226,250]" captionStart="FIGURE 1" captionStartId="1.[151,250,1300,1325]" captionTargetBox="[151,1436,308,1273]" captionTargetId="figure@1.[151,1436,308,1273]" captionTargetPageId="1" captionText="FIGURE 1. Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov., male:A, dorsal habitus; B, ventral habitus; C, lateral view of venter, with arrows pointing to the projection on coxae IV; D, male palp, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772815" httpUri="https://zenodo.org/record/3772815/files/figure.png" pageId="2" pageNumber="293">Figs 1D</figureCitation>
,
<figureCitation id="13722A7E3C3A2865FC601AB3FBA465CF" box="[1022,1059,226,250]" captionStart="FIGURE 2" captionStartId="2.[151,250,1637,1662]" captionTargetBox="[161,1422,393,1596]" captionTargetId="figure@2.[151,1436,379,1611]" captionTargetPageId="2" captionText="FIGURE 2. Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov.: A, male palp, ventral view; B, epigynum, ventral view; C, female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772817" httpUri="https://zenodo.org/record/3772817/files/figure.png" pageId="2" pageNumber="293">2A</figureCitation>
): tibia approximately two thirds of cymbium length; RTA with ventral branch subtriangular; tibia with short and curved ventral apophysis; cymbium with ventral retrolateral projection reduced and dorsal retrolateral projection rounded; short and laminar embolus, with bifid tip and short basal projection; robust median apophysis, with conical anterior and posterior projections.
</paragraph>
<caption id="DF3666733C3A2865FF091C34FD926397" ID-DOI="http://doi.org/10.5281/zenodo.3772817" ID-Zenodo-Dep="3772817" httpUri="https://zenodo.org/record/3772817/files/figure.png" pageId="2" pageNumber="293" startId="2.[151,250,1637,1662]" targetBox="[161,1422,393,1596]" targetPageId="2">
<paragraph id="8BF636FB3C3A2865FF091C34FD926397" blockId="2.[151,1436,1637,1698]" pageId="2" pageNumber="293">
<emphasis id="B93DEAE93C3A2865FF091C34FE94634B" bold="true" box="[151,275,1637,1662]" pageId="2" pageNumber="293">FIGURE 2.</emphasis>
<taxonomicName id="4C494D783C3A2865FE871C36FD4B634B" authority="(Mello-Leitao, 1941)" authorityName="Polotow &amp; Brescovit" authorityYear="2019" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1941" box="[281,716,1637,1662]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="guadalupei" status="comb. nov.">
<emphasis id="B93DEAE93C3A2865FE871C36FE76634B" box="[281,497,1638,1662]" italics="true" pageId="2" pageNumber="293">Isoctenus guadalupei</emphasis>
(
<bibRefCitation id="EFD84B0A3C3A2865FE601C34FD42634B" author="Mello-Leitao, C. F. de" box="[510,709,1637,1662]" pageId="2" pageNumber="293" pagination="199 - 225" refId="ref2105" refString="Mello-Leitao, C. F. de (1941) Las aranas de la provincia de Santa Fe colectadas por el Profesor Biraben. Revista del Museo de La Plata, New Series (Zoologia), 2, 199 - 225." type="journal article" year="1941">Mello-Leitão, 1941</bibRefCitation>
)
</taxonomicName>
<emphasis id="B93DEAE93C3A2865FD4D1C37FCC1634B" bold="true" box="[723,838,1638,1662]" pageId="2" pageNumber="293">
<taxonomicNameLabel id="A20E57923C3A2865FD4D1C37FCC1634B" box="[723,838,1638,1662]" pageId="2" pageNumber="293" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
: A, male palp, ventral view; B, epigynum, ventral view; C, female internal genitalia, dorsal view.
</paragraph>
</caption>
<paragraph id="8BF636FB3C3A2865FF591C83FEB062F7" blockId="2.[151,1437,1746,2022]" pageId="2" pageNumber="293">
<emphasis id="B93DEAE93C3A2865FF591C83FE9063DF" box="[199,279,1746,1770]" italics="true" pageId="2" pageNumber="293">Female</emphasis>
(MLP 15180). Coloration faded, ranging from light brown to beige in the whole specimen. Total length 11.60. Carapace 5.50 long and 4.20 wide. Clypeus 0.17 high. Eye diameters: AME 0.26, ALE 0.20, PME 0.31, PLE 0.27. Leg measurements: I: femur 5.20/ patella 2.50/ tibia 5.00/ metatarsus 4.20/ tarsus 1.80/ total 18.70; II: 4.70/ 2.60/ 4.40/ 3.80/ 1.60/ 17.10; III: 4.20/ 2.00/ 3.50/ 3.70/ 1.50/ 14.90; IV: 5.60/ -/ -/ -/ -/ -. Leg spination: tibia I-II v2-2-2-2-2, r0, p0, III-IV v2-2-2, r1-1, p1-1; metatarsus I-II v2-2-2, r0, p0, III v2-2-2, r1-1-1, p-1-1-1. Epigynum (
<figureCitation id="13722A7E3C3A2865FBDD1D33FB04624F" box="[1091,1155,1890,1914]" captionStart="FIGURE 2" captionStartId="2.[151,250,1637,1662]" captionTargetBox="[161,1422,393,1596]" captionTargetId="figure@2.[151,1436,379,1611]" captionTargetPageId="2" captionText="FIGURE 2. Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov.: A, male palp, ventral view; B, epigynum, ventral view; C, female internal genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772817" httpUri="https://zenodo.org/record/3772817/files/figure.png" pageId="2" pageNumber="293">Fig. 2</figureCitation>
B−C): subrounded median field; lateral spurs short, partially covered by the median field and with a pointed tip. Spermathecae with rounded head and large base.
</paragraph>
</subSubSection>
<subSubSection id="C35365703C3A2864FF591D9FFD9665E0" lastPageId="3" lastPageNumber="294" pageId="2" pageNumber="293" type="distribution">
<paragraph id="8BF636FB3C3A2864FF591D9FFD9665E0" blockId="2.[151,1437,1746,2022]" lastBlockId="3.[151,1436,153,213]" lastPageId="3" lastPageNumber="294" pageId="2" pageNumber="293">
<emphasis id="B93DEAE93C3A2865FF591D9FFEDD62D3" bold="true" box="[199,346,1998,2022]" pageId="2" pageNumber="293">Distribution.</emphasis>
<collectingRegion id="498DF8193C3A2865FEFE1D9FFE4862D3" box="[352,463,1998,2022]" country="Argentina" name="Corrientes" pageId="2" pageNumber="293">Corrientes</collectingRegion>
and
<collectingRegion id="498DF8193C3A2865FD9A1D9FFDE562D3" box="[516,610,1998,2022]" country="Argentina" name="Santa Fe" pageId="2" pageNumber="293">Santa Fé</collectingRegion>
Provinces, northeastern
<collectingCountry id="F35E766B3C3A2865FCF71D9FFC5462D3" box="[873,979,1998,2022]" name="Argentina" pageId="2" pageNumber="293">Argentina</collectingCountry>
(
<figureCitation id="13722A7E3C3A2865FC7C1D9FFBA362D3" box="[994,1060,1998,2022]" captionStart="FIGURE 3" captionStartId="3.[151,250,860,885]" captionTargetBox="[289,1289,234,833]" captionTargetId="figure@3.[289,1299,234,834]" captionTargetPageId="3" captionText="FIGURE 3. Collection localities of Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov. and I. ordinario Polotow &amp; Brescovit, 2009: A, map representing South America (partial), with Chacoan and Parana dominions colored; B, detail of Chacoan and Parana dominions with distributional records of Isoctenus guadalupei and I. ordinario." figureDoi="http://doi.org/10.5281/zenodo.3772819" httpUri="https://zenodo.org/record/3772819/files/figure.png" pageId="2" pageNumber="293">Fig. 3</figureCitation>
).
<taxonomicName id="4C494D783C3A2865FBA71D9EFA8D62D3" authorityName="Polotow &amp; Brescovit" authorityYear="2009" box="[1081,1290,1998,2022]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="293" phylum="Arthropoda" rank="species" species="ordinario">
<emphasis id="B93DEAE93C3A2865FBA71D9EFA8D62D3" box="[1081,1290,1998,2022]" italics="true" pageId="2" pageNumber="293">Isoctenus ordinario</emphasis>
</taxonomicName>
is commonly found in the
<collectingRegion id="498DF8193C3B2864FEBF1ACBFEED6584" box="[289,362,154,177]" country="Brazil" name="Parana" pageId="3" pageNumber="294">Parana</collectingRegion>
Dominion of
<collectingCountry id="F35E766B3C3B2864FD9C1AC8FDC46584" box="[514,579,153,177]" name="Brazil" pageId="3" pageNumber="294">Brazil</collectingCountry>
and
<collectingCountry id="F35E766B3C3B2864FDE81AC8FD646584" box="[630,739,153,177]" name="Argentina" pageId="3" pageNumber="294">Argentina</collectingCountry>
, while
<taxonomicName id="4C494D783C3B2864FCAF1ACBFC476584" authorityName="Polotow &amp; Brescovit" authorityYear="2019" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1941" box="[817,960,153,177]" class="Arachnida" family="Ctenidae" genus="Isoctenus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="guadalupei">
<emphasis id="B93DEAE93C3B2864FCAF1ACBFC476584" box="[817,960,153,177]" italics="true" pageId="3" pageNumber="294">I. guadalupei</emphasis>
</taxonomicName>
is only found in the Chacoan province (part of the Chacoan Dominion) (
<figureCitation id="13722A7E3C3B2864FE5D1AECFD8365E0" box="[451,516,189,213]" captionStart="FIGURE 3" captionStartId="3.[151,250,860,885]" captionTargetBox="[289,1289,234,833]" captionTargetId="figure@3.[289,1299,234,834]" captionTargetPageId="3" captionText="FIGURE 3. Collection localities of Isoctenus guadalupei (Mello-Leitão, 1941) comb. nov. and I. ordinario Polotow &amp; Brescovit, 2009: A, map representing South America (partial), with Chacoan and Parana dominions colored; B, detail of Chacoan and Parana dominions with distributional records of Isoctenus guadalupei and I. ordinario." figureDoi="http://doi.org/10.5281/zenodo.3772819" httpUri="https://zenodo.org/record/3772819/files/figure.png" pageId="3" pageNumber="294">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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