246 lines
29 KiB
XML
246 lines
29 KiB
XML
<document id="B9F474F6BD7DE557048FC8E5F4A1CC86" ID-CLB-Dataset="55724" ID-DOI="10.5281/zenodo.4507307" ID-GBIF-Dataset="72a1f456-f29c-471e-baf8-6c026cc900a1" ID-ISSN="0253-116X" ID-RefBankID="1643FFB2ED42FFC2FFE4FFE28E27FFAF" ID-Zenodo-Dep="4507307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612537541968" checkinUser="felipe" docAuthor="Assing, V." docDate="2013" docId="03E087C369657A31DDA82A1D06FEFDEC" docLanguage="en" docName="LinzerbiolBeitr.45.1.0171-0190.pdf" docOrigin="Linzer biologische Beiträge 45 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Rugilus saudicus Assing 2013, nov.sp." docType="treatment" docVersion="10" lastPageNumber="189" masterDocId="FFD9FFBB69747A23DD0B2B5B0548FFA4" masterDocTitle="A revision of Palaearctic and Oriental Rugilus. III. Five new species from the Palaearctic region and additional records (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="190" masterPageNumber="171" pageNumber="188" updateTime="1698867118016" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="9502FC9CC4198317D73805528DFC736A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4D0C30C149EF2E9D0830B4E162C0F49C">
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<mods:title id="0E5E0F24C7C9D659AB9F1F0D7DAC533E">A revision of Palaearctic and Oriental Rugilus. III. Five new species from the Palaearctic region and additional records (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2DB7E63EC11862792334E2C44F397F66" type="personal">
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<mods:role id="5881B64CEBC83EB4F7FE079B60CE9BCF">
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<mods:roleTerm id="67AF72032A7E0C3371D8BA60CE24348D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9DFF18EA98261A6837E50D2B9761E0AF">Assing, V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A9AC239724DF8DEC88AF3550F3975509">text</mods:typeOfResource>
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<mods:titleInfo id="CF52ED39898F9F1892C3C75352CE7719">
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<mods:title id="132D91A77FA195CAB7E8A04D53341B04">Linzer biologische Beiträge</mods:title>
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</mods:titleInfo>
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<mods:part id="F5C65A4FFBC0AC4A71A5538BD26493C1">
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<mods:date id="74EA3D549FAD10B9E6507FE8A08785DE">2013</mods:date>
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<mods:detail id="30053D7A0088F136446E2F9F42A0AECA" type="pubDate">
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<mods:number id="340137F8F9576FD19C26B71BE951D211">2013-07-31</mods:number>
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</mods:detail>
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<mods:detail id="2FE6474B7CCF6601FAC075C362D7C24D" type="volume">
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<mods:number id="FB8A8B7979A39109BC038F513738223C">45</mods:number>
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</mods:detail>
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<mods:detail id="78D0F69BB602D8D0D8AC26A442E4D6B3" type="issue">
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<mods:number id="9F44129FC0174307F4571D28B139C656">1</mods:number>
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</mods:detail>
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<mods:extent id="F129938C32A3CCA01098DC352DAE704D" unit="page">
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<mods:start id="B032EFCA9613D3FC161F38107F991767">171</mods:start>
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<mods:end id="E777262B17A7F658D8130E6289C9BB5E">190</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="C77A4FC0297D433ED119DC54D472DE30">journal article</mods:classification>
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<mods:identifier id="7C73C9A887BC4EA21A63588265CC0DEE" type="DOI">10.5281/zenodo.4507307</mods:identifier>
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<mods:identifier id="3AE9ED27160DD4A5C298E735E64CDC0C" type="ISSN">0253-116X</mods:identifier>
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<treatment id="03E087C369657A31DDA82A1D06FEFDEC" ID-DOI="http://doi.org/10.5281/zenodo.6521081" ID-GBIF-Taxon="195191796" ID-Zenodo-Dep="6521081" LSID="urn:lsid:plazi:treatment:03E087C369657A31DDA82A1D06FEFDEC" httpUri="http://treatment.plazi.org/id/03E087C369657A31DDA82A1D06FEFDEC" lastPageId="18" lastPageNumber="189" pageId="17" pageNumber="188">
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<subSubSection id="C353655E69657A32DDA82A1D04E1FEF8" box="[163,425,326,348]" pageId="17" pageNumber="188" type="nomenclature">
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<paragraph id="8BF636D569657A32DDA82A1D04E1FEF8" blockId="17.[163,556,326,348]" box="[163,425,326,348]" pageId="17" pageNumber="188">
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<heading id="D0BE81B969657A32DDA82A1D04E1FEF8" box="[163,425,326,348]" fontSize="8" level="2" pageId="17" pageNumber="188" reason="2">
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<emphasis id="B93DEAC769657A32DDA82A1D04E1FEF8" bold="true" box="[163,425,326,348]" pageId="17" pageNumber="188">
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<taxonomicName id="4C494D5669657A32DDA82A1D041EFEF8" ID-CoL="799CD" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[163,342,326,348]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="188" phylum="Arthropoda" rank="species" species="saudicus" status="sp. nov.">
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<emphasis id="B93DEAC769657A32DDA82A1D041EFEF8" bold="true" box="[163,342,326,348]" italics="true" pageId="17" pageNumber="188">Rugilus saudicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20E57BC69657A32DC572A1D04E1FEF8" box="[348,425,326,348]" pageId="17" pageNumber="188" rank="species">nov.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C353655E69657A32DCA42A1D0764FEF8" box="[431,556,326,348]" pageId="17" pageNumber="188" type="description">
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<paragraph id="8BF636D569657A32DCA42A1D0764FEF8" blockId="17.[163,556,326,348]" box="[431,556,326,348]" pageId="17" pageNumber="188">
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<heading id="D0BE81B969657A32DCA42A1D0764FEF8" box="[431,556,326,348]" fontSize="8" level="2" pageId="17" pageNumber="188" reason="2">
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(
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<figureCitation id="13722A5069657A32DCBD2A1D076DFEF8" box="[438,549,326,348]" captionStart="Figs 23-30" captionStartId="16.[163,203,1425,1445]" captionTargetBox="[168,1061,201,1403]" captionTargetId="figure-92@16.[163,1070,197,1407]" captionTargetPageId="16" captionText="Figs 23-30: Rugilus lucens ASSING (23-25) and R. saudicus nov.sp. (26-30): (23, 27) male sternite VIII; (24, 28) aedeagus in lateral view; (25, 29) aedeagus in ventral view; (26) forebody; (30) apical portion of aedeagus in lateral view. Scale bars: 26: 1.0 mm; 23, 27: 0.2 mm; 24-25, 28-30: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4507339" httpUri="https://zenodo.org/record/4507339/files/figure.png" pageId="17" pageNumber="188">Figs 26-30</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C353655E69657A32DDA82A360420FE57" pageId="17" pageNumber="188" type="materials_examined">
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<paragraph id="8BF636D569657A32DDA82A360420FE57" blockId="17.[163,1071,363,499]" pageId="17" pageNumber="188">
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Type material:
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<materialsCitation id="3B213C8869657A32DC902A36074FFE0A" ID-GBIF-Occurrence="3762949386" collectingDate="2011-05-15" collectionCode="KSMA" collectorName="Sharaf" country="Saudi Arabia" county="Dhi Ayn Arch." determinerName="V. Assing" elevation="740" latitude="19.929443" location="Al Mukhwah" longLatPrecision="21" longitude="41.44167" municipality="Al Mukhwah" pageId="17" pageNumber="188" specimenCount="1" specimenCount-male="1" stateProvince="Al Bahah" typeStatus="holotype">
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<typeStatus id="54F2887769657A32DC902A3604A7FE25" box="[411,495,365,385]" pageId="17" pageNumber="188" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D4FFD5C69657A32DCFC2A300742FE24" box="[503,522,363,384]" pageId="17" pageNumber="188" type="male">♂</specimenCount>
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: "
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<collectingCountry id="F35E764569657A32DF2B2A36078FFE25" box="[544,711,365,385]" name="Saudi Arabia" pageId="17" pageNumber="188">SAUDI ARABIA</collectingCountry>
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<quantity id="4CB19B3069657A32DFDB2A360647FE25" box="[720,783,365,385]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="17" pageNumber="188" unit="m" value="740.0">
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<elevation id="0064D1E669657A32DFDB2A360647FE25" box="[720,783,365,385]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="17" pageNumber="188" unit="m" value="740.0">740 m</elevation>
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</quantity>
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,
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<collectingRegion id="498DF83769657A32DE112A36063EFE25" box="[794,886,365,385]" country="Saudi Arabia" name="Al Bahah" pageId="17" pageNumber="188">Al Bahah</collectingRegion>
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,
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<collectingMunicipality id="6B92ACAF69657A32DE892A360148FE25" box="[898,1024,365,385]" pageId="17" pageNumber="188">Al Mukhwah</collectingMunicipality>
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,
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<collectingCounty id="62974E5969657A32D9072A36044EFE3C" pageId="17" pageNumber="188">Dhi Ayn Arch.</collectingCounty>
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vill.,
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<geoCoordinate id="EE7D501269657A32DC362ADF04E1FE3C" box="[317,425,388,408]" degrees="19" direction="north" minutes="55" orientation="latitude" pageId="17" pageNumber="188" precision="15" seconds="46" value="19.929443">19°55'46''N</geoCoordinate>
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,
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<geoCoordinate id="EE7D501269657A32DCBE2ADF0756FE3C" box="[437,542,388,408]" degrees="41" direction="east" minutes="26" orientation="longitude" pageId="17" pageNumber="188" precision="15" seconds="30" value="41.44167">41°26'30''E</geoCoordinate>
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,
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<date id="FFF7101569657A32DF202ADF07C5FE3C" box="[555,653,388,408]" pageId="17" pageNumber="188" value="2011-05-15">
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<collectingDate id="EFB3E9FD69657A32DF202ADF07C5FE3C" box="[555,653,388,408]" pageId="17" pageNumber="188" value="2011-05-15">15.V.2011</collectingDate>
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</date>
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, leg.
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<collectorName id="26BC530369657A32DFCA2ADF07B4FE3C" box="[705,764,388,408]" pageId="17" pageNumber="188">Sharaf</collectorName>
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/ Holotypus ♂
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<originalDetermination id="A8578E2969657A32DE9A2AD90590FE0A" pageId="17" pageNumber="188">
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<taxonomicName id="4C494D5669657A32DE9A2AD90165FE33" authority="Assing, 2013" authorityName="Assing" authorityYear="2013" box="[913,1069,386,407]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="188" phylum="Arthropoda" rank="species" species="saudicus" status="sp. nov.">
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<emphasis id="B93DEAC769657A32DE9A2AD90165FE33" box="[913,1069,386,407]" italics="true" pageId="17" pageNumber="188">Rugilus saudicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20E57BC69657A32DDA82AC10590FE0A" box="[163,216,410,430]" pageId="17" pageNumber="188" rank="species">sp. n.</taxonomicNameLabel>
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</originalDetermination>
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, det.
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<determinerName id="7E2ADA1169657A32DC052AC10424FE0A" box="[270,364,410,430]" pageId="17" pageNumber="188">V. Assing</determinerName>
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2013" (
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<collectionCode id="ED58AE1069657A32DCB12AC10749FE0A" box="[442,513,410,430]" pageId="17" pageNumber="188">KSMA</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B213C8869657A32DF182AC106A3FE0A" ID-GBIF-Occurrence="3762949328" box="[531,1003,409,435]" collectingDate="2011-05-15" collectionCode="KSMA" collectorName="Sharaf" country="Saudi Arabia" determinerName="V. Assing" elevation="740" latitude="19.929443" location="Dhi Ayn Arch." longLatPrecision="21" longitude="41.44167" municipality="Al Mukhwah" pageId="17" pageNumber="188" specimenCount="1" specimenCount-female="1" stateProvince="Al Bahah" typeStatus="paratype">
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<typeStatus id="54F2887769657A32DF182AC10727FE0A" box="[531,623,410,430]" pageId="17" pageNumber="188" type="paratype">Paratypes</typeStatus>
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: 1♂,
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<specimenCount id="9D4FFD5C69657A32DFAC2AC1078DFE17" box="[679,709,410,435]" pageId="17" pageNumber="188" type="female">1♀</specimenCount>
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: same data as holotype (
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<collectionCode id="ED58AE1069657A32DEB82AC106ABFE0A" box="[947,995,410,430]" pageId="17" pageNumber="188">cAss</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="3B213C8869657A32DEFC2AC107EAFE61" ID-GBIF-Occurrence="3762949376" collectingDate="2011-05-11" collectionCode="KSMA" collectorName="Sharaf" country="Saudi Arabia" determinerName="V. Assing" elevation="740" latitude="19.929443" location="Dhi Ayn Arch." longLatPrecision="21" longitude="41.44167" municipality="Al Mukhwah" pageId="17" pageNumber="188" specimenCount="2" specimenCount-female="2" stateProvince="Al Bahah" typeStatus="paratype">
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2♂♂,
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<specimenCount id="9D4FFD5C69657A32DDA82AEA059CFE6E" box="[163,212,433,458]" pageId="17" pageNumber="188" type="female">2♀♀</specimenCount>
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: same data as holotype, but
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<date id="FFF7101569657A32DCF22AEA0710FE61" box="[505,600,433,453]" pageId="17" pageNumber="188" value="2011-05-11">
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<collectingDate id="EFB3E9FD69657A32DCF22AEA0710FE61" box="[505,600,433,453]" pageId="17" pageNumber="188" value="2011-05-11">11.V.2011</collectingDate>
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</date>
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(
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<collectionCode id="ED58AE1069657A32DF602AEA07D3FE61" box="[619,667,433,453]" pageId="17" pageNumber="188">cAss</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="3B213C8869657A32DFBA2AEA042CFE57" ID-GBIF-Occurrence="3762949366" collectingDate="2011-09-23" collectorName="Leg. M. R. Sharaf" country="Saudi Arabia" county="Saudi Arabia" elevation="744" latitude="19.929583" location="Al Mukhwah" longLatPrecision="1" longitude="41.442863" municipality="Al Bahan" pageId="17" pageNumber="188" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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1♂,
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<specimenCount id="9D4FFD5C69657A32DFEA2AEA07B7FE6E" box="[737,767,433,458]" pageId="17" pageNumber="188" type="female">1♀</specimenCount>
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: "
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<collectingCounty id="62974E5969657A32DE122AEA06D2FE61" box="[793,922,433,453]" pageId="17" pageNumber="188">Saudi Arabia</collectingCounty>
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,
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<collectingMunicipality id="6B92ACAF69657A32DEA22AEA0140FE61" box="[937,1032,433,453]" pageId="17" pageNumber="188">Al Bahan</collectingMunicipality>
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,
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<location id="8E96600E69657A32D91C2AEA044AFE78" LSID="urn:lsid:plazi:treatment:03E087C369657A31DDA82A1D06FEFDEC:8E96600E69657A32D91C2AEA044AFE78" country="Saudi Arabia" county="Saudi Arabia" latitude="19.929583" longLatPrecision="1" longitude="41.442863" municipality="Al Bahan" name="Al Mukhwah" pageId="17" pageNumber="188">Al Mukhwah</location>
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,
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<location id="8E96600E69657A32DC1A2A9304EEFE78" LSID="urn:lsid:plazi:treatment:03E087C369657A31DDA82A1D06FEFDEC:8E96600E69657A32DC1A2A9304EEFE78" box="[273,422,456,476]" country="Saudi Arabia" county="Saudi Arabia" latitude="19.929583" longLatPrecision="1" longitude="41.442863" municipality="Al Bahan" name="Dhi Ayn Arch." pageId="17" pageNumber="188">Dhi Ayn Arch.</location>
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vill.,
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<date id="FFF7101569657A32DCEC2A930719FE78" box="[487,593,456,476]" pageId="17" pageNumber="188" value="2011-09-23">
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<collectingDate id="EFB3E9FD69657A32DCEC2A930719FE78" box="[487,593,456,476]" pageId="17" pageNumber="188" value="2011-09-23">23.IX.2011</collectingDate>
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</date>
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,
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<geoCoordinate id="EE7D501269657A32DF6A2A930796FE78" box="[609,734,456,476]" degrees="19" direction="north" minutes="55" orientation="latitude" pageId="17" pageNumber="188" precision="1" seconds="46.5" value="19.929583">19°55'46.5''N</geoCoordinate>
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,
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<geoCoordinate id="EE7D501269657A32DFE52A930620FE78" box="[750,872,456,476]" degrees="41" direction="east" minutes="26" orientation="longitude" pageId="17" pageNumber="188" precision="1" seconds="34.3" value="41.442863">41°26'34.3''E</geoCoordinate>
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,
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<quantity id="4CB19B3069657A32DE732A9306F2FE78" box="[888,954,456,476]" metricMagnitude="2" metricUnit="m" metricValue="7.44" pageId="17" pageNumber="188" unit="m" value="744.0">
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<elevation id="0064D1E669657A32DE732A9306F2FE78" box="[888,954,456,476]" metricMagnitude="2" metricUnit="m" metricValue="7.44" pageId="17" pageNumber="188" unit="m" value="744.0">744 m</elevation>
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</quantity>
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,
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<collectorName id="26BC530369657A32DEC12A9305A9FE57" pageId="17" pageNumber="188">Leg. M.R. Sharaf</collectorName>
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, 0046" (
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<collectionCode id="ED58AE1069657A32DC262A840416FE57" box="[301,350,479,499]" pageId="17" pageNumber="188">cAss</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C353655E69657A32DDA8295904F2FD93" pageId="17" pageNumber="188" type="etymology">
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<paragraph id="8BF636D569657A32DDA8295904F2FD93" blockId="17.[163,1070,514,567]" pageId="17" pageNumber="188">
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Etymology: The specific epithet (adjective) is derived from
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<collectingCountry id="F35E764569657A32DE51295906ACFDBC" box="[858,996,514,536]" name="Saudi Arabia" pageId="17" pageNumber="188">Saudi Arabia</collectingCountry>
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, where the species was discovered.
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</paragraph>
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</subSubSection>
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<subSubSection id="C353655E69657A32DDA8291D059EF9B4" pageId="17" pageNumber="188" type="description">
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<paragraph id="8BF636D569657A32DDA8291D07AFFD75" blockId="17.[163,1070,582,721]" pageId="17" pageNumber="188">
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Description: Small species; body length
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<quantity id="4CB19B3069657A32DF90291D065CFDF8" box="[667,788,582,604]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.9" metricValueMin="3.4" pageId="17" pageNumber="188" unit="mm" value="3.65" valueMax="3.9" valueMin="3.4">3.4-3.9 mm</quantity>
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; length of forebody
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<quantity id="4CB19B3069657A32DEED291D0585FDDD" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="17" pageNumber="188" unit="mm" value="2.2" valueMax="2.3" valueMin="2.1">2.1-2.3 mm</quantity>
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. Coloration: head and pronotum bright reddish; elytra reddish, posterior margins broadly yellow, anterior to this yellow margin usually more or less extensively and more or less strongly tranversely infuscate, occasionally leaving only the anterior third reddish; abdomen dark-brown; legs and antennae yellow.
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</paragraph>
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<paragraph id="8BF636D569657A32DDA829BA06B0FC5A" blockId="17.[163,1070,737,1022]" pageId="17" pageNumber="188">
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Head (
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<figureCitation id="13722A5069657A32DDE329BA047DFD53" box="[232,309,737,759]" captionStart="Figs 23-30" captionStartId="16.[163,203,1425,1445]" captionTargetBox="[168,1061,201,1403]" captionTargetId="figure-92@16.[163,1070,197,1407]" captionTargetPageId="16" captionText="Figs 23-30: Rugilus lucens ASSING (23-25) and R. saudicus nov.sp. (26-30): (23, 27) male sternite VIII; (24, 28) aedeagus in lateral view; (25, 29) aedeagus in ventral view; (26) forebody; (30) apical portion of aedeagus in lateral view. Scale bars: 26: 1.0 mm; 23, 27: 0.2 mm; 24-25, 28-30: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4507339" httpUri="https://zenodo.org/record/4507339/files/figure.png" pageId="17" pageNumber="188">Fig. 26</figureCitation>
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) distinctly transverse, 1.20-1.25 times as broad as long; posterior outline (behind eyes) smoothly and weakly convex in dorsal view, posterior angles completely obsolete; punctation rather coarse and moderately dense in anterior and antero-lateral portion, and sparse in postero-median portion; interstices without microsculpture and glossy. Eyes conspicuously large and bulging, longer than distance from posterior margin of eye to posterior constriction in dorsal view. Labrum with one distinct toothlike projection on either side of middle. Antenna with antennomere III approximately 2.5 times as long as broad, as long as, or slightly longer than II; IV approximately 1.5 times as long as broad and distinctly shorter than III; V weakly oblong; VI approximately a long as broad; VII-X increasingly transverse; X less than 1.5 times as broad as long.
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</paragraph>
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<paragraph id="8BF636D569657A32DDA82F5507BEFBFA" blockId="17.[163,1070,1037,1118]" pageId="17" pageNumber="188">
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Pronotum (
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<figureCitation id="13722A5069657A32DC1E2F560429FB87" box="[277,353,1037,1059]" captionStart="Figs 23-30" captionStartId="16.[163,203,1425,1445]" captionTargetBox="[168,1061,201,1403]" captionTargetId="figure-92@16.[163,1070,197,1407]" captionTargetPageId="16" captionText="Figs 23-30: Rugilus lucens ASSING (23-25) and R. saudicus nov.sp. (26-30): (23, 27) male sternite VIII; (24, 28) aedeagus in lateral view; (25, 29) aedeagus in ventral view; (26) forebody; (30) apical portion of aedeagus in lateral view. Scale bars: 26: 1.0 mm; 23, 27: 0.2 mm; 24-25, 28-30: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4507339" httpUri="https://zenodo.org/record/4507339/files/figure.png" pageId="17" pageNumber="188">Fig. 26</figureCitation>
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) small in relation to head, 1.05-1.10 times as long as broad and 0.65- 0.70 times as broad as head; midline broadly impunctate; laterally with rather sparse and fine punctation; interstices glossy, without microsculpture.
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</paragraph>
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<paragraph id="8BF636D569657A32DDA82F3606FCFB5C" blockId="17.[163,1070,1133,1272]" pageId="17" pageNumber="188">
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Elytra (
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<figureCitation id="13722A5069657A32DDFD2F36040FFB27" box="[246,327,1133,1155]" captionStart="Figs 23-30" captionStartId="16.[163,203,1425,1445]" captionTargetBox="[168,1061,201,1403]" captionTargetId="figure-92@16.[163,1070,197,1407]" captionTargetPageId="16" captionText="Figs 23-30: Rugilus lucens ASSING (23-25) and R. saudicus nov.sp. (26-30): (23, 27) male sternite VIII; (24, 28) aedeagus in lateral view; (25, 29) aedeagus in ventral view; (26) forebody; (30) apical portion of aedeagus in lateral view. Scale bars: 26: 1.0 mm; 23, 27: 0.2 mm; 24-25, 28-30: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4507339" httpUri="https://zenodo.org/record/4507339/files/figure.png" pageId="17" pageNumber="188">Fig. 26</figureCitation>
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) approximately as long as pronotum and with pronounced humeral angles; punctation fine and sparse; interstices without microsculpture. Hind wings apparently fully developed. Metatarsus slender, approximately 0.9 times as long as metatibia; metatarsomeres I-IV of decreasing length, but all oblong; metatarsomere I approximately as long as the combined length of II and III, or slightly longer.
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</paragraph>
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<paragraph id="8BF636D569657A32DDA82E53061AFAD2" blockId="17.[163,1071,1288,1398]" pageId="17" pageNumber="188">Abdomen broadest at segments V/VI, slightly narrower than elytra; anterior impressions of tergites III-V shallow and with dense, coarse punctation; remainder of tergal surfaces with moderately fine and moderately dense punctation; interstices without microsculpture; posterior margin of tergite VII with palisade fringe.</paragraph>
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<paragraph id="8BF636D569657A32DDA82EDE059EF9B4" blockId="17.[163,1070,1413,1552]" pageId="17" pageNumber="188">
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♂: sternite VII in postero-median portion without setae, posterior margin weakly concave; posterior excision of sternite VIII broad and not very deep (
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<figureCitation id="13722A5069657A32DE642EF906F5FA1C" box="[879,957,1442,1464]" captionStart="Figs 23-30" captionStartId="16.[163,203,1425,1445]" captionTargetBox="[168,1061,201,1403]" captionTargetId="figure-92@16.[163,1070,197,1407]" captionTargetPageId="16" captionText="Figs 23-30: Rugilus lucens ASSING (23-25) and R. saudicus nov.sp. (26-30): (23, 27) male sternite VIII; (24, 28) aedeagus in lateral view; (25, 29) aedeagus in ventral view; (26) forebody; (30) apical portion of aedeagus in lateral view. Scale bars: 26: 1.0 mm; 23, 27: 0.2 mm; 24-25, 28-30: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4507339" httpUri="https://zenodo.org/record/4507339/files/figure.png" pageId="17" pageNumber="188">Fig. 27</figureCitation>
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); aedeagus (
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<figureCitation id="13722A5069657A32DDA12EE40452FA71" box="[170,282,1471,1493]" captionStart="Figs 23-30" captionStartId="16.[163,203,1425,1445]" captionTargetBox="[168,1061,201,1403]" captionTargetId="figure-92@16.[163,1070,197,1407]" captionTargetPageId="16" captionText="Figs 23-30: Rugilus lucens ASSING (23-25) and R. saudicus nov.sp. (26-30): (23, 27) male sternite VIII; (24, 28) aedeagus in lateral view; (25, 29) aedeagus in ventral view; (26) forebody; (30) apical portion of aedeagus in lateral view. Scale bars: 26: 1.0 mm; 23, 27: 0.2 mm; 24-25, 28-30: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4507339" httpUri="https://zenodo.org/record/4507339/files/figure.png" pageId="17" pageNumber="188">Figs 28-30</figureCitation>
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) approximately
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<quantity id="4CB19B3069657A32DCCA2EE40754FA71" box="[449,540,1471,1493]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="17" pageNumber="188" unit="mm" value="0.38">0.38 mm</quantity>
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long; ventral process apically bent dorsad (hooked) in lateral view, on either side of ventral process with a moderately sclerotized lateral lobe.
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</paragraph>
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</subSubSection>
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<subSubSection id="C353655E69657A31DDA82D44049FFE68" lastPageId="18" lastPageNumber="189" pageId="17" pageNumber="188" type="discussion">
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<paragraph id="8BF636D569657A31DDA82D44049FFE68" blockId="17.[163,1070,1567,1589]" lastBlockId="18.[163,1072,203,460]" lastPageId="18" lastPageNumber="189" pageId="17" pageNumber="188">
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Comparative notes
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<taxonomicName id="4C494D5669657A32DF0F2D4407FFF990" authorityName="Assing" authorityYear="2013" box="[516,695,1567,1588]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="188" phylum="Arthropoda" rank="species" species="saudicus">
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<emphasis id="B93DEAC769657A32DF0F2D4407FFF990" box="[516,695,1567,1588]" italics="true" pageId="17" pageNumber="188">Rugilus saudicus</emphasis>
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</taxonomicName>
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|
is evidently closely related to
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<taxonomicName id="4C494D5669657A31D9132D7B05ACFF45" authorityName="ASSING" authorityYear="2012" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" lastPageId="18" lastPageNumber="189" order="Coleoptera" pageId="17" pageNumber="188" phylum="Arthropoda" rank="species" species="lucens">
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<emphasis id="B93DEAC769657A31D9132D7B05ACFF45" italics="true" lastPageId="18" lastPageNumber="189" pageId="17" pageNumber="188">R. lucens</emphasis>
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</taxonomicName>
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, as can be inferred from the similar external and male sexual characters. It is distinguished from
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<taxonomicName id="4C494D5669667A31DC612BB10480FF5A" authorityName="ASSING" authorityYear="2012" box="[362,456,233,255]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="189" phylum="Arthropoda" rank="species" species="lucens">
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<emphasis id="B93DEAC769667A31DC612BB10480FF5A" box="[362,456,233,255]" italics="true" pageId="18" pageNumber="189">R. lucens</emphasis>
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</taxonomicName>
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|
particularly by the different shape of the head (in
|
|
<taxonomicName id="4C494D5669667A31DEDA2BB10167FF5A" authorityName="ASSING" authorityYear="2012" box="[977,1071,233,255]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="189" phylum="Arthropoda" rank="species" species="lucens">
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<emphasis id="B93DEAC769667A31DEDA2BB10167FF5A" box="[977,1071,233,255]" italics="true" pageId="18" pageNumber="189">R. lucens</emphasis>
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</taxonomicName>
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|
relatively smaller, less transverse, and with moderately marked posterior angles), the sparser punctation of the head, the finer and much sparser punctation of the elytra, and the morphology of the aedeagus. The coloration is similar to that of
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<emphasis id="B93DEAC769667A31DE922A1A0166FEF2" box="[921,1070,321,342]" italics="true" pageId="18" pageNumber="189">R. clarissimus</emphasis>
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|
(BERNHAUER 1915) from the Afrotropical region, but that species is larger (
|
|
<quantity id="4CB19B3069667A31DEA32A05016BFED7" box="[936,1059,350,372]" metricMagnitude="-3" metricUnit="m" metricValue="4.199999999999999" metricValueMax="4.3" metricValueMin="4.1" pageId="18" pageNumber="189" unit="mm" value="4.199999999999999" valueMax="4.3" valueMin="4.1">4.1-4.3 mm</quantity>
|
|
), has a less transverse head, much smaller eyes (distinctly shorter than postocular region in dorsal view), and an aedeagus of different shape. For illustrations of the aedeagus of
|
|
<emphasis id="B93DEAC769667A31D9112AC2045EFE6F" italics="true" pageId="18" pageNumber="189">R. clarissimus</emphasis>
|
|
see
|
|
<bibRefCitation id="EFD84B2469667A31DC482AED049BFE68" author="FAGEL G" box="[323,467,438,460]" pageId="18" pageNumber="189" pagination="1 - 31" refId="ref11154" refString="FAGEL G. (1953): Contribution a la connaissance des Staphylinidae. XVIII. - Les Stilicus africains a edeage pubescent. - Institut royal des Sciences naturelles de Belgique Bulletin 29 (20): 1 - 31." type="journal article" year="1953">FAGEL (1953)</bibRefCitation>
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.
|
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</paragraph>
|
|
</subSubSection>
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<subSubSection id="C353655E69667A31DDA82A8006FEFDEC" pageId="18" pageNumber="189" type="distribution">
|
|
<paragraph id="8BF636D569667A31DDA82A8006FEFDEC" blockId="18.[163,1071,475,585]" pageId="18" pageNumber="189">
|
|
Distribution and natural history:The type locality is situated in
|
|
<collectingRegion id="498DF83769667A31DDA82AA3044CFDAA" box="[163,260,504,526]" country="Saudi Arabia" name="Al Bahah" pageId="18" pageNumber="189">Al Bahah</collectingRegion>
|
|
in southwestern
|
|
<collectingCountry id="F35E764569667A31DCBB2AA3077DFDAA" box="[432,565,504,526]" name="Saudi Arabia" pageId="18" pageNumber="189">Saudi Arabia</collectingCountry>
|
|
at an altitude of approximately
|
|
<specimenCount id="9D4FFD5C69667A31DE732AA306F7FDAA" box="[888,959,504,526]" pageId="18" pageNumber="189" type="generic">
|
|
<quantity id="4CB19B3069667A31DE732AA306F2FDAA" box="[888,954,504,526]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="18" pageNumber="189" unit="m" value="740.0">740 m</quantity>
|
|
.
|
|
</specimenCount>
|
|
The soil is compact clay and humid throughout the year due to irrigation of banana plantations. The specimens were sifted from leaf litter beneath
|
|
<emphasis id="B93DEAC769667A31DF7C296807F5FDEC" box="[631,701,563,584]" italics="true" pageId="18" pageNumber="189">Acacia</emphasis>
|
|
(SHARAF, pers. comm.).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
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|
</document> |