219 lines
26 KiB
XML
219 lines
26 KiB
XML
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<mods:title id="DE9DE11A18A69622A6BAA3F29490B370">Systematic revision of the freshwater snail Margarya Nevill, 1877 (Mollusca: Viviparidae) endemic to the ancient lakes of Yunnan, China, with description of new taxa</mods:title>
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<mods:namePart id="E6733E46F5BF94845955522DCEAD85EC">Zhang, Le-Jia</mods:namePart>
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<mods:namePart id="C6EB5E7E3984034699F74D69740850D6">Chen, Shi-Chao</mods:namePart>
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<mods:namePart id="91DC88A5CA4B8A243933CC38298D5629">Yang, Li-Te</mods:namePart>
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<mods:namePart id="E3F4D52A6AADCBB35E05E980A5E23A23">Jin, Lei</mods:namePart>
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<mods:namePart id="B83A08678DD4784DCF738A7238DE1ACA">Köhler, Frank</mods:namePart>
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<mods:title id="2491A1F5BCCD1DFAB734C988F3F6347A">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="76F67F793B7D6C54CC39C39C7688C9BB">2015</mods:date>
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<mods:number id="7F3AA1080CDB35585D4B667CF3B1936F">2015-07-21</mods:number>
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<mods:number id="F635E880134A3E028A6CA341EFD22DFD">174</mods:number>
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<paragraph id="8BF6368FFF96E264FC7686DFFA8BFBE9" blockId="22.[971,1296,1012,1034]" box="[971,1296,1012,1034]" pageId="22" pageNumber="782">
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<heading id="D0BE81E3FF96E264FC7686DFFA8BFBE9" box="[971,1296,1012,1034]" centered="true" fontSize="8" level="2" pageId="22" pageNumber="782" reason="8">
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<taxonomicName id="4C494D0CFF96E264FC7686DFFA8BFBE9" ID-CoL="7PJ47" authority="HE, 2013" authorityName="He" authorityYear="2013" box="[971,1296,1012,1034]" class="Gastropoda" family="Viviparidae" genus="Tchangmargarya" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="782" phylum="Mollusca" rank="genus">
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<emphasis id="B93DEA9DFF96E264FC7686DFFB06FBE8" box="[971,1181,1013,1034]" italics="true" pageId="22" pageNumber="782">TCHANGMARGARYA</emphasis>
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<bibRefCitation id="EFD84B7EFF96E264FB1886DFFA8BFBE9" author="He J" box="[1189,1296,1012,1034]" pageId="22" pageNumber="782" pagination="3 - 15" refId="ref23578" refString="He J. 2013. A conchological review of Margarya - complex (Gastropoda: Viviparidae). Shell Discoveries 2: 3 - 15." type="journal article" year="2013">HE, 2013</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C3536504FF96E265FC878131FB5BFCC0" lastPageId="23" lastPageNumber="783" pageId="22" pageNumber="782" type="materials_examined">
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<paragraph id="8BF6368FFF96E264FC878131FB5CFBAF" blockId="22.[826,1441,1051,1103]" pageId="22" pageNumber="782">
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<typeStatus id="54F2882DFF96E264FC878131FCEAFBD0" box="[826,881,1051,1072]" pageId="22" pageNumber="782">Type</typeStatus>
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species:
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<taxonomicName id="4C494D0CFF96E264FC668136FC3DFBAF" authority="Tchang & Tsi, 1949" authorityName="Tchang & Tsi" authorityYear="1949" class="Gastropoda" family="Viviparidae" genus="Margarya" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="782" phylum="Mollusca" rank="species" species="yangtsunghaiensis">
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<emphasis id="B93DEA9DFF96E264FC668136FAB3FBD0" box="[987,1320,1051,1073]" italics="true" pageId="22" pageNumber="782">Margarya yangtsunghaiensis</emphasis>
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<bibRefCitation id="EFD84B7EFF96E264FA8C8131FC3DFBAF" author="Tchang X & Tsi ZY" pageId="22" pageNumber="782" pagination="1 - 26" refId="ref25436" refString="Tchang X, Tsi ZY. 1949. A revision of the genus Margarya of the family Viviparidae. Contributions from the Institute of Zoology National Academy of Peiping 5: 1 - 26." type="journal article" year="1949">Tchang & Tsi, 1949</bibRefCitation>
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, by original designation.
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</paragraph>
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<emphasis id="B93DEA9DFF96E264FC878153FB82FB6D" box="[826,1049,1144,1166]" italics="true" pageId="22" pageNumber="782">Taxonomic remarks</emphasis>
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</paragraph>
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<paragraph id="8BF6368FFF96E264FC8781BDFBF3F955" blockId="22.[826,1442,1144,1717]" pageId="22" pageNumber="782">
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<bibRefCitation id="EFD84B7EFF96E264FC8781BDFB8EFB4D" author="Tchang X & Tsi ZY" box="[826,1045,1175,1197]" pageId="22" pageNumber="782" pagination="1 - 26" refId="ref25436" refString="Tchang X, Tsi ZY. 1949. A revision of the genus Margarya of the family Viviparidae. Contributions from the Institute of Zoology National Academy of Peiping 5: 1 - 26." type="journal article" year="1949">Tchang & Tsi (1949)</bibRefCitation>
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descibed
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<taxonomicName id="4C494D0CFF96E264FBC381B2FAFCFB4C" authorityName="Tchang & Tsi" authorityYear="1949" box="[1150,1383,1175,1197]" class="Gastropoda" family="Viviparidae" genus="Margarya" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="782" phylum="Mollusca" rank="species" species="yangtsunghaiensis">
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<emphasis id="B93DEA9DFF96E264FBC381B2FAFCFB4C" box="[1150,1383,1175,1197]" italics="true" pageId="22" pageNumber="782">M. yangtsunghaiensis</emphasis>
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</taxonomicName>
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from Lake Yangzonghai, and they pointed out that it greatly differs from any other
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<taxonomicName id="4C494D0CFF96E264FB9281FFFB75FB0A" box="[1071,1262,1236,1258]" class="Gastropoda" family="Viviparidae" genus="Margarya" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="782" phylum="Mollusca" rank="species" species="undetermined">
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<emphasis id="B93DEA9DFF96E264FB9281FFFB00FB0A" box="[1071,1179,1237,1258]" italics="true" pageId="22" pageNumber="782">Margarya</emphasis>
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species
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</taxonomicName>
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by having acute apex and extremely prominent regular nodules; however, they still treated it as a speices of
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<taxonomicName id="4C494D0CFF96E264FB738038FAA4FAC7" authorityName="Nevill" authorityYear="1877" box="[1230,1343,1298,1319]" class="Gastropoda" family="Viviparidae" genus="Margarya" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="782" phylum="Mollusca" rank="genus">
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<emphasis id="B93DEA9DFF96E264FB738038FAA4FAC7" box="[1230,1343,1298,1319]" italics="true" pageId="22" pageNumber="782">Margarya</emphasis>
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</taxonomicName>
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because of its large solid shell with ramp-like shoulder, elevat- ed keels, and distribution limited to
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<collectingRegion id="498DF86DFF96E264FB548065FADEFA84" box="[1257,1349,1359,1380]" country="China" name="Yunnan" pageId="22" pageNumber="782">Yunnan</collectingRegion>
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. All the reasearchers followed their conclusions until
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<bibRefCitation id="EFD84B7EFF96E264FAFD8044FC1BFA42" author="Du LN & Yang JX & Rintelen T & Chen YX & Aldridge D" pageId="22" pageNumber="782" pagination="2154 - 2162" refId="ref23356" refString="Du LN, Yang JX, Rintelen T, Chen YX, Aldridge D. 2013. Molecular phylogenetic evidence that the Chinese Viviparid genus Margarya (Gastropoda: Viviparidae) is polyphyletic. Chinese Science Bulletin 58: 2154 - 2162." type="journal article" year="2013">
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Du
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<emphasis id="B93DEA9DFF96E264FAD78044FA01FA62" box="[1386,1434,1389,1411]" italics="true" pageId="22" pageNumber="782">et al</emphasis>
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. (2013)
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</bibRefCitation>
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pointed out this species formed a monophyletic group and didn′ t cluster with any other
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<taxonomicName id="4C494D0CFF96E264FA908081FC16FA3F" class="Gastropoda" family="Viviparidae" genus="Margarya" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="782" phylum="Mollusca" rank="species" species="undetermined">
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<emphasis id="B93DEA9DFF96E264FA908081FA3BFA20" box="[1325,1440,1451,1472]" italics="true" pageId="22" pageNumber="782">Margarya</emphasis>
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species
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</taxonomicName>
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, doubting its delimitation of genus. Based on the morphological differences described by
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<bibRefCitation id="EFD84B7EFF96E264FA8C80C2FC2CF9FC" author="Tchang X & Tsi ZY" pageId="22" pageNumber="782" pagination="1 - 26" refId="ref25436" refString="Tchang X, Tsi ZY. 1949. A revision of the genus Margarya of the family Viviparidae. Contributions from the Institute of Zoology National Academy of Peiping 5: 1 - 26." type="journal article" year="1949">Tchang & Tsi (1949)</bibRefCitation>
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,
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<bibRefCitation id="EFD84B7EFF96E264FC75832DFBDAF9FC" author="He J" box="[968,1089,1542,1564]" pageId="22" pageNumber="782" pagination="3 - 15" refId="ref23578" refString="He J. 2013. A conchological review of Margarya - complex (Gastropoda: Viviparidae). Shell Discoveries 2: 3 - 15." type="journal article" year="2013">He (2013)</bibRefCitation>
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founded a new subgenus of
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<taxonomicName id="4C494D0CFF96E264FC87830FFC32F9DA" authorityName="Nevill" authorityYear="1877" box="[826,937,1573,1594]" class="Gastropoda" family="Viviparidae" genus="Margarya" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="782" phylum="Mollusca" rank="genus">
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<emphasis id="B93DEA9DFF96E264FC87830FFC32F9DA" box="[826,937,1573,1594]" italics="true" pageId="22" pageNumber="782">Margarya</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C494D0CFF96E264FC05830FFBE3F9DA" authorityName="He" authorityYear="2013" box="[952,1144,1573,1594]" class="Gastropoda" family="Viviparidae" genus="Tchangmargarya" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="782" phylum="Mollusca" rank="genus">
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<emphasis id="B93DEA9DFF96E264FC05830FFBE3F9DA" box="[952,1144,1573,1594]" italics="true" pageId="22" pageNumber="782">Tchangmargarya</emphasis>
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</taxonomicName>
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, for
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<taxonomicName id="4C494D0CFF96E264FB10830FFA3AF9DA" authorityName="Tchang & Tsi" authorityYear="1949" box="[1197,1441,1573,1594]" class="Gastropoda" family="Viviparidae" genus="Margarya" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="782" phylum="Mollusca" rank="species" species="yangtsunghaiensis">
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<emphasis id="B93DEA9DFF96E264FB10830FFA3AF9DA" box="[1197,1441,1573,1594]" italics="true" pageId="22" pageNumber="782">M. yangtsunghaiensis</emphasis>
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</taxonomicName>
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as the
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<typeStatus id="54F2882DFF96E264FC34836EFC21F9B9" box="[905,954,1604,1625]" pageId="22" pageNumber="782">type</typeStatus>
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species. Herein we elevated the subgenus
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<taxonomicName id="4C494D0CFF96E264FC878348FC61F997" authorityName="He" authorityYear="2013" box="[826,1018,1634,1655]" class="Gastropoda" family="Viviparidae" genus="Tchangmargarya" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="782" phylum="Mollusca" rank="genus">
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<emphasis id="B93DEA9DFF96E264FC878348FC61F997" box="[826,1018,1634,1655]" italics="true" pageId="22" pageNumber="782">Tchangmargarya</emphasis>
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</taxonomicName>
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to a full genus for phylogeny, and for extensive differences of shell, operculum, and anatomy, compared with
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<taxonomicName id="4C494D0CFF96E264FC4D838AFBFAF955" authorityName="Nevill" authorityYear="1877" box="[1008,1121,1696,1717]" class="Gastropoda" family="Viviparidae" genus="Margarya" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="782" phylum="Mollusca" rank="genus">
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<emphasis id="B93DEA9DFF96E264FC4D838AFBFAF955" box="[1008,1121,1696,1717]" italics="true" pageId="22" pageNumber="782">Margarya</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BF6368FFF96E264FC8783F5FC20F914" blockId="22.[826,1441,1759,1903]" box="[826,955,1759,1780]" pageId="22" pageNumber="782">
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||
<emphasis id="B93DEA9DFF96E264FC8783F5FC20F914" box="[826,955,1759,1780]" italics="true" pageId="22" pageNumber="782">Description</emphasis>
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||
</paragraph>
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||
<paragraph id="8BF6368FFF96E265FC8783D7FD6CFE31" blockId="22.[826,1441,1759,1903]" lastBlockId="23.[144,759,198,1015]" lastPageId="23" lastPageNumber="783" pageId="22" pageNumber="782">
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Shell large, up to nearly
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<quantity id="4CB19B6AFF96E264FBE083D7FB37F8F2" box="[1117,1196,1789,1810]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="782" unit="mm" value="70.0">70 mm</quantity>
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in height; very thick and solid; mostly with seven whorls, including two protoconch whorls, well-developed subsutural ramp or depression at upper portion of each whorl, except for the protoconch; whorls generally with between three and five prominent spiral keels, and enormous obvious axial ribs on each whorls, crossing into regular and elevated nodules, nodules small and dense on upper whorls, gradually becoming large and sparse on lower whorls, lower keel always covered by upper suture, and thus nearly invisible on the spiral whorls; two incomplete spiral ribs with elevated nodules extending onto umbilical area; aperture ovate; umbilicus small or absent.
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</paragraph>
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<paragraph id="8BF6368FFF97E265FF1584F3FE63FD68" blockId="23.[144,759,198,1015]" pageId="23" pageNumber="783">Operculum corneous, ovate, reddish to yellowish brown, with subcentral nucleus and concentric growth lines; nucleus with few grains or veins; opercular scar large, relatively rough, outside smooth, obviously thickened, but the margin of operculum thin; operculum slightly smaller than aperture.</paragraph>
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<paragraph id="8BF6368FFF97E265FF1587BBFE71FCE2" blockId="23.[144,759,198,1015]" pageId="23" pageNumber="783">Radula with about 100–140 rows of teeth, each with one central tooth, one lateral tooth, and two marginal teeth on either side of it; outer marginal teeth with between ten and 16 denticles.</paragraph>
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<paragraph id="8BF6368FFF97E265FF158621FB35FF1A" blockId="23.[144,759,198,1015]" lastBlockId="23.[806,1421,198,250]" pageId="23" pageNumber="783">
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Body and mantle colour of living animals transparent white with dense yellow or orange pigment spots above (
|
||
<figureCitation id="13722A0AFF97E265FF5E8662FEACFCBE" box="[227,311,840,862]" captionStart="Figure 7" captionStartId="10.[164,242,1502,1521]" captionTargetBox="[164,1445,194,1465]" captionTargetId="figure-150@10.[164,1445,194,1472]" captionTargetPageId="10" captionText="Figure 7. Skin colour and morpholgy of cephalic tentacles: A, Margarya melanioides; B, Margarya francheti; C, Margarya oxytropoides; D, Margarya monodi; E, Tchangmargarya yangtsunghaiensis (offered by Jin Chen); F, Anularya bicostata; G, head of male Margarya species; H, head of male Tchangmargarya species; I, head of male Anularya species. Abbreviations: ey, eye; lt, left tentacle; pb, proboscis; pn, penis (right tentacle)." figureDoi="http://doi.org/10.5281/zenodo.10107702" httpUri="https://zenodo.org/record/10107702/files/figure.png" pageId="23" pageNumber="783">Fig. 7E</figureCitation>
|
||
); foot comparatively small, with large opercular lobe, two cephalic tentacles of equal length, about 4.5–6.0 mm long, right tentacle thickened in males, modified into a penis (
|
||
<figureCitation id="13722A0AFF97E265FE5A868EFDA4FC5A" box="[487,575,932,954]" captionStart="Figure 7" captionStartId="10.[164,242,1502,1521]" captionTargetBox="[164,1445,194,1465]" captionTargetId="figure-150@10.[164,1445,194,1472]" captionTargetPageId="10" captionText="Figure 7. Skin colour and morpholgy of cephalic tentacles: A, Margarya melanioides; B, Margarya francheti; C, Margarya oxytropoides; D, Margarya monodi; E, Tchangmargarya yangtsunghaiensis (offered by Jin Chen); F, Anularya bicostata; G, head of male Margarya species; H, head of male Tchangmargarya species; I, head of male Anularya species. Abbreviations: ey, eye; lt, left tentacle; pb, proboscis; pn, penis (right tentacle)." figureDoi="http://doi.org/10.5281/zenodo.10107702" httpUri="https://zenodo.org/record/10107702/files/figure.png" pageId="23" pageNumber="783">Fig. 7H</figureCitation>
|
||
). Gonochoristic, prostate in male reddish yellow, about
|
||
<quantity id="4CB19B6AFF97E265FD8C86E9FDE7FC38" box="[561,636,963,984]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="783" unit="mm" value="20.0">20 mm</quantity>
|
||
long, testis auratus about
|
||
<quantity id="4CB19B6AFF97E265FE8186CBFE11FC17" box="[316,394,993,1015]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="783" unit="mm" value="25.0">25 mm</quantity>
|
||
long. Gravid females retaining one or two developing juveniles in uterus, with size at birth mostly exceeding
|
||
<quantity id="4CB19B6AFF97E265FBE485CEFB31FF1A" box="[1113,1194,228,250]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="783" unit="mm" value="10.0">10 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF6368FFF97E265FC9B8408FB86FED6" blockId="23.[806,1422,289,801]" box="[806,1053,289,311]" pageId="23" pageNumber="783">
|
||
<emphasis id="B93DEA9DFF97E265FC9B8408FB86FED6" box="[806,1053,289,311]" italics="true" pageId="23" pageNumber="783">Comparative remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF6368FFF97E265FC9B846AFB5BFCC0" blockId="23.[806,1422,289,801]" pageId="23" pageNumber="783">
|
||
<taxonomicName id="4C494D0CFF97E265FC9B846AFC61FEB5" authorityName="He" authorityYear="2013" box="[806,1018,320,341]" class="Gastropoda" family="Viviparidae" genus="Tchangmargarya" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="783" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93DEA9DFF97E265FC9B846AFC61FEB5" box="[806,1018,320,341]" italics="true" pageId="23" pageNumber="783">Tchangmargarya</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C494D0CFF97E265FB71846BFA16FEB6" authority="and" authorityName="AND" box="[1228,1421,320,342]" class="Gastropoda" family="Viviparidae" genus="Margarya" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="785" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93DEA9DFF97E265FB71846BFADDFEB6" box="[1228,1350,321,342]" italics="true" pageId="23" pageNumber="783">Margarya</emphasis>
|
||
and
|
||
</taxonomicName>
|
||
<taxonomicName id="4C494D0CFF97E265FC9B8475FC0AFE94" authority="Zhang & Chen & Yang & Jin & Köhler, 2015" authorityName="Zhang & Chen & Yang & Jin & Köhler" authorityYear="2015" box="[806,913,351,372]" class="Gastropoda" family="Viviparidae" genus="Anularya" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="783" phylum="Mollusca" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93DEA9DFF97E265FC9B8475FC0AFE94" box="[806,913,351,372]" italics="true" pageId="23" pageNumber="783">Anularya</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20E57E6FF97E265FC258475FC67FE94" box="[920,1020,351,372]" pageId="23" pageNumber="783" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
by two incomplete spiral ribs with elevated nodules in the umbilical area, an operculum smaller than the aperture, with an almost smooth nucleus, and with few grains or veins only, thickened outside of the opercular scar but with a thin margin. Extant
|
||
<emphasis id="B93DEA9DFF97E265FCC784D2FC79FDED" box="[890,994,504,525]" italics="true" pageId="23" pageNumber="783">Tulotoma</emphasis>
|
||
Haldeman, 1840 endemic to
|
||
<collectingRegion id="498DF86DFF97E265FA9984D2FA11FDEE" box="[1316,1418,504,526]" country="United States of America" name="Alabama" pageId="23" pageNumber="783">Alabama</collectingRegion>
|
||
,
|
||
<collectingCountry id="F35E761FFF97E265FC9B873CFCC6FDCB" box="[806,861,534,555]" name="United States of America" pageId="23" pageNumber="783">USA</collectingCountry>
|
||
, has nodules similar to
|
||
<taxonomicName id="4C494D0CFF97E265FBDE873CFA85FDCB" authorityName="He" authorityYear="2013" box="[1123,1310,534,555]" class="Gastropoda" family="Viviparidae" genus="Tchangmargarya" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="783" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93DEA9DFF97E265FBDE873CFA85FDCB" box="[1123,1310,534,555]" italics="true" pageId="23" pageNumber="783">Tchangmargarya</emphasis>
|
||
</taxonomicName>
|
||
; however, its shell is smaller, having no plate area on each whorl or two keels with nodules on the base of the body whorl, and an operculum that is hemicycle in shape. As for anatomical characters,
|
||
<taxonomicName id="4C494D0CFF97E265FBFE87BBFA90FD46" authorityName="He" authorityYear="2013" box="[1091,1291,657,678]" class="Gastropoda" family="Viviparidae" genus="Tchangmargarya" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="783" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93DEA9DFF97E265FBFE87BBFA90FD46" box="[1091,1291,657,678]" italics="true" pageId="23" pageNumber="783">Tchangmargarya</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="4C494D0CFF97E265FC9B879AFC0CFD25" box="[806,919,688,709]" class="Gastropoda" family="Viviparidae" genus="Margarya" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="783" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93DEA9DFF97E265FC9B879AFC0CFD25" box="[806,919,688,709]" italics="true" pageId="23" pageNumber="783">Margarya</emphasis>
|
||
</taxonomicName>
|
||
in having a small foot but large opercular lobe and similar colour pattern of body. It differs by its reddish yellow prostate and much shorter cephalic tentacles in males (
|
||
<figureCitation id="13722A0AFF97E265FBE68621FB2FFCC0" box="[1115,1204,779,801]" captionStart="Figure 7" captionStartId="10.[164,242,1502,1521]" captionTargetBox="[164,1445,194,1465]" captionTargetId="figure-150@10.[164,1445,194,1472]" captionTargetPageId="10" captionText="Figure 7. Skin colour and morpholgy of cephalic tentacles: A, Margarya melanioides; B, Margarya francheti; C, Margarya oxytropoides; D, Margarya monodi; E, Tchangmargarya yangtsunghaiensis (offered by Jin Chen); F, Anularya bicostata; G, head of male Margarya species; H, head of male Tchangmargarya species; I, head of male Anularya species. Abbreviations: ey, eye; lt, left tentacle; pb, proboscis; pn, penis (right tentacle)." figureDoi="http://doi.org/10.5281/zenodo.10107702" httpUri="https://zenodo.org/record/10107702/files/figure.png" pageId="23" pageNumber="783">Fig. 7H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3536504FF97E265FCD1867CFADCFC8A" box="[876,1351,854,875]" pageId="23" pageNumber="783" type="description">
|
||
<paragraph id="8BF6368FFF97E265FCD1867CFADCFC8A" blockId="23.[876,1351,854,908]" box="[876,1351,854,875]" pageId="23" pageNumber="783">
|
||
<taxonomicName id="4C494D0CFF97E265FCD1867CFADCFC8A" box="[876,1351,854,875]" pageId="23" pageNumber="783">
|
||
<emphasis id="B93DEA9DFF97E265FCD1867CFADCFC8A" bold="true" box="[876,1351,854,875]" italics="true" pageId="23" pageNumber="783">
|
||
<emphasis id="B93DEA9DFF97E265FCD1867CFBD2FC8A" bold="true" box="[876,1097,854,875]" italics="true" pageId="23" pageNumber="783">TCHANGMARGARYA</emphasis>
|
||
YANGTSUNGHAIENSIS
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |