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<mods:title id="7903AC0F8DFF74B917D6B5CE6614EF85">New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae)</mods:title>
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<mods:namePart id="4BCD89F139E97C55AA5C20AD6E0CE966">Brown, John W.</mods:namePart>
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<mods:date id="6CED8E2D03F3DC34C233A0E613E5F200">2019</mods:date>
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<treatment id="03E08794FFC2FFF8FF5B78ADFB0CFC29" ID-GBIF-Taxon="159608299" LSID="urn:lsid:plazi:treatment:03E08794FFC2FFF8FF5B78ADFB0CFC29" httpUri="http://treatment.plazi.org/id/03E08794FFC2FFF8FF5B78ADFB0CFC29" lastPageId="4" lastPageNumber="199" pageId="3" pageNumber="198">
<subSubSection id="C3536509FFC2FFFFFF5B78ADFE10FB0E" box="[151,497,1197,1224]" pageId="3" pageNumber="198" type="nomenclature">
<paragraph id="8BF63682FFC2FFFFFF5B78ADFE10FB0E" blockId="3.[151,497,1197,1224]" box="[151,497,1197,1224]" pageId="3" pageNumber="198">
<heading id="D0BE81EEFFC2FFFFFF5B78ADFE10FB0E" bold="true" box="[151,497,1197,1224]" fontSize="11" level="1" pageId="3" pageNumber="198" reason="1">
<emphasis id="B93DEA90FFC2FFFFFF5B78ADFE10FB0E" bold="true" box="[151,497,1197,1224]" pageId="3" pageNumber="198">
<taxonomicName id="4C494D01FFC2FFFFFF5B78ADFE83FB0E" ID-CoL="93NCJ" authority="Pogue" authorityName="Brown" authorityYear="2019" box="[151,354,1197,1224]" class="Insecta" family="Tortricidae" genus="Atroposia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93DEA90FFC2FFFFFF5B78ADFEEDFB01" bold="true" box="[151,268,1197,1223]" italics="true" pageId="3" pageNumber="198">Atroposia</emphasis>
Pogue
</taxonomicName>
,
<taxonomicNameLabel id="A20E57EBFFC2FFFFFEA378AEFE10FB0E" box="[367,497,1198,1224]" pageId="3" pageNumber="198" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3536509FFC2FFFFFF5B78F2FCE1FA81" pageId="3" pageNumber="198" type="materials_examined">
<paragraph id="8BF63682FFC2FFFFFF5B78F2FCC7FACC" blockId="3.[151,806,1265,1290]" box="[151,806,1265,1290]" pageId="3" pageNumber="198">
<typeStatus id="54F28820FFC2FFFFFF5B78F2FF28FACC" box="[151,201,1266,1290]" pageId="3" pageNumber="198">Type</typeStatus>
species:
<taxonomicName id="4C494D01FFC2FFFFFEEA78F2FD6EFACC" authority="Riley, 1881" authorityName="Riley" authorityYear="1881" box="[294,655,1265,1290]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="oenotherana">
<emphasis id="B93DEA90FFC2FFFFFEEA78F2FDF3FACC" box="[294,530,1266,1290]" italics="true" pageId="3" pageNumber="198">Conchylis oenotherana</emphasis>
<bibRefCitation id="EFD84B73FFC2FFFFFDD478F1FD6EFACC" author="Riley, C. V." box="[536,655,1265,1290]" pageId="3" pageNumber="198" pagination="316 - 324" refId="ref21341" refString="Riley, C. V. (1881) Descriptions of some new Tortricidae. Transactions of the St. Louis Academy of Sciences, 4, 316 - 324." type="journal article" year="1881">Riley, 1881</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
<paragraph id="8BF63682FFC2FFFFFF5B792FFCE1FA81" blockId="3.[151,1129,1326,1382]" box="[151,768,1326,1351]" pageId="3" pageNumber="198">
<taxonomicName id="4C494D01FFC2FFFFFF5B792FFEA1FA81" authority="Pogue" authorityName="Brown" authorityYear="2019" box="[151,320,1327,1351]" class="Insecta" family="Tortricidae" genus="Atroposia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93DEA90FFC2FFFFFF5B792FFF1BFA80" box="[151,250,1327,1350]" italics="true" pageId="3" pageNumber="198">Atroposia</emphasis>
Pogue
</taxonomicName>
and Mickevich, 1990: 322 (nomen nudum).
</paragraph>
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<subSubSection id="C3536509FFC2FFFFFF5B794DFCFCF941" pageId="3" pageNumber="198" type="reference_group">
<paragraph id="8BF63682FFC2FFFFFF5B794DFB88FAA0" blockId="3.[151,1129,1326,1382]" box="[151,1129,1357,1382]" pageId="3" pageNumber="198">
<treatmentCitationGroup id="AB5911ACFFC2FFFFFF5B794DFB88FAA0" box="[151,1129,1357,1382]" pageId="3" pageNumber="198">
<taxonomicName id="4C494D01FFC2FFFFFF5B794DFF1AFAA0" ID-CoL="93N3S" authorityName="Guenée" authorityYear="1845" box="[151,251,1357,1382]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="tribe" tribe="Cochylini">Cochylini</taxonomicName>
New Genus 1 [
<taxonomicName id="4C494D01FFC2FFFFFE56794EFDFBFAA0" authorityName="Riley" authorityYear="1881" box="[410,538,1358,1382]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="oenotherana">
<emphasis id="B93DEA90FFC2FFFFFE56794EFDFBFAA0" box="[410,538,1358,1382]" italics="true" pageId="3" pageNumber="198">oenotherana</emphasis>
</taxonomicName>
group]:
<treatmentCitation id="0AE81093FFC2FFFFFDBD794EFCC7FAA0" author="Brown, J. W." box="[625,806,1357,1382]" page="207" pageId="3" pageNumber="198" year="2005">
<bibRefCitation id="EFD84B73FFC2FFFFFDBD794EFCC7FAA0" author="Brown, J. W." box="[625,806,1357,1382]" pageId="3" pageNumber="198" refId="ref19539" refString="Brown, J. W. (2005) World catalogue of insects. Vol. 5. Tortricidae (Lepidoptera). Apollo Books, Stenstrup, 741 pp." type="book" year="2005">Brown 2005: 207</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE81093FFC2FFFFFCFC794DFB85FAA0" author="Metzler, E. H. &amp; Brown, J. W." box="[816,1124,1357,1382]" page="278" pageId="3" pageNumber="198" year="2014">
<bibRefCitation id="EFD84B73FFC2FFFFFCFC794DFB85FAA0" author="Metzler, E. H. &amp; Brown, J. W." box="[816,1124,1357,1382]" pageId="3" pageNumber="198" pagination="274 - 282" refId="ref20205" refString="Metzler, E. H. &amp; Brown, J. W. (2014) An updated check list the Cochylina (Tortricidae, Tortricinae, Euliini) of North America north of Mexico including Greenland, with comments on classification and identification. Journal of the Lepidopterists' Society, 68, 274 - 282. https: // doi. org / 10.18473 / lepi. v 68 i 4. a 5" type="journal article" year="2014">Metzler and Brown 2014: 278</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF63682FFC2FFFFFF5B7994FCFCF941" blockId="3.[151,1437,1428,2031]" pageId="3" pageNumber="198">
<emphasis id="B93DEA90FFC2FFFFFF5B7994FEE5FA68" box="[151,260,1428,1454]" italics="true" pageId="3" pageNumber="198">Atroposia</emphasis>
is proposed to accommodate the single species
<taxonomicName id="4C494D01FFC2FFFFFCEA7994FC55FA68" authorityName="Riley" authorityYear="1881" box="[806,948,1428,1454]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="oenotherana">
<emphasis id="B93DEA90FFC2FFFFFCEA7994FC55FA68" box="[806,948,1428,1454]" italics="true" pageId="3" pageNumber="198">oenotherana</emphasis>
</taxonomicName>
. Originally described in
<taxonomicName id="4C494D01FFC2FFFFFB037994FEE2FA14" authority="Sodoffsky, 1837" authorityName="Sodoffsky" authorityYear="1837" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC2FFFFFB037994FADEFA68" box="[1231,1343,1428,1454]" italics="true" pageId="3" pageNumber="198">Conchylis</emphasis>
Sodoffsky, 1837
</taxonomicName>
(an unjustified emendation of
<taxonomicName id="4C494D01FFC2FFFFFDAC79B8FD23FA14" ID-CoL="925PD" authorityName="Treitschke" authorityYear="1829" box="[608,706,1464,1490]" class="Insecta" family="Tortricidae" genus="Cochylis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC2FFFFFDAC79B8FD23FA14" box="[608,706,1464,1490]" italics="true" pageId="3" pageNumber="198">Cochylis</emphasis>
</taxonomicName>
), the species was assigned to
<taxonomicName id="4C494D01FFC2FFFFFBDB79B8FB3AFA14" authority="Hubner" authorityName="Hubner" authorityYear="1825" box="[1047,1243,1464,1490]" class="Insecta" family="Tortricidae" genus="Phalonia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC2FFFFFBDB79B8FB9FFA14" box="[1047,1150,1464,1490]" italics="true" pageId="3" pageNumber="198">Phalonia</emphasis>
Hübner
</taxonomicName>
, [1825], 1816 by
<bibRefCitation id="EFD84B73FFC2FFFFFF5B79DCFE65FA30" author="McDunnough, J." box="[151,388,1500,1526]" pageId="3" pageNumber="198" refId="ref20127" refString="McDunnough, J. (1939) Check List of the Lepidoptera of Canada and the United States of America. II. Microlepidoptera. Memoirs of the Southern California Academy of Sciences, 2, 171 pp." type="book" year="1939">McDunnough (1939)</bibRefCitation>
and placed in “incertae sedis” by
<bibRefCitation id="EFD84B73FFC2FFFFFCCF79DCFC47FA30" author="Powell, J. A." box="[771,934,1500,1526]" pageId="3" pageNumber="198" pagination="31 - 42" refId="ref20538" refString="Powell, J. A. (1983) Tortricoidea. In: Hodges, R. W. (Ed.), Check list of the Lepidoptera of America north of Mexico. E. W. Classey, Ltd. and Wedge Entomological Research Foundation, London, pp. 31 - 42." type="book chapter" year="1983">Powell (1983)</bibRefCitation>
.
<bibRefCitation id="EFD84B73FFC2FFFFFC7C79DDFBA9FA31" author="Pogue, M. G." box="[944,1096,1500,1527]" pageId="3" pageNumber="198" refId="ref20442" refString="Pogue, M. G. (1986) A generic revision of the Cochylidae (Lepidoptera) of North America. Ph. D. dissertation, University of Minnesota, Minneapolis, Minnesota, 280 pp." type="book" year="1986">Pogue (1986)</bibRefCitation>
proposed the genus
<emphasis id="B93DEA90FFC2FFFFFAE379DCFA7DFA30" box="[1327,1436,1500,1526]" italics="true" pageId="3" pageNumber="198">Atroposia</emphasis>
for the species in his unpublished Ph.D. dissertation;
<bibRefCitation id="EFD84B73FFC2FFFFFD3A7A00FC56F9DC" author="Razowski, J." box="[758,951,1536,1562]" pageId="3" pageNumber="198" pagination="107 - 163" refId="ref21015" refString="Razowski, J. (1997) Cochylini (Lepidoptera: Tortricidae) of Canada. Acta Zoologica Cracoviensia, 40, 107 - 163." type="journal article" year="1997">Razowski (1997)</bibRefCitation>
treated
<taxonomicName id="4C494D01FFC2FFFFFBD87A00FB43F9DC" authorityName="Riley" authorityYear="1881" box="[1044,1186,1536,1562]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="oenotherana">
<emphasis id="B93DEA90FFC2FFFFFBD87A00FB43F9DC" box="[1044,1186,1536,1562]" italics="true" pageId="3" pageNumber="198">oenotherana</emphasis>
</taxonomicName>
under “Unplaced
<taxonomicName id="4C494D01FFC2FFFFFABE7A00FF0AF9F8" ID-CoL="93N3S" authorityName="Guenée" authorityYear="1845" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="tribe" tribe="Cochylini">Cochylini</taxonomicName>
”; and
<bibRefCitation id="EFD84B73FFC2FFFFFEE27A25FE2AF9F8" author="Brown, J. W." box="[302,459,1571,1599]" pageId="3" pageNumber="198" refId="ref19539" refString="Brown, J. W. (2005) World catalogue of insects. Vol. 5. Tortricidae (Lepidoptera). Apollo Books, Stenstrup, 741 pp." type="book" year="2005">Brown (2005)</bibRefCitation>
and
<bibRefCitation id="EFD84B73FFC2FFFFFDCC7A24FCF7F9F8" author="Metzler, E. H. &amp; Brown, J. W." box="[512,790,1572,1599]" pageId="3" pageNumber="198" pagination="274 - 282" refId="ref20205" refString="Metzler, E. H. &amp; Brown, J. W. (2014) An updated check list the Cochylina (Tortricidae, Tortricinae, Euliini) of North America north of Mexico including Greenland, with comments on classification and identification. Journal of the Lepidopterists' Society, 68, 274 - 282. https: // doi. org / 10.18473 / lepi. v 68 i 4. a 5" type="journal article" year="2014">Metzler &amp; Brown (2014)</bibRefCitation>
treated it as “
<taxonomicName id="4C494D01FFC2FFFFFC607A24FBFCF9F8" ID-CoL="93N3S" authorityName="Guenée" authorityYear="1845" box="[940,1053,1572,1598]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="tribe" tribe="Cochylini">Cochylini</taxonomicName>
New Genus”. According to
<bibRefCitation id="EFD84B73FFC2FFFFFA9A7A25FF05F9A5" author="Pogue, M. G." pageId="3" pageNumber="198" refId="ref20442" refString="Pogue, M. G. (1986) A generic revision of the Cochylidae (Lepidoptera) of North America. Ph. D. dissertation, University of Minnesota, Minneapolis, Minnesota, 280 pp." type="book" year="1986">Pogue (1986)</bibRefCitation>
,
<emphasis id="B93DEA90FFC2FFFFFF3D7A48FEBFF9A4" box="[241,350,1608,1634]" italics="true" pageId="3" pageNumber="198">Atroposia</emphasis>
is most closely related to
<taxonomicName id="4C494D01FFC2FFFFFD447A48FCE9F9A4" ID-CoL="92N4C" authorityName="Razowski" authorityYear="1967" box="[648,776,1608,1634]" class="Insecta" family="Tortricidae" genus="Spinipogon" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC2FFFFFD447A48FCE9F9A4" box="[648,776,1608,1634]" italics="true" pageId="3" pageNumber="198">Spinipogon</emphasis>
</taxonomicName>
among North American genera. However, based on facies and genitalia,
<emphasis id="B93DEA90FFC2FFFFFEF97A6CFE43F940" box="[309,418,1644,1670]" italics="true" pageId="3" pageNumber="198">Atroposia</emphasis>
appears to be related to
<taxonomicName id="4C494D01FFC2FFFFFD787A6CFCF7F940" ID-CoL="925PD" authorityName="Treitschke" authorityYear="1829" box="[692,790,1644,1670]" class="Insecta" family="Tortricidae" genus="Cochylis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC2FFFFFD787A6CFCF7F940" box="[692,790,1644,1670]" italics="true" pageId="3" pageNumber="198">Cochylis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536509FFC2FFFFFF0B7A8FFC4DF860" pageId="3" pageNumber="198" type="diagnosis">
<paragraph id="8BF63682FFC2FFFFFF0B7A8FFC4DF860" blockId="3.[151,1437,1428,2031]" pageId="3" pageNumber="198">
<emphasis id="B93DEA90FFC2FFFFFF0B7A8FFEDBF96C" bold="true" box="[199,314,1679,1706]" pageId="3" pageNumber="198">Diagnosis</emphasis>
.
<emphasis id="B93DEA90FFC2FFFFFE847A90FE54F96C" box="[328,437,1680,1706]" italics="true" pageId="3" pageNumber="198">Atroposia</emphasis>
is easily distinguished from other genera by facies (
<figureCitation id="13722A07FFC2FFFFFC357A90FB85F96C" box="[1017,1124,1680,1706]" captionStart="FIGURES 110" captionStartId="5.[151,264,1832,1857]" captionTargetBox="[194,1394,166,1812]" captionTargetId="figure@5.[192,1396,166,1812]" captionTargetPageId="5" captionText="FIGURES 110. Adults of Cochylini genera. 1. Anacochylidia maderana, Madera Canyon, Santa Rita Mountains, Arizona (EME). 2. Cagiva cephalanthana, Dorman Lake, Oktibbeha Co., Mississippi (USNM). 3. Atroposia oenotherana, Houston, Harris Co., Texas (USNM). 4. Atroposia oenotherana, Radisson, Onondaga Co., New York (USNM). 5. Cybilla hubbardana, Shafter, Presidio Co., Texas (USNM). 6. Cybilla hubbardana, Lordsburg, Hidalgo Co., New Mexico (CNC). 7. Honca grandis, Chimney Gulch, Golden, Colorado (USNM). 8. Pogospinia floridana, Long Key State Park, Monroe Co., Florida (USNM). 9. Nycthia pimama, Portal, Cochise Co., Arizona (USNM). 10. Nycthia yuccatana, Falcon State Park, Starr Co., Texas (USNM)." figureDoi="http://doi.org/10.5281/zenodo.3442348" httpUri="https://zenodo.org/record/3442348/files/figure.png" pageId="3" pageNumber="198">Figs. 3, 4</figureCitation>
), which feature a somewhat two-toned forewing with a yellow-tan basal half and a pinkish red distal half with a variably developed yellowish spot. However, in some specimens both the basal and distal halves of the wing are overscaled with dark gray, reducing the two-toned pattern. Putative autapomorphies of
<emphasis id="B93DEA90FFC2FFFFFCE57AFCFC77F8D0" box="[809,918,1788,1814]" italics="true" pageId="3" pageNumber="198">Atroposia</emphasis>
include the unusual shape of the valva with a distinctly pointed costa and sacculus; the distal portion of the phallus, with a pair of long, pointed, flattened, lateral sheaths and three platelike internal sclerites; and the broad sterigma composed of four fused spheroid plates. The absence of the male hindwing costal roll in
<taxonomicName id="4C494D01FFC2FFFFFDA87B68FCF0F844" authorityName="Riley" authorityYear="1881" box="[612,785,1896,1922]" class="Insecta" family="Tortricidae" genus="Conchylis" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="oenotherana">
<emphasis id="B93DEA90FFC2FFFFFDA87B68FCF0F844" box="[612,785,1896,1922]" italics="true" pageId="3" pageNumber="198">A. oenotherana</emphasis>
</taxonomicName>
distinguishes it from most species of
<taxonomicName id="4C494D01FFC2FFFFFB747B68FAFBF844" authorityName="Treitschke" authorityYear="1829" box="[1208,1306,1896,1922]" class="Insecta" family="Tortricidae" genus="Cochylis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC2FFFFFB747B68FAFBF844" box="[1208,1306,1896,1922]" italics="true" pageId="3" pageNumber="198">Cochylis</emphasis>
</taxonomicName>
and related genera (e.g.,
<emphasis id="B93DEA90FFC2FFFFFEE57B8CFE21F860" box="[297,448,1932,1958]" italics="true" pageId="3" pageNumber="198">Cochylichroa</emphasis>
,
<taxonomicName id="4C494D01FFC2FFFFFE027B8CFDA8F860" authorityName="Obraztsov" authorityYear="1956" box="[462,585,1932,1958]" class="Insecta" family="Tortricidae" genus="Cochylidia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC2FFFFFE027B8CFDA8F860" box="[462,585,1932,1958]" italics="true" pageId="3" pageNumber="198">Cochylidia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C494D01FFC2FFFFFD9B7B8CFC7EF860" authority="Walsingham, 1897" authorityName="Walsingham" authorityYear="1897" box="[599,927,1932,1958]" class="Insecta" family="Tortricidae" genus="Thyraylia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEA90FFC2FFFFFD9B7B8CFD25F860" box="[599,708,1932,1958]" italics="true" pageId="3" pageNumber="198">Thyraylia</emphasis>
Walsingham, 1897
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3536509FFC2FFF8FF0B7BAFFC9CFC61" lastPageId="4" lastPageNumber="199" pageId="3" pageNumber="198" type="description">
<paragraph id="8BF63682FFC2FFF8FF0B7BAFFC9CFC61" blockId="3.[151,1437,1428,2031]" lastBlockId="4.[151,1437,153,1008]" lastPageId="4" lastPageNumber="199" pageId="3" pageNumber="198">
<emphasis id="B93DEA90FFC2FFFFFF0B7BAFFEB0F80C" bold="true" box="[199,337,1967,1994]" pageId="3" pageNumber="198">Description</emphasis>
. Head: Vertex and upper frons rough scaled, lower frons with appressed scales; ocellus present; sensory setae of antenna 1.01.2 times flagellomere diameter in male, shorter in female; labial palpus porrect, com- bined length of all segments 1.61.8 times diameter of compound eye, scaling of segment II concealing segment III; maxillary palpus 1-segmented. Thorax: Posterior crest present; lateral scale tufts of metanotum flat. Forewing in male slightly broadened distally with costa weakly convex and termen straight, moderately oblique, forewing not expanding distally in female, apex rounded in both; forewing length
<quantity id="4CB19B67FFC5FFF8FC1C7D04FBBBFED9" box="[976,1114,260,287]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.4" metricValueMin="3.8" pageId="4" pageNumber="199" unit="mm" value="4.6" valueMax="5.4" valueMin="3.8">3.85.4 mm</quantity>
, length 2.53.4 times width, slightly wider in female; without costal fold in male; Sc less than 0.5 wing length; R
<subScript id="17CD34C7FFC5FFF8FBBF7D3AFB9AFE8E" attach="left" box="[1139,1147,314,328]" fontSize="6" pageId="4" pageNumber="199">1</subScript>
originating at or beyond middle of discal cell; R
<subScript id="17CD34C7FFC5FFF8FE517D5EFE44FEAA" attach="left" box="[413,421,350,364]" fontSize="6" pageId="4" pageNumber="199">2</subScript>
originating nearer R
<subScript id="17CD34C7FFC5FFF8FD427D5EFD77FEAA" attach="left" box="[654,662,350,364]" fontSize="6" pageId="4" pageNumber="199">3</subScript>
than R
<subScript id="17CD34C7FFC5FFF8FD2B7D5EFD0EFEAA" attach="left" box="[743,751,350,364]" fontSize="6" pageId="4" pageNumber="199">1</subScript>
; R
<subScript id="17CD34C7FFC5FFF8FCDD7D5EFCF8FEAA" attach="left" box="[785,793,350,364]" fontSize="6" pageId="4" pageNumber="199">5</subScript>
ending at costa; M
<subScript id="17CD34C7FFC5FFF8FC3C7D5EFC19FEAA" attach="left" box="[1008,1016,350,364]" fontSize="6" pageId="4" pageNumber="199">3</subScript>
and CuA
<subScript id="17CD34C7FFC5FFF8FBAF7D5EFB8AFEAA" attach="left" box="[1123,1131,350,364]" fontSize="6" pageId="4" pageNumber="199">1</subScript>
separate; CuA
<subScript id="17CD34C7FFC5FFF8FADE7D5EFAFBFEAA" attach="left" box="[1298,1306,350,364]" fontSize="6" pageId="4" pageNumber="199">2</subScript>
originating about 0.75 length of discal cell; CuP absent; A1+2 stalked at 0.5 length. Hindwing length 2.52.8 times width; costa straight, without costal roll; apex rounded; termen straight, oblique; Sc+R
<subScript id="17CD34C7FFC5FFF8FC1D7DA6FC38FE72" attach="left" box="[977,985,422,436]" fontSize="6" pageId="4" pageNumber="199">1</subScript>
ca. 0.75 wing length, Rs and M
<subScript id="17CD34C7FFC5FFF8FAF27DA6FAA7FE72" attach="left" box="[1342,1350,422,436]" fontSize="6" pageId="4" pageNumber="199">1</subScript>
stalked ca. 0.65 length of M
<subScript id="17CD34C7FFC5FFF8FEB27DCAFE67FE1E" attach="left" box="[382,390,458,472]" fontSize="6" pageId="4" pageNumber="199">1</subScript>
; M
<subScript id="17CD34C7FFC5FFF8FE627DCAFE57FE1E" attach="left" box="[430,438,458,472]" fontSize="6" pageId="4" pageNumber="199">3</subScript>
and CuA
<subScript id="17CD34C7FFC5FFF8FDEF7DCAFDCAFE1E" attach="left" box="[547,555,458,472]" fontSize="6" pageId="4" pageNumber="199">1</subScript>
separate; CuA
<subScript id="17CD34C7FFC5FFF8FD187DCAFD3DFE1E" attach="left" box="[724,732,458,472]" fontSize="6" pageId="4" pageNumber="199">2</subScript>
originating 0.65 length of discal cell; male frenulum with one acanthus, female with two. Abdomen: Unmodified. Male genitalia (
<figureCitation id="13722A07FFC5FFF8FC547DDDFC0AFE31" box="[920,1003,477,503]" captionStart="FIGURES 1926" captionStartId="17.[151,264,1861,1886]" captionTargetBox="[151,1436,174,1805]" captionTargetId="figure@17.[151,1436,174,1805]" captionTargetPageId="17" captionText="FIGURES 1926. Male genitalia of Cochylini genera. 19. Anacochylidia maderana, Palmerlee, Arizona, USNM slide 23,359. 20. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,596. 21. Cagiva cephalanthana, Dorman Lake, Mississippi, USNM slide 150,530. 22. Cybilla hubbardana, Tucson, Arizona, USNM slide 29,397. 23. Honca grandis, Ogden Canyon, Utah, USNM slide 153,598. 24. Nycthia pimama, Blair Valley, California, USNM slide 144,042. 25. Nycthia yuccatana, Laguna Atacosa, Texas, USNM slide 120,405. 26. Pogospinia floridana, Key Largo, Florida, USNM slide 23,249." figureDoi="http://doi.org/10.5281/zenodo.3442352" httpUri="https://zenodo.org/record/3442352/files/figure.png" pageId="4" pageNumber="199">Fig. 20</figureCitation>
) with tegumen short, broad, convexly rounded at top; uncus absent; socius small, digitate, setose; transtilla a well-developed band, with triangulate to hooklike median process, keeled ventrally; valva highly modified, short, broad basally, with well-developed costal and saccular portions; costal portion broad at base, constricted apically into a robust, curved spine; saccular half with a short, blunt dorsal process and a longer, robust, pointed basal process; inner base of sacculus with elongate, pointed, extension confluent with arms of vinculum; vinculum arms free distally; juxta large, subrectangular, with a pair of dorso-lateral lobes. Phallus large, length about equal to basal width of valva, distally with three median platelike processes covered with minute spicules, subtended laterally by a pair of flattened, distally-pointed flanges extending ventro-terminally; vesica with minute cornuti. Female genitalia (
<figureCitation id="13722A07FFC5FFF8FC2B7EFDFBD8FCD1" box="[999,1081,765,791]" captionStart="FIGURES 3338" captionStartId="20.[151,264,1743,1768]" captionTargetBox="[151,1436,166,1723]" captionTargetId="figure@20.[151,1436,166,1723]" captionTargetPageId="20" captionText="FIGURES 3338. Female genitalia of Cochylini genera. 33. Anacochylidia maderana, Palmerlee,Arizona, USNM slide 23,358. 34. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,597. 35. Cagiva cephalanthana, Dorman, Mississippi, USNM slide 153,609. 36. Cybilla hubbardana (holotype), Tucson, Arizona, USNM slide 23833. 37. Honca grandis, Wellsville, Utah, USNM slide 153,599. 38. Nycthia pimama, Miller Canyon, Arizona, slide MGP 660." figureDoi="http://doi.org/10.5281/zenodo.3442356" httpUri="https://zenodo.org/record/3442356/files/figure.png" pageId="4" pageNumber="199">Fig. 34</figureCitation>
) with ovipositor short; papillae anales narrow, weakly sclerotized, setose, joined posteriorly; length of apophyses anteriores ca. 1.2 times length of apophyses posteriores; sterigma large, composed of four fused spheroid plates, sterigma not attached to apophyses anteriores; ductus bursae short, broad, undifferentiated from corpus bursae; corpus bursae a short convoluted sac lacking a signum; accessory bursa originating from ductus bursae.
</paragraph>
</subSubSection>
<subSubSection id="C3536509FFC5FFF8FF0B7FB1FB0CFC29" pageId="4" pageNumber="199" type="etymology">
<paragraph id="8BF63682FFC5FFF8FF0B7FB1FB0CFC29" blockId="4.[151,1437,153,1008]" pageId="4" pageNumber="199">
<emphasis id="B93DEA90FFC5FFF8FF0B7FB1FEA9FC0D" bold="true" box="[199,328,945,971]" pageId="4" pageNumber="199">Etymology</emphasis>
. The name comes from the Greek “atropos, meaning without turn. In Greek mythology, Atropos (or Aisa) was one of the three Moirai, the goddess of fate and destiny. The name is feminine in gender.
</paragraph>
</subSubSection>
</treatment>
</document>