496 lines
85 KiB
XML
496 lines
85 KiB
XML
<document id="326AAA4FDCB0F9AC72B4FE3C0482F0E6" ID-DOI="10.11646/zootaxa.3368.1.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5252292" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629789001705" checkinUser="felipe" docAuthor="Sørensen, Martin V., Rho, Hyun Soo, Min, Won-Gi, Kim, Dongsung & Chang, Cheon Young" docDate="2012" docId="03E08792E208FFA7FF36F9C0FA71FE65" docLanguage="en" docName="zt03368p196.pdf" docOrigin="Zootaxa 3368 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3368.1.8" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Echinoderes aspinosus Sørensen & Rho & Min & Kim & Chang 2012, sp. nov." docType="treatment" docVersion="4" lastPageNumber="169" masterDocId="FFD9FFEAE20CFFAFFFA1FF8DFFF0FF88" masterDocTitle="An exploration of Echinoderes (Kinorhyncha: Cyclorhagida) in Korean and neighboring waters, with the description of four new species and a redescription of E. tchefouensis Lou, 1934 *" masterLastPageNumber="196" masterPageNumber="161" pageNumber="165" updateTime="1699112222437" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7002065D48C2860237D9F069255ABCF7">An exploration of Echinoderes (Kinorhyncha: Cyclorhagida) in Korean and neighboring waters, with the description of four new species and a redescription of E. tchefouensis Lou, 1934 *</mods:title>
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<mods:roleTerm id="0BE88358C8E1274B995DAEF77EA26680">Author</mods:roleTerm>
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<mods:namePart id="052CACF9B85258FEB20DD72F882F3597">Sørensen, Martin V.</mods:namePart>
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<mods:affiliation id="CB81325251DA7F84ACACDBE58502C0BF">Zoological Museum, The Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, 2100 Copenhagen, Denmark</mods:affiliation>
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<mods:nameIdentifier id="1DD595C1FBAD5588AA8F4E3518674CEA" type="email">mvsorensen@snm.ku.dk</mods:nameIdentifier>
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<mods:roleTerm id="DA1BF264A0942074C6E5B5554144D8B6">Author</mods:roleTerm>
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<mods:namePart id="39F3D9B24F9356D6CF0E296000FBCBB9">Rho, Hyun Soo</mods:namePart>
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<mods:affiliation id="2F2F162A4F9313E6052652FC0B0AA3FD">Dokdo Research Center, Korea Ocean Research and Development Institute, Uljin 767 - 813, Korea</mods:affiliation>
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</mods:name>
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<mods:name id="03E75AB4BB425B68E1D6A096A41AABDB" type="personal">
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<mods:namePart id="60BD464ED268926347CE7C41A96EC0CE">Min, Won-Gi</mods:namePart>
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<mods:affiliation id="925DF37F2DF3DEE849AA13E392D0F883">Dokdo Research Center, Korea Ocean Research and Development Institute, Uljin 767 - 813, Korea</mods:affiliation>
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<mods:namePart id="D5D2E9E7F21BD253EF92C192C5B4DDB9">Kim, Dongsung</mods:namePart>
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<mods:affiliation id="CFF010309C0CA639D844AB802325558C">Marine Living Resources Research Department, Korea Ocean Research and Development Institute, Ansan 425 - 600, Korea Department of Biological Sciences, College of Natural Sciences, Daegu University, Gyeongsan 712 - 714, Korea</mods:affiliation>
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<mods:namePart id="3C68783709DDE25DA1E81E34997548D6">Chang, Cheon Young</mods:namePart>
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<mods:title id="B921CA79725CEF597CD8E46019C1A994">Zootaxa</mods:title>
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<mods:date id="29B363A46CE48F42489A3C153FDA233C">2012</mods:date>
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<mods:number id="83AFEB7E4D230065A8AFF50B3C174694">2012-07-04</mods:number>
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<mods:number id="30BD1A27D70ED88685BF2E71D8863277">3368</mods:number>
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<mods:url id="AEC7B0E09B5E1D53E85C0C252F42D909">https://biotaxa.org/Zootaxa/article/view/zootaxa.3368.1.8</mods:url>
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<mods:classification id="8E35A1C8FA341CADAFC13947F0017BDE">journal article</mods:classification>
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<mods:identifier id="F8A540B723B0DE2B4EEAB82151FAE7BC" type="DOI">10.11646/zootaxa.3368.1.8</mods:identifier>
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<mods:identifier id="2D563451F88E7F34F1F65668B2ACD50A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E08792E208FFA7FF36F9C0FA71FE65" ID-DOI="http://doi.org/10.5281/zenodo.5253988" ID-Zenodo-Dep="5253988" LSID="urn:lsid:plazi:treatment:03E08792E208FFA7FF36F9C0FA71FE65" httpUri="http://treatment.plazi.org/id/03E08792E208FFA7FF36F9C0FA71FE65" lastPageId="8" lastPageNumber="169" pageId="4" pageNumber="165">
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<subSubSection id="C353650FE208FFABFF36F9C0FDE1F9EF" box="[151,529,1613,1639]" pageId="4" pageNumber="165" type="nomenclature">
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<paragraph id="8BF63684E208FFABFF36F9C0FDE1F9EF" blockId="4.[151,529,1613,1672]" box="[151,529,1613,1639]" pageId="4" pageNumber="165">
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<heading id="D0BE81E8E208FFABFF36F9C0FDE1F9EF" bold="true" box="[151,529,1613,1639]" fontSize="11" level="1" pageId="4" pageNumber="165" reason="1">
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<emphasis id="B93DEA96E208FFABFF36F9C0FDE1F9EF" bold="true" box="[151,529,1613,1639]" pageId="4" pageNumber="165">
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<taxonomicName id="4C494D07E208FFABFF36F9C0FE5CF9EF" ID-CoL="38HMD" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[151,428,1613,1639]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="4" pageNumber="165" phylum="Kinorhyncha" rank="species" species="aspinosus" status="sp. nov.">
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<emphasis id="B93DEA96E208FFABFF36F9C0FE5CF9EF" bold="true" box="[151,428,1613,1639]" italics="true" pageId="4" pageNumber="165">Echinoderes aspinosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20E57EDE208FFABFE15F9C0FDE1F9EF" box="[436,529,1613,1639]" pageId="4" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
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<subSubSection id="C353650FE208FFABFF36F9FDFE7FF900" box="[151,399,1648,1672]" pageId="4" pageNumber="165" type="description">
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<paragraph id="8BF63684E208FFABFF36F9FDFE7FF900" blockId="4.[151,529,1613,1672]" box="[151,399,1648,1672]" pageId="4" pageNumber="165">
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(
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<figureCitation id="13722A01E208FFABFF3FF9FDFED8F900" box="[158,296,1648,1672]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="5.[151,244,1905,1926]" captionStartId-1="7.[151,244,1299,1320]" captionTargetBox-0="[205,1404,186,1891]" captionTargetBox-1="[188,1397,191,1275]" captionTargetId-0="figure-95@5.[185,1409,181,1896]" captionTargetId-1="figure-370@7.[186,1401,183,1278]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionText-0="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." captionText-1="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi-0="http://doi.org/10.5281/zenodo.5252298" figureDoi-1="http://doi.org/10.5281/zenodo.5252306" httpUri-0="https://zenodo.org/record/5252298/files/figure.png" httpUri-1="https://zenodo.org/record/5252306/files/figure.png" pageId="4" pageNumber="165">Figures 2–3</figureCitation>
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,
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<tableCitation id="C6CB033FE208FFABFE92F9FDFE77F900" box="[307,391,1648,1672]" captionStart="TABLE 2" captionStartId="6.[151,239,468,492]" captionText="TABLE 2. Measurements of female holotype and male allotype of Echinoderes aspinosus sp. nov. from Yeonpyeong Island (CYC-23). Abbreviations: LTS: lateral terminal spine; MSW-7: Maximum sternal width, measured on segment 7 in this species; S: segment lengths; SW-10, standard width, always measured on segment 10; TL: trunk length." pageId="4" pageNumber="165">Table 2</tableCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C353650FE208FFABFF36F93AFB62F8B4" pageId="4" pageNumber="165" type="diagnosis">
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<paragraph id="8BF63684E208FFABFF36F93AFB62F8B4" blockId="4.[151,1436,1719,2036]" pageId="4" pageNumber="165">
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<emphasis id="B93DEA96E208FFABFF36F93AFEE1F958" bold="true" box="[151,273,1719,1744]" inLineHeading="true" pageId="4" pageNumber="165" reason="1">Diagnosis.</emphasis>
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<taxonomicName id="4C494D07E208FFABFEB8F935FE54F958" authorityName="Claparede" authorityYear="1863" box="[281,420,1720,1744]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="4" pageNumber="165" phylum="Kinorhyncha" rank="genus">
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<emphasis id="B93DEA96E208FFABFEB8F935FE54F958" box="[281,420,1720,1744]" italics="true" pageId="4" pageNumber="165">Echinoderes</emphasis>
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</taxonomicName>
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without tubules or spines, except lateral terminal ones. Both sexes with rather thin lateral terminal spines; females furthermore with extremely thin lateral terminal accessory spines. Pectinate fringe on ventral side of segments 2 to 5 very strong and obliquely orientated, pointing towards the midventral line. Sieve plates on segment 9 elongate and narrow, reaching almost 50% of the total segment length.
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</paragraph>
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</subSubSection>
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<subSubSection id="C353650FE208FFABFF66F8C5FD83F87C" pageId="4" pageNumber="165" type="materials_examined">
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<paragraph id="8BF63684E208FFABFF66F8C5FD83F87C" blockId="4.[151,1436,1719,2036]" pageId="4" pageNumber="165">
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<emphasis id="B93DEA96E208FFABFF66F8C5FE86F8E8" bold="true" box="[199,374,1863,1888]" inLineHeading="true" pageId="4" pageNumber="165" reason="1">Type material.</emphasis>
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<materialsCitation id="3B213CD9E208FFABFEDEF8C5FC76F824" collectingDate="2001-03-31" collectionCode="NIBR" country="South Korea" latitude="37.650833" location="Yeonpyeong Island" longLatPrecision="11" longitude="125.6905" municipality="Korean" pageId="4" pageNumber="165" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54F28826E208FFABFEDEF8C5FE1CF8E8" box="[383,492,1864,1888]" pageId="4" pageNumber="165" type="holotype">Holotype</typeStatus>
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: adult female, collected on
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<date id="FFF71044E208FFABFC87F8C5FC27F8E8" box="[806,983,1864,1888]" pageId="4" pageNumber="165" value="2001-03-31">
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<collectingDate id="EFB3E9ACE208FFABFC87F8C5FC27F8E8" box="[806,983,1864,1888]" pageId="4" pageNumber="165" value="2001-03-31">31 March 2001</collectingDate>
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</date>
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, at station CYC-23,
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<location id="8E96605FE208FFABFB62F8C4FA6BF8E8" LSID="urn:lsid:plazi:treatment:03E08792E208FFA7FF36F9C0FA71FE65:8E96605FE208FFABFB62F8C4FA6BF8E8" box="[1219,1435,1864,1889]" country="South Korea" latitude="37.650833" longLatPrecision="11" longitude="125.6905" municipality="Korean" name="Yeonpyeong Island" pageId="4" pageNumber="165">Yeonpyeong Island</location>
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off the
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<collectingMunicipality id="6B92ACFEE208FFABFF4AF8FCFECDF800" box="[235,317,1905,1928]" pageId="4" pageNumber="165">Korean</collectingMunicipality>
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west coast, (
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<figureCitation id="13722A01E208FFABFE75F8FDFDDEF800" box="[468,558,1904,1928]" captionStart="FIGURE 1" captionStartId="1.[151,244,1947,1968]" captionTargetBox="[179,1407,1068,1890]" captionTargetId="figure-431@1.[179,1407,1068,1927]" captionTargetPageId="1" captionText="FIGURE 1. Maps showing collecting localities for: A, Echinoderes aspinosus sp. nov. and Echinoderes obtuspinosus sp. nov.; B, Echinoderes cernunnos sp. nov.; C, Echinoderes microaperturus sp. nov.; D, Echinoderes tchefouensis Lou, 1934. Type localities are marked with a ring around the dot." figureDoi="http://doi.org/10.5281/zenodo.5252296" httpUri="https://zenodo.org/record/5252296/files/figure.png" pageId="4" pageNumber="165">Fig. 1A</figureCitation>
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),
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<geoCoordinate id="EE7D5043E208FFABFDE7F8FDFD37F800" box="[582,711,1900,1928]" degrees="37" direction="north" minutes="39.05" orientation="latitude" pageId="4" pageNumber="165" precision="9" value="37.650833">
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37
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<superScript id="7C3C9BCCE208FFABFDC3F8E1FD9AF8F2" attach="none" box="[610,618,1900,1914]" fontSize="6" pageId="4" pageNumber="165">o</superScript>
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39.05’N
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</geoCoordinate>
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<geoCoordinate id="EE7D5043E208FFABFD6EF8FDFCADF800" box="[719,861,1900,1928]" degrees="125" direction="east" minutes="41.43" orientation="longitude" pageId="4" pageNumber="165" precision="9" value="125.6905">
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125
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<superScript id="7C3C9BCCE208FFABFD58F8E1FCF1F8F2" attach="none" box="[761,769,1900,1914]" fontSize="6" pageId="4" pageNumber="165">o</superScript>
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41.43’E
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</geoCoordinate>
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,
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<collectingCountry id="F35E7614E208FFABFCCAF8FCFC42F800" box="[875,946,1905,1928]" name="South Korea" pageId="4" pageNumber="165">Korea</collectingCountry>
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, from subtidal sand, mounted in glycerine, deposited at
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<collectionCode id="ED58AE41E208FFABFE8BF818FE9DF824" box="[298,365,1941,1964]" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="4" pageNumber="165">NIBR</collectionCode>
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under accession number INBRIV0000245081
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</materialsCitation>
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.
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<materialsCitation id="3B213CD9E208FFABFC32F819FD9FF87C" collectingDate="2001-03-31" collectionCode="NHMD, ZMUC, SEM" country="South Korea" latitude="37.650833" location="Yeonpyeong Island" longLatPrecision="11" longitude="125.6905" municipality="Korean" pageId="4" pageNumber="165" specimenCode="KIN-538" specimenCount="1" typeStatus="allotype">
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<typeStatus id="54F28826E208FFABFC32F819FC09F824" box="[915,1017,1940,1964]" pageId="4" pageNumber="165" type="allotype">Allotype</typeStatus>
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: adult male, same collecting data as holotype, mounted in glycerine, deposited at
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<collectionCode id="ED58AE41E208FFABFD07F834FD0BF858" box="[678,763,1977,2000]" pageId="4" pageNumber="165">NHMD</collectionCode>
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under accession number
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||
<collectionCode id="ED58AE41E208FFABFB89F835FB89F858" box="[1064,1145,1976,2000]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="165">ZMUC</collectionCode>
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<specimenCode id="DBEF9EFFE208FFABFB22F835FB1CF858" box="[1155,1260,1976,2000]" collectionCode="KIN-" pageId="4" pageNumber="165">KIN-538</specimenCode>
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. No specimens were available for examination with
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<collectionCode id="ED58AE41E208FFABFD93F851FD9FF87C" box="[562,623,2012,2036]" pageId="4" pageNumber="165">SEM</collectionCode>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<caption id="DF36660CE209FFAAFF36F8FCFBE0F856" ID-DOI="http://doi.org/10.5281/zenodo.5252298" ID-Zenodo-Dep="5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="5" pageNumber="166" startId="5.[151,244,1905,1926]" targetBox="[205,1404,186,1891]" targetPageId="5">
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<paragraph id="8BF63684E209FFAAFF36F8FCFBE0F856" blockId="5.[151,1436,1905,2014]" pageId="5" pageNumber="166">
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<emphasis id="B93DEA96E209FFAAFF36F8FCFEFFF80E" bold="true" box="[151,271,1905,1926]" pageId="5" pageNumber="166">FIGURE 2.</emphasis>
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Line art illustrations showing general habitus in
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<taxonomicName id="4C494D07E209FFAAFD5CF8FCFC2BF80E" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[765,987,1905,1926]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="5" pageNumber="166" phylum="Kinorhyncha" rank="species" species="aspinosus" status="sp. nov.">
|
||
<emphasis id="B93DEA96E209FFAAFD5CF8FCFC2BF80E" box="[765,987,1905,1926]" italics="true" pageId="5" pageNumber="166">Echinoderes aspinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E209FFAAFC45F8FCFBC4F80E" bold="true" box="[996,1076,1905,1926]" pageId="5" pageNumber="166">
|
||
<taxonomicNameLabel id="A20E57EDE209FFAAFC45F8FCFBC4F80E" box="[996,1076,1905,1926]" pageId="5" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C353650FE20AFFA9FF66FF15FBA5FF5C" pageId="6" pageNumber="167" type="etymology">
|
||
<paragraph id="8BF63684E20AFFA9FF66FF15FBA5FF5C" blockId="6.[151,1436,151,428]" pageId="6" pageNumber="167">
|
||
<emphasis id="B93DEA96E20AFFA9FF66FF15FEBDFF38" bold="true" box="[199,333,152,176]" inLineHeading="true" pageId="6" pageNumber="167" reason="1">Etymology.</emphasis>
|
||
The species name is composed of the Latin
|
||
<emphasis id="B93DEA96E20AFFA9FC9FFF14FCBCFF38" box="[830,844,153,176]" italics="true" pageId="6" pageNumber="167">a</emphasis>
|
||
- (no or lacking) and -
|
||
<emphasis id="B93DEA96E20AFFA9FBE2FF15FB70FF27" box="[1091,1152,152,175]" italics="true" pageId="6" pageNumber="167">spina</emphasis>
|
||
(spine), meaning the one without spines, with reference to the complete absence of spines on segments 1 to 10.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C353650FE20AFFA7FF66FF52FA71FE65" lastPageId="8" lastPageNumber="169" pageId="6" pageNumber="167" type="description">
|
||
<paragraph id="8BF63684E20AFFA9FF66FF52FBEDFE24" blockId="6.[151,1436,151,428]" pageId="6" pageNumber="167">
|
||
<emphasis id="B93DEA96E20AFFA9FF66FF52FEA8FF70" bold="true" box="[199,344,223,248]" inLineHeading="true" pageId="6" pageNumber="167" reason="1">Description.</emphasis>
|
||
Adult specimens consist of a head, a neck and eleven trunk segments (
|
||
<figureCitation id="13722A01E20AFFA9FB70FF52FAA6FF7F" box="[1233,1366,223,247]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="6" pageNumber="167">Figs 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20AFFA9FAC8FF52FA7FFF7F" box="[1385,1423,223,247]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="6" pageNumber="167">3D</figureCitation>
|
||
). Measurements and dimensions are given in
|
||
<tableCitation id="C6CB033FE20AFFA9FD25FE89FD0AFE94" box="[644,762,260,284]" captionStart="TABLE 2" captionStartId="6.[151,239,468,492]" captionText="TABLE 2. Measurements of female holotype and male allotype of Echinoderes aspinosus sp. nov. from Yeonpyeong Island (CYC-23). Abbreviations: LTS: lateral terminal spine; MSW-7: Maximum sternal width, measured on segment 7 in this species; S: segment lengths; SW-10, standard width, always measured on segment 10; TL: trunk length." pageId="6" pageNumber="167">Table 2. A</tableCitation>
|
||
table summarizing sensory spots, spines, and glandular cell outlet positions is not provided, since such structures apparently are absent (or could at least not be identified) on most of the animal. Because no specimens were available for SEM examinations, it was not possible to identify minor cuticular structures such as sensory spots and pore fields. Hence, not mentioning these structures in the description should not be understood as a positive confirmation of their absence.
|
||
</paragraph>
|
||
<caption id="DF36660CE20AFFA9FF36FE59FAC3FDA7" pageId="6" pageNumber="167" startId="6.[151,239,468,492]">
|
||
<paragraph id="8BF63684E20AFFA9FF36FE59FAC3FDA7" blockId="6.[151,1435,468,559]" pageId="6" pageNumber="167">
|
||
<emphasis id="B93DEA96E20AFFA9FF36FE59FEFCFE64" bold="true" box="[151,268,468,492]" pageId="6" pageNumber="167">TABLE 2.</emphasis>
|
||
Measurements of female holotype and male allotype of
|
||
<taxonomicName id="4C494D07E20AFFA9FCDBFE58FB81FE64" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[890,1137,469,492]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="6" pageNumber="167" phylum="Kinorhyncha" rank="species" species="aspinosus" status="sp. nov.">
|
||
<emphasis id="B93DEA96E20AFFA9FCDBFE58FB81FE64" box="[890,1137,469,492]" italics="true" pageId="6" pageNumber="167">Echinoderes aspinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E20AFFA9FBD8FE59FB21FE64" bold="true" box="[1145,1233,468,492]" pageId="6" pageNumber="167">
|
||
<taxonomicNameLabel id="A20E57EDE20AFFA9FBD8FE59FB21FE64" box="[1145,1233,468,492]" pageId="6" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Yeonpyeong Island (CYC-23). Abbreviations: LTS: lateral terminal spine; MSW-7: Maximum sternal width, measured on segment 7 in this species; S: segment lengths; SW-10, standard width, always measured on segment 10; TL: trunk length.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF63684E20AFFA9FF3EFDCCFBDBFB04" pageId="6" pageNumber="167">
|
||
<table id="F949C424E20A0050FF3EFDCCFB64FB04" box="[159,1172,577,1164]" gridcols="3" gridrows="19" pageId="6" pageNumber="167">
|
||
<tr id="357934C6E20A0050FF3EFDCCFB64FDDE" box="[159,1172,577,598]" gridrow="0" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFDCCFEFEFDDE" box="[159,270,577,598]" gridcol="0" gridrow="0" pageId="6" pageNumber="167">
|
||
<emphasis id="B93DEA96E20AFFA9FF3EFDCCFEF8FDDE" bold="true" box="[159,264,577,598]" pageId="6" pageNumber="167">Character</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE20A0050FD94FDCCFD11FDDE" box="[565,737,577,598]" gridcol="1" gridrow="0" pageId="6" pageNumber="167">
|
||
<emphasis id="B93DEA96E20AFFA9FD94FDCCFD11FDDE" bold="true" box="[565,737,577,598]" pageId="6" pageNumber="167">Length Holotype</emphasis>
|
||
</th>
|
||
<th id="76A85DBAE20A0050FC4CFDCCFB64FDDE" box="[1005,1172,577,598]" gridcol="2" gridrow="0" pageId="6" pageNumber="167">
|
||
<emphasis id="B93DEA96E20AFFA9FC4CFDCCFB64FDDE" bold="true" box="[1005,1172,577,598]" pageId="6" pageNumber="167">Length Allotype</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFDEAFB64FDF4" box="[159,1172,615,636]" gridrow="1" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFDEAFEFEFDF4" box="[159,270,615,636]" gridcol="0" gridrow="1" pageId="6" pageNumber="167">TL</th>
|
||
<td id="76A85DBAE20A0050FD94FDEAFD11FDF4" box="[565,737,615,636]" gridcol="1" gridrow="1" pageId="6" pageNumber="167">284 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFDEAFB64FDF4" box="[1005,1172,615,636]" gridcol="2" gridrow="1" pageId="6" pageNumber="167">263 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFD09FB64FD11" box="[159,1172,644,665]" gridrow="2" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFD09FEFEFD11" box="[159,270,644,665]" gridcol="0" gridrow="2" pageId="6" pageNumber="167">MSW-7</th>
|
||
<td id="76A85DBAE20A0050FD94FD09FD11FD11" box="[565,737,644,665]" gridcol="1" gridrow="2" pageId="6" pageNumber="167">72 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFD09FB64FD11" box="[1005,1172,644,665]" gridcol="2" gridrow="2" pageId="6" pageNumber="167">66 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFD2FFB64FD3F" box="[159,1172,674,695]" gridrow="3" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFD2FFEFEFD3F" box="[159,270,674,695]" gridcol="0" gridrow="3" pageId="6" pageNumber="167">MSW-7/TL</th>
|
||
<td id="76A85DBAE20A0050FD94FD2FFD11FD3F" box="[565,737,674,695]" gridcol="1" gridrow="3" pageId="6" pageNumber="167">25.4 %</td>
|
||
<td id="76A85DBAE20A0050FC4CFD2FFB64FD3F" box="[1005,1172,674,695]" gridcol="2" gridrow="3" pageId="6" pageNumber="167">25.1 %</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFD32FB64FD5C" box="[159,1172,703,724]" gridrow="4" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFD32FEFEFD5C" box="[159,270,703,724]" gridcol="0" gridrow="4" pageId="6" pageNumber="167">SW-10</th>
|
||
<td id="76A85DBAE20A0050FD94FD32FD11FD5C" box="[565,737,703,724]" gridcol="1" gridrow="4" pageId="6" pageNumber="167">57 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFD32FB64FD5C" box="[1005,1172,703,724]" gridcol="2" gridrow="4" pageId="6" pageNumber="167">50 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFD51FB64FD79" box="[159,1172,732,753]" gridrow="5" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFD51FEFEFD79" box="[159,270,732,753]" gridcol="0" gridrow="5" pageId="6" pageNumber="167">SW-10/TL</th>
|
||
<td id="76A85DBAE20A0050FD94FD51FD11FD79" box="[565,737,732,753]" gridcol="1" gridrow="5" pageId="6" pageNumber="167">20.1 %</td>
|
||
<td id="76A85DBAE20A0050FC4CFD51FB64FD79" box="[1005,1172,732,753]" gridcol="2" gridrow="5" pageId="6" pageNumber="167">19.0 %</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFD77FB64FC87" box="[159,1172,762,783]" gridrow="6" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFD77FEFEFC87" box="[159,270,762,783]" gridcol="0" gridrow="6" pageId="6" pageNumber="167">S1</th>
|
||
<td id="76A85DBAE20A0050FD94FD77FD11FC87" box="[565,737,762,783]" gridcol="1" gridrow="6" pageId="6" pageNumber="167">30 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFD77FB64FC87" box="[1005,1172,762,783]" gridcol="2" gridrow="6" pageId="6" pageNumber="167">29 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFC9AFB64FCA4" box="[159,1172,791,812]" gridrow="7" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFC9AFEFEFCA4" box="[159,270,791,812]" gridcol="0" gridrow="7" pageId="6" pageNumber="167">S2</th>
|
||
<td id="76A85DBAE20A0050FD94FC9AFD11FCA4" box="[565,737,791,812]" gridcol="1" gridrow="7" pageId="6" pageNumber="167">32 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFC9AFB64FCA4" box="[1005,1172,791,812]" gridcol="2" gridrow="7" pageId="6" pageNumber="167">30 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFCB9FB64FCC1" box="[159,1172,820,841]" gridrow="8" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFCB9FEFEFCC1" box="[159,270,820,841]" gridcol="0" gridrow="8" pageId="6" pageNumber="167">S3</th>
|
||
<td id="76A85DBAE20A0050FD94FCB9FD11FCC1" box="[565,737,820,841]" gridcol="1" gridrow="8" pageId="6" pageNumber="167">37 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFCB9FB64FCC1" box="[1005,1172,820,841]" gridcol="2" gridrow="8" pageId="6" pageNumber="167">33 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFCDFFB64FCEF" box="[159,1172,850,871]" gridrow="9" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFCDFFEFEFCEF" box="[159,270,850,871]" gridcol="0" gridrow="9" pageId="6" pageNumber="167">S4</th>
|
||
<td id="76A85DBAE20A0050FD94FCDFFD11FCEF" box="[565,737,850,871]" gridcol="1" gridrow="9" pageId="6" pageNumber="167">42 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFCDFFB64FCEF" box="[1005,1172,850,871]" gridcol="2" gridrow="9" pageId="6" pageNumber="167">37 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFCE2FB64FC0C" box="[159,1172,879,900]" gridrow="10" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFCE2FEFEFC0C" box="[159,270,879,900]" gridcol="0" gridrow="10" pageId="6" pageNumber="167">S5</th>
|
||
<td id="76A85DBAE20A0050FD94FCE2FD11FC0C" box="[565,737,879,900]" gridcol="1" gridrow="10" pageId="6" pageNumber="167">45 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFCE2FB64FC0C" box="[1005,1172,879,900]" gridcol="2" gridrow="10" pageId="6" pageNumber="167">41 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFC01FB64FC29" box="[159,1172,908,929]" gridrow="11" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFC01FEFEFC29" box="[159,270,908,929]" gridcol="0" gridrow="11" pageId="6" pageNumber="167">S6</th>
|
||
<td id="76A85DBAE20A0050FD94FC01FD11FC29" box="[565,737,908,929]" gridcol="1" gridrow="11" pageId="6" pageNumber="167">51 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFC01FB64FC29" box="[1005,1172,908,929]" gridcol="2" gridrow="11" pageId="6" pageNumber="167">47 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFC27FB64FC37" box="[159,1172,938,959]" gridrow="12" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFC27FEFEFC37" box="[159,270,938,959]" gridcol="0" gridrow="12" pageId="6" pageNumber="167">S7</th>
|
||
<td id="76A85DBAE20A0050FD94FC27FD11FC37" box="[565,737,938,959]" gridcol="1" gridrow="12" pageId="6" pageNumber="167">55 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFC27FB64FC37" box="[1005,1172,938,959]" gridcol="2" gridrow="12" pageId="6" pageNumber="167">52 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFC4AFB64FC54" box="[159,1172,967,988]" gridrow="13" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFC4AFEFEFC54" box="[159,270,967,988]" gridcol="0" gridrow="13" pageId="6" pageNumber="167">S8</th>
|
||
<td id="76A85DBAE20A0050FD94FC4AFD11FC54" box="[565,737,967,988]" gridcol="1" gridrow="13" pageId="6" pageNumber="167">61 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFC4AFB64FC54" box="[1005,1172,967,988]" gridcol="2" gridrow="13" pageId="6" pageNumber="167">59 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFC69FB64FC71" box="[159,1172,996,1017]" gridrow="14" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFC69FEFEFC71" box="[159,270,996,1017]" gridcol="0" gridrow="14" pageId="6" pageNumber="167">S9</th>
|
||
<td id="76A85DBAE20A0050FD94FC69FD11FC71" box="[565,737,996,1017]" gridcol="1" gridrow="14" pageId="6" pageNumber="167">60 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFC69FB64FC71" box="[1005,1172,996,1017]" gridcol="2" gridrow="14" pageId="6" pageNumber="167">58 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFB8FFB64FB9F" box="[159,1172,1026,1047]" gridrow="15" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFB8FFEFEFB9F" box="[159,270,1026,1047]" gridcol="0" gridrow="15" pageId="6" pageNumber="167">S10</th>
|
||
<td id="76A85DBAE20A0050FD94FB8FFD11FB9F" box="[565,737,1026,1047]" gridcol="1" gridrow="15" pageId="6" pageNumber="167">47 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFB8FFB64FB9F" box="[1005,1172,1026,1047]" gridcol="2" gridrow="15" pageId="6" pageNumber="167">47 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFB92FB64FBBC" box="[159,1172,1055,1076]" gridrow="16" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFB92FEFEFBBC" box="[159,270,1055,1076]" gridcol="0" gridrow="16" pageId="6" pageNumber="167">S11</th>
|
||
<td id="76A85DBAE20A0050FD94FB92FD11FBBC" box="[565,737,1055,1076]" gridcol="1" gridrow="16" pageId="6" pageNumber="167">33 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFB92FB64FBBC" box="[1005,1172,1055,1076]" gridcol="2" gridrow="16" pageId="6" pageNumber="167">28 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFBD7FB64FBE7" box="[159,1172,1114,1135]" gridrow="17" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFBD7FEFEFBE7" box="[159,270,1114,1135]" gridcol="0" gridrow="17" pageId="6" pageNumber="167">LTS</th>
|
||
<td id="76A85DBAE20A0050FD94FBD7FD11FBE7" box="[565,737,1114,1135]" gridcol="1" gridrow="17" pageId="6" pageNumber="167">156 µm</td>
|
||
<td id="76A85DBAE20A0050FC4CFBD7FB64FBE7" box="[1005,1172,1114,1135]" gridcol="2" gridrow="17" pageId="6" pageNumber="167">146 µm</td>
|
||
</tr>
|
||
<tr id="357934C6E20A0050FF3EFBFAFB64FB04" box="[159,1172,1143,1164]" gridrow="18" pageId="6" pageNumber="167">
|
||
<th id="76A85DBAE20A0050FF3EFBFAFEFEFB04" box="[159,270,1143,1164]" gridcol="0" gridrow="18" pageId="6" pageNumber="167">LTS/TL</th>
|
||
<td id="76A85DBAE20A0050FD94FBFAFD11FB04" box="[565,737,1143,1164]" gridcol="1" gridrow="18" pageId="6" pageNumber="167">54.9 %</td>
|
||
<td id="76A85DBAE20A0050FC4CFBFAFB64FB04" box="[1005,1172,1143,1164]" gridcol="2" gridrow="18" pageId="6" pageNumber="167">55.5%</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BF63684E20AFFA9FF66FB4DFEF6FAA8" blockId="6.[151,1436,1216,2032]" pageId="6" pageNumber="167">
|
||
The head consists of a retractable mouth cone and an introvert. Inner and outer armature could not be examined in detail. The pharynx of the
|
||
<typeStatus id="54F28826E20AFFA9FE79FB68FDCAFB75" box="[472,570,1253,1277]" pageId="6" pageNumber="167" type="holotype">holotype</typeStatus>
|
||
was strongly protruded though, revealing the presence of a pharyngeal crown (
|
||
<figureCitation id="13722A01E20AFFA9FF3EFA85FF09FAA8" box="[159,249,1288,1312]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="6" pageNumber="167">Fig. 3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF63684E20AFFA9FF66FAA0FB29FA38" blockId="6.[151,1436,1216,2032]" pageId="6" pageNumber="167">
|
||
The neck consists of 16 placids, all measuring 18 µm in length and 10 µm in width at bases (
|
||
<figureCitation id="13722A01E20AFFA9FB19FAA0FAE3FACD" box="[1208,1299,1325,1349]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="6" pageNumber="167">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20AFFA9FABFFAA0FAB0FACD" box="[1310,1344,1325,1349]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="6" pageNumber="167">3B</figureCitation>
|
||
), except midventral placid that measures 16 µm in width (
|
||
<figureCitation id="13722A01E20AFFA9FD11FADDFCF9FAE0" box="[688,777,1360,1384]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="6" pageNumber="167">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20AFFA9FCB4FADDFCC8FAE0" box="[789,824,1360,1384]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="6" pageNumber="167">3C</figureCitation>
|
||
). If the midventral placid is number 1, and other placids are numbered clockwise from this one, placids number 2 and 16 have broad trichoscalid plates, whereas smaller, but rather elongate trichoscalid plates are associated with placids number 6, 8, 10 and 12 (
|
||
<figureCitation id="13722A01E20AFFA9FBE3FA15FB6EFA38" box="[1090,1182,1432,1456]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="6" pageNumber="167">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20AFFA9FB09FA15FB3CFA38" box="[1192,1228,1432,1456]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="6" pageNumber="167">3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF63684E20AFFA9FF66FA30FA7EF995" blockId="6.[151,1436,1216,2032]" pageId="6" pageNumber="167">
|
||
Segment 1 consists of one complete cuticular ring (
|
||
<figureCitation id="13722A01E20AFFA9FCAAFA30FC7AFA5D" box="[779,906,1469,1493]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="6" pageNumber="167">Figs 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20AFFA9FC37FA30FC2BFA5D" box="[918,987,1469,1493]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="6" pageNumber="167">3B–C</figureCitation>
|
||
). Sensory spots or pore fields could not be identified with certainty. Cuticular hairs emerge through rounded perforation sites, and are densely scattered over the posterior half of the segment, down to the IJ-line. Posterior margin with well-developed pectinate fringe.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E20AFFA9FF66F9A5FD25F925" blockId="6.[151,1436,1216,2032]" pageId="6" pageNumber="167">
|
||
Segment 2 consists of one complete cuticular ring. Cuticular hairs are distributed in a median belt around the segment, limited posteriorly by the IJ-line. Posterior segment margin with regular pectinate fringe on dorsal side, but a conspicuously strong fringe ventrally; fringe tips within the ventromedial areas are obliquely orientated, pointing towards the midventral line (
|
||
<figureCitation id="13722A01E20AFFA9FD9CF918FD68F925" box="[573,664,1685,1709]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="6" pageNumber="167">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20AFFA9FD05F918FD38F925" box="[676,712,1685,1709]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="6" pageNumber="167">3C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF63684E20AFFA9FF66F935FC23F80D" blockId="6.[151,1436,1216,2032]" pageId="6" pageNumber="167">
|
||
Segment 3, and following 8 segments, consist of one tergal and two sternal plates (
|
||
<figureCitation id="13722A01E20AFFA9FB3EF935FB0EF958" box="[1183,1278,1720,1744]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="6" pageNumber="167">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20AFFA9FAAFF935FAC1F958" box="[1294,1329,1720,1744]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="6" pageNumber="167">3B</figureCitation>
|
||
). Spines, tubules, glandular cell outlets
|
||
<typeStatus id="54F28826E20AFFA9FE4FF950FDEEF97D" box="[494,542,1757,1781]" pageId="6" pageNumber="167">type</typeStatus>
|
||
1 or 2 are not present. Cuticular hairs are densely distributed in a median belt around the segment, except in paraventral areas, limited posteriorly by the IJ-line. Posterior margin with regular pectinate fringe on dorsal side, and a conspicuously strong one on ventral side; fringe tips on sternal plates of segments 3 to 5 are obliquely orientated, pointing towards the midventral line (
|
||
<figureCitation id="13722A01E20AFFA9FBB3F8C5FB9FF8E8" box="[1042,1135,1864,1888]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="6" pageNumber="167">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20AFFA9FBDAF8C5FB6EF8E8" box="[1147,1182,1864,1888]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="6" pageNumber="167">3E</figureCitation>
|
||
); fringe tips on sternal plates of following segment not oblique and less prominent (
|
||
<figureCitation id="13722A01E20AFFA9FC9CF8E0FC6AF80D" box="[829,922,1901,1925]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="6" pageNumber="167">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20AFFA9FC04F8E0FC36F80D" box="[933,966,1901,1925]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="6" pageNumber="167">3F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF63684E20AFFA9FF66F81DFD9BF878" blockId="6.[151,1436,1216,2032]" pageId="6" pageNumber="167">
|
||
Segment 9 similar to preceding segments, but with weaker pectinate fringe, and a pair of very conspicuous sieve plates (
|
||
<figureCitation id="13722A01E20AFFA9FE86F838FE73F845" box="[295,387,1973,1997]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="6" pageNumber="167">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20AFFA9FE2FF838FE5FF845" box="[398,431,1973,1997]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="6" pageNumber="167">3F</figureCitation>
|
||
). The sieve plates are elongate, 27 µm long, i.e. almost 50% of segment length, and rather narrow, but broadens slightly in each end.
|
||
</paragraph>
|
||
<caption id="DF36660CE20BFFA8FF36FA9EFDFEFA50" ID-DOI="http://doi.org/10.5281/zenodo.5252306" ID-Zenodo-Dep="5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="7" pageNumber="168" startId="7.[151,244,1299,1320]" targetBox="[188,1397,191,1275]" targetPageId="7">
|
||
<paragraph id="8BF63684E20BFFA8FF36FA9EFDFEFA50" blockId="7.[151,1436,1299,1496]" pageId="7" pageNumber="168">
|
||
<emphasis id="B93DEA96E20BFFA8FF36FA9EFEFDFAA0" bold="true" box="[151,269,1299,1320]" pageId="7" pageNumber="168">FIGURE 3.</emphasis>
|
||
Light micrographs showing details in neck and trunk morphology of
|
||
<taxonomicName id="4C494D07E20BFFA8FC13FA99FB7FFAA1" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[946,1167,1300,1321]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="168" phylum="Kinorhyncha" rank="species" species="aspinosus" status="sp. nov.">
|
||
<emphasis id="B93DEA96E20BFFA8FC13FA99FB7FFAA1" box="[946,1167,1300,1321]" italics="true" pageId="7" pageNumber="168">Echinoderes aspinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E20BFFA8FB34FA9EFB13FAA0" bold="true" box="[1173,1251,1299,1320]" pageId="7" pageNumber="168">
|
||
<taxonomicNameLabel id="A20E57EDE20BFFA8FB34FA9EFB13FAA0" box="[1173,1251,1299,1320]" pageId="7" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF63684E20BFFA8FF66F9A4FADEF9ED" blockId="7.[151,1436,1577,2033]" pageId="7" pageNumber="168">
|
||
Segment 10 generally as preceding segments, but with the posterior margins of the sternal plates forming a broadly lobed midventral extension that almost covers the ventral side of the terminal segment (
|
||
<figureCitation id="13722A01E20BFFA8FB69F9C0FAD0F9ED" box="[1224,1312,1613,1637]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="7" pageNumber="168">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF63684E20BFFA8FF66F9FCFCE0F8B5" blockId="7.[151,1436,1577,2033]" pageId="7" pageNumber="168">
|
||
Segment 11 with long and rather thin lateral terminal spines. Females furthermore with pair of very thin lateral terminal accessory spines (
|
||
<figureCitation id="13722A01E20BFFA8FE63F918FDCFF925" box="[450,575,1685,1709]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="7" pageNumber="168">Figs 2A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20BFFA8FDEBF918FD80F925" box="[586,624,1685,1709]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="7" pageNumber="168">3G</figureCitation>
|
||
). Males with three pairs of flexible penile spines; dorsalmost spine thick, ca. 30 µm long, median penile spine slightly thinner and pointed, ca. 26 µm, ventralmost penile spine thicker, ca. 41 µm (
|
||
<figureCitation id="13722A01E20BFFA8FF59F950FE8BF97D" box="[248,379,1757,1781]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="7" pageNumber="168">Figs 2C–D</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20BFFA8FE2AF950FE41F97D" box="[395,433,1757,1781]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="7" pageNumber="168">3H</figureCitation>
|
||
). Cuticular hairs are not present. Tergal extensions are short, with interrupted mesial margins (
|
||
<figureCitation id="13722A01E20BFFA8FF5EF88CFEA8F891" box="[255,344,1793,1817]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="7" pageNumber="168">Fig. 3G</figureCitation>
|
||
); sternal extensions short and lobed with weak fringes. A pair of long (23 µm) setae emerges from the outer margins of the tergal extensions (
|
||
<figureCitation id="13722A01E20BFFA8FDD5F8A8FD21F8B5" box="[628,721,1829,1853]" captionStart="FIGURE 2" captionStartId="5.[151,244,1905,1926]" captionTargetBox="[205,1404,186,1891]" captionTargetId="figure-95@5.[185,1409,181,1896]" captionTargetPageId="5" captionText="FIGURE 2. Line art illustrations showing general habitus in Echinoderes aspinosus sp. nov.: A, female, dorsal view; B, female, ventral view; C, segments 10 and 11 in male, dorsal view; D, segments 10 and 11 in male, ventral view. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; pe, penile spines; pf, pectinate fringe; s, segment followed by segment number; set, seta; si, sieve plate; te, tergal extension." figureDoi="http://doi.org/10.5281/zenodo.5252298" httpUri="https://zenodo.org/record/5252298/files/figure.png" pageId="7" pageNumber="168">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="13722A01E20BFFA8FD7CF8A8FCF3F8B5" box="[733,771,1829,1853]" captionStart="FIGURE 3" captionStartId="7.[151,244,1299,1320]" captionTargetBox="[188,1397,191,1275]" captionTargetId="figure-370@7.[186,1401,183,1278]" captionTargetPageId="7" captionText="FIGURE 3. Light micrographs showing details in neck and trunk morphology of Echinoderes aspinosus sp. nov., displaying female holotype, NIBR Acc. No. INBRIV0000245081 (A, C–G) and male allotype, ZMUC KIN-538 (B, H): A, detail of extended mouth cone; B, neck and segments 1 and 2, dorsal view; C, neck and segments 1 and 2, ventral view; D, ventral overview of holotype; E, Segments 2 to 5, ventral view; F, Segments 8 and 9, ventral view; G, Segment 11 and terminal spines in female; H, Segment 11 and terminal spines in male. Abbreviations: ltas, lateral terminal accessory spine; lts, lateral terminal spine; mdp, middorsal placid; mvp, midventral placid; oos, outer oral style; pc, pharyngeal crown; pe, penile spine; pf, pectinate fringe; se, sternal extension; set, seta; te, tergal extension; trp, trichoscalid plate." figureDoi="http://doi.org/10.5281/zenodo.5252306" httpUri="https://zenodo.org/record/5252306/files/figure.png" pageId="7" pageNumber="168">3G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF63684E20BFFA8FF66F8C4FB15F879" blockId="7.[151,1436,1577,2033]" pageId="7" pageNumber="168">
|
||
<emphasis id="B93DEA96E20BFFA8FF66F8C4FCC1F8E9" bold="true" box="[199,817,1864,1889]" inLineHeading="true" pageId="7" pageNumber="168" reason="1">Notes on diagnostic features and potential relatives.</emphasis>
|
||
Due to its complete lack of tubules and spines (except terminal ones),
|
||
<taxonomicName id="4C494D07E20BFFA8FEEAF8E0FDBEF80D" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[331,590,1901,1925]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="168" phylum="Kinorhyncha" rank="species" species="aspinosus" status="sp. nov.">
|
||
<emphasis id="B93DEA96E20BFFA8FEEAF8E0FDBEF80D" box="[331,590,1901,1925]" italics="true" pageId="7" pageNumber="168">Echinoderes aspinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E20BFFA8FDF9F8E0FD44F80D" bold="true" box="[600,692,1901,1925]" pageId="7" pageNumber="168">
|
||
<taxonomicNameLabel id="A20E57EDE20BFFA8FDF9F8E0FD44F80D" box="[600,692,1901,1925]" pageId="7" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
cannot be confused with any other species of
|
||
<taxonomicName id="4C494D07E20BFFA8FB73F8E0FA87F80D" authorityName="Butschli" authorityYear="1876" box="[1234,1399,1901,1925]" class="Cyclorhagida" family="Echinoderidae" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="168" phylum="Kinorhyncha" rank="family">Echinoderidae</taxonomicName>
|
||
. In addition to this negative character, the species can also be recognized by its strong, obliquely orientated pectinate fringe on the ventral side of segments 2 to 5. This character has to our knowledge not been reported from any other kinorhynch species. Another noteworthy character, not unique though, is the enlarged sieve plates.
|
||
</paragraph>
|
||
<paragraph id="8BF63684E204FFA7FF66FF1AFA71FE65" blockId="8.[151,1436,151,493]" pageId="8" pageNumber="169">
|
||
The only species that potentially could be confused with
|
||
<taxonomicName id="4C494D07E204FFA7FCE5FF14FC21FF27" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[836,977,152,176]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="aspinosus" status="sp. nov.">
|
||
<emphasis id="B93DEA96E204FFA7FCE5FF14FC21FF27" box="[836,977,152,176]" italics="true" pageId="8" pageNumber="169">E. aspinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E204FFA7FC78FF15FBC2FF38" bold="true" box="[985,1074,152,176]" pageId="8" pageNumber="169">
|
||
<taxonomicNameLabel id="A20E57EDE204FFA7FC78FF15FBC2FF38" box="[985,1074,152,176]" pageId="8" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, are those of the
|
||
<taxonomicName id="4C494D07E204FFA7FB53FF14FAA1FF38" authorityName="Higgins" authorityYear="1977" box="[1266,1361,152,176]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="coulli">
|
||
<emphasis id="B93DEA96E204FFA7FB53FF14FAA1FF38" box="[1266,1361,152,176]" italics="true" pageId="8" pageNumber="169">E. coulli</emphasis>
|
||
</taxonomicName>
|
||
-group that also tend to have few, small or completely reduced spines and tubules (
|
||
<bibRefCitation id="EFD84B75E204FFA7FBA0FF36FB02FF5B" author="Ostmann, A. & Nordhaus, I. & Sorensen, M. V." box="[1025,1266,187,211]" pageId="8" pageNumber="169" pagination="71 - 91" refId="ref19819" refString="Ostmann, A., Nordhaus, I. & Sorensen, M. V. (2012). First recording of kinorhynchs from Java, with the description of a new brackish water species from a mangrove-fringed lagoon. Marine Biodiversity, 42, 71 - 91." type="journal article" year="2012">
|
||
Ostmann
|
||
<emphasis id="B93DEA96E204FFA7FBD0FF31FB5CFF5B" box="[1137,1196,187,211]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
). This speciesgroup includes, besides
|
||
<taxonomicName id="4C494D07E204FFA7FE0AFF52FD49FF7E" authority="Higgins, 1977" authorityName="Higgins" authorityYear="1977" box="[427,697,222,246]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="coulli">
|
||
<emphasis id="B93DEA96E204FFA7FE0AFF52FDFDFF7E" box="[427,525,222,246]" italics="true" pageId="8" pageNumber="169">E. coulli</emphasis>
|
||
<bibRefCitation id="EFD84B75E204FFA7FDB7FF53FD49FF7E" author="Higgins, R. P." box="[534,697,222,246]" pageId="8" pageNumber="169" pagination="340 - 354" refId="ref19306" refString="Higgins, R. P. (1977) Two new species of Echinoderes (Kinorhyncha) from South Carolina. Transactions of the American Microscopical Society, 96, 340 - 354." type="journal article" year="1977">Higgins, 1977</bibRefCitation>
|
||
</taxonomicName>
|
||
, the following five species:
|
||
<taxonomicName id="4C494D07E204FFA7FBA0FF52FA85FF7E" authority="Omer-Cooper, 1957" authorityName="Omer-Cooper" authorityYear="1957" box="[1025,1397,222,246]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="maxwelli">
|
||
<emphasis id="B93DEA96E204FFA7FBA0FF52FB78FF7E" box="[1025,1160,222,246]" italics="true" pageId="8" pageNumber="169">E. maxwelli</emphasis>
|
||
<bibRefCitation id="EFD84B75E204FFA7FB30FF53FA85FF7E" author="Omer-Cooper, J." box="[1169,1397,222,246]" pageId="8" pageNumber="169" pagination="213 - 216" refId="ref19785" refString="Omer-Cooper, J. (1957) Deux nouvelle especes de Kinorhyncha en provenance de L'Afrique du Sud. Bulletin mensuel de la Societe Linneenne de Lyon, 26, 213 - 216." type="journal article" year="1957">Omer-Cooper, 1957</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C494D07E204FFA7FA25FF52FE88FE91" authority="Brown, 1999" authorityName="Brown" authorityYear="1999" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="teretis">
|
||
<emphasis id="B93DEA96E204FFA7FA25FF52FF2CFE91" italics="true" pageId="8" pageNumber="169">E. teretis</emphasis>
|
||
Brown,
|
||
<quantity id="4CB19B61E204FFA7FEE1FE8CFE88FE91" box="[320,376,257,281]" metricMagnitude="1" metricUnit="m" metricValue="5.07746" pageId="8" pageNumber="169" unit="in" value="1999.0">1999</quantity>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EFD84B75E204FFA7FE3FFE8CFD19FE91" author="Adrianov, A. V. & Malakhov, V. V." box="[414,745,257,281]" pageId="8" pageNumber="169" refId="ref18869" refString="Adrianov, A. V. & Malakhov, V. V. (1999) Cephalorhyncha of the World Ocean. KMK Scientific Press, Moscow, 328 pp." type="book" year="1999">Adrianov & Malakhov, 1999</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C494D07E204FFA7FD54FE8FFCC8FE91" authorityName="Lundbye, Rho & Sorensen" authorityYear="2011" box="[757,824,258,281]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="rex">
|
||
<emphasis id="B93DEA96E204FFA7FD54FE8FFCC8FE91" box="[757,824,258,281]" italics="true" pageId="8" pageNumber="169">E. rex</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C494D07E204FFA7FCE9FE8FFA68FE91" authority="Ostmann, Nordhaus & Sorensen, 2012" authorityName="Ostmann, Nordhaus & Sorensen" authorityYear="2012" box="[840,1432,257,281]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="applicitus">
|
||
<emphasis id="B93DEA96E204FFA7FCE9FE8FFC27FE91" box="[840,983,257,281]" italics="true" pageId="8" pageNumber="169">E. applicitus</emphasis>
|
||
<bibRefCitation id="EFD84B75E204FFA7FC7EFE8CFA68FE91" author="Ostmann, A. & Nordhaus, I. & Sorensen, M. V." box="[991,1432,257,281]" pageId="8" pageNumber="169" pagination="71 - 91" refId="ref19819" refString="Ostmann, A., Nordhaus, I. & Sorensen, M. V. (2012). First recording of kinorhynchs from Java, with the description of a new brackish water species from a mangrove-fringed lagoon. Marine Biodiversity, 42, 71 - 91." type="journal article" year="2012">Ostmann, Nordhaus & Sørensen, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C494D07E204FFA7FF69FEA8FD7FFEB4" authority="Yamasaki & Kajihara, 2012" authorityName="Yamasaki & Kajihara" authorityYear="2012" box="[200,655,292,317]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="ohtsukai">
|
||
<emphasis id="B93DEA96E204FFA7FF69FEA8FEB7FEB5" box="[200,327,293,317]" italics="true" pageId="8" pageNumber="169">E. ohtsukai</emphasis>
|
||
<bibRefCitation id="EFD84B75E204FFA7FEF1FEA9FD7FFEB4" author="Yamasaki, H. & Kajihara, H." box="[336,655,292,316]" pageId="8" pageNumber="169" pagination="109 - 118" refId="ref20390" refString="Yamasaki, H. & Kajihara, H. (2012) A New Brackish-water Species of Echinoderes (Kinorhyncha: Cyclorhagida) from the Seto Inland Sea, Japan. Species Diversity, 17, 109 - 118." type="journal article" year="2012">Yamasaki & Kajihara, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
. Common for all these six species are, besides the reduced spines in terms of number and size, that they all possess a somewhat enlarged sieve plate, and that females do not possess lateral terminal accessory spines (see
|
||
<bibRefCitation id="EFD84B75E204FFA7FDF7FEE6FD06FE0B" author="Higgins, R. P." box="[598,758,363,387]" pageId="8" pageNumber="169" pagination="340 - 354" refId="ref19306" refString="Higgins, R. P. (1977) Two new species of Echinoderes (Kinorhyncha) from South Carolina. Transactions of the American Microscopical Society, 96, 340 - 354." type="journal article" year="1977">Higgins 1977</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD84B75E204FFA7FCA7FEE6FC08FE0B" author="Lundbye, H. & Rho, H. S. & Sorensen, M. V." box="[774,1016,363,387]" pageId="8" pageNumber="169" pagination="41 - 51" refId="ref19650" refString="Lundbye, H., Rho, H. S. & Sorensen, M. V. (2011) Echinoderes rex n. sp. (Kinorhyncha: Cyclorhagida), the largest Echinoderes species found so far. Scientia Marina, 75, 41 - 51." type="journal article" year="2011">
|
||
Lundbye
|
||
<emphasis id="B93DEA96E204FFA7FCD4FEE1FC42FE0B" box="[885,946,363,387]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD84B75E204FFA7FBA9FEE6FB0DFE0B" author="Ostmann, A. & Nordhaus, I. & Sorensen, M. V." box="[1032,1277,363,387]" pageId="8" pageNumber="169" pagination="71 - 91" refId="ref19819" refString="Ostmann, A., Nordhaus, I. & Sorensen, M. V. (2012). First recording of kinorhynchs from Java, with the description of a new brackish water species from a mangrove-fringed lagoon. Marine Biodiversity, 42, 71 - 91." type="journal article" year="2012">
|
||
Ostmann
|
||
<emphasis id="B93DEA96E204FFA7FBD8FEE1FB46FE0B" box="[1145,1206,363,387]" italics="true" pageId="8" pageNumber="169">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD84B75E204FFA7FAACFEE6FECEFE2E" author="Yamasaki, H. & Kajihara, H." pageId="8" pageNumber="169" pagination="109 - 118" refId="ref20390" refString="Yamasaki, H. & Kajihara, H. (2012) A New Brackish-water Species of Echinoderes (Kinorhyncha: Cyclorhagida) from the Seto Inland Sea, Japan. Species Diversity, 17, 109 - 118." type="journal article" year="2012">Yamasaki & Kajihara 2012</bibRefCitation>
|
||
). An enlarged sieve plate is found in
|
||
<taxonomicName id="4C494D07E204FFA7FD53FE02FC72FE2E" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[754,898,399,422]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="aspinosus" status="sp. nov.">
|
||
<emphasis id="B93DEA96E204FFA7FD53FE02FC72FE2E" box="[754,898,399,422]" italics="true" pageId="8" pageNumber="169">E. aspinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E204FFA7FC2DFE03FC18FE2E" bold="true" box="[908,1000,398,422]" pageId="8" pageNumber="169">
|
||
<taxonomicNameLabel id="A20E57EDE204FFA7FC2DFE03FC18FE2E" box="[908,1000,398,422]" pageId="8" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also, and even though females of the species have lateral terminal accessory spines, it is noteworthy that they are very weakly developed. These similarities could indicate a closer relationship between
|
||
<taxonomicName id="4C494D07E204FFA7FCADFE58FC69FE64" authority="Sørensen & Rho & Min & Kim & Chang, 2012" authorityName="Sørensen & Rho & Min & Kim & Chang" authorityYear="2012" box="[780,921,469,492]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="aspinosus" status="sp. nov.">
|
||
<emphasis id="B93DEA96E204FFA7FCADFE58FC69FE64" box="[780,921,469,492]" italics="true" pageId="8" pageNumber="169">E. aspinosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93DEA96E204FFA7FC01FE58FC0AFE65" bold="true" box="[928,1018,469,493]" pageId="8" pageNumber="169">
|
||
<taxonomicNameLabel id="A20E57EDE204FFA7FC01FE58FC0AFE65" box="[928,1018,469,493]" pageId="8" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and species of the
|
||
<taxonomicName id="4C494D07E204FFA7FB71FE58FADFFE65" authorityName="Higgins" authorityYear="1977" box="[1232,1327,469,493]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="169" phylum="Kinorhyncha" rank="species" species="coulli">
|
||
<emphasis id="B93DEA96E204FFA7FB71FE58FADFFE65" box="[1232,1327,469,493]" italics="true" pageId="8" pageNumber="169">E. coulli</emphasis>
|
||
</taxonomicName>
|
||
-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |