282 lines
38 KiB
XML
282 lines
38 KiB
XML
<document id="31BC6988E8FD5D0502D9B857EEFBD5C6" ID-CLB-Dataset="32636" ID-DOI="10.11646/zootaxa.4294.4.1" ID-GBIF-Dataset="f3e2e226-a609-453b-995f-b5bee31a5642" ID-ISSN="1175-5326" ID-Zenodo-Dep="832774" ID-ZooBank="AEE0A680-3206-4AFA-968D-5EC5313738D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500555101147" checkinUser="plazi" docAuthor="Huerta, Herón & Grogan, William L." docDate="2017" docId="03E0878EFF95F462FF39FBD70798C58E" docLanguage="en" docName="zootaxa.4294.4.1.pdf" docOrigin="Zootaxa 4294 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stilobezzia (Acanthohelea) spinellii Huerta & Grogan 2017, new species" docType="treatment" docVersion="8" lastPageNumber="411" masterDocId="FFD9FFF6FF92F468FFAEFFCD0477C423" masterDocTitle="New species and new records of predaceous midges in the genera, Schizonyxhelea Clastrier and Stilobezzia Kieffer from Mexico (Diptera: Ceratopogonidae)" masterLastPageNumber="418" masterPageNumber="401" pageNumber="408" updateTime="1698465852412" updateUser="ExternalLinkService">
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<mods:title id="B8F8CA1CE4BC82B97BFFE80D0C3AE36D">New species and new records of predaceous midges in the genera, Schizonyxhelea Clastrier and Stilobezzia Kieffer from Mexico (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="8FC994F51B79BC76F73D673E9D8FA347">Huerta, Herón</mods:namePart>
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<mods:namePart id="17A4F580D0CE8BB2946F81A0A372541D">Grogan, William L.</mods:namePart>
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<mods:title id="023AF3B5C12EBD80FEFA8EE208BB96E2">Zootaxa</mods:title>
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<mods:date id="416F65FC20D9BBFBF5EF33B3FA506F8B">2017</mods:date>
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<mods:number id="3A643DFF326A66D77EB4BC6BCCC15780">4294</mods:number>
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<mods:identifier id="6BCEAE794EAB45804130F7F64ACBE8A5" type="DOI">10.11646/zootaxa.4294.4.1</mods:identifier>
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<treatment id="03E0878EFF95F462FF39FBD70798C58E" ID-DOI="http://doi.org/10.5281/zenodo.5189446" ID-GBIF-Taxon="132550624" ID-Zenodo-Dep="5189446" LSID="urn:lsid:plazi:treatment:03E0878EFF95F462FF39FBD70798C58E" httpUri="http://treatment.plazi.org/id/03E0878EFF95F462FF39FBD70798C58E" lastPageId="10" lastPageNumber="411" pageId="7" pageNumber="408">
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<subSubSection id="C3536513FF95F46FFF39FBD70557C076" pageId="7" pageNumber="408" type="nomenclature">
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<paragraph id="8BF63698FF95F46FFF39FBD706A0C017" blockId="7.[151,727,1050,1109]" box="[151,727,1050,1076]" pageId="7" pageNumber="408">
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<heading id="D0BE81F4FF95F46FFF39FBD706A0C017" bold="true" box="[151,727,1050,1076]" fontSize="11" level="1" pageId="7" pageNumber="408" reason="1">
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<emphasis id="B93DEA8AFF95F46FFF39FBD706A0C017" bold="true" box="[151,727,1050,1076]" pageId="7" pageNumber="408">
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<taxonomicName id="4C494D1BFF95F46FFF39FBD70637C017" ID-CoL="9HWRT" authority="Huerta & Grogan, 2017" authorityName="Huerta & Grogan" authorityYear="2017" box="[151,576,1050,1076]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="spinellii" status="sp. nov." subGenus="Acanthohelea">
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<emphasis id="B93DEA8AFF95F46FFF39FBD70637C017" bold="true" box="[151,576,1050,1076]" italics="true" pageId="7" pageNumber="408">Stilobezzia (Acanthohelea) spinellii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20E57F1FF95F46FFDE9FBD706A0C017" box="[583,727,1050,1076]" pageId="7" pageNumber="408" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF63698FF95F46FFF39FBF10557C076" blockId="7.[151,727,1050,1109]" box="[151,288,1084,1109]" pageId="7" pageNumber="408">(Figs. 8–24)</paragraph>
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</subSubSection>
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<subSubSection id="C3536513FF95F46FFF39FB490702C10E" pageId="7" pageNumber="408" type="diagnosis">
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<paragraph id="8BF63698FF95F46FFF39FB490702C10E" blockId="7.[151,1437,1156,2021]" pageId="7" pageNumber="408">
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<emphasis id="B93DEA8AFF95F46FFF39FB490566C0BE" bold="true" box="[151,273,1156,1181]" pageId="7" pageNumber="408">Diagnosis.</emphasis>
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Males are the only extant New World species of
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<emphasis id="B93DEA8AFF95F46FFCEFFB480738C0BF" box="[833,847,1157,1180]" italics="true" pageId="7" pageNumber="408">S</emphasis>
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. (
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<taxonomicName id="4C494D1BFF95F46FFCC9FB490076C0BE" authorityName="Kieffer, Debenhamia Wirth & Grogan" authorityYear="1917" box="[871,1025,1156,1181]" class="Insecta" family="Ceratopogonidae" genus="Acanthohelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
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<emphasis id="B93DEA8AFF95F46FFCC9FB490076C0BE" box="[871,1025,1156,1181]" italics="true" pageId="7" pageNumber="408">Acanthohelea</emphasis>
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</taxonomicName>
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) with sternite 9 having a deep V- or U-shaped posteromedian excavation; and parameres separate, subcylindrical, sub-parallel, extreme bases pointed, distal 1/3 tapered with elongate apicolateral concavities and blunt tips with a few tiny spicules. Females with dark brown legs except bases of femora and bases, apices of tibiae with pale rings; antennal flagellum with bases of flagellomeres pale; and wing with a darkly infuscated membrane.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3536513FF95F460FF69FAF507A2C3C8" lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408" type="description">
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<paragraph id="8BF63698FF95F460FF69FAF50026C4F6" blockId="7.[151,1437,1156,2021]" lastBlockId="8.[151,1437,151,725]" lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408">
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<emphasis id="B93DEA8AFF95F46FFF69FAF5057CC172" bold="true" box="[199,267,1336,1361]" pageId="7" pageNumber="408">Male.</emphasis>
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Head (Fig. 8). Dark brown. Antennal flagellum with flagellomeres 1–10 pale, 11–13 (Fig. 10) dark brown; plume pale, well developed; antennal ratio 1.07–1.33 (1.20, n=4). Palpus (Fig. 12) brown; segment 3 slender with small shallow subapical sensory pit; palpal ratio 2.50–2.60 (2.50, n= 4). Clypeus with 3–4 lateral, 2 subapical setae. Thorax (Fig. 14). Scutum dark brown, medial vittae, humeral pits and prescutellar depression paler; scutellum dark brown with 6 large setae; postscutellum, pleurae dark brown. Legs (
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<figureCitation id="13722A1DFF95F46FFB2CFA0500ABC1C2" box="[1154,1244,1480,1505]" captionStart="FIGURES 15 – 19" captionStartId="9.[151,264,1462,1485]" captionTargetBox="[177,1417,224,1418]" captionTargetId="figure@9.[151,1436,193,1442]" captionTargetPageId="9" captionText="FIGURES 15 – 19. Stilobezzia (Acanthohelea) spinellii n. sp. Male: 15 a – c – 16. Female: 17 – 19. 15, 18. Legs, lateral view. a. Fore leg. b. Mid leg. c. Hind leg. 16 – 17. Wings. 19. Abdominal segments 8 – 10 and spermathecae, ventral view. Scales: 0.1 mm. Abbreviations: A = anal vein; c = costal vein; CuA 1 and CuA 2 = cubital veins, anterior branches; CuP = cubital vein, posterior branch; M 1 and M 2 = medial veins; r-m = radial-medial crossvein; R 1 = radial vein, first longitudinal vein; R 3 = radial vein, second longitudinal vein. Scales: 0.1 mm." httpUri="https://zenodo.org/record/832778/files/figure.png" pageId="7" pageNumber="408">Figs. 15</figureCitation>
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a–c) dark brown; femora with narrow pale basal band, fore femur paler than mid, hind femora; hind femur more massive than fore, mid femora; tibiae with basal, apical pale bands which are more distinctive on hind tibia; hind tibia with dorsal row of long setae; fore, mid tarsi pale brown, hind tarsus darker brown. Wing (
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<figureCitation id="13722A1DFF95F46FFBB7F9F90018C26E" box="[1049,1135,1588,1613]" captionStart="FIGURES 15 – 19" captionStartId="9.[151,264,1462,1485]" captionTargetBox="[177,1417,224,1418]" captionTargetId="figure@9.[151,1436,193,1442]" captionTargetPageId="9" captionText="FIGURES 15 – 19. Stilobezzia (Acanthohelea) spinellii n. sp. Male: 15 a – c – 16. Female: 17 – 19. 15, 18. Legs, lateral view. a. Fore leg. b. Mid leg. c. Hind leg. 16 – 17. Wings. 19. Abdominal segments 8 – 10 and spermathecae, ventral view. Scales: 0.1 mm. Abbreviations: A = anal vein; c = costal vein; CuA 1 and CuA 2 = cubital veins, anterior branches; CuP = cubital vein, posterior branch; M 1 and M 2 = medial veins; r-m = radial-medial crossvein; R 1 = radial vein, first longitudinal vein; R 3 = radial vein, second longitudinal vein. Scales: 0.1 mm." httpUri="https://zenodo.org/record/832778/files/figure.png" pageId="7" pageNumber="408">Fig. 16</figureCitation>
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) membrane, veins darkly infuscated; short anterior, longer posterior intercalary veins in distal portion of cell r3; 2nd radial cell 2.30 x longer than 1st; cubital fork at level of base of r-m crossvein; macrotrichia on costa, Rs, distal margins of M1, and cells r3 and m1; wing length 1.47–1.50 (1.48, n = 4) mm, width 0.47–0.49 (0.48, n= 4) mm; costal ratio 0.76–0.78 (0.77, n= 4). Haltere pale. Abdomen (
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<figureCitation id="13722A1DFF95F46FFE61F9020653C2CB" box="[463,548,1743,1768]" captionStart="FIGURES 20 – 23" captionStartId="10.[151,264,1788,1811]" captionTargetBox="[193,1420,485,1747]" captionTargetId="figure@10.[151,1436,468,1776]" captionTargetPageId="10" captionText="FIGURES 20 – 23. Stilobezzia (Acanthohelea) spinellii n. sp. Male. 20. Abdomen and genitalia, ventral view. 21. Genitalia, ventral view. 22. Aedeagus, ventral view. 23. Parameres, ventral view. Scales: 20 = 0.1 mm; 21 = 0.064 mm; 22, 23 = 0.04 mm." httpUri="https://zenodo.org/record/832780/files/figure.png" pageId="7" pageNumber="408">Fig. 20</figureCitation>
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). Dark brown. Genitalia (
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<figureCitation id="13722A1DFF95F46FFCEFF90207BDC2CB" box="[833,970,1743,1768]" captionStart="FIGURES 20 – 23" captionStartId="10.[151,264,1788,1811]" captionTargetBox="[193,1420,485,1747]" captionTargetId="figure@10.[151,1436,468,1776]" captionTargetPageId="10" captionText="FIGURES 20 – 23. Stilobezzia (Acanthohelea) spinellii n. sp. Male. 20. Abdomen and genitalia, ventral view. 21. Genitalia, ventral view. 22. Aedeagus, ventral view. 23. Parameres, ventral view. Scales: 20 = 0.1 mm; 21 = 0.064 mm; 22, 23 = 0.04 mm." httpUri="https://zenodo.org/record/832780/files/figure.png" pageId="7" pageNumber="408">Figs. 20–21</figureCitation>
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). Tergite 9 tapered on distal 1/3, posterior margin rounded, extending to apices of parameres; sternite 9 2.8 x broader than long, posteromedian excavation very deep, V- or U-shaped; sternite 10 narrow, not produced beyond bases of cerci. Gonocoxite stout, 1.60 x longer than greatest breadth, proximal ½ broad, inner margin with row of 5 long setae, distal half tapered, apex rounded. Gonostylus 0.80 length of gonocoxite; proximal 2/3 nearly straight; distal 1/3 curved, tapering to slightly pointed tip. Parameres (
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<figureCitation id="13722A1DFF95F46FFEFDF8490592C3BF" box="[339,485,1924,1949]" captionStart="FIGURES 20 – 23" captionStartId="10.[151,264,1788,1811]" captionTargetBox="[193,1420,485,1747]" captionTargetId="figure@10.[151,1436,468,1776]" captionTargetPageId="10" captionText="FIGURES 20 – 23. Stilobezzia (Acanthohelea) spinellii n. sp. Male. 20. Abdomen and genitalia, ventral view. 21. Genitalia, ventral view. 22. Aedeagus, ventral view. 23. Parameres, ventral view. Scales: 20 = 0.1 mm; 21 = 0.064 mm; 22, 23 = 0.04 mm." httpUri="https://zenodo.org/record/832780/files/figure.png" pageId="7" pageNumber="408">Figs. 21, 23</figureCitation>
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) separate, sub-cylindrical, sub-parallel; basal section ovoidal with extreme base pointed; proximal 2/3 broadening distally; distal 1/3 tapered with elongate apicolateral concavities and blunt tips with a few tiny spicules. Aedeagus (
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<figureCitation id="13722A1DFF95F46FFD93F80106B1C3C7" box="[573,710,1996,2021]" captionStart="FIGURES 20 – 23" captionStartId="10.[151,264,1788,1811]" captionTargetBox="[193,1420,485,1747]" captionTargetId="figure@10.[151,1436,468,1776]" captionTargetPageId="10" captionText="FIGURES 20 – 23. Stilobezzia (Acanthohelea) spinellii n. sp. Male. 20. Abdomen and genitalia, ventral view. 21. Genitalia, ventral view. 22. Aedeagus, ventral view. 23. Parameres, ventral view. Scales: 20 = 0.1 mm; 21 = 0.064 mm; 22, 23 = 0.04 mm." httpUri="https://zenodo.org/record/832780/files/figure.png" pageId="7" pageNumber="408">Figs. 21–22</figureCitation>
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) composed of 2 sinuous, stout, sclerites; basal arm moderately slender, heavily sclerotized with recurved apices; mid portion broad, wing-like, with heavily sclerotized apicomesal sections; distal portion slender, with convergent pointed tips that overlap.
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</paragraph>
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<paragraph id="8BF63698FF9AF460FF69FF12055DC6F6" blockId="8.[151,1437,151,725]" pageId="8" pageNumber="409">
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<emphasis id="B93DEA8AFF9AF460FF69FF120569C4DB" bold="true" box="[199,286,223,248]" pageId="8" pageNumber="409">Female</emphasis>
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. Similar to male with the following notable sexual differences. Head (Fig. 9). Eyes larger, more broadly contiguous. Antennal flagellomeres (Figs. 11a–b) greatly elongate, dark brown, bases paler; antennal ratio 1.06–1.07 (1.06, n=2). Palpus (Fig. 13) brown; segment 3 longer, slenderer with slightly deeper pit; palpal ratio 3.50–3.70 (
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<geoCoordinate id="EE7D505FFF9AF460FEB9FE810515C547" box="[279,354,332,357]" direction="north" orientation="latitude" pageId="8" pageNumber="409" precision="555" value="3.6">3.60 n</geoCoordinate>
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=2). Mandible with 6 teeth. Thorax. Scutellum with 6 large setae. Legs (
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<figureCitation id="13722A1DFF9AF460FB35FE810081C547" box="[1179,1270,332,357]" captionStart="FIGURES 15 – 19" captionStartId="9.[151,264,1462,1485]" captionTargetBox="[177,1417,224,1418]" captionTargetId="figure@9.[151,1436,193,1442]" captionTargetPageId="9" captionText="FIGURES 15 – 19. Stilobezzia (Acanthohelea) spinellii n. sp. Male: 15 a – c – 16. Female: 17 – 19. 15, 18. Legs, lateral view. a. Fore leg. b. Mid leg. c. Hind leg. 16 – 17. Wings. 19. Abdominal segments 8 – 10 and spermathecae, ventral view. Scales: 0.1 mm. Abbreviations: A = anal vein; c = costal vein; CuA 1 and CuA 2 = cubital veins, anterior branches; CuP = cubital vein, posterior branch; M 1 and M 2 = medial veins; r-m = radial-medial crossvein; R 1 = radial vein, first longitudinal vein; R 3 = radial vein, second longitudinal vein. Scales: 0.1 mm." httpUri="https://zenodo.org/record/832778/files/figure.png" pageId="8" pageNumber="409">Figs. 18</figureCitation>
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a–c) similar to male; hind tibial comb with 8 large spines; prothoracic tarsal ratio 2.20 (n=2); mesothoracic tarsal ratio 2.60–2.80 (2.70, n=2); metathoracic tarsal ratio 2.20 (n=2); claws shorter than their respective tarsomeres 5, talon of fore leg twice as long as basal tooth, talon of mid leg
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<date id="FFF71058FF9AF460FD0DFE7506BFC5EC" box="[675,712,440,464]" pageId="8" pageNumber="409">4 x</date>
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longer than basal tooth, talon of hind leg
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<date id="FFF71058FF9AF460FB02FE7500A6C5EC" box="[1196,1233,440,464]" pageId="8" pageNumber="409">3 x</date>
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longer than basal tooth. Wing (
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<figureCitation id="13722A1DFF9AF460FE85FE11050CC5D7" box="[299,379,476,501]" captionStart="FIGURES 15 – 19" captionStartId="9.[151,264,1462,1485]" captionTargetBox="[177,1417,224,1418]" captionTargetId="figure@9.[151,1436,193,1442]" captionTargetPageId="9" captionText="FIGURES 15 – 19. Stilobezzia (Acanthohelea) spinellii n. sp. Male: 15 a – c – 16. Female: 17 – 19. 15, 18. Legs, lateral view. a. Fore leg. b. Mid leg. c. Hind leg. 16 – 17. Wings. 19. Abdominal segments 8 – 10 and spermathecae, ventral view. Scales: 0.1 mm. Abbreviations: A = anal vein; c = costal vein; CuA 1 and CuA 2 = cubital veins, anterior branches; CuP = cubital vein, posterior branch; M 1 and M 2 = medial veins; r-m = radial-medial crossvein; R 1 = radial vein, first longitudinal vein; R 3 = radial vein, second longitudinal vein. Scales: 0.1 mm." httpUri="https://zenodo.org/record/832778/files/figure.png" pageId="8" pageNumber="409">Fig. 17</figureCitation>
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) darkly infuscated; 2nd radial cell
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<date id="FFF71058FF9AF460FCADFE110752C5D7" box="[771,805,476,500]" pageId="8" pageNumber="409">3 x</date>
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longer than 1st; macrotrichia sparse on costa, R1, R2, R3, M1, M2, abundant on distal portions of cells r3, m1; wing length 1.75–1.77 (1.76, n=2) mm, width 0.62–0.65 (0.63, n=2) mm; costal ratio 0.78–0.79 (0.78, n=2). Abdomen (
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<figureCitation id="13722A1DFF9AF460FCB0FDE40704C661" box="[798,883,553,578]" captionStart="FIGURES 15 – 19" captionStartId="9.[151,264,1462,1485]" captionTargetBox="[177,1417,224,1418]" captionTargetId="figure@9.[151,1436,193,1442]" captionTargetPageId="9" captionText="FIGURES 15 – 19. Stilobezzia (Acanthohelea) spinellii n. sp. Male: 15 a – c – 16. Female: 17 – 19. 15, 18. Legs, lateral view. a. Fore leg. b. Mid leg. c. Hind leg. 16 – 17. Wings. 19. Abdominal segments 8 – 10 and spermathecae, ventral view. Scales: 0.1 mm. Abbreviations: A = anal vein; c = costal vein; CuA 1 and CuA 2 = cubital veins, anterior branches; CuP = cubital vein, posterior branch; M 1 and M 2 = medial veins; r-m = radial-medial crossvein; R 1 = radial vein, first longitudinal vein; R 3 = radial vein, second longitudinal vein. Scales: 0.1 mm." httpUri="https://zenodo.org/record/832778/files/figure.png" pageId="8" pageNumber="409">Fig. 19</figureCitation>
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). Brown. Sternite 9 dark brown; anterior margin straight, posteromedian excavation broad, V-shaped, extending 1/3 of total sternite length. Sternite 10 short with 4 pairs of large setae; cercus rounded. Two elongate ovoid, heavily sclerotized spermathecae with hyaline punctuations, measuring 0.075 by
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||
<quantity id="4CB19B7DFF9AF460FD96FD5406D1C693" box="[568,678,665,689]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="409" unit="mm" value="0.32">0.32 mm</quantity>
|
||
, 0.055 by
|
||
<quantity id="4CB19B7DFF9AF460FC9CFD5407DBC693" box="[818,940,665,689]" metricMagnitude="-5" metricUnit="m" metricValue="4.2" pageId="8" pageNumber="409" unit="mm" value="0.042">0.042 mm</quantity>
|
||
with short necks, and a 3rd rudimentary spermatheca.
|
||
</paragraph>
|
||
<paragraph id="8BF63698FF9AF460FF39F85A07A2C3C8" blockId="8.[151,1437,1943,2027]" pageId="8" pageNumber="409">
|
||
<emphasis id="B93DEA8AFF9AF460FF39F85A0530C38E" bold="true" box="[151,327,1943,1966]" pageId="8" pageNumber="409">FIGURES 8–14.</emphasis>
|
||
<taxonomicName id="4C494D1BFF9AF460FEE3F85506C2C38D" authority="Huerta & Grogan, 2017" authorityName="Huerta & Grogan" authorityYear="2017" box="[333,693,1944,1966]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="spinellii" status="sp. nov." subGenus="Acanthohelea">
|
||
<emphasis id="B93DEA8AFF9AF460FEE3F85506C2C38D" box="[333,693,1944,1966]" italics="true" pageId="8" pageNumber="409">Stilobezzia (Acanthohelea) spinellii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20E57F1FF9AF460FD15F8550683C38E" box="[699,756,1944,1965]" pageId="8" pageNumber="409" rank="species">
|
||
<emphasis id="B93DEA8AFF9AF460FD15F8550683C38E" bold="true" box="[699,756,1944,1965]" pageId="8" pageNumber="409">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Male: 8, 10, 12, 14. Female: 9, 11a–b, 13. 8–9. Head, frontal view. 10–11b. Flagellomeres XI–XIII, lateral view. 11a. Flagellomeres I–X, lateral view. 12–13. Palpus, lateral view. 14. Thorax, lateral view. Scales: 8, 9, 14 =
|
||
<quantity id="4CB19B7DFF9AF460FE63F8180669C3C8" box="[461,542,2005,2027]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="409" unit="mm" value="0.1">0.1 mm</quantity>
|
||
; 10, 11a–b =
|
||
<quantity id="4CB19B7DFF9AF460FD08F8180767C3C8" box="[678,784,2005,2027]" metricMagnitude="-5" metricUnit="m" metricValue="6.4" pageId="8" pageNumber="409" unit="mm" value="0.064">0.064 mm</quantity>
|
||
; 12, 13 =
|
||
<quantity id="4CB19B7DFF9AF460FCDAF81807A6C3C8" box="[884,977,2005,2027]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="409" unit="mm" value="0.04">0.04 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF366610FF9BF461FF39FA7B0600C26E" httpUri="https://zenodo.org/record/832778/files/figure.png" pageId="9" pageNumber="410" targetBox="[177,1417,224,1418]" targetPageId="9">
|
||
<paragraph id="8BF63698FF9BF461FF39FA7B0600C26E" blockId="9.[151,1436,1462,1613]" pageId="9" pageNumber="410">
|
||
<emphasis id="B93DEA8AFF9BF461FF39FA7B0522C1EF" bold="true" box="[151,341,1462,1485]" pageId="9" pageNumber="410">FIGURES 15–19.</emphasis>
|
||
<taxonomicName id="4C494D1BFF9BF461FEF0FA7A06BDC1EE" authority="Huerta & Grogan, 2017" authorityName="Huerta & Grogan" authorityYear="2017" box="[350,714,1463,1485]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="spinellii" status="sp. nov." subGenus="Acanthohelea">
|
||
<emphasis id="B93DEA8AFF9BF461FEF0FA7A06BDC1EE" box="[350,714,1463,1485]" italics="true" pageId="9" pageNumber="410">Stilobezzia (Acanthohelea) spinellii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20E57F1FF9BF461FD7CFA7A0779C1EF" box="[722,782,1463,1484]" pageId="9" pageNumber="410" rank="species">
|
||
<emphasis id="B93DEA8AFF9BF461FD7CFA7A0779C1EF" bold="true" box="[722,782,1463,1484]" pageId="9" pageNumber="410">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Male: 15a–c–16. Female: 17–19. 15, 18. Legs, lateral view. a. Fore leg. b. Mid leg. c. Hind leg. 16–17. Wings. 19. Abdominal segments 8–10 and spermathecae, ventral view. Scales: 0.1 mm. Abbreviations: A = anal vein; c = costal vein; CuA1 and CuA2 = cubital veins, anterior branches; CuP = cubital vein, posterior branch; M1 and M2 = medial veins; r-m = radial-medial crossvein; R1 = radial vein, first longitudinal vein; R3 = radial vein, second longitudinal vein. Scales: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3536513FF9BF461FF69F9B406ECC2B1" box="[199,667,1657,1682]" pageId="9" pageNumber="410" type="distribution">
|
||
<paragraph id="8BF63698FF9BF461FF69F9B406ECC2B1" blockId="9.[151,1436,1657,2006]" box="[199,667,1657,1682]" pageId="9" pageNumber="410">
|
||
<emphasis id="B93DEA8AFF9BF461FF69F9B40517C2B1" bold="true" box="[199,352,1657,1682]" pageId="9" pageNumber="410">Distribution.</emphasis>
|
||
<collectingRegion id="498DF87AFF9BF461FEC9F9B405CBC2B1" box="[359,444,1657,1682]" country="Mexico" name="Mexico" pageId="9" pageNumber="410">
|
||
<collectingCountry id="F35E7608FF9BF461FEC9F9B405CBC2B1" box="[359,444,1657,1682]" name="Mexico" pageId="9" pageNumber="410">Mexico</collectingCountry>
|
||
</collectingRegion>
|
||
(
|
||
<collectingRegion id="498DF87AFF9BF461FE65F9B70655C2B1" box="[459,546,1658,1682]" country="Mexico" name="Oaxaca" pageId="9" pageNumber="410">Oaxaca</collectingRegion>
|
||
) (
|
||
<figureCitation id="13722A1DFF9BF461FD97F9B406F9C2B1" box="[569,654,1657,1682]" captionStart="FIGURE 24" captionStartId="12.[151,250,1623,1645]" captionTargetBox="[195,1386,588,1581]" captionTargetId="figure@12.[151,1436,555,1601]" captionTargetPageId="12" captionText="FIGURE 24. Distribution of Schizonyxhelea zoologica n. sp. (solid rectangles), and Stilobezzia (Acanthohelea) spinellii n. sp. (solid circles) in Mexico." httpUri="https://zenodo.org/record/832782/files/figure.png" pageId="9" pageNumber="410">Fig. 24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3536513FF9BF461FF69F95005DDC2DD" pageId="9" pageNumber="410" type="etymology">
|
||
<paragraph id="8BF63698FF9BF461FF69F95005DDC2DD" blockId="9.[151,1436,1657,2006]" pageId="9" pageNumber="410">
|
||
<emphasis id="B93DEA8AFF9BF461FF69F9500539C295" bold="true" box="[199,334,1693,1718]" pageId="9" pageNumber="410">Etymology.</emphasis>
|
||
We are pleased to name this new species after our good friend and colleague Gustavo R. Spinelli, Museo de La Plata,
|
||
<collectingCountry id="F35E7608FF9BF461FED3F90C0598C2F9" box="[381,495,1729,1754]" name="Argentina" pageId="9" pageNumber="410">Argentina</collectingCountry>
|
||
, in recognition of his numerous important contributions to our knowledge of New World
|
||
<taxonomicName id="4C494D1BFF9BF461FF4DF92805D1C2DD" authorityName="Newman" authorityYear="1834" box="[227,422,1765,1790]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3536513FF9BF461FF69F8C40499C3AD" pageId="9" pageNumber="410" type="materials_examined">
|
||
<paragraph id="8BF63698FF9BF461FF69F8C40499C3AD" blockId="9.[151,1436,1657,2006]" pageId="9" pageNumber="410">
|
||
<materialsCitation id="3B213CC5FF9BF461FF69F8C405B3C349" ID-GBIF-Occurrence="1576193804" collectingDate="2009-07-23" collectingDateMax="2009-07-29" collectingDateMin="2009-07-23" collectionCode="CDC, CAIM" collectorName="B. Salceda-Sanchez & A. Rodriguez-Atanacio & J. Ordonez-Alvarez" country="Mexico" location="Santo Domingo Petapa" municipality="Localidad La Esperanza" pageId="9" pageNumber="410" specimenCount="3" stateProvince="Oaxaca" typeStatus="holotype">
|
||
<emphasis id="B93DEA8AFF9BF461FF69F8C4050FC301" bold="true" box="[199,376,1801,1826]" pageId="9" pageNumber="410">
|
||
<typeStatus id="54F2883AFF9BF461FF69F8C40575C301" box="[199,258,1801,1826]" pageId="9" pageNumber="410">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="54F2883AFF9BF461FE2DF8C4059CC301" box="[387,491,1801,1826]" pageId="9" pageNumber="410" type="holotype">Holotype</typeStatus>
|
||
male,
|
||
<typeStatus id="54F2883AFF9BF461FD90F8C406EFC301" box="[574,664,1801,1826]" pageId="9" pageNumber="410" type="allotype">allotype</typeStatus>
|
||
female,
|
||
<collectingCountry id="F35E7608FF9BF461FCAEF8C70706C301" box="[768,881,1802,1826]" name="Mexico" pageId="9" pageNumber="410">MEXICO</collectingCountry>
|
||
,
|
||
<collectingRegion id="498DF87AFF9BF461FC2EF8C707A0C302" box="[896,983,1802,1825]" country="Mexico" name="Oaxaca" pageId="9" pageNumber="410">
|
||
<emphasis id="B93DEA8AFF9BF461FC2EF8C707A0C302" box="[896,983,1802,1825]" italics="true" pageId="9" pageNumber="410">Oaxaca</emphasis>
|
||
</collectingRegion>
|
||
,
|
||
<location id="8E966043FF9BF461FC47F8C70081C302" LSID="urn:lsid:plazi:treatment:03E0878EFF95F462FF39FBD70798C58E:8E966043FF9BF461FC47F8C70081C302" box="[1001,1270,1801,1826]" country="Mexico" municipality="Localidad La Esperanza" name="Santo Domingo Petapa" pageId="9" pageNumber="410" stateProvince="Oaxaca">Santo Domingo Petapa</location>
|
||
,
|
||
<collectingMunicipality id="6B92ACE2FF9BF461FAAAF8C4057AC366" pageId="9" pageNumber="410">Localidad La Esperanza</collectingMunicipality>
|
||
,
|
||
<collectingDate id="EFB3E9B0FF9BF461FEB4F8E005BCC365" box="[282,459,1837,1862]" pageId="9" pageNumber="410" value="2009-07-23" valueMax="2009-07-29" valueMin="2009-07-23">
|
||
<date id="FFF71058FF9BF461FEB4F8E005BCC365" box="[282,459,1837,1862]" pageId="9" pageNumber="410" value="2009-07-23" valueMax="2009-07-29" valueMin="2009-07-23">23–29-jul-2009</date>
|
||
</collectingDate>
|
||
, cerca a la localidad, trampa
|
||
<collectionCode id="ED58AE5DFF9BF461FCB1F8E3072CC365" box="[799,859,1838,1862]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12786" name="Changdu Institute for Drug Control" pageId="9" pageNumber="410">CDC</collectionCode>
|
||
,
|
||
<collectorName id="26BC534EFF9BF461FCC7F8E3003BC365" box="[873,1100,1837,1862]" pageId="9" pageNumber="410">B. Salceda-Sánchez</collectorName>
|
||
,
|
||
<collectorName id="26BC534EFF9BF461FBF4F8E30115C365" box="[1114,1378,1837,1862]" pageId="9" pageNumber="410">A. Rodríguez-Atanacio</collectorName>
|
||
&
|
||
<collectorName id="26BC534EFF9BF461FA24F8E3052EC349" pageId="9" pageNumber="410">J. Ordoñez-Álvarez</collectorName>
|
||
(
|
||
<collectionCode id="ED58AE5DFF9BF461FEC2F89F05CCC349" box="[364,443,1874,1898]" country="Mexico" httpUri="http://grbio.org/cool/z08b-033r" name="Collection of Aquatic Important Microorganisms" pageId="9" pageNumber="410">CAIM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B213CC5FF9BF461FE7CF89F049DC3AD" ID-GBIF-Occurrence="1576193821" collectingDate="2009-07-23" collectingDateMax="2009-07-29" collectingDateMin="2009-07-23" collectionCode="CAIM, FSCA" collectorName="B. Salceda-Sanchez & A. Rodriguez-Atanacio & J. Ordonez-Alvarez" country="Mexico" location="Santo Domingo Petapa" municipality="Localidad La Esperanza" pageId="9" pageNumber="410" specimenCount="8" specimenCount-female="1" specimenCount-male="7" stateProvince="Oaxaca" typeStatus="paratype">
|
||
<typeStatus id="54F2883AFF9BF461FE7CF89F0635C349" box="[466,578,1874,1898]" pageId="9" pageNumber="410" type="paratype">Paratypes</typeStatus>
|
||
,
|
||
<specimenCount id="9D4FFD11FF9BF461FDFFF89C06D8C349" box="[593,687,1873,1898]" pageId="9" pageNumber="410" type="male">5 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D4FFD11FF9BF461FD10F89F0751C349" box="[702,806,1873,1898]" pageId="9" pageNumber="410" type="female">1 female</specimenCount>
|
||
, same data as holotype (
|
||
<collectionCode id="ED58AE5DFF9BF461FBE2F89F00EDC349" box="[1100,1178,1874,1898]" country="Mexico" httpUri="http://grbio.org/cool/z08b-033r" name="Collection of Aquatic Important Microorganisms" pageId="9" pageNumber="410">CAIM</collectionCode>
|
||
;
|
||
<specimenCount id="9D4FFD11FF9BF461FB04F89F0173C349" box="[1194,1284,1873,1898]" pageId="9" pageNumber="410" type="male">2 males</specimenCount>
|
||
deposited in
|
||
<collectionCode id="ED58AE5DFF9BF461FF39F8BB0495C3AD" box="[151,226,1910,1934]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="9" pageNumber="410">FSCA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3536513FF9BF462FF69F8540798C58E" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410" type="discussion">
|
||
<paragraph id="8BF63698FF9BF461FF69F85401EBC3F6" blockId="9.[151,1436,1657,2006]" pageId="9" pageNumber="410">
|
||
<emphasis id="B93DEA8AFF9BF461FF69F854053BC391" bold="true" box="[199,332,1945,1970]" pageId="9" pageNumber="410">Discussion.</emphasis>
|
||
The aedeagus and parameres of males of this new species are similar to those of males of
|
||
<taxonomicName id="4C494D1BFF9BF461FA29F85706DDC3F5" authority="Lane & Forattini" authorityName="Lane & Forattini" authorityYear="1956" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="atrichopogon" subGenus="Acanthohelea">
|
||
<emphasis id="B93DEA8AFF9BF461FA29F85701E2C392" box="[1415,1429,1946,1969]" italics="true" pageId="9" pageNumber="410">S</emphasis>
|
||
.
|
||
<emphasis id="B93DEA8AFF9BF461FF39F870054DC3F5" box="[151,314,1981,2006]" italics="true" pageId="9" pageNumber="410">(Acanthohelea</emphasis>
|
||
)
|
||
<emphasis id="B93DEA8AFF9BF461FEE5F8700595C3F5" box="[331,482,1981,2006]" italics="true" pageId="9" pageNumber="410">atrichopogon</emphasis>
|
||
Lane & Forattini
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C494D1BFF9BF461FD18F873006CC3F5" authority="Lane & Forattini" authorityName="Lane & Forattini" authorityYear="1956" box="[694,1051,1981,2006]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="tibialis" subGenus="Acanthohelea">
|
||
<emphasis id="B93DEA8AFF9BF461FD18F8730724C3F5" box="[694,851,1981,2006]" italics="true" pageId="9" pageNumber="410">S. (A.) tibialis</emphasis>
|
||
Lane & Forattini
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C494D1BFF9BF461FBF6F87301EBC3F6" authority="Grogan et al." authorityName="Grogan et al." box="[1112,1436,1981,2006]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="thomasi" subGenus="Acanthohelea">
|
||
<emphasis id="B93DEA8AFF9BF461FBF6F87300E1C3F6" box="[1112,1174,1982,2005]" italics="true" pageId="9" pageNumber="410">S. (A.</emphasis>
|
||
)
|
||
<emphasis id="B93DEA8AFF9BF461FB09F8700177C3F5" box="[1191,1280,1981,2006]" italics="true" pageId="9" pageNumber="410">thomasi</emphasis>
|
||
Grogan
|
||
<emphasis id="B93DEA8AFF9BF461FACDF87201E2C3F5" box="[1379,1429,1981,2006]" italics="true" pageId="9" pageNumber="410">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BF63698FF98F462FF39FF550129C53E" blockId="10.[151,1436,151,429]" pageId="10" pageNumber="411">
|
||
<bibRef id="52E9A26EFF98F462FF39FF550129C53E" pageId="10" pageNumber="411" type="book chapter">
|
||
(
|
||
<year id="FFE177CBFF98F462FF31FF5504ABC493" box="[159,220,152,176]" pageId="10" pageNumber="411">2013</year>
|
||
).
|
||
<title id="F94DDDAEFF98F462FF41FF550737C493" box="[239,832,151,176]" jopScore="0.05557152" pageId="10" pageNumber="411" type="_" typeScore="71">However, the most notable differences between these</title>
|
||
<pagination id="B280C86CFF98F462FCE9FF550722C493" box="[839,853,152,176]" pageId="10" pageNumber="411" type="pagination">3</pagination>
|
||
species are the shapes and orientations of the distalmost portions of their parameres.
|
||
<journalOrPublisher id="51883C54FF98F462FDB5FF70012DC53E" jopScore="0.6547942" pageId="10" pageNumber="411">
|
||
For example, in
|
||
<emphasis id="B93DEA8AFF98F462FD76FF7007B6C4F6" box="[728,961,188,213]" italics="true" pageId="10" pageNumber="411">S. (A.) atrichopogon</emphasis>
|
||
they are bifurcate, with greatly divergent apices; in
|
||
<emphasis id="B93DEA8AFF98F462FEA6FF2D05D4C4DB" box="[264,419,223,248]" italics="true" pageId="10" pageNumber="411">S. (A.) tibialis</emphasis>
|
||
they are rounded; in
|
||
<emphasis id="B93DEA8AFF98F462FD3EFF2D06BAC4D4" box="[656,717,224,247]" italics="true" pageId="10" pageNumber="411">S. (A.</emphasis>
|
||
)
|
||
<emphasis id="B93DEA8AFF98F462FD73FF120741C4DB" box="[733,822,223,248]" italics="true" pageId="10" pageNumber="411">thomasi</emphasis>
|
||
they are rounded with a hyaline pointed process that is adpressed to the ventral surface; whereas in this new Mexican species, they are blunt, with tiny apical spines
|
||
</journalOrPublisher>
|
||
.
|
||
</bibRef>
|
||
</paragraph>
|
||
<paragraph id="8BF63698FF98F462FF69FEEA0798C58E" blockId="10.[151,1436,151,429]" pageId="10" pageNumber="411">
|
||
Females of this new species are very similar to females of
|
||
<taxonomicName id="4C494D1BFF98F462FCFAFEE5004EC563" authorityName="Lane & Forattini" authorityYear="1956" box="[852,1081,295,320]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="atrichopogon" subGenus="Acanthohelea">
|
||
<emphasis id="B93DEA8AFF98F462FCFAFEE5004EC563" box="[852,1081,295,320]" italics="true" pageId="10" pageNumber="411">S. (A.) atrichopogon</emphasis>
|
||
</taxonomicName>
|
||
, but in that species, the legs are yellow with slightly darker femorotibial joints. The legs of this new species are also similar to those of females of
|
||
<taxonomicName id="4C494D1BFF98F462FF39FEBD0544C5AB" authorityName="Lane & Forattini" authorityYear="1956" box="[151,307,367,392]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="tibialis" subGenus="Acanthohelea">
|
||
<emphasis id="B93DEA8AFF98F462FF39FEBD0544C5AB" box="[151,307,367,392]" italics="true" pageId="10" pageNumber="411">S. (A.) tibialis</emphasis>
|
||
</taxonomicName>
|
||
. However, the wing of
|
||
<taxonomicName id="4C494D1BFF98F462FD93FEBD06ADC5AB" authorityName="Lane & Forattini" authorityYear="1956" box="[573,730,367,392]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="tibialis" subGenus="Acanthohelea">
|
||
<emphasis id="B93DEA8AFF98F462FD93FEBD06ADC5AB" box="[573,730,367,392]" italics="true" pageId="10" pageNumber="411">S. (A.) tibialis</emphasis>
|
||
</taxonomicName>
|
||
has an irregular spot at the level of the r-m crossvein, whereas in females of this new Mexican species, the entire wing is darkly infuscated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |