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<document id="D10FCB62591D4BB405EB55C41C185977" ID-CLB-Dataset="22182" ID-DOI="10.11646/zootaxa.4774.1.1" ID-GBIF-Dataset="269cf599-428d-464d-8f21-994d69110c5c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3825140" ID-ZooBank="B0C4A2F8-F2AB-4147-BB12-63720EEF2516" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="admin" checkinTime="1589391149229" checkinUser="plazi" docAuthor="Ekins, Merrick, Erpenbeck, Dirk &amp; Hooper, John N. A." docDate="2020" docId="03D287B691C23B97FF7EFD62FC75FD36" docLanguage="en" docName="zootaxa.4774.1.1.imf" docOrigin="Zootaxa 4774 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Lycopodina nikitawimandi Ekins &amp; Erpenbeck &amp; Hooper 2020, sp. nov." docType="treatment" docUuid="F5AAEA15-DBAF-4A21-A2B9-E336733E613D" docUuidSource="ZooBank" docVersion="8" lastPageNumber="144" masterDocId="FFEBFFCE914E3B18FFE9FFBBFFF8FF9D" masterDocTitle="Carnivorous sponges from the Australian Bathyal and Abyssal zones collected during the RV Investigator 2017 Expedition" masterLastPageNumber="159" masterPageNumber="1" pageNumber="141" updateTime="1720375898568" updateUser="admin">
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<mods:title id="77095ADBDCD89175EC5EB2E44DC3955F">Carnivorous sponges from the Australian Bathyal and Abyssal zones collected during the RV Investigator 2017 Expedition</mods:title>
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<mods:namePart id="79D1C5972AAA40F2408E026215820223">Ekins, Merrick</mods:namePart>
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<mods:namePart id="36394ABA70C31962AF7934AF03973270">Erpenbeck, Dirk</mods:namePart>
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<mods:namePart id="FD7E314247E4BD9BAA876DBCF83579B0">Hooper, John N. A.</mods:namePart>
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<treatment id="03D287B691C23B97FF7EFD62FC75FD36" ID-DOI="http://doi.org/10.5281/zenodo.3846429" ID-GBIF-Taxon="164131173" ID-Zenodo-Dep="3846429" LSID="urn:lsid:zoobank.org:act:F5AAEA15-DBAF-4A21-A2B9-E336733E613D" httpUri="http://treatment.plazi.org/id/03D287B691C23B97FF7EFD62FC75FD36" lastPageId="143" lastPageNumber="144" pageId="140" pageNumber="141">
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<paragraph id="8BC436A091C23B94FF7EFD62FDC7FD6E" blockId="140.[151,988,729,827]" box="[151,575,729,756]" pageId="140" pageNumber="141">
<heading id="D08C81CC91C23B94FF7EFD62FDC7FD6E" bold="true" box="[151,575,729,756]" fontSize="11" level="1" pageId="140" pageNumber="141" reason="1">
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<taxonomicName id="4C7B4D2391C23B94FF7EFD62FE21FD69" ID-CoL="86X39" authority="Ekins &amp; Erpenbeck &amp; Hooper, 2020" authorityName="Ekins &amp; Erpenbeck &amp; Hooper" authorityYear="2020" box="[151,473,729,756]" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="140" pageNumber="141" phylum="Porifera" rank="species" species="nikitawimandi" status="sp. nov.">
<emphasis id="B90FEAB291C23B94FF7EFD62FE21FD69" bold="true" box="[151,473,729,756]" italics="true" pageId="140" pageNumber="141">Lycopodina nikitawimandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57C991C23B94FE08FD62FDC7FD6E" box="[481,575,729,755]" pageId="140" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C361652B91C23B94FF7EFD46FC24FCA6" pageId="140" pageNumber="141" type="description">
<paragraph id="8BC436A091C23B94FF7EFD46FDFFFC8A" blockId="140.[151,988,729,827]" box="[151,519,765,791]" pageId="140" pageNumber="141">
<figureCitation id="13402A2591C23B94FF7EFD46FEF6FC8A" box="[151,270,765,791]" captionStart="FIGURE 27" captionStartId="141.[151,250,1078,1103]" captionTargetBox="[189,1398,181,1053]" captionTargetId="figure@141.[189,1398,181,1053]" captionTargetPageId="141" captionText="FIGURE 27. Lycopodina nikitawimandi sp. nov. QM G337534. A. Holotype QM G337534. B. Palmate anisochelae. C. Large mycalostyle 1. D. Magnified ends of the mycalostyle 1 illustrated in C. E. Small mycalostyle 2. F. Magnified ends of the smaller mycalostyle 2 illustrated in E." figureDoi="http://doi.org/10.5281/zenodo.3825198" httpUri="https://zenodo.org/record/3825198/files/figure.png" pageId="140" pageNumber="141">Figures 27</figureCitation>
&amp;
<figureCitation id="13402A2591C23B94FEDBFD46FEA8FC8A" box="[306,336,765,791]" captionStart="FIGURE 28" captionStartId="142.[151,250,1651,1676]" captionTargetBox="[151,1437,181,1623]" captionTargetId="figure@142.[151,1437,181,1625]" captionTargetPageId="142" captionText="FIGURE 28. Lycopodina nikitawimandi sp. nov. A. Preserved specimens of paratypes QM G337510 and QM G337534 still attached to the hexactinellid skeleton substrate.Arrows pointing to carnivorous sponges. B. Copepod trapped in isochelae, arrow pointing to the copepod. C. Bottlebrush body of the holotype QM G337534, showing the many filaments and the anisochelae in the ectosome. D. Close-up of the filaments branching from the stem. E. Filaments showing the ectosomal clusters of anisochelae and the underlying mycalostyles. F. Root ball and stem showing the hispid surface from the small mycalostyles." figureDoi="http://doi.org/10.5281/zenodo.3825200" httpUri="https://zenodo.org/record/3825200/files/figure.png" pageId="140" pageNumber="141">28</figureCitation>
,
<tableCitation id="C6F9031B91C23B94FEB2FD46FE3FFC8A" box="[347,455,765,791]" captionStart="TABLE 14" captionStartId="124.[151,239,487,513]" captionTargetBox="[162,2024,564,1671]" captionTargetPageId="124" captionText="TABLE 14. Comparative morphological and distributional data for all known species of Lycopodina." httpUri="http://table.plazi.org/id/DF04662891323B64FF7EFE5CFB29FD9C" pageId="140" pageNumber="141" tableUuid="DF04662891323B64FF7EFE5CFB29FD9C">Tables 14</tableCitation>
&amp;
<tableCitation id="C6F9031B91C23B94FE02FD46FDFFFC8A" box="[491,519,765,791]" captionStart="TABLE 16" captionStartId="141.[151,239,1223,1247]" captionTargetBox="[163,1326,1299,1946]" captionText="TABLE 16. A comparison of spicule measurements (µm) between the different specimens of Lycopodina nikitawimandi sp. nov." httpUri="http://table.plazi.org/id/DF04662891C33B95FF7EFB7CFF15FA9E" pageId="140" pageNumber="141" tableUuid="DF04662891C33B95FF7EFB7CFF15FA9E">16</tableCitation>
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<emphasis id="B90FEAB291C23B94FF7EFCD3FE80FC1E" bold="true" box="[151,376,872,899]" pageId="140" pageNumber="141">Material examined</emphasis>
:
<materialsCitation id="3B133CFD91C23B94FE6FFCD2FC53FC56" ID-GBIF-Occurrence="2617493337" collectingDate="2017-05-19" collectionCode="QM, RV" collectorName="Box Corer &amp; Coll. Merrick Ekins &amp; Sample" country="Australia" county="Station" elevation="2793" latitude="-41.720695" location="Freycinet Peninsular" longLatPrecision="1" longitude="149.1252" municipality="Tasman Sea" pageId="140" pageNumber="141" specimenCode="G337534, V03" specimenCount="1" stateProvince="Tasmania" typeStatus="holotype">
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<specimenCode id="DBDD9EDB91C23B94FDCFFCD3FD77FC1E" box="[550,655,872,899]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="140" pageNumber="141" type="Herbarium">G337534</specimenCode>
off
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,
<collectingMunicipality id="6BA0ACDA91C23B94FC42FCD2FBCAFC1E" box="[939,1074,873,899]" pageId="140" pageNumber="141">Tasman Sea</collectingMunicipality>
,
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11,
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,
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,
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,
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,
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,
<collectorName id="268E537691C23B94FD80FC36FD19FC3A" box="[617,737,909,935]" pageId="140" pageNumber="141">Box Corer</collectorName>
,
<collectorName id="268E537691C23B94FD02FC36FC32FC3A" box="[747,970,909,935]" pageId="140" pageNumber="141">Coll. Merrick Ekins</collectorName>
on
<collectionCode id="ED6AAE6591C23B94FC1DFC35FBEDFC3A" box="[1012,1045,910,935]" country="Italy" name="Collection of Leptospira Strains" pageId="140" pageNumber="141">
<emphasis id="B90FEAB291C23B94FC1DFC35FBEDFC3A" box="[1012,1045,910,935]" italics="true" pageId="140" pageNumber="141">RV</emphasis>
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<emphasis id="B90FEAB291C23B94FBF5FC35FB5BFC3A" box="[1052,1187,910,935]" italics="true" pageId="140" pageNumber="141">Investigator</emphasis>
, Cruise IN2017_
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,
<collectorName id="268E537691C23B94FF7EFC0AFF13FC56" box="[151,235,945,971]" pageId="140" pageNumber="141">Sample</collectorName>
11-110.2,
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<collectingDate id="EF81E98891C23B94FE8BFC0AFE2EFC56" box="[354,470,945,971]" pageId="140" pageNumber="141" value="2017-05-19">19/v/2017</collectingDate>
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, growing on a skeleton of a hexactinellid.
</materialsCitation>
</paragraph>
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<materialsCitation id="3B133CFD91C23B94FF2EFC6EFA6FFC72" ID-GBIF-Occurrence="2617493302" box="[199,1431,980,1007]" collectionCode="QM" collectorName="Sample" pageId="140" pageNumber="141" specimenCode="G337510" specimenCount="2" typeStatus="paratype">
<typeStatus id="54C0880291C23B94FF2EFC6EFEC0FC72" box="[199,312,981,1007]" pageId="140" pageNumber="141" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="ED6AAE6591C23B94FEADFC6EFE89FC72" box="[324,369,981,1007]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="140" pageNumber="141" type="Museum">QM</collectionCode>
<specimenCode id="DBDD9EDB91C23B94FE91FC6FFE19FC72" box="[376,481,980,1007]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="140" pageNumber="141" type="Herbarium">G337510</specimenCode>
same collection details and hexactinellid skeleton as for
<typeStatus id="54C0880291C23B94FB8FFC6EFB33FC72" box="[1126,1227,981,1007]" pageId="140" pageNumber="141" type="holotype">holotype</typeStatus>
,
<collectorName id="268E537691C23B94FB3FFC6EFAD2FC72" box="[1238,1322,981,1007]" pageId="140" pageNumber="141">Sample</collectorName>
11-110.1
</materialsCitation>
;
<materialsCitation id="3B133CFD91C23B94FF7EFC42FB4DFB8E" ID-GBIF-Occurrence="2617493377" box="[151,1205,1016,1043]" collectionCode="QM" collectorName="Sample" pageId="140" pageNumber="141" specimenCode="G337537" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED6AAE6591C23B94FF7EFC42FF3CFB8E" box="[151,196,1017,1043]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="140" pageNumber="141" type="Museum">QM</collectionCode>
<specimenCode id="DBDD9EDB91C23B94FF22FC43FECBFB8E" box="[203,307,1016,1043]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="140" pageNumber="141" type="Herbarium">G337537</specimenCode>
same collection details and hexactinellid skeleton as
<typeStatus id="54C0880291C23B94FC6FFC42FC13FB8E" box="[902,1003,1017,1043]" pageId="140" pageNumber="141" type="holotype">holotype</typeStatus>
,
<collectorName id="268E537691C23B94FC1CFC42FBB1FB8E" box="[1013,1097,1017,1043]" pageId="140" pageNumber="141">Sample</collectorName>
11-110.3
</materialsCitation>
;
<materialsCitation id="3B133CFD91C23B94FB29FC42FC2BFBAA" ID-GBIF-Occurrence="2617493315" collectionCode="QM" collectorName="Sample" pageId="140" pageNumber="141" specimenCode="G337549" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED6AAE6591C23B94FB29FC42FB15FB8E" box="[1216,1261,1017,1043]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="140" pageNumber="141" type="Museum">QM</collectionCode>
<specimenCode id="DBDD9EDB91C23B94FB1DFC43FAA5FB8E" box="[1268,1373,1016,1043]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="140" pageNumber="141" type="Herbarium">G337549</specimenCode>
same collection details and hexactinellid skeleton as
<typeStatus id="54C0880291C23B94FD4AFBA6FCF0FBAA" box="[675,776,1053,1079]" pageId="140" pageNumber="141" type="holotype">holotype</typeStatus>
,
<collectorName id="268E537691C23B94FCFAFBA6FC9FFBAA" box="[787,871,1053,1079]" pageId="140" pageNumber="141">Sample</collectorName>
11-110.4
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;
<materialsCitation id="3B133CFD91C23B94FC37FBA6FC6EFBC6" ID-GBIF-Occurrence="2617493318" collectionCode="QM" collectorName="Sample" pageId="140" pageNumber="141" specimenCode="G337197" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED6AAE6591C23B94FC37FBA6FBF3FBAA" box="[990,1035,1053,1079]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="140" pageNumber="141" type="Museum">QM</collectionCode>
<specimenCode id="DBDD9EDB91C23B94FBFBFBA6FB82FBAA" box="[1042,1146,1053,1079]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="140" pageNumber="141" type="Herbarium">G337197</specimenCode>
same collection details as
<typeStatus id="54C0880291C23B94FF7EFBFAFF01FBC6" box="[151,249,1089,1115]" pageId="140" pageNumber="141" type="holotype">holotype</typeStatus>
but on a different hexactinellid skeleton,
<collectorName id="268E537691C23B94FD3AFBFAFCDFFBC6" box="[723,807,1089,1115]" pageId="140" pageNumber="141">Sample</collectorName>
11-120.1
</materialsCitation>
;
<materialsCitation id="3B133CFD91C23B94FC4AFBFAFC6AFB3E" ID-GBIF-Occurrence="2617493367" collectionCode="QM" collectorName="Brenke Epibenthic Sledge &amp; Coll. Merrick Ekins" country="Australia" county="Station" elevation="4414" latitude="-30.289305" location="Central" longLatPrecision="1" longitude="153.8438" municipality="Tasman Sea" pageId="140" pageNumber="141" specimenCode="G337514" specimenCount="1" stateProvince="New South Wales" typeStatus="paratype">
<collectionCode id="ED6AAE6591C23B94FC4AFBFAFC28FBC6" box="[931,976,1089,1115]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="140" pageNumber="141" type="Museum">QM</collectionCode>
<specimenCode id="DBDD9EDB91C23B94FC31FBFBFBB9FBC6" box="[984,1089,1088,1115]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="140" pageNumber="141" type="Herbarium">G337514</specimenCode>
off
<location id="8EA4607B91C23B94FB98FBFAFB3BFBC6" LSID="urn:lsid:plazi:treatment:03D287B691C23B97FF7EFD62FC75FD36:8EA4607B91C23B94FB98FBFAFB3BFBC6" box="[1137,1219,1089,1115]" country="Australia" county="Station" latitude="-30.289305" longLatPrecision="1" longitude="153.8438" municipality="Tasman Sea" name="Central" pageId="140" pageNumber="141" stateProvince="New South Wales">Central</location>
<collectingRegion id="49BFF84291C23B94FB22FBFAFA61FBC6" box="[1227,1433,1089,1115]" country="Australia" name="New South Wales" pageId="140" pageNumber="141">New South Wales</collectingRegion>
,
<collectingMunicipality id="6BA0ACDA91C23B94FF7EFBDEFEE7FBE2" box="[151,287,1125,1151]" pageId="140" pageNumber="141">Tasman Sea</collectingMunicipality>
,
<collectingCounty id="62A54E2C91C23B94FEC3FBDEFE81FBE2" box="[298,377,1125,1151]" pageId="140" pageNumber="141">Station</collectingCounty>
89,
<collectingRegion id="49BFF84291C23B94FE40FBDEFD8AFBE2" box="[425,626,1125,1151]" country="Australia" name="New South Wales" pageId="140" pageNumber="141">New South Wales</collectingRegion>
,
<collectingCountry id="F36C763091C23B94FD92FBDEFD1DFBE2" box="[635,741,1125,1151]" name="Australia" pageId="140" pageNumber="141">Australia</collectingCountry>
, 30° 15 47.9”–
<geoCoordinate id="EE4F506791C23B94FC7AFBDEFBC5FBE2" box="[915,1085,1124,1151]" degrees="30" direction="south" minutes="17" orientation="latitude" pageId="140" pageNumber="141" precision="1" seconds="21.5" value="-30.289305">30° 17 21.5” S</geoCoordinate>
, 153° 51 31.3”–
<geoCoordinate id="EE4F506791C23B94FB15FBDFFF52FB3E" degrees="153" direction="east" minutes="50" orientation="longitude" pageId="140" pageNumber="141" precision="1" seconds="37.7" value="153.8438">153° 50 37.7” E</geoCoordinate>
, 4436
<quantity id="4C839B4591C23B94FF15FB32FEABFB3E" box="[252,339,1161,1187]" metricMagnitude="3" metricUnit="m" metricValue="4.414" pageId="140" pageNumber="141" unit="m" value="4414.0">
<elevation id="0056D19391C23B94FF15FB32FEABFB3E" box="[252,339,1161,1187]" metricMagnitude="3" metricUnit="m" metricValue="4.414" pageId="140" pageNumber="141" unit="m" value="4414.0">4414 m</elevation>
</quantity>
,
<collectorName id="268E537691C23B94FEB6FB32FD7EFB3E" box="[351,646,1161,1187]" pageId="140" pageNumber="141">Brenke Epibenthic Sledge</collectorName>
,
<collectorName id="268E537691C23B94FD78FB32FC88FB3E" box="[657,880,1161,1187]" pageId="140" pageNumber="141">Coll. Merrick Ekins</collectorName>
on
</materialsCitation>
<collectionCode id="ED6AAE6591C23B94FC70FB31FC42FB3E" box="[921,954,1162,1187]" country="Italy" name="Collection of Leptospira Strains" pageId="140" pageNumber="141">
<emphasis id="B90FEAB291C23B94FC70FB31FC42FB3E" box="[921,954,1162,1187]" italics="true" pageId="140" pageNumber="141">RV</emphasis>
</collectionCode>
<emphasis id="B90FEAB291C23B94FC28FB31FBB0FB3E" box="[961,1096,1162,1187]" italics="true" pageId="140" pageNumber="141">Investigator</emphasis>
, Cruise IN2017_
<materialsCitation id="3B133CFD91C23B94FAE7FB32FD85FB5A" ID-GBIF-Occurrence="2617493312" collectingDate="2017-06-06" collectionCode="V" country="Australia" location="Sample" pageId="140" pageNumber="141" specimenCode="V03" specimenCount="1" stateProvince="New South Wales" typeStatus="paratype">
<specimenCode id="DBDD9EDB91C23B94FAE7FB32FAC5FB3E" box="[1294,1341,1161,1187]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="140" pageNumber="141" type="Museum">V03</specimenCode>
,
<location id="8EA4607B91C23B94FAA1FB32FA64FB3E" LSID="urn:lsid:plazi:treatment:03D287B691C23B97FF7EFD62FC75FD36:8EA4607B91C23B94FAA1FB32FA64FB3E" box="[1352,1436,1161,1187]" country="Australia" name="Sample" pageId="140" pageNumber="141" stateProvince="New South Wales">Sample</location>
89-129,
<date id="FFC5106091C23B94FF1CFB16FE9BFB5A" box="[245,355,1197,1223]" pageId="140" pageNumber="141" value="2017-06-06">
<collectingDate id="EF81E98891C23B94FF1CFB16FE9BFB5A" box="[245,355,1197,1223]" pageId="140" pageNumber="141" value="2017-06-06">6/vi/2017</collectingDate>
</date>
, growing on worm tubes
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361652B91C23B94FF2EFB6AFBF8FB76" box="[199,1024,1233,1259]" pageId="140" pageNumber="141" type="etymology">
<paragraph id="8BC436A091C23B94FF2EFB6AFBF8FB76" blockId="140.[151,1437,872,1836]" box="[199,1024,1233,1259]" pageId="140" pageNumber="141">
<emphasis id="B90FEAB291C23B94FF2EFB6AFEB0FB76" bold="true" box="[199,328,1233,1259]" pageId="140" pageNumber="141">Etymology</emphasis>
: Named for the son of the first author, Nikita Wimandi Ekins.
</paragraph>
</subSubSection>
<subSubSection id="C361652B91C23B94FF2EFB4FFA8EFA92" box="[199,1398,1268,1295]" pageId="140" pageNumber="141" type="distribution">
<paragraph id="8BC436A091C23B94FF2EFB4FFA8EFA92" blockId="140.[151,1437,872,1836]" box="[199,1398,1268,1295]" pageId="140" pageNumber="141">
<emphasis id="B90FEAB291C23B94FF2EFB4FFEA1FA92" bold="true" box="[199,345,1268,1295]" pageId="140" pageNumber="141">Distribution</emphasis>
. Central East coast and central
<collectingRegion id="49BFF84291C23B94FD52FB4EFCD1FA92" box="[699,809,1269,1295]" country="Australia" name="Tasmania" pageId="140" pageNumber="141">Tasmania</collectingRegion>
, Tasman Sea,
<collectingCountry id="F36C763091C23B94FC2FFB4EFBC8FA92" box="[966,1072,1269,1295]" name="Australia" pageId="140" pageNumber="141">Australia</collectingCountry>
, at bathyal to abyssal depths.
</paragraph>
</subSubSection>
<subSubSection id="C361652B91C23B97FF2EFAA3FDE0FF66" lastPageId="143" lastPageNumber="144" pageId="140" pageNumber="141" type="description">
<paragraph id="8BC436A091C23B94FF2EFAA3FEA3FAAE" blockId="140.[151,1437,872,1836]" box="[199,347,1304,1331]" pageId="140" pageNumber="141">
<emphasis id="B90FEAB291C23B94FF2EFAA3FEA3FAAE" bold="true" box="[199,347,1304,1331]" pageId="140" pageNumber="141">Description:</emphasis>
</paragraph>
<paragraph id="8BC436A091C23B94FF2EFA86FDD2F996" blockId="140.[151,1437,872,1836]" pageId="140" pageNumber="141">
<emphasis id="B90FEAB291C23B94FF2EFA86FEA0FACA" box="[199,344,1341,1367]" italics="true" pageId="140" pageNumber="141">Growth form</emphasis>
: This sponge is an erect stipitate sponge with filaments radiating in all directions projecting from the cylindrical stalk (
<figureCitation id="13402A2591C23B94FE69FADAFDECFAE6" box="[384,532,1377,1403]" captionStart="FIGURE 27" captionStartId="141.[151,250,1078,1103]" captionTargetBox="[189,1398,181,1053]" captionTargetId="figure@141.[189,1398,181,1053]" captionTargetPageId="141" captionText="FIGURE 27. Lycopodina nikitawimandi sp. nov. QM G337534. A. Holotype QM G337534. B. Palmate anisochelae. C. Large mycalostyle 1. D. Magnified ends of the mycalostyle 1 illustrated in C. E. Small mycalostyle 2. F. Magnified ends of the smaller mycalostyle 2 illustrated in E." figureDoi="http://doi.org/10.5281/zenodo.3825198" httpUri="https://zenodo.org/record/3825198/files/figure.png" pageId="140" pageNumber="141">Figures 27 A</figureCitation>
,
<figureCitation id="13402A2591C23B94FDF6FADAFDAFFAE6" box="[543,599,1377,1403]" captionStart="FIGURE 28" captionStartId="142.[151,250,1651,1676]" captionTargetBox="[151,1437,181,1623]" captionTargetId="figure@142.[151,1437,181,1625]" captionTargetPageId="142" captionText="FIGURE 28. Lycopodina nikitawimandi sp. nov. A. Preserved specimens of paratypes QM G337510 and QM G337534 still attached to the hexactinellid skeleton substrate.Arrows pointing to carnivorous sponges. B. Copepod trapped in isochelae, arrow pointing to the copepod. C. Bottlebrush body of the holotype QM G337534, showing the many filaments and the anisochelae in the ectosome. D. Close-up of the filaments branching from the stem. E. Filaments showing the ectosomal clusters of anisochelae and the underlying mycalostyles. F. Root ball and stem showing the hispid surface from the small mycalostyles." figureDoi="http://doi.org/10.5281/zenodo.3825200" httpUri="https://zenodo.org/record/3825200/files/figure.png" pageId="140" pageNumber="141">28 C</figureCitation>
). The body is
<quantity id="4C839B4591C23B94FD1AFADAFCB9FAE6" box="[755,833,1377,1403]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="140" pageNumber="141" unit="mm" value="28.0">28 mm</quantity>
in length and
<quantity id="4C839B4591C23B94FC37FADAFBC3FAE6" box="[990,1083,1377,1403]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="140" pageNumber="141" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
in width. The filaments are
<quantity id="4C839B4591C23B94FA9BFADAFF3BFA02" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="140" pageNumber="141" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
in length and between
<quantity id="4C839B4591C23B94FE23FA3FFDB4FA02" box="[458,588,1412,1439]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" pageId="140" pageNumber="141" unit="mm" value="150.0" valueMax="250.0" valueMin="50.0">50250 µm</quantity>
in width, and cover approximately 80% of the upper stem. The sponge has a spherical basal region of
<quantity id="4C839B4591C23B94FE2CFA12FDFEFA5E" box="[453,518,1449,1475]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="140" pageNumber="141" unit="mm" value="3.0">3 mm</quantity>
diameter, with protruding smaller mycalostyles (
<figureCitation id="13402A2591C23B94FBC4FA12FB4DFA5E" box="[1069,1205,1449,1475]" captionStart="FIGURE 28" captionStartId="142.[151,250,1651,1676]" captionTargetBox="[151,1437,181,1623]" captionTargetId="figure@142.[151,1437,181,1625]" captionTargetPageId="142" captionText="FIGURE 28. Lycopodina nikitawimandi sp. nov. A. Preserved specimens of paratypes QM G337510 and QM G337534 still attached to the hexactinellid skeleton substrate.Arrows pointing to carnivorous sponges. B. Copepod trapped in isochelae, arrow pointing to the copepod. C. Bottlebrush body of the holotype QM G337534, showing the many filaments and the anisochelae in the ectosome. D. Close-up of the filaments branching from the stem. E. Filaments showing the ectosomal clusters of anisochelae and the underlying mycalostyles. F. Root ball and stem showing the hispid surface from the small mycalostyles." figureDoi="http://doi.org/10.5281/zenodo.3825200" httpUri="https://zenodo.org/record/3825200/files/figure.png" pageId="140" pageNumber="141">Figure 28 F</figureCitation>
). This basal holdfast region grows within a hard or consolidated substrate. Many specimens appear to be a 2 dimensional feather due to damage sustained during collection.
</paragraph>
<paragraph id="8BC436A091C23B94FF2EF9AEFD55F9B2" blockId="140.[151,1437,872,1836]" box="[199,685,1557,1583]" pageId="140" pageNumber="141">
<emphasis id="B90FEAB291C23B94FF2EF9AEFEEEF9B2" box="[199,278,1557,1583]" italics="true" pageId="140" pageNumber="141">Colour</emphasis>
: Pale cream on deck and in ethanol.
</paragraph>
<paragraph id="8BC436A091C23B94FF2EF982FF41F9EA" blockId="140.[151,1437,872,1836]" pageId="140" pageNumber="141">
<emphasis id="B90FEAB291C23B94FF2EF982FE59F9CE" box="[199,417,1593,1619]" italics="true" pageId="140" pageNumber="141">Ectosomal skeleton</emphasis>
: The ectosomal skeleton is thin and membranous and contains the anisochelae (
<figureCitation id="13402A2591C23B94FAC4F982FF54F9EA" captionStart="FIGURE 28" captionStartId="142.[151,250,1651,1676]" captionTargetBox="[151,1437,181,1623]" captionTargetId="figure@142.[151,1437,181,1625]" captionTargetPageId="142" captionText="FIGURE 28. Lycopodina nikitawimandi sp. nov. A. Preserved specimens of paratypes QM G337510 and QM G337534 still attached to the hexactinellid skeleton substrate.Arrows pointing to carnivorous sponges. B. Copepod trapped in isochelae, arrow pointing to the copepod. C. Bottlebrush body of the holotype QM G337534, showing the many filaments and the anisochelae in the ectosome. D. Close-up of the filaments branching from the stem. E. Filaments showing the ectosomal clusters of anisochelae and the underlying mycalostyles. F. Root ball and stem showing the hispid surface from the small mycalostyles." figureDoi="http://doi.org/10.5281/zenodo.3825200" httpUri="https://zenodo.org/record/3825200/files/figure.png" pageId="140" pageNumber="141">Figure 28 E</figureCitation>
).
</paragraph>
<paragraph id="8BC436A091C23B94FF2EF93AFE18F922" blockId="140.[151,1437,872,1836]" pageId="140" pageNumber="141">
<emphasis id="B90FEAB291C23B94FF2EF93AFE51F906" box="[199,425,1665,1691]" italics="true" pageId="140" pageNumber="141">Endosomal skeleton</emphasis>
: The axis of the peduncle and the filaments consist of bundles of mycalostyles longitudinally arranged (
<figureCitation id="13402A2591C23B94FEAEF91EFE2CF922" box="[327,468,1701,1727]" captionStart="FIGURE 28" captionStartId="142.[151,250,1651,1676]" captionTargetBox="[151,1437,181,1623]" captionTargetId="figure@142.[151,1437,181,1625]" captionTargetPageId="142" captionText="FIGURE 28. Lycopodina nikitawimandi sp. nov. A. Preserved specimens of paratypes QM G337510 and QM G337534 still attached to the hexactinellid skeleton substrate.Arrows pointing to carnivorous sponges. B. Copepod trapped in isochelae, arrow pointing to the copepod. C. Bottlebrush body of the holotype QM G337534, showing the many filaments and the anisochelae in the ectosome. D. Close-up of the filaments branching from the stem. E. Filaments showing the ectosomal clusters of anisochelae and the underlying mycalostyles. F. Root ball and stem showing the hispid surface from the small mycalostyles." figureDoi="http://doi.org/10.5281/zenodo.3825200" httpUri="https://zenodo.org/record/3825200/files/figure.png" pageId="140" pageNumber="141">Figure 28 D</figureCitation>
).
</paragraph>
<paragraph id="8BC436A091C23B94FF2EF972FD29F8B1" blockId="140.[151,1437,872,1836]" pageId="140" pageNumber="141">
<emphasis id="B90FEAB291C23B94FF2EF972FEA5F97E" box="[199,349,1737,1763]" italics="true" pageId="140" pageNumber="141">Megascleres:</emphasis>
Large mycalostyles only occurring in the main axis (10401910 x
<quantity id="4C839B4591C23B94FBA4F972FB3AF97E" box="[1101,1218,1737,1763]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.7" metricValueMin="1.2" pageId="140" pageNumber="141" unit="mm" value="24.5" valueMax="37.0" valueMin="12.0">1237 µm</quantity>
, n=132) (
<figureCitation id="13402A2591C23B94FAC7F972FA64F97E" box="[1326,1436,1737,1763]" captionStart="FIGURE 27" captionStartId="141.[151,250,1078,1103]" captionTargetBox="[189,1398,181,1053]" captionTargetId="figure@141.[189,1398,181,1053]" captionTargetPageId="141" captionText="FIGURE 27. Lycopodina nikitawimandi sp. nov. QM G337534. A. Holotype QM G337534. B. Palmate anisochelae. C. Large mycalostyle 1. D. Magnified ends of the mycalostyle 1 illustrated in C. E. Small mycalostyle 2. F. Magnified ends of the smaller mycalostyle 2 illustrated in E." figureDoi="http://doi.org/10.5281/zenodo.3825198" httpUri="https://zenodo.org/record/3825198/files/figure.png" pageId="140" pageNumber="141">Figure 27</figureCitation>
CD). Smaller mycalostyles consistent throughout the sponge including the filaments and basal holdfast (209992 x
<quantity id="4C839B4591C23B94FF45F8AAFEEBF8B6" box="[172,275,1809,1835]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.9" metricValueMin="0.3" pageId="140" pageNumber="141" unit="mm" value="11.0" valueMax="19.0" valueMin="3.0">319 µm</quantity>
, n=250) (
<figureCitation id="13402A2591C23B94FE96F8AAFE15F8B6" box="[383,493,1809,1835]" captionStart="FIGURE 27" captionStartId="141.[151,250,1078,1103]" captionTargetBox="[189,1398,181,1053]" captionTargetId="figure@141.[189,1398,181,1053]" captionTargetPageId="141" captionText="FIGURE 27. Lycopodina nikitawimandi sp. nov. QM G337534. A. Holotype QM G337534. B. Palmate anisochelae. C. Large mycalostyle 1. D. Magnified ends of the mycalostyle 1 illustrated in C. E. Small mycalostyle 2. F. Magnified ends of the smaller mycalostyle 2 illustrated in E." figureDoi="http://doi.org/10.5281/zenodo.3825198" httpUri="https://zenodo.org/record/3825198/files/figure.png" pageId="140" pageNumber="141">Figure 27</figureCitation>
EF) (see
<tableCitation id="C6F9031B91C23B94FD8FF8AAFD30F8B1" box="[614,712,1809,1836]" captionStart="TABLE 16" captionStartId="141.[151,239,1223,1247]" captionTargetBox="[163,1326,1299,1946]" captionText="TABLE 16. A comparison of spicule measurements (µm) between the different specimens of Lycopodina nikitawimandi sp. nov." httpUri="http://table.plazi.org/id/DF04662891C33B95FF7EFB7CFF15FA9E" pageId="140" pageNumber="141" tableUuid="DF04662891C33B95FF7EFB7CFF15FA9E">Table 16</tableCitation>
)
</paragraph>
<caption id="DF04662891C33B95FF7EFB8DFE3FFB0A" ID-DOI="http://doi.org/10.5281/zenodo.3825198" ID-Zenodo-Dep="3825198" httpUri="https://zenodo.org/record/3825198/files/figure.png" pageId="141" pageNumber="142" startId="141.[151,250,1078,1103]" targetBox="[189,1398,181,1053]" targetPageId="141">
<paragraph id="8BC436A091C33B95FF7EFB8DFE3FFB0A" blockId="141.[151,1437,1078,1175]" pageId="141" pageNumber="142">
<emphasis id="B90FEAB291C33B95FF7EFB8DFED8FBD2" bold="true" box="[151,288,1078,1103]" pageId="141" pageNumber="142">FIGURE 27.</emphasis>
<taxonomicName id="4C7B4D2391C33B95FECFFB8CFDCDFBD2" authority="Ekins &amp; Erpenbeck &amp; Hooper, 2020" authorityName="Ekins &amp; Erpenbeck &amp; Hooper" authorityYear="2020" box="[294,565,1079,1103]" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="141" pageNumber="142" phylum="Porifera" rank="species" species="nikitawimandi" status="sp. nov.">
<emphasis id="B90FEAB291C33B95FECFFB8CFDCDFBD2" box="[294,565,1079,1103]" italics="true" pageId="141" pageNumber="142">Lycopodina nikitawimandi</emphasis>
</taxonomicName>
<emphasis id="B90FEAB291C33B95FDD2FB8CFD74FBD2" bold="true" box="[571,652,1079,1103]" pageId="141" pageNumber="142">
<taxonomicNameLabel id="A23C57C991C33B95FDD2FB8CFD74FBD2" box="[571,652,1079,1103]" pageId="141" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
QM G337534. A. Holotype QM G337534. B. Palmate anisochelae. C. Large mycalostyle 1. D. Magnified ends of the mycalostyle 1 illustrated in C. E. Small mycalostyle 2. F. Magnified ends of the smaller mycalostyle 2 illustrated in E.
</paragraph>
</caption>
<caption id="DF04662891C33B95FF7EFB7CFF15FA9E" ID-Table-UUID="DF04662891C33B95FF7EFB7CFF15FA9E" httpUri="http://table.plazi.org/id/DF04662891C33B95FF7EFB7CFF15FA9E" pageId="141" pageNumber="142" startId="141.[151,239,1223,1247]" targetBox="[163,1326,1299,1946]" targetIsTable="true" targetPageId="141">
<paragraph id="8BC436A091C33B95FF7EFB7CFF15FA9E" blockId="141.[151,1437,1223,1283]" pageId="141" pageNumber="142">
<emphasis id="B90FEAB291C33B95FF7EFB7CFEEFFB42" bold="true" box="[151,279,1223,1247]" pageId="141" pageNumber="142">TABLE 16.</emphasis>
A comparison of spicule measurements (µm) between the different specimens of
<taxonomicName id="4C7B4D2391C33B95FB69FB7CFA65FB42" authority="Ekins &amp; Erpenbeck &amp; Hooper, 2020" authorityName="Ekins &amp; Erpenbeck &amp; Hooper" authorityYear="2020" box="[1152,1437,1223,1247]" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="141" pageNumber="142" phylum="Porifera" rank="species" species="nikitawimandi" status="sp. nov.">Lycopodina nikitawimandi</taxonomicName>
<emphasis id="B90FEAB291C33B95FF7EFB50FF15FA9E" bold="true" box="[151,237,1259,1283]" pageId="141" pageNumber="142">sp. nov.</emphasis>
</paragraph>
</caption>
<paragraph id="8BC436A091C33B95FF4AFAA8FB68F807" pageId="141" pageNumber="142">
<table id="F97BC40091C3C4E7FF4AFAA8FAD6F807" box="[163,1326,1299,1946]" gridcols="4" gridrows="7" pageId="141" pageNumber="142">
<tr id="354B34E291C3C4E7FF4AFAA8FAD6FAB1" box="[163,1326,1299,1324]" gridrow="0" pageId="141" pageNumber="142">
<th id="769A5D9E91C3C4E7FF4AFAA8FE6DFAB1" box="[163,405,1299,1324]" gridcol="0" gridrow="0" pageId="141" pageNumber="142">Specimen</th>
<th id="769A5D9E91C3C4E7FE0CFAA8FD74FAB1" box="[485,652,1299,1324]" gridcol="1" gridrow="0" pageId="141" pageNumber="142">Anisochelae</th>
<th id="769A5D9E91C3C4E7FCCBFAA8FC1FFAB1" box="[802,999,1299,1324]" gridcol="2" gridrow="0" pageId="141" pageNumber="142">Large mycalostyles</th>
<th id="769A5D9E91C3C4E7FB80FAA8FAD6FAB1" box="[1129,1326,1299,1324]" gridcol="3" gridrow="0" pageId="141" pageNumber="142">Small mycalostyles</th>
</tr>
<tr id="354B34E291C3C4E7FF4AFAF3FAD6FA03" box="[163,1326,1352,1438]" gridrow="1" pageId="141" pageNumber="142">
<th id="769A5D9E91C3C4E7FF4AFAF3FE6DFA03" box="[163,405,1352,1438]" gridcol="0" gridrow="1" pageId="141" pageNumber="142">Holotype QM G337534</th>
<td id="769A5D9E91C3C4E7FE0CFAF3FD74FA03" box="[485,652,1352,1438]" gridcol="1" gridrow="1" pageId="141" pageNumber="142">12.5-(15.6)-17.8 x 3.3-(4.0)-4.8, n=65</td>
<td id="769A5D9E91C3C4E7FCCBFAF3FC1FFA03" box="[802,999,1352,1438]" gridcol="2" gridrow="1" pageId="141" pageNumber="142">1080-(1400)-1660 x 11.9-(20.1)-33.3, n=54</td>
<td id="769A5D9E91C3C4E7FB80FAF3FAD6FA03" box="[1129,1326,1352,1438]" gridcol="3" gridrow="1" pageId="141" pageNumber="142">476-(615)-824 x 4.4-(9.6)-17.9, n=92</td>
</tr>
<tr id="354B34E291C3C4E7FF4AFA16FAD6F999" box="[163,1326,1453,1540]" gridrow="2" pageId="141" pageNumber="142">
<th id="769A5D9E91C3C4E7FF4AFA16FE6DF999" box="[163,405,1453,1540]" gridcol="0" gridrow="2" pageId="141" pageNumber="142">Paratype QM G337510</th>
<td id="769A5D9E91C3C4E7FE0CFA16FD74F999" box="[485,652,1453,1540]" gridcol="1" gridrow="2" pageId="141" pageNumber="142">11.0-(14.1)-16.5 x 2.8-(3.9)-4.9, n=63</td>
<td id="769A5D9E91C3C4E7FCCBFA16FC1FF999" box="[802,999,1453,1540]" gridcol="2" gridrow="2" pageId="141" pageNumber="142">1040-(1482)-1910 x 15.2-(24.4)-36.6, n=38</td>
<td id="769A5D9E91C3C4E7FB80FA16FAD6F999" box="[1129,1326,1453,1540]" gridcol="3" gridrow="2" pageId="141" pageNumber="142">209-(557)-777 x 6.7-(12.3)-18.6, n=34</td>
</tr>
<tr id="354B34E291C3C4E7FF4AF9A8FAD6F9F4" box="[163,1326,1555,1641]" gridrow="3" pageId="141" pageNumber="142">
<th id="769A5D9E91C3C4E7FF4AF9A8FE6DF9F4" box="[163,405,1555,1641]" gridcol="0" gridrow="3" pageId="141" pageNumber="142">Paratype QM G337537</th>
<td id="769A5D9E91C3C4E7FE0CF9A8FD74F9F4" box="[485,652,1555,1641]" gridcol="1" gridrow="3" pageId="141" pageNumber="142">11.9-(13.7)-15.7 x 2.8-(3.7)-4.9, n=26</td>
<td id="769A5D9E91C3C4E7FCCBF9A8FC1FF9F4" box="[802,999,1555,1641]" gridcol="2" gridrow="3" pageId="141" pageNumber="142">1130-(1387)-1650 x 14.9-(20.3)-35.7, n=15</td>
<td id="769A5D9E91C3C4E7FB80F9A8FAD6F9F4" box="[1129,1326,1555,1641]" gridcol="3" gridrow="3" pageId="141" pageNumber="142">340-(555)-650 x 7.3-(10.7)-14.9, n=37</td>
</tr>
<tr id="354B34E291C3C4E7FF4AF9C3FAD6F952" box="[163,1326,1656,1743]" gridrow="4" pageId="141" pageNumber="142">
<th id="769A5D9E91C3C4E7FF4AF9C3FE6DF952" box="[163,405,1656,1743]" gridcol="0" gridrow="4" pageId="141" pageNumber="142">Paratype QM G337549</th>
<td id="769A5D9E91C3C4E7FE0CF9C3FD74F952" box="[485,652,1656,1743]" gridcol="1" gridrow="4" pageId="141" pageNumber="142">10.4-(12.8)-15.6 x 2.8-(3.6)-4.9, n=29</td>
<td id="769A5D9E91C3C4E7FCCBF9C3FC1FF952" box="[802,999,1656,1743]" gridcol="2" gridrow="4" pageId="141" pageNumber="142">1070-(1235)-1400 x 13.5-(21.9)-37.2, n=5</td>
<td id="769A5D9E91C3C4E7FB80F9C3FAD6F952" box="[1129,1326,1656,1743]" gridcol="3" gridrow="4" pageId="141" pageNumber="142">376-(512)-667 x 4.7-(10.7)-16.5, n=29</td>
</tr>
<tr id="354B34E291C3C4E7FF4AF965FAD6F8A9" box="[163,1326,1758,1844]" gridrow="5" pageId="141" pageNumber="142">
<th id="769A5D9E91C3C4E7FF4AF965FE6DF8A9" box="[163,405,1758,1844]" gridcol="0" gridrow="5" pageId="141" pageNumber="142">Paratype QM G337197</th>
<td id="769A5D9E91C3C4E7FE0CF965FD74F8A9" box="[485,652,1758,1844]" gridcol="1" gridrow="5" pageId="141" pageNumber="142">12.4-(14.4)-17.6 x 3.2-(4.1)-6.0, n=33</td>
<td id="769A5D9E91C3C4E7FCCBF965FC1FF8A9" box="[802,999,1758,1844]" gridcol="2" gridrow="5" pageId="141" pageNumber="142">1120-(1487)-1770 x 17.9-(21.8)-28.3, n=20</td>
<td id="769A5D9E91C3C4E7FB80F965FAD6F8A9" box="[1129,1326,1758,1844]" gridcol="3" gridrow="5" pageId="141" pageNumber="142">331-(583)-992 x 5.0-(12.1)-17.5, n=56</td>
</tr>
<tr id="354B34E291C3C4E7FF4AF8FFFAD6F807" box="[163,1326,1860,1946]" gridrow="6" pageId="141" pageNumber="142">
<th id="769A5D9E91C3C4E7FF4AF8FFFE6DF807" box="[163,405,1860,1946]" gridcol="0" gridrow="6" pageId="141" pageNumber="142">Paratype QM G337514</th>
<td id="769A5D9E91C3C4E7FE0CF8FFFD74F807" box="[485,652,1860,1946]" gridcol="1" gridrow="6" pageId="141" pageNumber="142">9.3-(11.4)-13.0 x 2.1-(3.2)-3.9, n=42</td>
<td id="769A5D9E91C3C4E7FCCBF8FFFC1FF807" box="[802,999,1860,1946]" gridcol="2" gridrow="6" pageId="141" pageNumber="142">-*</td>
<td id="769A5D9E91C3C4E7FB80F8FFFAD6F807" box="[1129,1326,1860,1946]" gridcol="3" gridrow="6" pageId="141" pageNumber="142">378-(508)-638 x 3.1-(6.8)-9.7, n=2</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC436A091C33B95FF7EF81CFEC2F87E" blockId="141.[151,1436,1959,2019]" pageId="141" pageNumber="142">
<tableNote id="769D372E91C33B95FF7EF81CFEC2F87E" pageId="141" pageNumber="142" targetBox="[163,1326,1299,1946]" targetPageId="141">* Most of the specimen disappeared off the SEM stub before the measurements of the large styles, which were present, could be made.</tableNote>
</paragraph>
<caption id="DF04662891C03B96FF7EF9C8FAFDF881" ID-DOI="http://doi.org/10.5281/zenodo.3825200" ID-Zenodo-Dep="3825200" httpUri="https://zenodo.org/record/3825200/files/figure.png" pageId="142" pageNumber="143" startId="142.[151,250,1651,1676]" targetBox="[151,1437,181,1623]" targetPageId="142">
<paragraph id="8BC436A091C03B96FF7EF9C8FAFDF881" blockId="142.[151,1437,1651,1820]" pageId="142" pageNumber="143">
<emphasis id="B90FEAB291C03B96FF7EF9C8FED9F911" bold="true" box="[151,289,1651,1676]" pageId="142" pageNumber="143">FIGURE 28.</emphasis>
<taxonomicName id="4C7B4D2391C03B96FEC0F9CFFDC0F911" authority="Ekins &amp; Erpenbeck &amp; Hooper, 2020" authorityName="Ekins &amp; Erpenbeck &amp; Hooper" authorityYear="2020" box="[297,568,1652,1676]" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="142" pageNumber="143" phylum="Porifera" rank="species" species="nikitawimandi" status="sp. nov.">
<emphasis id="B90FEAB291C03B96FEC0F9CFFDC0F911" box="[297,568,1652,1676]" italics="true" pageId="142" pageNumber="143">Lycopodina nikitawimandi</emphasis>
</taxonomicName>
<emphasis id="B90FEAB291C03B96FDA9F9CFFD6AF911" bold="true" box="[576,658,1652,1676]" pageId="142" pageNumber="143">
<taxonomicNameLabel id="A23C57C991C03B96FDA9F9CFFD6AF911" box="[576,658,1652,1676]" pageId="142" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Preserved specimens of paratypes QM G337510 and QM G337534 still attached to the hexactinellid skeleton substrate. Arrows pointing to carnivorous sponges. B. Copepod trapped in isochelae, arrow pointing to the copepod. C. Bottlebrush body of the holotype QM G337534, showing the many filaments and the anisochelae in the ectosome. D. Close-up of the filaments branching from the stem. E. Filaments showing the ectosomal clusters of anisochelae and the underlying mycalostyles. F. Root ball and stem showing the hispid surface from the small mycalostyles.
</paragraph>
</caption>
<paragraph id="8BC436A091C03B96FF2EF8F1FC39F84C" blockId="142.[151,1436,1866,2001]" pageId="142" pageNumber="143">
<emphasis id="B90FEAB291C03B96FF2EF8F1FEAFF8F9" box="[199,343,1866,1892]" italics="true" pageId="142" pageNumber="143">Microscleres</emphasis>
: Palmate anisochelae with the frontal upper alae nearly fully detached from the two lateral alae, and the three lower alae nearly completely fused to each other and the fimbria, with the frontal lower alae bearing three terminal spines and each of the lateral alae with two terminal spines (
<quantity id="4C839B4591C03B96FC0DF829FBB3F830" box="[996,1099,1938,1965]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.8" metricValueMin="0.9" pageId="142" pageNumber="143" unit="mm" value="13.5" valueMax="18.0" valueMin="9.0">918 µm</quantity>
(length) x
<quantity id="4C839B4591C03B96FB21F829FAE7F830" box="[1224,1311,1938,1965]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="142" pageNumber="143" unit="mm" value="4.0" valueMax="6.0" valueMin="2.0">26 µm</quantity>
(large alae width),
<quantity id="4C839B4591C03B96FF07F80DFEBDF84C" box="[238,325,1974,2001]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="142" pageNumber="143" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 µm</quantity>
(small alae width), n=258) (
<figureCitation id="13402A2591C03B96FD6DF80DFCF4F84C" box="[644,780,1974,2001]" captionStart="FIGURE 27" captionStartId="141.[151,250,1078,1103]" captionTargetBox="[189,1398,181,1053]" captionTargetId="figure@141.[189,1398,181,1053]" captionTargetPageId="141" captionText="FIGURE 27. Lycopodina nikitawimandi sp. nov. QM G337534. A. Holotype QM G337534. B. Palmate anisochelae. C. Large mycalostyle 1. D. Magnified ends of the mycalostyle 1 illustrated in C. E. Small mycalostyle 2. F. Magnified ends of the smaller mycalostyle 2 illustrated in E." figureDoi="http://doi.org/10.5281/zenodo.3825198" httpUri="https://zenodo.org/record/3825198/files/figure.png" pageId="142" pageNumber="143">Figure 27 B</figureCitation>
) (see
<tableCitation id="C6F9031B91C03B96FCA6F80DFC4BF84C" box="[847,947,1974,2001]" captionStart="TABLE 16" captionStartId="141.[151,239,1223,1247]" captionTargetBox="[163,1326,1299,1946]" captionText="TABLE 16. A comparison of spicule measurements (µm) between the different specimens of Lycopodina nikitawimandi sp. nov." httpUri="http://table.plazi.org/id/DF04662891C33B95FF7EFB7CFF15FA9E" pageId="142" pageNumber="143" tableUuid="DF04662891C33B95FF7EFB7CFF15FA9E">Table 16</tableCitation>
).
</paragraph>
<paragraph id="8BC436A091C13B97FF2EFF22FDE0FF66" blockId="143.[151,1437,152,683]" pageId="143" pageNumber="144">
<emphasis id="B90FEAB291C13B97FF2EFF22FE7FFF2E" bold="true" box="[199,391,153,179]" pageId="143" pageNumber="144">Molecular data:</emphasis>
The 28S sequences of QM G337197 and QM G337534 are provided in the Sponge Barcoding Database under accession numbers SBD#2304, SBD#2305 respectively and the molecular difference to other congenerics displayed in
<figureCitation id="13402A2591C13B97FE5AFF5AFDEBFF66" box="[435,531,225,251]" captionStart="FIGURE 3" captionStartId="21.[151,250,1733,1758]" captionTargetBox="[428,1156,182,1704]" captionTargetId="figure@21.[428,1159,181,1708]" captionTargetPageId="21" captionText="FIGURE 3. Maximum-likelihood phylogram of the DNA sequences of this study in comparison with other, published cornivorous sponges. Sequences from taxa of this study are in bold. Number at the branches are RAxML rapid bootstrap values. Numbers following the taxa are either QM registration numbers (QM G3xxxxx) or NCBI Genbank accession numbers. Scale bare are substitutions per site. Please note that the reconstruction does not represent a phylogeny as due to the variability of the 28SrRNA gene C-Region barcoding marker no unambiguous alignment is possible and clipping ambiguous sites would remove diagnostic characters for taxon distinction. The tree therefore indicates molecular differences rather than a robust phylogeny." figureDoi="http://doi.org/10.5281/zenodo.3825146" httpUri="https://zenodo.org/record/3825146/files/figure.png" pageId="143" pageNumber="144">Figure 3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361652B91C13B97FF2EFEBEFC75FD36" pageId="143" pageNumber="144" type="discussion">
<paragraph id="8BC436A091C13B97FF2EFEBEFC75FD36" blockId="143.[151,1437,152,683]" pageId="143" pageNumber="144">
<emphasis id="B90FEAB291C13B97FF2EFEBEFECCFE82" bold="true" box="[199,308,261,287]" pageId="143" pageNumber="144">Remarks</emphasis>
: Of the 29 described species of
<taxonomicName id="4C7B4D2391C13B97FD73FEBEFCE5FE82" authorityName="Lundbeck" authorityYear="1905" box="[666,797,261,287]" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="143" pageNumber="144" phylum="Porifera" rank="genus">
<emphasis id="B90FEAB291C13B97FD73FEBEFCE5FE82" box="[666,797,261,287]" italics="true" pageId="143" pageNumber="144">Lycopodina</emphasis>
</taxonomicName>
six have stipitate growth form, numerous large filaments along most of the stem, an enlarged basal attachment (where known), and only simple spiculation of styles or mycalostyles as structural megascleres, and palmate anisochelae as microscleres (
<tableCitation id="C6F9031B91C13B97FBCAFEF6FB74FEFA" box="[1059,1164,333,359]" captionStart="TABLE 14" captionStartId="124.[151,239,487,513]" captionTargetBox="[162,2024,564,1671]" captionTargetPageId="124" captionText="TABLE 14. Comparative morphological and distributional data for all known species of Lycopodina." httpUri="http://table.plazi.org/id/DF04662891323B64FF7EFE5CFB29FD9C" pageId="143" pageNumber="144" tableUuid="DF04662891323B64FF7EFE5CFB29FD9C">Table 14</tableCitation>
).
<taxonomicName id="4C7B4D2391C13B97FB4AFEF6FD6DFE16" authority="Goodwin, Berman, Downey &amp; Hendry, 2017" authorityName="Goodwin, Berman, Downey &amp; Hendry" authorityYear="2017" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="143" pageNumber="144" phylum="Porifera" rank="species" species="drakensis">
<emphasis id="B90FEAB291C13B97FB4AFEF6FA64FEFA" box="[1187,1436,333,359]" italics="true" pageId="143" pageNumber="144">Lycopodina drakensis</emphasis>
<bibRefCitation id="EFEA4B5191C13B97FF7EFECAFD6DFE16" author="Goodwin, Berman, Downey &amp; Hendry" box="[151,661,369,395]" firstAuthor="Goodwin" pageId="143" pageNumber="144" pagination="37 - 64" refId="ref73334" refString="Goodwin, C. E., Berman, J., Downey, R. V. &amp; Hendry, K. R. (2017) Carnivorous sponges (Porifera: Demospongiae: Poecilosclerida: Cladorhizidae) from the Drake Passage (Southern Ocean) with a description of eight new species and a review of the family Cladorhizidae in the Southern Ocean. Invertebrate Systematics, 31 (1), 37 - 64. https: // doi. org / 10.1071 / IS 16020" type="journal article" year="2017">Goodwin, Berman, Downey &amp; Hendry, 2017</bibRefCitation>
</taxonomicName>
, also has forceps microscleres, and the shorter mycalostyles occur in the stem and the longest ones in the filaments, the opposite of those in
<taxonomicName id="4C7B4D2391C13B97FC77FE2DFBA2FE32" authority="Ekins &amp; Erpenbeck &amp; Hooper, 2020" authorityName="Ekins &amp; Erpenbeck &amp; Hooper" authorityYear="2020" box="[926,1114,405,431]" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="143" pageNumber="144" phylum="Porifera" rank="species" species="nikitawimandi" status="sp. nov.">
<emphasis id="B90FEAB291C13B97FC77FE2DFBA2FE32" box="[926,1114,405,431]" italics="true" pageId="143" pageNumber="144">L. nikitawimandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57C991C13B97FB89FE2EFB41FE32" box="[1120,1209,405,431]" pageId="143" pageNumber="144" rank="species">
<emphasis id="B90FEAB291C13B97FB89FE2EFB4BFE32" bold="true" box="[1120,1203,405,431]" pageId="143" pageNumber="144">sp. nov</emphasis>
.
</taxonomicNameLabel>
<taxonomicName id="4C7B4D2391C13B97FB56FE2EFE6DFE4E" authority="(Levinsen, 1887)" baseAuthorityName="Levinsen" baseAuthorityYear="1887" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="143" pageNumber="144" phylum="Porifera" rank="species" species="lycopodium">
<emphasis id="B90FEAB291C13B97FB56FE2EFF37FE4E" italics="true" pageId="143" pageNumber="144">Lycopodina lycopodium</emphasis>
(
<bibRefCitation id="EFEA4B5191C13B97FF37FE02FE75FE4E" author="Levinsen" box="[222,397,441,467]" firstAuthor="Levinsen" pageId="143" pageNumber="144" pagination="339 - 372" refId="ref75332" refString="Levinsen, G. M. R. (1887) Kara-Havets Svampe (Porifera). Dijmphna-Togtets zoologisk-botaniske Udbytte, 1, 339 - 372, pls. XXIX-XXXI." type="journal article" year="1887">Levinsen, 1887</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C7B4D2391C13B97FE25FE01FCE6FE4E" authority="(Lambe, 1893)" baseAuthorityName="Lambe" baseAuthorityYear="1893" box="[460,798,441,467]" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="143" pageNumber="144" phylum="Porifera" rank="species" species="occidentalis">
<emphasis id="B90FEAB291C13B97FE25FE01FD88FE4E" box="[460,624,441,467]" italics="true" pageId="143" pageNumber="144">L. occidentalis</emphasis>
(
<bibRefCitation id="EFEA4B5191C13B97FD96FE02FCEEFE4E" author="Lambe" box="[639,790,441,467]" firstAuthor="Lambe" pageId="143" pageNumber="144" pagination="25 - 43" refId="ref74856" refString="Lambe, L. M. (1893 [1894]) Sponges from the Pacific coast of Canada. Proceedings and Transactions of the Royal Society of Canada, 11 (4), 25 - 43, pls. II-IV." type="journal article" year="1893">Lambe, 1893</bibRefCitation>
)
</taxonomicName>
have a similar distribution of short and long styles in the stem, body, filaments and basal attachment, but these are generally smaller than those of the new species, and both species also have forceps microscleres.
<taxonomicName id="4C7B4D2391C13B97FDB5FDBAFBFEFD86" authority="(Levinsen, 1887)" baseAuthorityName="Levinsen" baseAuthorityYear="1887" box="[604,1030,513,539]" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="143" pageNumber="144" phylum="Porifera" rank="species" species="robusta">
<emphasis id="B90FEAB291C13B97FDB5FDBAFCC5FD86" box="[604,829,513,539]" italics="true" pageId="143" pageNumber="144">Lycopodina robusta</emphasis>
(
<bibRefCitation id="EFEA4B5191C13B97FCA5FDBAFC05FD86" author="Levinsen" box="[844,1021,513,539]" firstAuthor="Levinsen" pageId="143" pageNumber="144" pagination="339 - 372" refId="ref75332" refString="Levinsen, G. M. R. (1887) Kara-Havets Svampe (Porifera). Dijmphna-Togtets zoologisk-botaniske Udbytte, 1, 339 - 372, pls. XXIX-XXXI." type="journal article" year="1887">Levinsen, 1887</bibRefCitation>
)
</taxonomicName>
has significantly smaller styles and also possesses forceps,
<taxonomicName id="4C7B4D2391C13B97FE49FD9DFD05FDA2" authority="Hestetun et al., 2017" authorityName="Hestetun" authorityYear="2017" box="[416,765,549,575]" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="143" pageNumber="144" phylum="Porifera" rank="species" species="tendali">
<emphasis id="B90FEAB291C13B97FE49FD9DFDF4FDA2" box="[416,524,549,575]" italics="true" pageId="143" pageNumber="144">L. tendali</emphasis>
<bibRefCitation id="EFEA4B5191C13B97FDFAFD9EFD05FDA2" author="Hestetun, J. T. &amp; Rapp, H. T. &amp; Xavier, J." box="[531,765,549,575]" pageId="143" pageNumber="144" pagination="166 - 189" refId="ref73911" refString="Hestetun, J. T., Rapp, H. T. &amp; Xavier, J. (2017 a) Carnivorous sponges (Porifera, Cladorhizidae) from the Southwest Indian Ocean Ridge seamounts. Deep Sea Research Part II: Topical Studies in Oceanography, 137, 166 - 189. https: // doi. org / 10.1016 / j. dsr 2.2016.03.004" type="journal article" year="2017">
Hestetun
<emphasis id="B90FEAB291C13B97FD97FD9DFD4FFDA2" box="[638,695,549,575]" italics="true" pageId="143" pageNumber="144">et al.</emphasis>
, 2017
</bibRefCitation>
</taxonomicName>
has larger styles that are uniformly distributed and forceps, and
<taxonomicName id="4C7B4D2391C13B97FF21FDF1FD81FDFE" authority="(Van Soest &amp; Baker, 2011)" baseAuthorityName="van Soest &amp; Baker" baseAuthorityYear="2011" box="[200,633,585,611]" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="143" pageNumber="144" phylum="Porifera" rank="species" species="vaceleti">
<emphasis id="B90FEAB291C13B97FF21FDF1FEC6FDFE" box="[200,318,585,611]" italics="true" pageId="143" pageNumber="144">L. vaceleti</emphasis>
(
<bibRefCitation id="EFEA4B5191C13B97FEA7FDF2FD89FDFE" author="Van Soest &amp; Baker" box="[334,625,585,611]" firstAuthor="Van Soest" pageId="143" pageNumber="144" pagination="495 - 501" refId="ref77556" refString="Van Soest, R. W. M. &amp; Baker, B. J. (2011), A new carnivorous shallow-water sponge from McMurdo Sound, Antarctica (Porifera, Poecilosclerida). Marine Biodiversity, 41 (4), 495 - 501. https: // doi. org / 10.1007 / s 12526 - 010 - 0076 - 6" type="journal article" year="2011">Van Soest &amp; Baker, 2011</bibRefCitation>
)
</taxonomicName>
has similar mycalostyles in the stem as
<taxonomicName id="4C7B4D2391C13B97FBACFDF1FAFCFDFE" authority="Ekins &amp; Erpenbeck &amp; Hooper, 2020" authorityName="Ekins &amp; Erpenbeck &amp; Hooper" authorityYear="2020" box="[1093,1284,585,611]" class="Demospongiae" family="Cladorhizidae" genus="Lycopodina" kingdom="Animalia" order="Poecilosclerida" pageId="143" pageNumber="144" phylum="Porifera" rank="species" species="nikitawimandi" status="sp. nov.">
<emphasis id="B90FEAB291C13B97FBACFDF1FAFCFDFE" box="[1093,1284,585,611]" italics="true" pageId="143" pageNumber="144">L. nikitawimandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57C991C13B97FAE4FDF2FA90FDFE" box="[1293,1384,585,611]" pageId="143" pageNumber="144" rank="species">
<emphasis id="B90FEAB291C13B97FAE4FDF2FA9AFDFE" bold="true" box="[1293,1378,585,611]" pageId="143" pageNumber="144">sp. nov</emphasis>
.
</taxonomicNameLabel>
, but also has a second category of smaller styles in the filaments and body, two categories of anisochelae, one of which has a unique sigma-like shape, and forceps microscleres (
<tableCitation id="C6F9031B91C13B97FCF3FD2AFC78FD36" box="[794,896,657,683]" captionStart="TABLE 14" captionStartId="124.[151,239,487,513]" captionTargetBox="[162,2024,564,1671]" captionTargetPageId="124" captionText="TABLE 14. Comparative morphological and distributional data for all known species of Lycopodina." httpUri="http://table.plazi.org/id/DF04662891323B64FF7EFE5CFB29FD9C" pageId="143" pageNumber="144" tableUuid="DF04662891323B64FF7EFE5CFB29FD9C">Table 14</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>