336 lines
32 KiB
XML
336 lines
32 KiB
XML
<document id="4678B7AD4EAD0A60F6C9D707114F5A87" ID-DOI="10.5281/zenodo.5178341" ID-ISSN="1942-1354" ID-Zenodo-Dep="5178341" ID-ZooBank="158EEA07-5357-4F7B-B8DF-C1B77EFE4DCB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628330624617" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2013" docId="03B487C3FFFBFF84FF6FDFFDFBA6F96C" docLanguage="en" docName="InsectaMundi.0325.1-25.pdf" docOrigin="Insecta Mundi 2013 (325)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Trogellus (Trogoides) hawksi Paulsen, 2013, new species" docType="treatment" docUuid="BC9B29B9-2B8F-4FDA-AD98-686C44BFD88A" docUuidSource="ZooBank" docVersion="4" lastPageNumber="12" masterDocId="FF8DFFBBFFF6FF89FFAFDF3EFF98FF90" masterDocTitle="A new genus for the Neotropical species of Aesalus Fabricius, with descriptions of eight new species (Coleoptera: Lucanidae: Aesalinae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="12" updateTime="1699044708161" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="5878E681D31D004F7F95107C7D28D4B4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="829DFB5B20778F68CA8E5DF80F289AC1">
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<mods:title id="FA2EE90219774605EF63AB8E7D4C278F">A new genus for the Neotropical species of Aesalus Fabricius, with descriptions of eight new species (Coleoptera: Lucanidae: Aesalinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="7A647F4AEAE16633C070AB7942D05B7D" type="personal">
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<mods:role id="5A82F4CAE0449B89AC897D7AB2F3B37D">
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<mods:roleTerm id="FD92AE53F95E3AA1B54B9E3D5CB6D6B6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="84E72963BD1596071019D9A92BF55DC1">Paulsen, M. J.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="564BF35B4F72A181BADD23BDADDCD0D6">text</mods:typeOfResource>
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<mods:titleInfo id="495684AB7E30E18572DE0ECB8BEDFEF2">
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<mods:title id="0F56EC219C47A262DC8AF675471AC929">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="C8B113D8F741E45B3C8FDFF565C110DC">
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<mods:date id="D737351A092651B0AF6877447904AE01">2013</mods:date>
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<mods:detail id="271A2ED1BB97995B853BB428AD5A6609" type="pubDate">
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<mods:number id="97755F33CA89F202FCFEE95148E33B12">2013-10-11</mods:number>
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</mods:detail>
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<mods:detail id="D8BE694FBEB2C3EF9A72D3CAA5B25AE1" type="volume">
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<mods:number id="A06FE645EC2145061930FAD215B730B2">2013</mods:number>
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</mods:detail>
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<mods:detail id="D2EB7100BDE38D2CEEBA2E55E5E12496" type="issue">
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<mods:number id="AC136B1E036BC041287DF0A21EF43554">325</mods:number>
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</mods:detail>
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<mods:extent id="A5DA01FAEEE801BB8E290788DFB35F77" unit="page">
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<mods:start id="F33483E16657A86D0F1C312CA37D8DAE">1</mods:start>
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<mods:end id="880FEE1E4C5DC1E52470F2250069E186">25</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="3CD60897748A34E4C952EF163E36643C">journal article</mods:classification>
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<mods:identifier id="804339A77F28E7E2E2EE43C82DE4D5ED" type="DOI">10.5281/zenodo.5178341</mods:identifier>
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<mods:identifier id="BE9341D120F151661D49DD0E00D5B605" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="661F536B3F9FC9493A28CFA5E5829BE1" type="Zenodo-Dep">5178341</mods:identifier>
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<mods:identifier id="90673450B687AE883010DE7C72AAF7FD" type="ZooBank">158EEA07-5357-4F7B-B8DF-C1B77EFE4DCB</mods:identifier>
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</mods:mods>
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<treatment id="03B487C3FFFBFF84FF6FDFFDFBA6F96C" LSID="urn:lsid:zoobank.org:act:BC9B29B9-2B8F-4FDA-AD98-686C44BFD88A" httpUri="http://treatment.plazi.org/id/03B487C3FFFBFF84FF6FDFFDFBA6F96C" lastPageNumber="12" pageId="13" pageNumber="12">
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<subSubSection id="C307655EFFFBFF84FF6FDFFDFCF4FF4B" box="[192,876,195,219]" pageId="13" pageNumber="12" type="nomenclature">
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<paragraph id="8BA236D5FFFBFF84FF6FDFFDFCF4FF4B" blockId="13.[192,876,195,219]" box="[192,876,195,219]" pageId="13" pageNumber="12">
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<heading id="D0EA81B9FFFBFF84FF6FDFFDFCF4FF4B" bold="true" box="[192,876,195,219]" fontSize="10" level="2" pageId="13" pageNumber="12" reason="3">
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<emphasis id="B969EAC7FFFBFF84FF6FDFFDFCFDFF4B" bold="true" box="[192,869,195,219]" pageId="13" pageNumber="12">
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<taxonomicName id="4C1D4D56FFFBFF84FF6FDFFDFDD0FF4B" ID-CoL="598HM" authorityName="Paulsen" authorityYear="2013" box="[192,584,195,219]" class="Insecta" family="Lucanidae" genus="Trogellus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="hawksi" subGenus="Trogoides">
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<emphasis id="B969EAC7FFFBFF84FF6FDFFDFED9FF4B" bold="true" box="[192,321,195,219]" italics="true" pageId="13" pageNumber="12">Trogellus</emphasis>
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||
(
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<emphasis id="B969EAC7FFFBFF84FEFFDFFDFE4DFF4B" bold="true" box="[336,469,195,219]" italics="true" pageId="13" pageNumber="12">Trogoides</emphasis>
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)
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<emphasis id="B969EAC7FFFBFF84FE4ADFFDFDD0FF4B" bold="true" box="[485,584,195,219]" italics="true" pageId="13" pageNumber="12">hawksi</emphasis>
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</taxonomicName>
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Paulsen,
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<taxonomicNameLabel id="A25A57BCFFFBFF84FD69DFFDFCFDFF4B" box="[710,869,195,219]" pageId="13" pageNumber="12" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C307655EFFFBFF84FF6FDE3DFEA6FE6B" pageId="13" pageNumber="12" type="materials_examined">
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<paragraph id="8BA236D5FFFBFF84FF6FDE3DFCD7FECB" blockId="13.[192,1440,259,507]" pageId="13" pageNumber="12">
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<emphasis id="B969EAC7FFFBFF84FF6FDE3DFE12FE8B" bold="true" box="[192,394,259,283]" pageId="13" pageNumber="12">Type Material.</emphasis>
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<materialsCitation id="3B753C88FFFBFF84FE3EDE3DFCD7FECB" collectingDate="1987-06-07" collectionCode="TAMU, ENTO" collectorName="W. Jotolito & A. Mendoza" location="Mex" pageId="13" pageNumber="12" specimenCode="X0074298" specimenCount="2" stateProvince="Chiapas" typeStatus="holotype">
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<typeStatus id="54A68877FFFBFF84FE3EDE3DFD98FE8B" box="[401,512,259,283]" pageId="13" pageNumber="12" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="ED0CAE10FFFBFF84FDFEDE3DFD30FE8B" box="[593,680,259,283]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;M University" pageId="13" pageNumber="12" type="Herbarium">TAMU</collectionCode>
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) labeled: a) “
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<location id="8EC2600EFFFBFF84FCE9DE3DFC19FE8B" LSID="urn:lsid:plazi:treatment:03B487C3FFFBFF84FF6FDFFDFBA6F96C:8EC2600EFFFBFF84FCE9DE3DFC19FE8B" box="[838,897,259,283]" name="Mex" pageId="13" pageNumber="12" stateProvince="Chiapas">Mex</location>
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||
:
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<collectingRegion id="49D9F837FFFBFF84FC23DE3DFC69FE8B" box="[908,1009,259,283]" country="Mexico" name="Chiapas" pageId="13" pageNumber="12">Chiapas</collectingRegion>
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, 3 / km.
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<collectorName id="26E85303FFFBFF84FBF9DE3AFB4EFE8B" box="[1110,1238,259,283]" pageId="13" pageNumber="12">W. Jotolito</collectorName>
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||
/
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<date id="FFA31015FFFBFF84FB44DE3DFAC7FE8B" box="[1259,1375,259,283]" pageId="13" pageNumber="12" value="1987-06-07">
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<collectingDate id="EFE7E9FDFFFBFF84FB44DE3DFAC7FE8B" box="[1259,1375,259,283]" pageId="13" pageNumber="12" value="1987-06-07">vi-7-1987</collectingDate>
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</date>
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, D.B. / Thomas &
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<collectorName id="26E85303FFFBFF84FEF2DE1DFE6EFEAB" box="[349,502,291,315]" pageId="13" pageNumber="12">A. Mendoza</collectorName>
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||
”; b) “TAMU –
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<collectionCode id="ED0CAE10FFFBFF84FD1DDE1DFC9BFEAB" box="[690,771,291,315]" pageId="13" pageNumber="12">ENTO</collectionCode>
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/
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||
<specimenCode id="DBBB9EAEFFFBFF84FCB1DE1DFC03FEAB" box="[798,923,291,315]" collectionCode="ENTO" pageId="13" pageNumber="12">X0074298</specimenCode>
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” barcode label; c) red paper: “
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<taxonomicName id="4C1D4D56FFFBFF84FAB5DE1DFA17FEAB" authorityName="Paulsen" authorityYear="2013" box="[1306,1423,291,315]" class="Insecta" family="Lucanidae" genus="Trogellus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B969EAC7FFFBFF84FAB5DE1DFA17FEAB" box="[1306,1423,291,315]" italics="true" pageId="13" pageNumber="12">Trogellus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C1D4D56FFFBFF84FF6FDE7DFE8FFECB" authorityName="Paulsen" authorityYear="2013" box="[192,279,323,347]" class="Insecta" family="Lucanidae" genus="Trogellus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="hawksi">
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<emphasis id="B969EAC7FFFBFF84FF6FDE7DFE8FFECB" box="[192,279,323,347]" italics="true" pageId="13" pageNumber="12">hawksi</emphasis>
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</taxonomicName>
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[male symbol] / Paulsen, 2013 /
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<typeStatus id="54A68877FFFBFF84FD32DE7DFCD8FECB" box="[669,832,323,347]" pageId="13" pageNumber="12" type="holotype">HOLOTYPE</typeStatus>
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||
”.
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</materialsCitation>
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||
</paragraph>
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<paragraph id="8BA236D5FFFBFF84FF5FDE5DFEA6FE6B" blockId="13.[192,1440,259,507]" pageId="13" pageNumber="12">
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||
<materialsCitation id="3B753C88FFFBFF84FF5FDE5DFEF5FE0B" collectionCode="WBWC" collectorName="W. B. Warner" country="Mexico" location="Parque Laguna" pageId="13" pageNumber="12" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
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One
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<typeStatus id="54A68877FFFBFF84FE85DE5AFE0EFEEC" box="[298,406,356,380]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
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female (
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<collectionCode id="ED0CAE10FFFBFF84FE52DE5DFDC1FEEB" box="[509,601,355,379]" pageId="13" pageNumber="12">WBWC</collectionCode>
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) labeled: a) “
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<collectingCountry id="F30A7645FFFBFF84FD58DE5DFCA5FEEB" box="[759,829,355,379]" name="Mexico" pageId="13" pageNumber="12">MEX</collectingCountry>
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: Edo.
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<collectingRegion id="49D9F837FFFBFF84FC29DE5DFC70FEEB" box="[902,1000,355,379]" country="Mexico" name="Chiapas" pageId="13" pageNumber="12">Chiapas</collectingRegion>
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||
/
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||
<location id="8EC2600EFFFBFF84FC51DE5AFB22FEEC" LSID="urn:lsid:plazi:treatment:03B487C3FFFBFF84FF6FDFFDFBA6F96C:8EC2600EFFFBFF84FC51DE5AFB22FEEC" box="[1022,1210,356,380]" country="Mexico" name="Parque Laguna" pageId="13" pageNumber="12" stateProvince="Chiapas">Parque Laguna</location>
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||
/ Belgica, VI-4-87 /
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<collectorName id="26E85303FFFBFF84FF6FDEBAFEFAFE0B" box="[192,354,387,411]" pageId="13" pageNumber="12">W.B. Warner</collectorName>
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||
”
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B753C88FFFBFF84FED2DEBDFBB1FE0B" box="[381,1065,387,411]" collectorName="Det. W. B. Warner" country="Mexico" determinerName="Det. W. B. Warner" pageId="13" pageNumber="12" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
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b) handwritten “
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||
<taxonomicName id="4C1D4D56FFFBFF84FDFDDEBDFD28FE0B" authorityName="Fabricius" authorityYear="1801" box="[594,688,387,411]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B969EAC7FFFBFF84FDFDDEBDFD28FE0B" box="[594,688,387,411]" italics="true" pageId="13" pageNumber="12">Aesalus</emphasis>
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||
</taxonomicName>
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||
/
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<emphasis id="B969EAC7FFFBFF84FD61DEBAFD68FE0B" box="[718,752,388,411]" italics="true" pageId="13" pageNumber="12">sp.</emphasis>
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||
/
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||
<determinerName id="7E7EDA11FFFBFF84FCA2DEBAFC70FE0B" box="[781,1000,388,411]" pageId="13" pageNumber="12">
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<collectorName id="26E85303FFFBFF84FCA2DEBAFC70FE0B" box="[781,1000,388,411]" pageId="13" pageNumber="12">Det. W.B. Warner</collectorName>
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</determinerName>
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‘92”
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</materialsCitation>
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;
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<materialsCitation id="3B753C88FFFBFF84FB96DEBDFEA2FE6B" collectingDate="2012-06-09" collectingDateMax="2012-06-13" collectingDateMin="2012-06-09" collectionCode="MJPC" collectorName="J. B. Heppner" country="Mexico" county="Finca" location="Volcan Atitlan" municipality="Tarrales" pageId="13" pageNumber="12" specimenCount="2" stateProvince="Chiapas" typeStatus="paratype">
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c) yellow paper: “
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<taxonomicName id="4C1D4D56FFFBFF84FAB6DEBDFA16FE0B" authorityName="Paulsen" authorityYear="2013" box="[1305,1422,387,411]" class="Insecta" family="Lucanidae" genus="Trogellus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B969EAC7FFFBFF84FAB6DEBDFA16FE0B" box="[1305,1422,387,411]" italics="true" pageId="13" pageNumber="12">Trogellus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C1D4D56FFFBFF84FF6FDE9DFE8FFE2B" authorityName="Paulsen" authorityYear="2013" box="[192,279,419,443]" class="Insecta" family="Lucanidae" genus="Trogellus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="hawksi">
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<emphasis id="B969EAC7FFFBFF84FF6FDE9DFE8FFE2B" box="[192,279,419,443]" italics="true" pageId="13" pageNumber="12">hawksi</emphasis>
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||
</taxonomicName>
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[female symbol] / Paulsen, 2013 /
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<typeStatus id="54A68877FFFBFF84FD1DDE9DFCD4FE2B" box="[690,844,419,443]" pageId="13" pageNumber="12" type="paratype">PARATYPE</typeStatus>
|
||
”. One
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||
<typeStatus id="54A68877FFFBFF84FC36DE9AFB9DFE2C" box="[921,1029,420,444]" pageId="13" pageNumber="12" type="paratype">paratype</typeStatus>
|
||
female (
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||
<collectionCode id="ED0CAE10FFFBFF84FBC3DE9DFB23FE2B" box="[1132,1211,419,443]" pageId="13" pageNumber="12">MJPC</collectionCode>
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||
) labeled: a) “GUA- TEMALA: Such- / itepéquez,
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||
<collectingCounty id="62C34E59FFFBFF84FD9EDEFDFDEDFE4B" box="[561,629,451,475]" pageId="13" pageNumber="12">Finca</collectingCounty>
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||
/
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||
<collectingMunicipality id="6BC6ACAFFFFBFF84FD3EDEFDFD63FE4B" box="[657,763,451,475]" pageId="13" pageNumber="12">Tarrales</collectingMunicipality>
|
||
,
|
||
<location id="8EC2600EFFFBFF84FCA5DEFDFC23FE4B" LSID="urn:lsid:plazi:treatment:03B487C3FFFBFF84FF6FDFFDFBA6F96C:8EC2600EFFFBFF84FCA5DEFDFC23FE4B" box="[778,955,451,475]" country="Mexico" county="Finca" municipality="Tarrales" name="Volcan Atitlan" pageId="13" pageNumber="12" stateProvince="Chiapas">Volcan Atitlan</location>
|
||
/
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||
<date id="FFA31015FFFBFF84FC78DEFDFBE4FE4B" box="[983,1148,451,475]" pageId="13" pageNumber="12" value="2012-06-09" valueMax="2012-06-13" valueMin="2012-06-09">
|
||
<collectingDate id="EFE7E9FDFFFBFF84FC78DEFDFBE4FE4B" box="[983,1148,451,475]" pageId="13" pageNumber="12" value="2012-06-09" valueMax="2012-06-13" valueMin="2012-06-09">9–13-VI-2012</collectingDate>
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||
</date>
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||
, 1340m / collector:
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||
<collectorName id="26E85303FFFBFF84FADDDEFAFEB7FE6B" pageId="13" pageNumber="12">J.B. Heppner</collectorName>
|
||
”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655EFFFBFF84FF6FDD1DFEEAFBCB" pageId="13" pageNumber="12" type="description">
|
||
<paragraph id="8BA236D5FFFBFF84FF6FDD1DFD1AFCCB" blockId="13.[192,1441,547,859]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FF6FDD1DFEF3FDAB" bold="true" box="[192,363,547,571]" pageId="13" pageNumber="12">Description.</emphasis>
|
||
<typeStatus id="54A68877FFFBFF84FEDBDD1DFE7BFDAB" box="[372,483,547,571]" pageId="13" pageNumber="12" type="holotype">Holotype</typeStatus>
|
||
male.
|
||
<emphasis id="B969EAC7FFFBFF84FD98DD1DFD38FDAB" bold="true" box="[567,672,547,571]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FD98DD1DFD00FDAB" bold="true" box="[567,664,547,571]" italics="true" pageId="13" pageNumber="12">Length</emphasis>
|
||
:
|
||
</emphasis>
|
||
<quantity id="4CE59B30FFFBFF84FD06DD1DFC92FDAB" box="[681,778,547,571]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="13" pageNumber="12" unit="mm" value="4.7">4.7 mm</quantity>
|
||
.
|
||
<emphasis id="B969EAC7FFFBFF84FCB9DD1DFCE9FDAB" bold="true" box="[790,881,547,571]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FCB9DD1DFCF0FDAB" bold="true" box="[790,872,547,571]" italics="true" pageId="13" pageNumber="12">Width</emphasis>
|
||
:
|
||
</emphasis>
|
||
<quantity id="4CE59B30FFFBFF84FCD5DD1DFC43FDAB" box="[890,987,547,571]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="12" unit="mm" value="2.5">2.5 mm</quantity>
|
||
.
|
||
<emphasis id="B969EAC7FFFBFF84FC48DD1DFBA0FDAB" bold="true" box="[999,1080,547,571]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FC48DD1DFBB7FDAB" bold="true" box="[999,1071,547,571]" italics="true" pageId="13" pageNumber="12">Head</emphasis>
|
||
:
|
||
</emphasis>
|
||
Clypeus triangular, apex not distinctly produced. Canthus long, length approximately 2× width. Mandibles without external tooth at angulation.
|
||
<emphasis id="B969EAC7FFFBFF84FEFADD5DFE70FDEB" bold="true" box="[341,488,611,635]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FEFADD5DFE47FDEA" bold="true" box="[341,479,611,634]" italics="true" pageId="13" pageNumber="12">Pronotum</emphasis>
|
||
:
|
||
</emphasis>
|
||
Surface densely punctate, with distinct tubercles posterolaterally at base of bristles, punctures large, ocellate.
|
||
<emphasis id="B969EAC7FFFBFF84FDCFDDBDFCB8FD0B" bold="true" box="[608,800,643,667]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FDCFDDBDFC8FFD0A" bold="true" box="[608,791,643,666]" italics="true" pageId="13" pageNumber="12">Metasternum</emphasis>
|
||
:
|
||
</emphasis>
|
||
Anterolateral punctures large.
|
||
<emphasis id="B969EAC7FFFBFF84FB30DDBDFB67FD0B" bold="true" box="[1183,1279,643,667]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FB30DDBDFB6FFD0B" bold="true" box="[1183,1271,643,667]" italics="true" pageId="13" pageNumber="12">Elytra</emphasis>
|
||
:
|
||
</emphasis>
|
||
Surface with blunt bristles, bristles moderately long (distinctly longer than 4
|
||
<superScript id="7C689B9DFFFBFF84FC18DD9CFC5EFD20" attach="left" box="[951,966,674,688]" fontSize="6" pageId="13" pageNumber="12">th</superScript>
|
||
tarsomere).
|
||
<emphasis id="B969EAC7FFFBFF84FBF1DD9DFB3AFD2B" bold="true" box="[1118,1186,675,699]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FBF1DD9DFB02FD2A" bold="true" box="[1118,1178,675,698]" italics="true" pageId="13" pageNumber="12">Legs</emphasis>
|
||
:
|
||
</emphasis>
|
||
Protibiae tridentate, teeth small, margin between teeth nearly flat. Mesotibia with 4 external teeth (proximal tooth smaller) and acute apex. Metatibia straight, with 1 small median tooth and 2–3 smaller teeth proximally; apex acute.
|
||
<emphasis id="B969EAC7FFFBFF84FEBFDC3DFE7AFC8B" bold="true" box="[272,482,771,795]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FEBFDC3DFE41FC8B" bold="true" box="[272,473,771,795]" italics="true" pageId="13" pageNumber="12">Male genitalia</emphasis>
|
||
:
|
||
</emphasis>
|
||
Parameres broad and elongate, over 1/2 as long as median lobe, located on side of median lobe; median lobe symmetrical, with large triangular lateral processes at distal third, processes with distal margin straight (
|
||
<figureCitation id="13262A50FFFBFF84FDB3DC7DFDEDFCCB" box="[540,629,835,859]" captionStart="Figures 18–23" captionStartId="10.[192,287,989,1011]" captionTargetBox="[207,1428,205,954]" captionTargetId="figure-250@10.[192,1440,192,960]" captionTargetPageId="10" captionText="Figures 18–23. Male genitalia of Trogellus species, dorsal view. 18. T. herrmanni n. sp. 19) T. ticiticus n. sp. 20) T. neotropicalis. 21) T. chapinitus n. sp. 22) T. hawksi n. sp. 23) T. trogoides. The male genitalia of T. maesi n. sp. are unknown." figureDoi="http://doi.org/10.5281/zenodo.5178353" httpUri="https://zenodo.org/record/5178353/files/figure.png" pageId="13" pageNumber="12">Fig. 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA236D5FFFBFF84FF6FDCBDFB7FFC4B" blockId="13.[192,1440,899,987]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FF6FDCBDFD17FC0B" bold="true" box="[192,655,899,923]" pageId="13" pageNumber="12">
|
||
<typeStatus id="54A68877FFFBFF84FF6FDCBDFED2FC0B" box="[192,330,899,923]" pageId="13" pageNumber="12" type="paratype">Paratypes</typeStatus>
|
||
vary as follows:
|
||
<emphasis id="B969EAC7FFFBFF84FD88DCBDFD1FFC0B" bold="true" box="[551,647,899,923]" italics="true" pageId="13" pageNumber="12">Length</emphasis>
|
||
:
|
||
</emphasis>
|
||
<quantity id="4CE59B30FFFBFF84FD39DCBDFCBDFC0B" box="[662,805,899,923]" metricMagnitude="-3" metricUnit="m" metricValue="4.949999999999999" metricValueMax="5.8" metricValueMin="4.1" pageId="13" pageNumber="12" unit="mm" value="4.949999999999999" valueMax="5.8" valueMin="4.1">4.1–5.8 mm</quantity>
|
||
.
|
||
<emphasis id="B969EAC7FFFBFF84FC81DCBDFC10FC0B" bold="true" box="[814,904,899,923]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FC81DCBDFC18FC0B" bold="true" box="[814,896,899,923]" italics="true" pageId="13" pageNumber="12">Width</emphasis>
|
||
:
|
||
</emphasis>
|
||
<quantity id="4CE59B30FFFBFF84FC21DCBDFB86FC0B" box="[910,1054,899,923]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.1" metricValueMin="2.3" pageId="13" pageNumber="12" unit="mm" value="2.7" valueMax="3.1" valueMin="2.3">2.3–3.1 mm</quantity>
|
||
. Female with external margin of protibiae slightly concave between teeth and mesotibial external tooth smaller.
|
||
<emphasis id="B969EAC7FFFBFF84FB25DC9DFB48FC2B" bold="true" box="[1162,1232,931,955]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FB25DC9DFB5FFC2A" bold="true" box="[1162,1223,931,954]" italics="true" pageId="13" pageNumber="12">Legs</emphasis>
|
||
:
|
||
</emphasis>
|
||
Metatibiae of females with tumid apex.
|
||
<emphasis id="B969EAC7FFFBFF84FE4EDCFDFD48FC4B" bold="true" box="[481,720,963,987]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FE4EDCFDFD50FC4B" bold="true" box="[481,712,963,987]" italics="true" pageId="13" pageNumber="12">Female genitalia</emphasis>
|
||
:
|
||
</emphasis>
|
||
Styli moderately elongate, strongly hooked.
|
||
</paragraph>
|
||
<paragraph id="8BA236D5FFFBFF84FF6FDB3DFEEAFBCB" blockId="13.[192,1441,1027,1115]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FF6FDB3DFE44FB8B" bold="true" box="[192,476,1027,1051]" pageId="13" pageNumber="12">Non-type specimens.</emphasis>
|
||
One male (CMNC) labeled: a) “
|
||
<collectingCountry id="F30A7645FFFBFF84FCF5DB3DFC2EFB8B" box="[858,950,1027,1051]" name="Mexico" pageId="13" pageNumber="12">Mexico</collectingCountry>
|
||
, V.C. Volcan / San Martin, E. slope / B. & B. Valentine”; b) “beating el. / 49-5000’ / 30-VII-59”; c) “[
|
||
<taxonomicName id="4C1D4D56FFFBFF84FC2ADB1DFC7BFBAB" authorityName="Fabricius" authorityYear="1801" box="[901,995,1059,1083]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B969EAC7FFFBFF84FC2ADB1DFC7BFBAB" box="[901,995,1059,1083]" italics="true" pageId="13" pageNumber="12">Aesalus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C1D4D56FFFBFF84FC53DB1DFB00FBAB" box="[1020,1176,1059,1083]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="neotropicalis">
|
||
<emphasis id="B969EAC7FFFBFF84FC53DB1DFB00FBAB" box="[1020,1176,1059,1083]" italics="true" pageId="13" pageNumber="12">neotropicalis</emphasis>
|
||
</taxonomicName>
|
||
/ Bates] / DET. / H.F. Howden [74]”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655EFFFBFF84FF6FDBBDFD46FB6B" pageId="13" pageNumber="12" type="distribution">
|
||
<paragraph id="8BA236D5FFFBFF84FF6FDBBDFA38FB2B" blockId="13.[192,1440,1154,1212]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FF6FDBBDFCBCFB0A" bold="true" box="[192,804,1154,1179]" pageId="13" pageNumber="12">
|
||
Distribution.
|
||
<collectingCountry id="F30A7645FFFBFF84FE2DDBBDFDDCFB0B" box="[386,580,1155,1179]" name="Guatemala" pageId="13" pageNumber="12">GUATEMALA</collectingCountry>
|
||
:
|
||
<collectingRegion id="49D9F837FFFBFF84FDF9DBBCFCBCFB0A" box="[598,804,1154,1178]" country="Guatemala" name="Suchitepequez" pageId="13" pageNumber="12">Suchitepéquez</collectingRegion>
|
||
</emphasis>
|
||
(1): Finca los Tarrales/Volcan Atitlan.
|
||
<emphasis id="B969EAC7FFFBFF84FAB2DBBDFEB5FB2B" bold="true" pageId="13" pageNumber="12">
|
||
<collectingCountry id="F30A7645FFFBFF84FAB2DBBDFA03FB0B" box="[1309,1435,1155,1179]" name="Mexico" pageId="13" pageNumber="12">MEXICO</collectingCountry>
|
||
:
|
||
<collectingRegion id="49D9F837FFFBFF84FF6FDB9DFEB5FB2B" box="[192,301,1187,1211]" country="Mexico" name="Chiapas" pageId="13" pageNumber="12">Chiapas</collectingRegion>
|
||
</emphasis>
|
||
(2): Jotolito; Parque Laguna Belgica.
|
||
<emphasis id="B969EAC7FFFBFF84FD4BDB9DFCC4FB2B" bold="true" box="[740,860,1187,1211]" pageId="13" pageNumber="12">
|
||
<collectingRegion id="49D9F837FFFBFF84FD4BDB9DFCC4FB2B" box="[740,860,1187,1211]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="13" pageNumber="12">Veracruz</collectingRegion>
|
||
</emphasis>
|
||
(1): Volcan San Martin Tuxtla, SE slope (CMNC);
|
||
</paragraph>
|
||
<paragraph id="8BA236D5FFFBFF84FF6FDBDDFD46FB6B" blockId="13.[192,734,1251,1275]" box="[192,734,1251,1275]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FF6FDBDDFD98FB6B" bold="true" box="[192,512,1251,1275]" pageId="13" pageNumber="12">Temporal Distribution.</emphasis>
|
||
June (3), July (1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655EFFFBFF84FF6FDA1DFBACF9EB" pageId="13" pageNumber="12" type="diagnosis">
|
||
<paragraph id="8BA236D5FFFBFF84FF6FDA1DFB76FAEB" blockId="13.[192,1441,1315,1659]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FF6FDA1DFE49FAAB" bold="true" box="[192,465,1315,1339]" pageId="13" pageNumber="12">Diagnosis/Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="54A68877FFFBFF84FDA2DA1DFDEEFAAB" box="[525,630,1315,1339]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
is distinguished from the other Mexican species (
|
||
<taxonomicName id="4C1D4D56FFFBFF84FB6ADA1AFAC8FAAB" box="[1221,1360,1315,1339]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="trogoides">
|
||
<emphasis id="B969EAC7FFFBFF84FB6ADA1AFAC8FAAB" box="[1221,1360,1315,1339]" italics="true" pageId="13" pageNumber="12">T. trogoides</emphasis>
|
||
</taxonomicName>
|
||
) by its distinctly tuberculate head and pronotum; elongate, narrow ocular canthus; and much longer parameres. Specimens of
|
||
<taxonomicName id="4C1D4D56FFFBFF84FECADA5AFD9BFAEB" authorityName="Paulsen" authorityYear="2013" box="[357,515,1379,1403]" class="Insecta" family="Lucanidae" genus="Trogellus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="chapinitus">
|
||
<emphasis id="B969EAC7FFFBFF84FECADA5AFD9BFAEB" box="[357,515,1379,1403]" italics="true" pageId="13" pageNumber="12">T. chapinitus</emphasis>
|
||
</taxonomicName>
|
||
have smaller, weakly ocellate punctures on the metasternum.
|
||
</paragraph>
|
||
<paragraph id="8BA236D5FFFBFF84FF5FDABDFBACF9EB" blockId="13.[192,1441,1315,1659]" pageId="13" pageNumber="12">
|
||
An additional specimen from
|
||
<collectingRegion id="49D9F837FFFBFF84FDF9DABAFD5BFA0B" box="[598,707,1412,1435]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="13" pageNumber="12">Veracruz</collectingRegion>
|
||
is tentatively identified as
|
||
<taxonomicName id="4C1D4D56FFFBFF84FBA4DABAFBE7FA0B" authorityName="Paulsen" authorityYear="2013" box="[1035,1151,1411,1435]" class="Insecta" family="Lucanidae" genus="Trogellus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="hawksi">
|
||
<emphasis id="B969EAC7FFFBFF84FBA4DABAFBE7FA0B" box="[1035,1151,1411,1435]" italics="true" pageId="13" pageNumber="12">T. hawksi</emphasis>
|
||
</taxonomicName>
|
||
because of the elongate ocular canthi and strongly tuberculate pronotum, but it is excluded from the type series. The
|
||
<collectingRegion id="49D9F837FFFBFF84FA9CDA9AFA38FA2B" box="[1331,1440,1444,1467]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="13" pageNumber="12">Veracruz</collectingRegion>
|
||
specimen is from the remote mountains of Tuxtla, an isolated montane area that is home to several endemic species, including two species and five subspecies of endemic birds (
|
||
<bibRefCitation id="EF8C4B24FFFBFF84FBEDDADDFA87FA6B" author="Winker, K. & R. J. Oehlenschlager & M. A. Ramos & R. M. Zink & J. H. Rappole & D. W. Warner" box="[1090,1311,1507,1531]" pageId="13" pageNumber="12" refId="ref14788" refString="Winker, K., R. J. Oehlenschlager, M. A. Ramos, R. M. Zink, J. H. Rappole, and D. W. Warner. 1992." type="book" year="1992">Winker et al. 1992</bibRefCitation>
|
||
). The male genitalia are similar to those of the
|
||
<typeStatus id="54A68877FFFBFF84FDC6D93DFD4DF98B" box="[617,725,1539,1563]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
, but the parameres appear slightly larger, and the vestiture of the elytra appears coarser. A series of specimens from the locality would indicate whether the population is
|
||
<taxonomicName id="4C1D4D56FFFBFF84FEA1D97AFE18F9CB" authorityName="Paulsen" authorityYear="2013" box="[270,384,1603,1627]" class="Insecta" family="Lucanidae" genus="Trogellus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="hawksi">
|
||
<emphasis id="B969EAC7FFFBFF84FEA1D97AFE18F9CB" box="[270,384,1603,1627]" italics="true" pageId="13" pageNumber="12">T. hawksi</emphasis>
|
||
</taxonomicName>
|
||
or an undescribed species. The specimen was identified and listed as
|
||
<emphasis id="B969EAC7FFFBFF84FB03D97DFB5EF9CA" box="[1196,1222,1603,1626]" italics="true" pageId="13" pageNumber="12">A.</emphasis>
|
||
near
|
||
<taxonomicName id="4C1D4D56FFFBFF84FAA8D97DFA38F9CB" box="[1287,1440,1603,1627]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="neotropicalis">
|
||
<emphasis id="B969EAC7FFFBFF84FAA8D97DFA38F9CB" box="[1287,1440,1603,1627]" italics="true" pageId="13" pageNumber="12">neotropicalis</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF8C4B24FFFBFF84FF4BD95DFDCBF9EB" author="Howden, H. F. & J. F. Lawrence" box="[228,595,1635,1659]" pageId="13" pageNumber="12" pagination="1505 - 1510" refId="ref14122" refString="Howden, H. F., and J. F. Lawrence. 1974. The New World Aesalinae, with notes on the North American lucanid subfamilies (Coleoptera, Lucanidae). Canadian Journal of Zoology 52: 1505 - 1510." type="journal article" year="1974">Howden and Lawrence (1974)</bibRefCitation>
|
||
, but as
|
||
<taxonomicName id="4C1D4D56FFFBFF84FD1FD95DFCF5F9EB" box="[688,877,1635,1659]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="neotropicalis">
|
||
<emphasis id="B969EAC7FFFBFF84FD1FD95DFCF5F9EB" box="[688,877,1635,1659]" italics="true" pageId="13" pageNumber="12">A. neotropicalis</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF8C4B24FFFBFF84FC37D95AFBA8F9EB" author="Maes, J. - M." box="[920,1072,1635,1659]" pageId="13" pageNumber="12" pagination="1 - 121" refId="ref14272" refString="Maes, J. - M. 1992. Lista de los Lucanidae (Coleoptera) del mundo. Revista Nicaraguense de Entomologia 22 A: 1 - 121." type="journal article" year="1992">Maes (1992)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655EFFFBFF84FF6FD99DFBA6F96C" pageId="13" pageNumber="12" type="etymology">
|
||
<paragraph id="8BA236D5FFFBFF84FF6FD99DFBA6F96C" blockId="13.[192,1440,1699,1788]" pageId="13" pageNumber="12">
|
||
<emphasis id="B969EAC7FFFBFF84FF6FD99DFECFF92B" bold="true" box="[192,343,1699,1723]" pageId="13" pageNumber="12">Etymology.</emphasis>
|
||
The species is named in honor of my friend and colleague, David C. Hawks, University of
|
||
<collectingRegion id="49D9F837FFFBFF84FF6FD9FDFEA5F94B" box="[192,317,1731,1755]" country="United States of America" name="California" pageId="13" pageNumber="12">California</collectingRegion>
|
||
, Riverside,
|
||
<collectingRegion id="49D9F837FFFBFF84FE69D9FDFDDBF94B" box="[454,579,1731,1755]" country="United States of America" name="California" pageId="13" pageNumber="12">California</collectingRegion>
|
||
, for his unceasing efforts to bring some order to the chaos of scarabaeoid classification and the tremendous help he has provided me over the years.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |