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<document id="E67FB449561899024FE4E466E9C5CE98" ID-CLB-Dataset="7439" ID-DOI="10.11646/zootaxa.4934.1.1" ID-GBIF-Dataset="300bac18-55f5-4e34-90e1-7fccd02d68c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4558796" ID-ZooBank="79895443-4597-42A5-AF8A-023EACB20E10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614157470936" checkinUser="plazi" docAuthor="Archibald, Bruce, Cannings, Robert A., Erickson, Robert J., Bybee, Seth M. &amp; Mathewes, Rolf W." docDate="2021" docId="03B487C2007BFFFCFF5BF9D3FA8F17D7" docLanguage="en" docName="zootaxa.4934.1.1.pdf" docOrigin="Zootaxa 4934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dysagrionites Archibald &amp; Cannings" docType="treatment" docVersion="7" lastPageNumber="108" masterDocId="FF8DFFBA0012FF97FFCCFFE5FFAE1215" masterDocTitle="The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America" masterLastPageNumber="133" masterPageNumber="1" pageNumber="106" updateTime="1698931040708" updateUser="ExternalLinkService">
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<mods:title id="B5D1F758A86EE9A2493A076085A888C5">The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America</mods:title>
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<mods:namePart id="503983913BB290A5B60005675F15A89C">Archibald, Bruce</mods:namePart>
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<mods:namePart id="02290AF41A30B485ED71B52DC758E609">Cannings, Robert A.</mods:namePart>
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<mods:namePart id="5EDBFD82BBCF1601671A4A9970C7A437">Erickson, Robert J.</mods:namePart>
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<mods:nameIdentifier id="10B8F4B8EF732BA717717FBFF27E3033" type="email">1990platypus@gmail.com</mods:nameIdentifier>
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<mods:namePart id="955793725306E097E1E11CFEF0F9CB23">Bybee, Seth M.</mods:namePart>
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<mods:namePart id="FDDF6BB8B8FB9E89F9E0AFE6C5BCB070">Mathewes, Rolf W.</mods:namePart>
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<mods:title id="1CD26D893259E6A690E9DD4FE67EFBB0">Zootaxa</mods:title>
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<mods:date id="E188C442AE1A542BBD833D890F672E53">2021</mods:date>
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<treatment id="03B487C2007BFFFCFF5BF9D3FA8F17D7" ID-DOI="http://doi.org/10.5281/zenodo.4672724" ID-GBIF-Taxon="180881790" ID-Zenodo-Dep="4672724" LSID="urn:lsid:plazi:treatment:03B487C2007BFFFCFF5BF9D3FA8F17D7" httpUri="http://treatment.plazi.org/id/03B487C2007BFFFCFF5BF9D3FA8F17D7" lastPageId="107" lastPageNumber="108" pageId="105" pageNumber="106">
<subSubSection id="C307655F007BFFFEFF5BF9D3FE2D1445" box="[151,387,1590,1616]" pageId="105" pageNumber="106" type="nomenclature">
<paragraph id="8BA236D4007BFFFEFF5BF9D3FE2D1445" blockId="105.[151,387,1590,1652]" box="[151,387,1590,1616]" pageId="105" pageNumber="106">
<heading id="D0EA81B8007BFFFEFF5BF9D3FE2D1445" bold="true" box="[151,387,1590,1616]" fontSize="11" level="2" pageId="105" pageNumber="106" reason="4">
<emphasis id="B969EAC6007BFFFEFF5BF9D3FE2D1445" bold="true" box="[151,387,1590,1616]" italics="true" pageId="105" pageNumber="106">
<taxonomicName id="4C1D4D57007BFFFEFF5BF9D3FE931445" authority="Archibald &amp; Cannings" authorityName="Archibald &amp; Cannings" box="[151,317,1590,1616]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="105" pageNumber="106" phylum="Arthropoda" rank="genus">Dysagrionites</taxonomicName>
sp. B
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C307655F007BFFFEFF5BF9BFFF401461" box="[151,238,1626,1652]" pageId="105" pageNumber="106" type="description">
<paragraph id="8BA236D4007BFFFEFF5BF9BFFF401461" blockId="105.[151,387,1590,1652]" box="[151,238,1626,1652]" pageId="105" pageNumber="106">
<figureCitation id="13262A51007BFFFEFF5BF9BFFF441461" box="[151,234,1626,1652]" captionStart="FIGURE 64" captionStartId="107.[151,250,1090,1114]" captionTargetBox="[157,1426,181,1030]" captionTargetId="figure-17@107.[151,1436,181,1066]" captionTargetPageId="107" captionText="FIGURE 64. Dysagrionites sp. B, RBCM P1554. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558956" httpUri="https://zenodo.org/record/4558956/files/figure.png" pageId="105" pageNumber="106">Fig. 64</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C307655F007BFFFEFF5BF978FD3A14EA" pageId="105" pageNumber="106" type="materials_examined">
<paragraph id="8BA236D4007BFFFEFF5BF978FD3A14EA" blockId="105.[151,1437,1693,2043]" pageId="105" pageNumber="106">
<emphasis id="B969EAC6007BFFFEFF5BF978FEA814A2" bold="true" box="[151,262,1693,1719]" pageId="105" pageNumber="106">Material.</emphasis>
<materialsCitation id="3B753C89007BFFFEFEDCF978FD3E14EA" ID-GBIF-Occurrence="3069296371" collectingDate="2002-08-10" collectionCode="RP" location="Museum" municipality="Royal" pageId="105" pageNumber="106" specimenCode="RP3-4" specimenCount="1" stateProvince="British Columbia">
RBCM P1554 (
<figureCitation id="13262A51007BFFFEFE08F978FDB414AD" box="[452,538,1693,1720]" captionStart="FIGURE 64" captionStartId="107.[151,250,1090,1114]" captionTargetBox="[157,1426,181,1030]" captionTargetId="figure-17@107.[151,1436,181,1066]" captionTargetPageId="107" captionText="FIGURE 64. Dysagrionites sp. B, RBCM P1554. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558956" httpUri="https://zenodo.org/record/4558956/files/figure.png" pageId="105" pageNumber="106">Fig. 64</figureCitation>
), the proximal half of a wing, damaged, apparently torn along or near
<specimenCode id="DBBB9EAF007BFFFEFA82F978FA3614A2" box="[1358,1432,1693,1719]" collectionCode="RP" pageId="105" pageNumber="106">RP3-4</specimenCode>
, which is missing or obscured. Collected at Driftwood Canyon Provincial Park by SBA,
<date id="FFA31014007BFFFEFBBDF924FB5514CE" box="[1137,1275,1729,1755]" pageId="105" pageNumber="106" value="2002-08-10">
<collectingDate id="EFE7E9FC007BFFFEFBBDF924FB5514CE" box="[1137,1275,1729,1755]" pageId="105" pageNumber="106" value="2002-08-10">10.viii.2002</collectingDate>
</date>
, housed in the
<collectingMunicipality id="6BC6ACAE007BFFFEFF5BF900FF7414EA" box="[151,218,1765,1791]" pageId="105" pageNumber="106">Royal</collectingMunicipality>
<collectingRegion id="49D9F836007BFFFEFF2DF900FE0C14EA" box="[225,418,1765,1791]" country="Canada" name="British Columbia" pageId="105" pageNumber="106">British Columbia</collectingRegion>
<location id="8EC2600F007BFFFEFE66F900FDA214EA" LSID="urn:lsid:plazi:treatment:03B487C2007BFFFCFF5BF9D3FA8F17D7:8EC2600F007BFFFEFE66F900FDA214EA" box="[426,524,1765,1791]" municipality="Royal" name="Museum" pageId="105" pageNumber="106" stateProvince="British Columbia">Museum</location>
collections
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C307655F007BFFFCFF0BF8ECFE9816D3" lastPageId="107" lastPageNumber="108" pageId="105" pageNumber="106" type="description">
<paragraph id="8BA236D4007BFFFEFF0BF8ECFC3815EE" blockId="105.[151,1437,1693,2043]" pageId="105" pageNumber="106">
<emphasis id="B969EAC6007BFFFEFF0BF8ECFD881536" bold="true" box="[199,550,1801,1827]" pageId="105" pageNumber="106">Description of RBCM P1554.</emphasis>
A somewhat damaged proximal portion of a wing. Length, width undeterminable by preservation. As in diagnosis and the following. Crossveins in postnodal, postsubnodal spaces aligned in preserved region. IR1 originates two, maybe three cells distal to origin of RP2. RP2 originates 2.5 cells distal to origin of IR2. IR2 originates at subnodus. RP3-4 is missing, wing damaged. MA linear to level of origin of IR1, not preserved distal to this. MP linear, rather straight from quadrangle to level of origin of IR2, not preserved distal to this. CuA slightly zigzagged distal to level of subnodus, little preserved distal to this. CuAA space at least three cells wide by level of subnodus, at least two briefly proximal to this.
</paragraph>
<caption id="DF62665C0078FFFDFF5BF880FDDA15B4" ID-DOI="http://doi.org/10.5281/zenodo.4558953" ID-Zenodo-Dep="4558953" httpUri="https://zenodo.org/record/4558953/files/figure.png" pageId="106" pageNumber="107" startId="106.[151,250,1893,1917]" targetBox="[162,1423,181,1856]" targetPageId="106">
<paragraph id="8BA236D40078FFFDFF5BF880FDDA15B4" blockId="106.[151,1437,1893,1953]" pageId="106" pageNumber="107">
<emphasis id="B969EAC60078FFFDFF5BF880FE8E1568" bold="true" box="[151,288,1893,1917]" pageId="106" pageNumber="107">FIGURE 63.</emphasis>
<taxonomicName id="4C1D4D570078FFFDFEEAF883FDAE1568" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[294,512,1893,1917]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="delinei">
<emphasis id="B969EAC60078FFFDFEEAF883FDAE1568" box="[294,512,1893,1917]" italics="true" pageId="106" pageNumber="107">Dysagrionites delinei</emphasis>
</taxonomicName>
paratype 1, UWBM 77612. A, photograph; B, drawing;
<taxonomicName id="4C1D4D570078FFFDFBF2F883FB621568" box="[1086,1228,1894,1917]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="106" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60078FFFDFBF2F883FB621568" box="[1086,1228,1894,1917]" italics="true" pageId="106" pageNumber="107">Dysagrionites</emphasis>
</taxonomicName>
sp. A, SR 96-12-03: C, photograph; D, drawing. All to scale, 5 mm.
</paragraph>
</caption>
<caption id="DF62665C0079FFFCFF5BFBA7FB28164C" ID-DOI="http://doi.org/10.5281/zenodo.4558956" ID-Zenodo-Dep="4558956" box="[151,1158,1090,1114]" httpUri="https://zenodo.org/record/4558956/files/figure.png" pageId="107" pageNumber="108" startId="107.[151,250,1090,1114]" targetBox="[157,1426,181,1030]" targetPageId="107">
<paragraph id="8BA236D40079FFFCFF5BFBA7FB28164C" blockId="107.[151,1158,1090,1114]" box="[151,1158,1090,1114]" pageId="107" pageNumber="108">
<emphasis id="B969EAC60079FFFCFF5BFBA7FE8E164F" bold="true" box="[151,288,1090,1114]" pageId="107" pageNumber="108">FIGURE 64.</emphasis>
<taxonomicName id="4C1D4D570079FFFCFEEAFBA6FE1A164F" box="[294,436,1091,1114]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="107" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60079FFFCFEEAFBA6FE1A164F" box="[294,436,1091,1114]" italics="true" pageId="107" pageNumber="108">Dysagrionites</emphasis>
</taxonomicName>
sp. B, RBCM P1554. A, photograph; B, drawing. Both to scale, 5 mm.
</paragraph>
</caption>
<paragraph id="8BA236D40079FFFCFF0BFB6DFE9816D3" blockId="107.[151,1437,1160,1475]" pageId="107" pageNumber="108">
<emphasis id="B969EAC60079FFFCFF0BFB6DFED616B7" bold="true" box="[199,376,1160,1186]" pageId="107" pageNumber="108">Range and age</emphasis>
. Okanagan Highlands shale at Driftwood Canyon Provincial Park,
<collectingRegion id="49D9F8360079FFFCFBBCFB6DFA9A16B7" box="[1136,1332,1160,1186]" country="Canada" name="British Columbia" pageId="107" pageNumber="108">British Columbia</collectingRegion>
,
<collectingCountry id="F30A76440079FFFCFA8CFB6DFA3616B7" box="[1344,1432,1160,1186]" name="Canada" pageId="107" pageNumber="108">Canada</collectingCountry>
; mid-Ypresian.
</paragraph>
</subSubSection>
<subSubSection id="C307655F0079FFFCFF0BFB35FA8F17D7" pageId="107" pageNumber="108" type="discussion">
<paragraph id="8BA236D40079FFFCFF0BFB35FA8F17D7" blockId="107.[151,1437,1160,1475]" pageId="107" pageNumber="108">
<emphasis id="B969EAC60079FFFCFF0BFB35FEE216FF" bold="true" box="[199,332,1232,1258]" pageId="107" pageNumber="108">Discussion.</emphasis>
This damaged proximal half of a wing may be separated from those of other species of
<taxonomicName id="4C1D4D570079FFFCFAE0FB35FF7A171B" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="107" pageNumber="108" phylum="Arthropoda" rank="subFamily" subFamily="Dysagrioninae">Dysagrioninae</taxonomicName>
except
<taxonomicName id="4C1D4D570079FFFCFEE1FB11FD87171B" box="[301,553,1268,1294]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="107" pageNumber="108" phylum="Arthropoda" rank="species" species="madseni">
<emphasis id="B969EAC60079FFFCFEE1FB11FD87171B" box="[301,553,1268,1294]" italics="true" pageId="107" pageNumber="108">Primorilestes madseni</emphasis>
</taxonomicName>
by infuscate membrane probably from the level of Ax1 (possibly proximal to this?) to the furthest preserved potion distal to the origin of IR1. The portion preserved is most easily distinguished from
<taxonomicName id="4C1D4D570079FFFCFF18FAD8FEE51743" box="[212,331,1340,1366]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="107" pageNumber="108" phylum="Arthropoda" rank="species" species="madseni">
<emphasis id="B969EAC60079FFFCFF18FAD8FEE51743" box="[212,331,1340,1366]" italics="true" pageId="107" pageNumber="108">P. madseni</emphasis>
</taxonomicName>
by being narrower between the quadrangle and the posterior margin.Although it is distinct by these traits, we do not name this species by the very damaged and incomplete condition of its only known fossil. It is of particular interest as the only known odonate fossil of the Okanagan Highlands from Driftwood Canyon, indeed, the only one north of McAbee, some 600 kilometres to the southeast of Driftwood Canyon Provincial Park.
</paragraph>
</subSubSection>
</treatment>
</document>