458 lines
54 KiB
XML
458 lines
54 KiB
XML
<document id="71E7CC3EE5655CFC1BDBE8F8C423B11F" ID-DOI="10.5281/zenodo.199004" ID-GBIF-Dataset="1967cb33-23fe-4cfa-9804-ff8420277315" ID-ISSN="1175-5326" ID-Zenodo-Dep="199004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460320311368" checkinUser="plazi" docAuthor="Kok, Philippe J. R., Macculloch, Ross D., Lathrop, Amy, Willaert, Bert & Bossuyt, Franky" docDate="2010" docId="03B487A4FFA4FF986CB07C8EFDC8FAF7" docLanguage="en" docName="zt02660p032.pdf" docOrigin="Zootaxa 2660" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anomaloglossus" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FF8DFFDCFFACFF926C27780CFF93FFF9" masterDocTitle="A new species of Anomaloglossus (Anura: Aromobatidae) from the Pakaraima Mountains of Guyana" masterLastPageNumber="32" masterPageNumber="18" pageNumber="26" updateTime="1698256450532" updateUser="plazi">
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<mods:titleInfo id="A6D62EFBA9BBA3B094DB956F1116CAF8">
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<mods:title id="35B93608EF76967D58E7D7067BC5A660">A new species of Anomaloglossus (Anura: Aromobatidae) from the Pakaraima Mountains of Guyana</mods:title>
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<mods:namePart id="AB4014E3E62367FF04825E247A7C7948">Kok, Philippe J. R.</mods:namePart>
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<mods:roleTerm id="8AE6DB421F75F71103E8B3788FD8657D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="442A2D2CC18FA87815EA2DA034E55CFB">Macculloch, Ross D.</mods:namePart>
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<mods:namePart id="4D0474BB097FF3A3EB042CDC3899163D">Lathrop, Amy</mods:namePart>
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<mods:namePart id="143F097F750EA2CB1548399667126BE1">Willaert, Bert</mods:namePart>
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<mods:name id="26894C42CB8FE527EC3B84E0E1C2DEA4" type="personal">
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<mods:namePart id="31CD0A0C4CD654B340FAB79310780ACE">Bossuyt, Franky</mods:namePart>
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<mods:title id="A3B51F2D2A160E6F4163E1583B6B92FE">Zootaxa</mods:title>
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<mods:part id="9DD15960EFA0FCF600B0A998C3E79B46">
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<mods:date id="857E225A5ACC1CFB4FF9741B1B1132CA">2010</mods:date>
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<mods:identifier id="947F9DBB05551A966B39B76297869E5B" type="DOI">10.5281/zenodo.199004</mods:identifier>
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<treatment id="03B487A4FFA4FF986CB07C8EFDC8FAF7" ID-DOI="http://doi.org/10.5281/zenodo.5680003" ID-GBIF-Taxon="119402338" ID-Zenodo-Dep="5680003" LSID="urn:lsid:plazi:treatment:03B487A4FFA4FF986CB07C8EFDC8FAF7" httpUri="http://treatment.plazi.org/id/03B487A4FFA4FF986CB07C8EFDC8FAF7" lastPageId="10" lastPageNumber="28" pageId="8" pageNumber="26">
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<subSubSection id="C3076539FFA4FF9A6CB07C8EFD08FB65" box="[151,667,1154,1180]" pageId="8" pageNumber="26" type="nomenclature">
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<paragraph id="8BA236B2FFA4FF9A6CB07C8EFD08FB65" blockId="8.[151,1438,1154,2002]" box="[151,667,1154,1180]" pageId="8" pageNumber="26">
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<heading id="D0EA81DEFFA4FF9A6CB07C8EFD08FB65" bold="true" box="[151,667,1154,1180]" fontSize="11" level="1" pageId="8" pageNumber="26" reason="1">
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<emphasis id="B969EAA0FFA4FF9A6CB07C8EFD08FB65" bold="true" box="[151,667,1154,1180]" pageId="8" pageNumber="26">
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Eastern Pantepui
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<taxonomicName id="4C1D4D31FFA4FF9A6D507C8EFDADFB65" ID-CoL="WS2" box="[375,574,1154,1180]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="genus">
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<emphasis id="B969EAA0FFA4FF9A6D507C8EFDADFB65" bold="true" box="[375,574,1154,1180]" italics="true" pageId="8" pageNumber="26">Anomaloglossus</emphasis>
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</taxonomicName>
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species
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3076539FFA4FF986CE17CA4FDC8FAF7" lastPageId="10" lastPageNumber="28" pageId="8" pageNumber="26" type="reference_group">
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<paragraph id="8BA236B2FFA4FF9A6CE17CA4FB86FA85" blockId="8.[151,1438,1154,2002]" pageId="8" pageNumber="26">
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<taxonomicName id="4C1D4D31FFA4FF9A6CE17CA4FDAEFB38" ID-CoL="6775Q" box="[198,573,1192,1217]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="megacephalus">
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<emphasis id="B969EAA0FFA4FF9A6CE17CA4FDAEFB38" box="[198,573,1192,1217]" italics="true" pageId="8" pageNumber="26">Anomaloglossus megacephalus</emphasis>
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</taxonomicName>
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can be distinguished from
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<taxonomicName id="4C1D4D31FFA4FF9A6FB97CA4FB81FB38" ID-CoL="6774J" box="[926,1042,1192,1217]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="beebei">
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<emphasis id="B969EAA0FFA4FF9A6FB97CA4FB81FB38" box="[926,1042,1192,1217]" italics="true" pageId="8" pageNumber="26">A. beebei</emphasis>
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</taxonomicName>
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by (characters of
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<taxonomicName id="4C1D4D31FFA4FF9A69247CA4FAE5FB38" ID-CoL="6774J" box="[1283,1398,1192,1217]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="beebei">
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<emphasis id="B969EAA0FFA4FF9A69247CA4FAE5FB38" box="[1283,1398,1192,1217]" italics="true" pageId="8" pageNumber="26">A. beebei</emphasis>
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</taxonomicName>
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in parentheses, see also
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<bibRefCitation id="EF8C4B43FFA4FF9A6DB27CC1FDCEFB1E" author="Kok" box="[405,605,1229,1255]" pageId="8" pageNumber="26" refString="Kok, P. J. R., MacCulloch, R. D., Gaucher, P., Poelman, E. H., Bourne, G. R., Lathrop, A. & Lenglet, G. L. (2006 b) A new species of Colostethus (Anura, Dendrobatidae) from French Guiana with a redescription of Colostethus beebei (Noble, 1923) from its type locality. Phyllomedusa, 5, 43 - 66." type="journal article" year="2006" yearSuffix="b">
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Kok
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<emphasis id="B969EAA0FFA4FF9A6DF77CC1FD97FB1F" box="[464,516,1229,1254]" italics="true" pageId="8" pageNumber="26">et al</emphasis>
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. 2006b
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</bibRefCitation>
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and
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<bibRefCitation id="EF8C4B43FFA4FF9A6EB07CC1FC44FB1E" author="Kok" box="[663,983,1229,1255]" pageId="8" pageNumber="26" refString="Kok, P. J. R. & Kalamandeen, M. (2008) Introduction to the taxonomy of the amphibians of Kaieteur National Park, Guyana. Abc Taxa, 5, 1 - 278." type="journal article" year="2008">Kok & Kalamandeen 2008</bibRefCitation>
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) its larger size, female SVL max
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<quantity id="4CE59B57FFA4FF9A694F7CC1FF55FAF5" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="8" pageNumber="26" unit="mm" value="28.3">28.3 mm</quantity>
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[n=3] in
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<taxonomicName id="4C1D4D31FFA4FF9A6D677CFFFD84FAF5" ID-CoL="6775Q" box="[320,535,1267,1292]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="megacephalus">
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<emphasis id="B969EAA0FFA4FF9A6D677CFFFD84FAF5" box="[320,535,1267,1292]" italics="true" pageId="8" pageNumber="26">A. megacephalus</emphasis>
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</taxonomicName>
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(
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<quantity id="4CE59B57FFA4FF9A6E0F7CFEFD0AFAF5" box="[552,665,1266,1292]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="8" pageNumber="26" unit="mm" value="18.7">18.7 mm</quantity>
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in
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<taxonomicName id="4C1D4D31FFA4FF9A6EE37CFFFCA9FAF5" ID-CoL="6774J" box="[708,826,1267,1292]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="beebei">
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<emphasis id="B969EAA0FFA4FF9A6EE37CFFFCA9FAF5" box="[708,826,1267,1292]" italics="true" pageId="8" pageNumber="26">A. beebei</emphasis>
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</taxonomicName>
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||
, n=27), fingers with keel-like lateral folds, best developed preaxially on Fingers II and III (fringes not folded), toes with folded flaplike fringing (fringes not folded), throat in adult female blotched (immaculate), palm dark brown to black (yellowish), distinct dark bands on thigh and shank (absent), dark interorbital V-shaped band (absent).
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||
</paragraph>
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<paragraph id="8BA236B2FFA4FF9A6CE17D84FC0FF9EB" blockId="8.[151,1438,1154,2002]" pageId="8" pageNumber="26">
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<taxonomicName id="4C1D4D31FFA4FF9A6CE17D84FDD2FA58" ID-CoL="6775Q" box="[198,577,1416,1441]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="megacephalus">
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||
<emphasis id="B969EAA0FFA4FF9A6CE17D84FDD2FA58" box="[198,577,1416,1441]" italics="true" pageId="8" pageNumber="26">Anomaloglossus megacephalus</emphasis>
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</taxonomicName>
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can be distinguished from
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<taxonomicName id="4C1D4D31FFA4FF9A6F9F7D84FAFFFA5B" ID-CoL="5V74K" authority="Barrio-Amoros, 2006" authorityName="Barrio-Amoros" authorityYear="2006" box="[952,1388,1416,1442]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="breweri">
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||
<emphasis id="B969EAA0FFA4FF9A6F9F7D84FBADFA58" box="[952,1086,1416,1441]" italics="true" pageId="8" pageNumber="26">A. breweri</emphasis>
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||
(
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||
<bibRefCitation id="EF8C4B43FFA4FF9A687F7D84FAF0FA5B" author="Barrio-Amoros" box="[1112,1379,1416,1442]" pageId="8" pageNumber="26" refString="Barrio-Amoros, C. L. (2006) A new dendrobatid frog (Anura: Dendrobatidae: Colostethus) from Aprada tepui, southern Venezuela. Zootaxa, 1110, 59 - 68." type="journal article" year="2006">Barrio-Amorós, 2006</bibRefCitation>
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)
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||
</taxonomicName>
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||
by (characters of
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<taxonomicName id="4C1D4D31FFA4FF9A6D677DA1FE28FA3F" ID-CoL="5V74K" box="[320,443,1453,1478]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="breweri">
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<emphasis id="B969EAA0FFA4FF9A6D677DA1FE28FA3F" box="[320,443,1453,1478]" italics="true" pageId="8" pageNumber="26">A. breweri</emphasis>
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||
</taxonomicName>
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in parentheses) its larger size, female SVL max
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||
<quantity id="4CE59B57FFA4FF9A6FDD7DA1FBF1FA3E" box="[1018,1122,1453,1479]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="8" pageNumber="26" unit="mm" value="28.3">28.3 mm</quantity>
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||
[n=3] in
|
||
<taxonomicName id="4C1D4D31FFA4FF9A68F47DA1FA0FFA3F" ID-CoL="6775Q" box="[1235,1436,1453,1478]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA4FF9A68F47DA1FA0FFA3F" box="[1235,1436,1453,1478]" italics="true" pageId="8" pageNumber="26">A. megacephalus</emphasis>
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||
</taxonomicName>
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||
(
|
||
<quantity id="4CE59B57FFA4FF9A6CB87DDEFE9FFA15" box="[159,268,1490,1516]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="8" pageNumber="26" unit="mm" value="23.8">23.8 mm</quantity>
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in
|
||
<taxonomicName id="4C1D4D31FFA4FF9A6D137DDFFE21FA15" ID-CoL="5V74K" box="[308,434,1491,1516]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="breweri">
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<emphasis id="B969EAA0FFA4FF9A6D137DDFFE21FA15" box="[308,434,1491,1516]" italics="true" pageId="8" pageNumber="26">A. breweri</emphasis>
|
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</taxonomicName>
|
||
, n=1), throat in adult female blotched (immaculate), median lingual process short, wider than long, tapered (distinctly longer than wide, tip pointed).
|
||
</paragraph>
|
||
<paragraph id="8BA236B2FFA4FF9A6CE27E11FB71F935" blockId="8.[151,1438,1154,2002]" pageId="8" pageNumber="26">
|
||
<taxonomicName id="4C1D4D31FFA4FF9A6CE27E11FDCAF9CF" ID-CoL="6775Q" box="[197,601,1565,1590]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA4FF9A6CE27E11FDCAF9CF" box="[197,601,1565,1590]" italics="true" pageId="8" pageNumber="26">Anomaloglossus megacephalus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C1D4D31FFA4FF9A6FE77E11FBB6F9CF" ID-CoL="5V762" box="[960,1061,1565,1590]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="kaiei">
|
||
<emphasis id="B969EAA0FFA4FF9A6FE77E11FBB6F9CF" box="[960,1061,1565,1590]" italics="true" pageId="8" pageNumber="26">A. kaiei</emphasis>
|
||
</taxonomicName>
|
||
by (characters of
|
||
<taxonomicName id="4C1D4D31FFA4FF9A69327E11FAE8F9CF" ID-CoL="5V762" box="[1301,1403,1565,1590]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="kaiei">
|
||
<emphasis id="B969EAA0FFA4FF9A69327E11FAE8F9CF" box="[1301,1403,1565,1590]" italics="true" pageId="8" pageNumber="26">A. kaiei</emphasis>
|
||
</taxonomicName>
|
||
in parentheses, see also
|
||
<bibRefCitation id="EF8C4B43FFA4FF9A6D837E4EFD67F9A5" author="Kok" box="[420,756,1602,1628]" pageId="8" pageNumber="26" refString="Kok, P. J. R. & Kalamandeen, M. (2008) Introduction to the taxonomy of the amphibians of Kaieteur National Park, Guyana. Abc Taxa, 5, 1 - 278." type="journal article" year="2008">Kok & Kalamandeen 2008</bibRefCitation>
|
||
) its larger size, female SVL max
|
||
<quantity id="4CE59B57FFA4FF9A68BB7E4EFA99F9A5" box="[1180,1290,1602,1628]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="8" pageNumber="26" unit="mm" value="28.3">28.3 mm</quantity>
|
||
[n=3] in
|
||
<taxonomicName id="4C1D4D31FFA4FF9A69A57E4FFED3F978" ID-CoL="6775Q" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA4FF9A69A57E4FFED3F978" italics="true" pageId="8" pageNumber="26">A. megacephalus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CE59B57FFA4FF9A6D777E64FE28F97B" box="[336,443,1640,1666]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="8" pageNumber="26" unit="mm" value="19.8">19.8 mm</quantity>
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in
|
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<taxonomicName id="4C1D4D31FFA4FF9A6DC47E64FDAEF978" ID-CoL="5V762" box="[483,573,1640,1665]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="kaiei">
|
||
<emphasis id="B969EAA0FFA4FF9A6DC47E64FDAEF978" box="[483,573,1640,1665]" italics="true" pageId="8" pageNumber="26">A. kaiei</emphasis>
|
||
</taxonomicName>
|
||
, n=25), fingers with keel-like lateral folds, best developed preaxially on Fingers II and III (fringes not folded), toes moderately webbed (basally webbed), dorsolateral stripe absent (present), oblique lateral stripe present (absent), throat in adult female blotched (immaculate).
|
||
</paragraph>
|
||
<paragraph id="8BA236B2FFA4FF9A6CE27ED4FE94F82B" blockId="8.[151,1438,1154,2002]" pageId="8" pageNumber="26">
|
||
<taxonomicName id="4C1D4D31FFA4FF9A6CE27ED4FDA5F908" ID-CoL="6775Q" box="[197,566,1752,1777]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA4FF9A6CE27ED4FDA5F908" box="[197,566,1752,1777]" italics="true" pageId="8" pageNumber="26">Anomaloglossus megacephalus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C1D4D31FFA4FF9A6F5E7ED4FAB7F90B" ID-CoL="5V75R" authority="La Marca, 1998" authorityName="La Marca" authorityYear="1998" box="[889,1316,1752,1778]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="murisipanensis">
|
||
<emphasis id="B969EAA0FFA4FF9A6F5E7ED4FBDEF908" box="[889,1101,1752,1777]" italics="true" pageId="8" pageNumber="26">A. murisipanensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8C4B43FFA4FF9A687A7ED4FA8FF90B" author="La" box="[1117,1308,1752,1778]" pageId="8" pageNumber="26" refString="La Marca, E. (1998 " 1996 ") Ranas del genero Colostethus (Amphibia: Anura: Dendrobatidae) de la Guyana Venezolana con la descripcion de siete especies nuevas. Publicaciones de la Asociacion de Amigos de Donana, 9, 1 - 64." type="journal article" year="1998">La Marca, 1998</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in having (characters of
|
||
<taxonomicName id="4C1D4D31FFA4FF9A6D667EF1FD86F8EF" ID-CoL="5V75R" box="[321,533,1789,1814]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="murisipanensis">
|
||
<emphasis id="B969EAA0FFA4FF9A6D667EF1FD86F8EF" box="[321,533,1789,1814]" italics="true" pageId="8" pageNumber="26">A. murisipanensis</emphasis>
|
||
</taxonomicName>
|
||
in parentheses, all based on the preserved
|
||
<typeStatus id="54A68810FFA4FF9A68357EF1FBEFF8EE" box="[1042,1148,1789,1815]" pageId="8" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
, which is a juvenile and the only known specimen) fingers with keel-like lateral folds, best developed preaxially on Fingers II and III (fringes barely detectable), more webbing on toes, symmetrical cloacal tubercles present (absent), tarsal keel weakly to distinctly curved, slightly tuberclelike (straight, not tuberclelike), oblique lateral stripe always present, even if broken in spots (not distinguishable), no white stripe between naris and eye in preservative (present).
|
||
</paragraph>
|
||
<caption id="DF62663AFFA5FF9B6CB07DEEFEDEF920" httpUri="https://zenodo.org/record/199009/files/figure.png" pageId="9" pageNumber="27" targetBox="[151,1436,194,1484]" targetPageId="9">
|
||
<paragraph id="8BA236B2FFA5FF9B6CB07DEEFEDEF920" blockId="9.[151,1437,1506,1753]" pageId="9" pageNumber="27">
|
||
<emphasis id="B969EAA0FFA5FF9B6CB07DEEFE8FFA03" bold="true" box="[151,284,1506,1530]" pageId="9" pageNumber="27">FIGURE 5.</emphasis>
|
||
Comparison of
|
||
<taxonomicName id="4C1D4D31FFA5FF9B6DEB7DEEFC8DFA00" ID-CoL="6775Q" box="[460,798,1506,1529]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="megacephalus" status="sp. nov.">
|
||
<emphasis id="B969EAA0FFA5FF9B6DEB7DEEFC8DFA00" box="[460,798,1506,1529]" italics="true" pageId="9" pageNumber="27">Anomaloglossus megacephalus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57DBFFA5FF9B6F017DEEFCEEFA03" box="[806,893,1506,1530]" pageId="9" pageNumber="27" rank="species">
|
||
<emphasis id="B969EAA0FFA5FF9B6F017DEEFCEEFA03" bold="true" box="[806,893,1506,1530]" pageId="9" pageNumber="27">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and the similar
|
||
<taxonomicName id="4C1D4D31FFA5FF9B68097DEEFB2FFA00" ID-CoL="6775P" box="[1070,1212,1506,1529]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA5FF9B68097DEEFB2FFA00" box="[1070,1212,1506,1529]" italics="true" pageId="9" pageNumber="27">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D31FFA5FF9B68D47DEEFAE0FA00" ID-CoL="66V5R" box="[1267,1395,1506,1529]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="parkerae">
|
||
<emphasis id="B969EAA0FFA5FF9B68D47DEEFAE0FA00" box="[1267,1395,1506,1529]" italics="true" pageId="9" pageNumber="27">A. parkerae</emphasis>
|
||
</taxonomicName>
|
||
. A: Dorsal view of preserved specimens of (from left to right)
|
||
<taxonomicName id="4C1D4D31FFA5FF9B6F347E0EFC5FF9E0" ID-CoL="6775Q" box="[787,972,1538,1561]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="megacephalus" status="sp. nov.">
|
||
<emphasis id="B969EAA0FFA5FF9B6F347E0EFC5FF9E0" box="[787,972,1538,1561]" italics="true" pageId="9" pageNumber="27">A. megacephalus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57DBFFA5FF9B6FF47E0EFBB9F9E3" box="[979,1066,1538,1562]" pageId="9" pageNumber="27" rank="species">
|
||
<emphasis id="B969EAA0FFA5FF9B6FF47E0EFBB9F9E3" bold="true" box="[979,1066,1538,1562]" pageId="9" pageNumber="27">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(IRSNB 1986, 27.3 mm SVL),
|
||
<taxonomicName id="4C1D4D31FFA5FF9B69A27E0EFE95F9C0" ID-CoL="6775P" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA5FF9B69A27E0EFE95F9C0" italics="true" pageId="9" pageNumber="27">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
(MHNLS 17401, 23.3 mm SVL) and
|
||
<taxonomicName id="4C1D4D31FFA5FF9B6E837E2EFCB0F9C0" ID-CoL="66V5R" box="[676,803,1570,1593]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="parkerae">
|
||
<emphasis id="B969EAA0FFA5FF9B6E837E2EFCB0F9C0" box="[676,803,1570,1593]" italics="true" pageId="9" pageNumber="27">A. parkerae</emphasis>
|
||
</taxonomicName>
|
||
(MHNLS 11089, 23.2 mm SVL). B: Comparison between midline distance from snout tip to an imaginary line between anterior arm insertions and the same distance reported posteriorly from the same imaginary line in
|
||
<taxonomicName id="4C1D4D31FFA5FF9B6EA07E6EFCD6F980" ID-CoL="6775Q" box="[647,837,1634,1657]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="megacephalus" status="sp. nov.">
|
||
<emphasis id="B969EAA0FFA5FF9B6EA07E6EFCD6F980" box="[647,837,1634,1657]" italics="true" pageId="9" pageNumber="27">A. megacephalus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25A57DBFFA5FF9B6F6A7E6EFC34F983" box="[845,935,1634,1658]" pageId="9" pageNumber="27" rank="species">
|
||
<emphasis id="B969EAA0FFA5FF9B6F6A7E6EFC34F983" bold="true" box="[845,935,1634,1658]" pageId="9" pageNumber="27">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C1D4D31FFA5FF9B6FF87E6EFBE1F980" ID-CoL="6775P" box="[991,1138,1634,1657]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA5FF9B6FF87E6EFBE1F980" box="[991,1138,1634,1657]" italics="true" pageId="9" pageNumber="27">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
. In most
|
||
<taxonomicName id="4C1D4D31FFA5FF9B68F97E6EFA0FF980" ID-CoL="6775Q" box="[1246,1436,1634,1657]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA5FF9B68F97E6EFA0FF980" box="[1246,1436,1634,1657]" italics="true" pageId="9" pageNumber="27">A. megacephalus</emphasis>
|
||
</taxonomicName>
|
||
specimens (left, IRSNB 1986, 27.3 mm SVL) the distance reaches or surpasses the anterior insertion of thigh, whereas in all specimens of
|
||
<taxonomicName id="4C1D4D31FFA5FF9B6D767EAEFE71F940" ID-CoL="6775P" box="[337,482,1698,1721]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA5FF9B6D767EAEFE71F940" box="[337,482,1698,1721]" italics="true" pageId="9" pageNumber="27">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
examined (right, MHNLS 17401, 23.3 mm SVL), the distance fails to reach the thigh. Photos by PJRK.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA236B2FFA5FF9B6CE27F04FAEFF825" blockId="9.[151,1436,1799,2012]" pageId="9" pageNumber="27">
|
||
<taxonomicName id="4C1D4D31FFA5FF9B6CE27F04FDAAF8D8" ID-CoL="6775Q" box="[197,569,1800,1825]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA5FF9B6CE27F04FDAAF8D8" box="[197,569,1800,1825]" italics="true" pageId="9" pageNumber="27">Anomaloglossus megacephalus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C1D4D31FFA5FF9B6F587F04FB82F8D8" ID-CoL="66V63" box="[895,1041,1800,1825]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="praderioi">
|
||
<emphasis id="B969EAA0FFA5FF9B6F587F04FB82F8D8" box="[895,1041,1800,1825]" italics="true" pageId="9" pageNumber="27">A. praderioi</emphasis>
|
||
</taxonomicName>
|
||
by (characters of
|
||
<taxonomicName id="4C1D4D31FFA5FF9B68CC7F04FAEFF8D8" ID-CoL="66V63" box="[1259,1404,1800,1825]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="praderioi">
|
||
<emphasis id="B969EAA0FFA5FF9B68CC7F04FAEFF8D8" box="[1259,1404,1800,1825]" italics="true" pageId="9" pageNumber="27">A. praderioi</emphasis>
|
||
</taxonomicName>
|
||
in parentheses, see also
|
||
<bibRefCitation id="EF8C4B43FFA5FF9B6DA87F21FD96F8BE" author="Kok" box="[399,517,1837,1863]" pageId="9" pageNumber="27" refString="Kok, P. J. R. (2010) A redescription of Anomaloglossus praderioi (La Marca, 1998) (Anura: Aromobatidae: Anomaloglossinae), with description of its tadpole and call. Papeis Avulsos de Zoologia, 50 (4), 51 - 68." type="journal article" year="2010">Kok 2010</bibRefCitation>
|
||
) its larger size, female SVL max
|
||
<quantity id="4CE59B57FFA5FF9B6FAE7F21FC62F8BE" box="[905,1009,1837,1863]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="9" pageNumber="27" unit="mm" value="28.3">28.3 mm</quantity>
|
||
(n=3) in
|
||
<taxonomicName id="4C1D4D31FFA5FF9B687B7F21FAB6F8BF" ID-CoL="6775Q" box="[1116,1317,1837,1862]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA5FF9B687B7F21FAB6F8BF" box="[1116,1317,1837,1862]" italics="true" pageId="9" pageNumber="27">A. megacephalus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CE59B57FFA5FF9B69147F21FA0FF8BE" box="[1331,1436,1837,1863]" metricMagnitude="-2" metricUnit="m" metricValue="2.27" pageId="9" pageNumber="27" unit="mm" value="22.7">22.7 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C1D4D31FFA5FF9B6C927F5EFED7F892" ID-CoL="66V63" box="[181,324,1874,1899]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="praderioi">
|
||
<emphasis id="B969EAA0FFA5FF9B6C927F5EFED7F892" box="[181,324,1874,1899]" italics="true" pageId="9" pageNumber="27">A. praderioi</emphasis>
|
||
</taxonomicName>
|
||
, n=1), Finger IV longer than I (equal), tip of Finger IV always surpassing the base of the distal subarticular tubercle on Finger III when fingers adpressed (barely reaching the base), toes moderately webbed with folded flaplike fringing (basally webbed, folded flaplike fringing not present on all toes), dorsolateral stripe absent (present), oblique lateral stripe present (absent), throat in adult female blotched (immaculate).
|
||
</paragraph>
|
||
<paragraph id="8BA236B2FFA6FF986CE27894FF6EFED8" blockId="10.[151,1437,152,1294]" pageId="10" pageNumber="28">
|
||
<taxonomicName id="4C1D4D31FFA6FF986CE27894FDA6FF48" ID-CoL="6775Q" box="[197,565,152,177]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA6FF986CE27894FDA6FF48" box="[197,565,152,177]" italics="true" pageId="10" pageNumber="28">Anomaloglossus megacephalus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C1D4D31FFA6FF986F527894FC66FF48" ID-CoL="676TR" box="[885,1013,152,177]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="roraima">
|
||
<emphasis id="B969EAA0FFA6FF986F527894FC66FF48" box="[885,1013,152,177]" italics="true" pageId="10" pageNumber="28">A. roraima</emphasis>
|
||
</taxonomicName>
|
||
in having (characters of
|
||
<taxonomicName id="4C1D4D31FFA6FF98693B7894FA0FFF48" ID-CoL="676TR" box="[1308,1436,152,177]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="roraima">
|
||
<emphasis id="B969EAA0FFA6FF98693B7894FA0FFF48" box="[1308,1436,152,177]" italics="true" pageId="10" pageNumber="28">A. roraima</emphasis>
|
||
</taxonomicName>
|
||
in parentheses) fingers with keel-like lateral folds, best developed preaxially on Fingers II and III (fringes barely distinct, not distinctly folded), all toes with folded flaplike fringing (absent), webbing on toes moderate (absent).
|
||
</paragraph>
|
||
<paragraph id="8BA236B2FFA6FF986CE27921FE78FE4F" blockId="10.[151,1437,152,1294]" pageId="10" pageNumber="28">
|
||
<taxonomicName id="4C1D4D31FFA6FF986CE27921FDCAFEBF" ID-CoL="6775Q" box="[197,601,301,326]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA6FF986CE27921FDCAFEBF" box="[197,601,301,326]" italics="true" pageId="10" pageNumber="28">Anomaloglossus megacephalus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C1D4D31FFA6FF986F997921FA8AFEBF" ID-CoL="6775R" authority="Gorzula, 1990" authorityName="Gorzula" authorityYear="1990" box="[958,1305,300,326]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="rufulus">
|
||
<emphasis id="B969EAA0FFA6FF986F997921FBD3FEBF" box="[958,1088,301,326]" italics="true" pageId="10" pageNumber="28">A. rufulus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8C4B43FFA6FF9868757920FA83FEBF" author="Gorzula" box="[1106,1296,300,326]" pageId="10" pageNumber="28" refString="Gorzula, S. (1990 " 1988 ") Una nueva especie de Dendrobates (Amphibia: Dendrobatidae) del Macizo de Chimanta, Estado Bolivar, Venezuela. Memoria de la Sociedad de Ciencias Naturales La Salle, 48, 143 - 149." type="journal article" year="1990">Gorzula, 1990</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in having (characters of
|
||
<taxonomicName id="4C1D4D31FFA6FF986D64795EFE29FE92" ID-CoL="6775R" box="[323,442,338,363]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="rufulus">
|
||
<emphasis id="B969EAA0FFA6FF986D64795EFE29FE92" box="[323,442,338,363]" italics="true" pageId="10" pageNumber="28">A. rufulus</emphasis>
|
||
</taxonomicName>
|
||
in parentheses) webbing on toes moderate (rudimentary), and most remarkably by ventral colour pattern; light with a few dark brown blotches in
|
||
<taxonomicName id="4C1D4D31FFA6FF986FAA797BFBCAFE69" ID-CoL="6775Q" box="[909,1113,375,400]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA6FF986FAA797BFBCAFE69" box="[909,1113,375,400]" italics="true" pageId="10" pageNumber="28">A. megacephalus</emphasis>
|
||
</taxonomicName>
|
||
(dark brown marbled with white blotches in
|
||
<taxonomicName id="4C1D4D31FFA6FF986D417990FE49FE4C" ID-CoL="6775R" box="[358,474,412,437]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="rufulus">
|
||
<emphasis id="B969EAA0FFA6FF986D417990FE49FE4C" box="[358,474,412,437]" italics="true" pageId="10" pageNumber="28">A. rufulus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA236B2FFA6FF986CE279CEFC12FC63" blockId="10.[151,1437,152,1294]" pageId="10" pageNumber="28">
|
||
The new species superficially most resembles
|
||
<taxonomicName id="4C1D4D31FFA6FF986F3079CEFC3DFE22" ID-CoL="66V5R" box="[791,942,450,475]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parkerae">
|
||
<emphasis id="B969EAA0FFA6FF986F3079CEFC3DFE22" box="[791,942,450,475]" italics="true" pageId="10" pageNumber="28">A. parkerae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D31FFA6FF986F9879CEFBFBFE22" ID-CoL="6775P" box="[959,1128,450,475]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA6FF986F9879CEFBFBFE22" box="[959,1128,450,475]" italics="true" pageId="10" pageNumber="28">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
(with which it has been confused), and
|
||
<taxonomicName id="4C1D4D31FFA6FF986D6879EBFB34FDF8" ID-CoL="5V766" authority="Barrio-Amoros, Fuentes-Ramos & Rivas-Fuenmayor, 2004" authorityName="Barrio-Amoros, Fuentes-Ramos & Rivas-Fuenmayor" authorityYear="2004" box="[335,1191,487,513]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="triunfo">
|
||
<emphasis id="B969EAA0FFA6FF986D6879EBFE51FDF9" box="[335,450,487,512]" italics="true" pageId="10" pageNumber="28">A. triunfo</emphasis>
|
||
(Barrio-Amorós,
|
||
<bibRefCitation id="EF8C4B43FFA6FF986EBD79EBFB0CFDF8" author="Fuentes-Ramos" box="[666,1183,487,513]" pageId="10" pageNumber="28" refString="Barrio-Amoros, C. L., Fuentes-Ramos, O. & Rivas, G. (2004) Two new species of Colostethus (Anura: Dendrobatidae) from the Venezuelan Guayana. Salamandra, 40, 183 - 200." type="journal article" year="2004">Fuentes-Ramos & Rivas-Fuenmayor, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, in sharing a similar dorsal pattern. It should be noted that
|
||
<taxonomicName id="4C1D4D31FFA6FF986E557A00FD7AFDDC" ID-CoL="5V766" box="[626,745,524,549]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="triunfo">
|
||
<emphasis id="B969EAA0FFA6FF986E557A00FD7AFDDC" box="[626,745,524,549]" italics="true" pageId="10" pageNumber="28">A. triunfo</emphasis>
|
||
</taxonomicName>
|
||
is a possible synonym of
|
||
<taxonomicName id="4C1D4D31FFA6FF9868087A00FB47FDDC" ID-CoL="6775P" box="[1071,1236,524,549]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA6FF9868087A00FB47FDDC" box="[1071,1236,524,549]" italics="true" pageId="10" pageNumber="28">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
(Kok & Barrio- Amorós unpubl. data, see also comments by
|
||
<bibRefCitation id="EF8C4B43FFA6FF986EFD7A3DFB89FDB2" author="Myers" box="[730,1050,561,587]" pageId="10" pageNumber="28" refString="Myers, C. W. & Donnelly, M. A. (2008) The summit herpetofauna of Auyantepui, Venezuela: report from the Robert G. Goelet American Museum - Terramar Expedition. Bulletin of the American Museum of Natural History, 308, 1 - 147." type="journal article" year="2008">Myers & Donnelly 2008</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C1D4D31FFA6FF9868167A3DFAEDFDB3" ID-CoL="66V5R" box="[1073,1406,561,586]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parkerae">
|
||
<emphasis id="B969EAA0FFA6FF9868167A3DFAEDFDB3" box="[1073,1406,561,586]" italics="true" pageId="10" pageNumber="28">Anomaloglossus parkerae</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C1D4D31FFA6FF986DA37A5BFDB7FD89" ID-CoL="6775P" box="[388,548,599,624]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA6FF986DA37A5BFDB7FD89" box="[388,548,599,624]" italics="true" pageId="10" pageNumber="28">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D31FFA6FF986E467A5BFD46FD89" ID-CoL="5V766" box="[609,725,599,624]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="triunfo">
|
||
<emphasis id="B969EAA0FFA6FF986E467A5BFD46FD89" box="[609,725,599,624]" italics="true" pageId="10" pageNumber="28">A. triunfo</emphasis>
|
||
</taxonomicName>
|
||
only by a few characters such as size, definition of dorsal pattern, and condition of finger and toe fringes (
|
||
<bibRefCitation id="EF8C4B43FFA6FF986EF87A77FB9DFD6C" author="Myers" box="[735,1038,635,661]" pageId="10" pageNumber="28" refString="Myers, C. W. & Donnelly, M. A. (2008) The summit herpetofauna of Auyantepui, Venezuela: report from the Robert G. Goelet American Museum - Terramar Expedition. Bulletin of the American Museum of Natural History, 308, 1 - 147." type="journal article" year="2008">Myers & Donnelly 2008</bibRefCitation>
|
||
). Adults of
|
||
<taxonomicName id="4C1D4D31FFA6FF9868BB7A70FAF9FD6C" ID-CoL="6775Q" box="[1180,1386,636,661]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA6FF9868BB7A70FAF9FD6C" box="[1180,1386,636,661]" italics="true" pageId="10" pageNumber="28">A. megacephalus</emphasis>
|
||
</taxonomicName>
|
||
can notably be distinguished from those three similar taxa in having a more slender body (
|
||
<taxonomicName id="4C1D4D31FFA6FF9868F57AADFE92FD26" ID-CoL="66V5R" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parkerae">
|
||
<emphasis id="B969EAA0FFA6FF9868F57AADFE92FD26" italics="true" pageId="10" pageNumber="28">Anomaloglossus parkerae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D31FFA6FF986D287ACAFE3AFD26" ID-CoL="6775P" box="[271,425,710,735]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA6FF986D287ACAFE3AFD26" box="[271,425,710,735]" italics="true" pageId="10" pageNumber="28">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D31FFA6FF986DC57ACAFDC0FD26" ID-CoL="5V766" box="[482,595,710,735]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="triunfo">
|
||
<emphasis id="B969EAA0FFA6FF986DC57ACAFDC0FD26" box="[482,595,710,735]" italics="true" pageId="10" pageNumber="28">A. triunfo</emphasis>
|
||
</taxonomicName>
|
||
are distinctly stockier frogs), a comparatively longer snout and a larger, somewhat more massive head [See
|
||
<figureCitation id="13262A37FFA6FF986E187AE7FD16FCFC" box="[575,645,747,773]" captionStart="FIGURE 5" captionStartId="9.[151,256,1506,1530]" captionTargetBox="[151,1436,194,1484]" captionTargetId="figure@9.[151,1436,194,1484]" captionTargetPageId="9" captionText="FIGURE 5. Comparison of Anomaloglossus megacephalus sp. nov. and the similar A. tepuyensis and A. parkerae. A: Dorsal view of preserved specimens of (from left to right) A. megacephalus sp. nov. (IRSNB 1986, 27.3 mm SVL), A. tepuyensis (MHNLS 17401, 23.3 mm SVL) and A. parkerae (MHNLS 11089, 23.2 mm SVL). B: Comparison between midline distance from snout tip to an imaginary line between anterior arm insertions and the same distance reported posteriorly from the same imaginary line in A. megacephalus sp. nov. and A. tepuyensis. In most A. megacephalus specimens (left, IRSNB 1986, 27.3 mm SVL) the distance reaches or surpasses the anterior insertion of thigh, whereas in all specimens of A. tepuyensis examined (right, MHNLS 17401, 23.3 mm SVL), the distance fails to reach the thigh. Photos by PJRK." httpUri="https://zenodo.org/record/199009/files/figure.png" pageId="10" pageNumber="28">Fig. 5</figureCitation>
|
||
for close comparison between
|
||
<taxonomicName id="4C1D4D31FFA6FF986FDD7AE7FB57FCFD" ID-CoL="6775Q" box="[1018,1220,747,772]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA6FF986FDD7AE7FB57FCFD" box="[1018,1220,747,772]" italics="true" pageId="10" pageNumber="28">A. megacephalus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D31FFA6FF9868D87AE7FA0FFCFD" ID-CoL="6775P" box="[1279,1436,747,772]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA6FF9868D87AE7FA0FFCFD" box="[1279,1436,747,772]" italics="true" pageId="10" pageNumber="28">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D31FFA6FF986CED7B1DFEC4FCD3" ID-CoL="66V5R" box="[202,343,785,810]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parkerae">
|
||
<emphasis id="B969EAA0FFA6FF986CED7B1DFEC4FCD3" box="[202,343,785,810]" italics="true" pageId="10" pageNumber="28">A. parkerae</emphasis>
|
||
</taxonomicName>
|
||
; compare also with Duellman’s (1997: 10) colour plate of
|
||
<taxonomicName id="4C1D4D31FFA6FF98683F7B1DFB36FCD3" ID-CoL="66V5R" box="[1048,1189,785,810]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parkerae">
|
||
<emphasis id="B969EAA0FFA6FF98683F7B1DFB36FCD3" box="[1048,1189,785,810]" italics="true" pageId="10" pageNumber="28">A. parkerae</emphasis>
|
||
</taxonomicName>
|
||
, fig. 30 (showing
|
||
<taxonomicName id="4C1D4D31FFA6FF9869A47B1DFE97FCB6" ID-CoL="66V5R" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parkerae">
|
||
<emphasis id="B969EAA0FFA6FF9869A47B1DFE97FCB6" italics="true" pageId="10" pageNumber="28">A. parkerae</emphasis>
|
||
</taxonomicName>
|
||
) in
|
||
<bibRefCitation id="EF8C4B43FFA6FF986D107B3AFDA3FCA9" box="[311,560,822,848]" pageId="10" pageNumber="28" refString="Lotters, S., Jungfer, K. H., Henkel, W. F. & Schmidt, W. (2007) Poison Frogs: Biology, Species & Captive Husbandry. Chimaira, Frankfurt aim Main, Germany, 668 pp." type="book">
|
||
Lötters
|
||
<emphasis id="B969EAA0FFA6FF986DB17B3AFE47FCB6" box="[406,468,822,847]" italics="true" pageId="10" pageNumber="28">et al.</emphasis>
|
||
(2007)
|
||
</bibRefCitation>
|
||
, and figs. 13, 15 and 16 illustrating
|
||
<taxonomicName id="4C1D4D31FFA6FF986FCB7B3AFB1DFCB6" ID-CoL="6775P" box="[1004,1166,822,847]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA6FF986FCB7B3AFB1DFCB6" box="[1004,1166,822,847]" italics="true" pageId="10" pageNumber="28">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EF8C4B43FFA6FF9868917B3AFF7FFC8C" author="Myers" pageId="10" pageNumber="28" refString="Myers, C. W. & Donnelly, M. A. (2008) The summit herpetofauna of Auyantepui, Venezuela: report from the Robert G. Goelet American Museum - Terramar Expedition. Bulletin of the American Museum of Natural History, 308, 1 - 147." type="journal article" year="2008">Myers & Donnelly (2008)</bibRefCitation>
|
||
; see also below for further comments], and in having symmetrical cloacal tubercles (absent in all females of
|
||
<taxonomicName id="4C1D4D31FFA6FF986D3E7B8CFE37FC60" ID-CoL="66V5R" box="[281,420,896,921]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parkerae">
|
||
<emphasis id="B969EAA0FFA6FF986D3E7B8CFE37FC60" box="[281,420,896,921]" italics="true" pageId="10" pageNumber="28">A. parkerae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D31FFA6FF986D957B8CFDDEFC60" ID-CoL="6775P" box="[434,589,896,921]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA6FF986D957B8CFDDEFC60" box="[434,589,896,921]" italics="true" pageId="10" pageNumber="28">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D31FFA6FF986EA27B8CFD65FC60" ID-CoL="5V766" box="[645,758,896,921]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="triunfo">
|
||
<emphasis id="B969EAA0FFA6FF986EA27B8CFD65FC60" box="[645,758,896,921]" italics="true" pageId="10" pageNumber="28">A. triunfo</emphasis>
|
||
</taxonomicName>
|
||
examined).
|
||
</paragraph>
|
||
<paragraph id="8BA236B2FFA6FF986CE27BA9FDC8FAF7" blockId="10.[151,1437,152,1294]" pageId="10" pageNumber="28">
|
||
Direct comparison of specimens shows a difference in head size in
|
||
<taxonomicName id="4C1D4D31FFA6FF9868307BAAFA0EFC46" ID-CoL="6775Q" box="[1047,1437,934,959]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA6FF9868307BAAFA0EFC46" box="[1047,1437,934,959]" italics="true" pageId="10" pageNumber="28">Anomaloglossus megacephalus</emphasis>
|
||
</taxonomicName>
|
||
compared to the similar
|
||
<taxonomicName id="4C1D4D31FFA6FF986D9F7BC7FDD0FC1D" ID-CoL="66V5R" box="[440,579,971,996]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parkerae">
|
||
<emphasis id="B969EAA0FFA6FF986D9F7BC7FDD0FC1D" box="[440,579,971,996]" italics="true" pageId="10" pageNumber="28">A. parkerae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D31FFA6FF986E747BC7FD7DFC1D" ID-CoL="6775P" box="[595,750,971,996]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA6FF986E747BC7FD7DFC1D" box="[595,750,971,996]" italics="true" pageId="10" pageNumber="28">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D31FFA6FF986F0E7BC7FC09FC1D" ID-CoL="5V766" box="[809,922,971,996]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="triunfo">
|
||
<emphasis id="B969EAA0FFA6FF986F0E7BC7FC09FC1D" box="[809,922,971,996]" italics="true" pageId="10" pageNumber="28">A. triunfo</emphasis>
|
||
</taxonomicName>
|
||
, as confirmed by plotting of morphometric proportions. Comparison of HL3 with BEL indicates that the new species proportionally has a longer head than
|
||
<taxonomicName id="4C1D4D31FFA6FF986CF77C19FEF9FBD7" ID-CoL="6775P" box="[208,362,1045,1070]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA6FF986CF77C19FEF9FBD7" box="[208,362,1045,1070]" italics="true" pageId="10" pageNumber="28">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13262A37FFA6FF986D5E7C19FE53FBD6" box="[377,448,1045,1071]" captionStart="FIGURE 6" captionStartId="11.[151,258,1400,1424]" captionTargetBox="[151,1436,595,1379]" captionTargetId="figure@11.[151,1436,595,1379]" captionTargetPageId="11" captionText="FIGURE 6. Graph showing correlation between HL 3 and BEL with standard errors for adult female specimens of Anomaloglossus tepuyensis (green triangles) and A. megacephalus sp. nov. (orange circles). Abbreviations are defined in the text. The graph clearly shows that the head of A. megacephalus sp. nov. is proportionally longer that the head of A. tepuyensis." httpUri="https://zenodo.org/record/199010/files/figure.png" pageId="10" pageNumber="28">Fig. 6</figureCitation>
|
||
). In order to distinguish between those similar species, the midline distance from the tip of the snout to an imaginary line between anterior insertions of upper arms is measured, and that distance is then extended posteriorly from the same imaginary line. In two of the three available
|
||
<taxonomicName id="4C1D4D31FFA6FF9868A87C6CFACAFB80" ID-CoL="6775Q" box="[1167,1369,1120,1145]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA6FF9868A87C6CFACAFB80" box="[1167,1369,1120,1145]" italics="true" pageId="10" pageNumber="28">A. megacephalus</emphasis>
|
||
</taxonomicName>
|
||
adult specimens the distance reaches or surpasses the anterior insertion of thigh, whereas in all adult specimens of
|
||
<taxonomicName id="4C1D4D31FFA6FF986CB07CA6FEB3FB3A" ID-CoL="66V5R" box="[151,288,1194,1219]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="parkerae">
|
||
<emphasis id="B969EAA0FFA6FF986CB07CA6FEB3FB3A" box="[151,288,1194,1219]" italics="true" pageId="10" pageNumber="28">A. parkerae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1D4D31FFA6FF986D087CA6FE59FB3A" ID-CoL="6775P" box="[303,458,1194,1219]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="tepuyensis">
|
||
<emphasis id="B969EAA0FFA6FF986D087CA6FE59FB3A" box="[303,458,1194,1219]" italics="true" pageId="10" pageNumber="28">A. tepuyensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1D4D31FFA6FF986E247CA6FDE0FB3A" ID-CoL="5V766" box="[515,627,1194,1219]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="triunfo">
|
||
<emphasis id="B969EAA0FFA6FF986E247CA6FDE0FB3A" box="[515,627,1194,1219]" italics="true" pageId="10" pageNumber="28">A. triunfo</emphasis>
|
||
</taxonomicName>
|
||
examined (n = 20), the distance fails to reach the thigh insertion (see
|
||
<figureCitation id="13262A37FFA6FF986CB07CC3FF4FFB10" box="[151,220,1231,1257]" captionStart="FIGURE 5" captionStartId="9.[151,256,1506,1530]" captionTargetBox="[151,1436,194,1484]" captionTargetId="figure@9.[151,1436,194,1484]" captionTargetPageId="9" captionText="FIGURE 5. Comparison of Anomaloglossus megacephalus sp. nov. and the similar A. tepuyensis and A. parkerae. A: Dorsal view of preserved specimens of (from left to right) A. megacephalus sp. nov. (IRSNB 1986, 27.3 mm SVL), A. tepuyensis (MHNLS 17401, 23.3 mm SVL) and A. parkerae (MHNLS 11089, 23.2 mm SVL). B: Comparison between midline distance from snout tip to an imaginary line between anterior arm insertions and the same distance reported posteriorly from the same imaginary line in A. megacephalus sp. nov. and A. tepuyensis. In most A. megacephalus specimens (left, IRSNB 1986, 27.3 mm SVL) the distance reaches or surpasses the anterior insertion of thigh, whereas in all specimens of A. tepuyensis examined (right, MHNLS 17401, 23.3 mm SVL), the distance fails to reach the thigh. Photos by PJRK." httpUri="https://zenodo.org/record/199009/files/figure.png" pageId="10" pageNumber="28">Fig. 5</figureCitation>
|
||
B). It is noteworthy to mention that the only specimen of
|
||
<taxonomicName id="4C1D4D31FFA6FF986FAB7CDCFBC6FB10" ID-CoL="6775Q" box="[908,1109,1232,1257]" class="Amphibia" family="Aromobatidae" genus="Anomaloglossus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="megacephalus">
|
||
<emphasis id="B969EAA0FFA6FF986FAB7CDCFBC6FB10" box="[908,1109,1232,1257]" italics="true" pageId="10" pageNumber="28">A. megacephalus</emphasis>
|
||
</taxonomicName>
|
||
in which HL3 is not longer than BEL is in poor preservation state.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |