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<document id="1DEADA2FB2D15F7A5CEC7232559222F3" ID-DOI="10.5281/zenodo.11513175" ID-ISSN="0068-547X" ID-Zenodo-Dep="11513175" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1717720859923" checkinUser="felipe" docAuthor="Griswold, Charles E." docDate="2022" docId="03B487984B60FFB2293F90B9E331F479" docLanguage="en" docName="ProcCalifAcadSci.67.14.329-355.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (14)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Vytfutia halandrefana Griswold 2022, new species" docType="treatment" docVersion="3" lastPageNumber="337" masterDocId="FF8DFFE04B68FFBB29449062E261F56A" masterDocTitle="The Lace Web Spider Genus Vytfutia Deeleman-Reinhold (Araneae, Phyxelididae) in the Indo-Pacific Region" masterLastPageNumber="355" masterPageNumber="329" pageNumber="337" updateTime="1717721805203" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="10778753FE02369FC4C1B3B6C8586785">
<mods:title id="BB0C9957F3D7D1131A754293828A5BC2">The Lace Web Spider Genus Vytfutia Deeleman-Reinhold (Araneae, Phyxelididae) in the Indo-Pacific Region</mods:title>
</mods:titleInfo>
<mods:name id="95A688C3D1B9A56AE94CD8D0F1B8A42D" type="personal">
<mods:role id="70B64D71D46F4E06B51114674FC593D7">
<mods:roleTerm id="537AE4EF274952665D588FE50B59FEF0">Author</mods:roleTerm>
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<mods:namePart id="ABEFD35B2C0F80357753AEC5E97CD491">Griswold, Charles E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C1456FAAA9CD5999E8B8AC1A9E7606B5">text</mods:typeOfResource>
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<mods:title id="5AF16CD92C07346512FDA25758D9845F">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="CAC8761948F1D14644D26C3117BF9EC4">
<mods:date id="58C3D4EF28C75845209426457C63B73F">2022</mods:date>
<mods:detail id="D2AB2580FF381F921DCDE271C6066808" type="pubDate">
<mods:number id="7245EEFB1ACE528D9065DD3962D62D79">2022-07-29</mods:number>
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<mods:number id="4AA7A70BE691522D19C118450167FFF9">67</mods:number>
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<mods:number id="BB9757DB46CF0549AB988A4C85755546">14</mods:number>
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<mods:classification id="BDD6C582C1A574B3AA2C588669FFE739">journal article</mods:classification>
<mods:identifier id="64F56C10F2C658C809DEB94620B9E5F3" type="DOI">10.5281/zenodo.11513175</mods:identifier>
<mods:identifier id="FA466D2544D06976D3A1921512D99FE8" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="01396B9E8EB843C50188B60D9CA4C7DC" type="Zenodo-Dep">11513175</mods:identifier>
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<treatment id="03B487984B60FFB2293F90B9E331F479" ID-DOI="http://doi.org/10.5281/zenodo.11513027" ID-Zenodo-Dep="11513027" LSID="urn:lsid:plazi:treatment:03B487984B60FFB2293F90B9E331F479" httpUri="http://treatment.plazi.org/id/03B487984B60FFB2293F90B9E331F479" lastPageId="9" lastPageNumber="337" pageId="8" pageNumber="337">
<subSubSection id="C30765054B60FFB3293F90B9E010F599" box="[123,625,219,243]" pageId="8" pageNumber="337" type="nomenclature">
<paragraph id="8BA2368E4B60FFB3293F90B9E010F599" blockId="8.[123,1174,219,1725]" box="[123,625,219,243]" pageId="8" pageNumber="337">
<heading id="D0EA81E24B60FFB3293F90B9E010F599" box="[123,625,219,243]" fontSize="10" level="2" pageId="8" pageNumber="337" reason="4">
<emphasis id="B969EA9C4B60FFB3293F90B9E010F599" bold="true" box="[123,625,219,243]" pageId="8" pageNumber="337">
<taxonomicName id="4C1D4D0D4B60FFB3293F90B9E380F599" ID-CoL="C8MQC" authority="Griswold" authorityName="Griswold" authorityYear="2022" box="[123,481,219,243]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="337" phylum="Arthropoda" rank="species" species="halandrefana" status="sp. nov.">
<emphasis id="B969EA9C4B60FFB3293F90B9E30EF599" bold="true" box="[123,367,219,243]" italics="true" pageId="8" pageNumber="337">Vytfutia halandrefana</emphasis>
Griswold
</taxonomicName>
,
<taxonomicNameLabel id="A25A57E74B60FFB328A990BEE010F599" box="[493,625,220,243]" pageId="8" pageNumber="337" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30765054B60FFB3293F909EE645F492" pageId="8" pageNumber="337" type="description">
<paragraph id="8BA2368E4B60FFB3293F909EE313F47B" blockId="8.[123,1174,219,1725]" box="[123,370,252,273]" pageId="8" pageNumber="337">
<figureCitation id="13262A0B4B60FFB3293F909EE369F47B" box="[123,264,252,273]" captionStart-0="FIGURES" captionStart-1="FIGURES" captionStartId-0="18.[155,167,1051,1070]" captionStartId-1="18.[155,167,1684,1703]" captionTargetBox-0="[216,1080,216,1035]" captionTargetBox-1="[224,1059,1157,1665]" captionTargetId-0="figure-19@18.[216,1080,216,1035]" captionTargetId-1="figure-120@18.[215,1081,1140,1672]" captionTargetPageId-0="18" captionTargetPageId-1="18" captionText-0="FIGURES 20–23. Vytfutia halandrefana new species, female, Berenty, Madagascar, CASENT9012513, habitus photos. 20. Dorsal. 21. Ventral. 22. Left metatarsus-tarsus IV and apex of abdomen with spinnerets, ventral. 23. Epigynal region, dorsal. Scale bars 20–22 = 0.5 mm, 23 = 0.25 mm." captionText-1="FIGURES 24, 25. Vytfutia halandrefana new species, female genitalia drawings, Tsingy de Bemaraha, Madagascar, CASENT9008848. 24. Epigynum, ventral. 25. Vulva, left half, dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.11513185" figureDoi-1="http://doi.org/10.5281/zenodo.11513187" httpUri-0="https://zenodo.org/record/11513185/files/figure.png" httpUri-1="https://zenodo.org/record/11513187/files/figure.png" pageId="8" pageNumber="337">Figures 20–25</figureCitation>
,
<figureCitation id="13262A0B4B60FFB32856909EE331F47B" box="[274,336,252,273]" captionStart="FIGURES" captionStartId="24.[155,167,765,784]" captionTargetBox="[120,1173,216,746]" captionTargetId="figure-19@24.[120,1176,216,747]" captionTargetPageId="24" captionText="FIGURES 50–55. Vytfutia halandrefana new species, Berenty, CASENT9012420, female legs and palps, SEM. 50. Metatarsus IV, retrolateral, showing calamistrum. 51. Metatarsus IV, retrolateral, showing calamistral setae. 52. Tarsus IV, apical, showing STC and ITC. 53. Palpal femur, anterolateral, showing thorns (arrow). 54. Palpal femur, anterolateral thorn, close up. 55. Tarsus IV, retrolateral showing STC and ITC. Scale bars: 50, 300 µm; 51, 20 µm, 52, 55, 50 µm, 53, 200 µm, 54, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.11513205" httpUri="https://zenodo.org/record/11513205/files/figure.png" pageId="8" pageNumber="337">50–55</figureCitation>
,
<figureCitation id="13262A0B4B60FFB3281E909EE313F47B" box="[346,370,252,273]" captionStart="FIGURE" captionStartId="26.[155,167,1071,1090]" captionTargetBox="[120,1176,216,1056]" captionTargetId="figure-19@26.[120,1176,216,1056]" captionTargetPageId="26" captionText="FIGURE 62. Vytfutia spp. Distributions on map of Eurasia and Australasia. Filled circle. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. Triangle. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia. Diamond. Vytfutia halandrefana new species, Madagascar. Star. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.11513211" httpUri="https://zenodo.org/record/11513211/files/figure.png" pageId="8" pageNumber="337">62</figureCitation>
</paragraph>
<paragraph id="8BA2368E4B60FFB329EF9142E645F492" blockId="8.[123,1174,219,1725]" pageId="8" pageNumber="337">
<emphasis id="B969EA9C4B60FFB329EF9142E297F45D" bold="true" box="[171,246,288,311]" pageId="8" pageNumber="337">Types:</emphasis>
<materialsCitation id="3B753CD34B60FFB329B99142E0D6F4FD" collectingDate="2002-02-03" collectingDateMax="2002-02-08" collectingDateMin="2002-02-03" collectionCode="CAS" country="Madagascar" county="Provence" elevation="34" latitude="-24.956945" location="Mandrare River" longLatPrecision="21" longitude="46.27139" municipality="Reserve Prive Berenty" pageId="8" pageNumber="337" specimenCode="CASENT9012513" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<typeStatus id="54A6882C4B60FFB329B99142E301F45D" box="[253,352,288,311]" pageId="8" pageNumber="337" type="holotype">Holotype</typeStatus>
female (
<specimenCode id="DBBB9EF54B60FFB328FA9142E0E6F45D" box="[446,647,288,311]" collectionCode="CASENT" pageId="8" pageNumber="337">CASENT9012513</specimenCode>
) from gallery forest along the
<location id="8EC260554B60FFB32A8B9142E618F45D" LSID="urn:lsid:plazi:treatment:03B487984B60FFB2293F90B9E331F479:8EC260554B60FFB32A8B9142E618F45D" box="[975,1145,288,311]" country="Madagascar" county="Provence" latitude="-24.956945" longLatPrecision="21" longitude="46.27139" municipality="Reserve Prive Berenty" name="Mandrare River" pageId="8" pageNumber="337" stateProvince="Toliara">Mandraré River</location>
in the
<location id="8EC260554B60FFB329EF9122E316F43D" LSID="urn:lsid:plazi:treatment:03B487984B60FFB2293F90B9E331F479:8EC260554B60FFB329EF9122E316F43D" box="[171,375,320,343]" country="Madagascar" county="Provence" latitude="-24.956945" longLatPrecision="21" longitude="46.27139" municipality="Reserve Prive Berenty" name="Foret de Bealoka" pageId="8" pageNumber="337" stateProvince="Toliara">Forêt de Bealoka</location>
,
<collectingMunicipality id="6BC6ACF44B60FFB328CF9122E0EEF43D" box="[395,655,320,343]" pageId="8" pageNumber="337">Réserve Privé Berenty</collectingMunicipality>
,
<collectingRegion id="49D9F86C4B60FFB32BE59122E08DF43D" box="[673,748,320,343]" country="Madagascar" name="Toliara" pageId="8" pageNumber="337">Toliara</collectingRegion>
<collectingCounty id="62C34E024B60FFB32BB99122E102F43D" box="[765,867,320,343]" pageId="8" pageNumber="337">Provence</collectingCounty>
,
<collectingCountry id="F30A761E4B60FFB32A329122E198F43D" box="[886,1017,320,343]" name="Madagascar" pageId="8" pageNumber="337">Madagascar</collectingCountry>
, (
<geoCoordinate id="EE2950494B60FFB32D519122E6F1F43D" box="[1045,1168,320,343]" degrees="24" direction="south" minutes="57" orientation="latitude" pageId="8" pageNumber="337" precision="15" seconds="25" value="-24.956945">24°57ʹ25ʺS</geoCoordinate>
,
<geoCoordinate id="EE2950494B60FFB3293F9102E299F41D" box="[123,248,352,375]" degrees="46" direction="east" minutes="16" orientation="longitude" pageId="8" pageNumber="337" precision="15" seconds="17" value="46.27139">46°16ʹ17ʺE</geoCoordinate>
, elev.
<quantity id="4CE59B6B4B60FFB328049102E312F41D" box="[320,371,352,375]" metricMagnitude="1" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="337" unit="m" value="34.0">
<elevation id="0030D1BD4B60FFB328049102E312F41D" box="[320,371,352,375]" metricMagnitude="1" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="337" unit="m" value="34.0">34m</elevation>
</quantity>
), collected
<date id="FFA3104E4B60FFB328B79102E0A2F41D" box="[499,707,352,375]" pageId="8" pageNumber="337" value="2002-02-03" valueMax="2002-02-08" valueMin="2002-02-03">
<collectingDate id="EFE7E9A64B60FFB328B79102E0A2F41D" box="[499,707,352,375]" pageId="8" pageNumber="337" value="2002-02-03" valueMax="2002-02-08" valueMin="2002-02-03">3–8 February 2002</collectingDate>
</date>
by the Fisher-Griswold Arthropod Survey Team, collection code BLF5320, deposited in
<collectionCode id="ED0CAE4B4B60FFB32BC491E2E0D6F4FD" box="[640,695,384,407]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="337" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B753CD34B60FFB32B8191E2E641F492" collectingDate="2001-11-06" collectingDateMax="2001-11-10" collectingDateMin="2001-11-06" collectionCode="CAS" country="Madagascar" county="Provence" elevation="50" latitude="-19.141945" location="Tombeau Vazimba" longLatPrecision="21" longitude="44.828056" municipality="Parc National Tsingy de Bemaraha" pageId="8" pageNumber="337" specimenCode="CASENT9008848" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<typeStatus id="54A6882C4B60FFB32B8191E2E143F4FD" box="[709,802,384,407]" pageId="8" pageNumber="337" type="paratype">Paratype</typeStatus>
female (
<specimenCode id="DBBB9EF54B60FFB32AC391E2E631F4FD" box="[903,1104,384,407]" collectionCode="CASENT" pageId="8" pageNumber="337">CASENT9008848</specimenCode>
) from tropical dry forest at
<location id="8EC260554B60FFB3282291C3E053F4D2" LSID="urn:lsid:plazi:treatment:03B487984B60FFB2293F90B9E331F479:8EC260554B60FFB3282291C3E053F4D2" box="[358,562,417,440]" country="Madagascar" county="Provence" latitude="-19.141945" longLatPrecision="21" longitude="44.828056" municipality="Parc National Tsingy de Bemaraha" name="Tombeau Vazimba" pageId="8" pageNumber="337" stateProvince="Toliara">Tombeau Vazimba</location>
,
<location id="8EC260554B60FFB32B0491C3E0CCF4D2" LSID="urn:lsid:plazi:treatment:03B487984B60FFB2293F90B9E331F479:8EC260554B60FFB32B0491C3E0CCF4D2" box="[576,685,417,440]" country="Madagascar" county="Provence" latitude="-19.141945" longLatPrecision="21" longitude="44.828056" municipality="Parc National Tsingy de Bemaraha" name="Bekopaka" pageId="8" pageNumber="337" stateProvince="Toliara">Bekopaka</location>
,
<collectingMunicipality id="6BC6ACF44B60FFB32BFF91C3E65CF4D2" box="[699,1085,417,440]" pageId="8" pageNumber="337">Parc National Tsingy de Bemaraha</collectingMunicipality>
,
<collectingRegion id="49D9F86C4B60FFB32D0E91C3E6F4F4D2" box="[1098,1173,417,440]" country="Madagascar" name="Toliara" pageId="8" pageNumber="337">Toliara</collectingRegion>
<collectingCounty id="62C34E024B60FFB3293F91A3E280F4B2" box="[123,225,449,472]" pageId="8" pageNumber="337">Provence</collectingCounty>
,
<collectingCountry id="F30A761E4B60FFB329AF91A3E30FF4B2" box="[235,366,449,472]" name="Madagascar" pageId="8" pageNumber="337">Madagascar</collectingCountry>
, (
<geoCoordinate id="EE2950494B60FFB328C491A3E38FF4B2" box="[384,494,449,472]" degrees="19" direction="south" minutes="8" orientation="latitude" pageId="8" pageNumber="337" precision="15" seconds="31" value="-19.141945">19°8ʹ31ʺS</geoCoordinate>
,
<geoCoordinate id="EE2950494B60FFB328BE91A3E016F4B2" box="[506,631,449,472]" degrees="44" direction="east" minutes="49" orientation="longitude" pageId="8" pageNumber="337" precision="15" seconds="41" value="44.828056">44°49ʹ41ʺE</geoCoordinate>
, elev.
<quantity id="4CE59B6B4B60FFB32BFD91A3E08DF4B2" box="[697,748,449,472]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="337" unit="m" value="50.0">
<elevation id="0030D1BD4B60FFB32BFD91A3E08DF4B2" box="[697,748,449,472]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="337" unit="m" value="50.0">50m</elevation>
</quantity>
), collected
<date id="FFA3104E4B60FFB32A2191A3E62CF4B2" box="[869,1101,449,472]" pageId="8" pageNumber="337" value="2001-11-06" valueMax="2001-11-10" valueMin="2001-11-06">
<collectingDate id="EFE7E9A64B60FFB32A2191A3E62CF4B2" box="[869,1101,449,472]" pageId="8" pageNumber="337" value="2001-11-06" valueMax="2001-11-10" valueMin="2001-11-06">6–10 November 2001</collectingDate>
</date>
by the Fisher-Griswold Arthropod Survey Team, collection code BLF4230, deposited in
<collectionCode id="ED0CAE4B4B60FFB32AAD9183E641F492" box="[1001,1056,481,504]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="337" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30765054B60FFB329EF9263E290F732" pageId="8" pageNumber="337" type="etymology">
<paragraph id="8BA2368E4B60FFB329EF9263E290F732" blockId="8.[123,1174,219,1725]" pageId="8" pageNumber="337">
<emphasis id="B969EA9C4B60FFB329EF9263E34EF772" bold="true" box="[171,303,513,536]" pageId="8" pageNumber="337">Etymology:</emphasis>
The specific name is from the Malagasy words for spider,
<emphasis id="B969EA9C4B60FFB32AA39263E677F772" box="[999,1046,513,536]" italics="true" pageId="8" pageNumber="337">hala</emphasis>
, and west,
<emphasis id="B969EA9C4B60FFB3293F9243E289F752" box="[123,232,545,568]" italics="true" pageId="8" pageNumber="337">andrefana</emphasis>
, commemorating the distribution of this westernmost
<taxonomicName id="4C1D4D0D4B60FFB32A249243E671F752" box="[864,1040,545,568]" pageId="8" pageNumber="337">
<emphasis id="B969EA9C4B60FFB32A249243E1D3F752" box="[864,946,545,568]" italics="true" pageId="8" pageNumber="337">Vytfutia</emphasis>
species
</taxonomicName>
; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C30765054B60FFB329EF9203E1B3F0B6" pageId="8" pageNumber="337" type="diagnosis">
<paragraph id="8BA2368E4B60FFB329EF9203E3FBF793" blockId="8.[123,1174,219,1725]" pageId="8" pageNumber="337">
<emphasis id="B969EA9C4B60FFB329EF9203E343F712" bold="true" box="[171,290,609,632]" pageId="8" pageNumber="337">Diagnosis:</emphasis>
Females with epigynum (
<figureCitation id="13262A0B4B60FFB32B1C9203E0BDF712" box="[600,732,609,632]" captionStart-0="FIGURES" captionStart-1="FIGURES" captionStartId-0="18.[155,167,1051,1070]" captionStartId-1="18.[155,167,1684,1703]" captionTargetBox-0="[216,1080,216,1035]" captionTargetBox-1="[224,1059,1157,1665]" captionTargetId-0="figure-19@18.[216,1080,216,1035]" captionTargetId-1="figure-120@18.[215,1081,1140,1672]" captionTargetPageId-0="18" captionTargetPageId-1="18" captionText-0="FIGURES 20–23. Vytfutia halandrefana new species, female, Berenty, Madagascar, CASENT9012513, habitus photos. 20. Dorsal. 21. Ventral. 22. Left metatarsus-tarsus IV and apex of abdomen with spinnerets, ventral. 23. Epigynal region, dorsal. Scale bars 20–22 = 0.5 mm, 23 = 0.25 mm." captionText-1="FIGURES 24, 25. Vytfutia halandrefana new species, female genitalia drawings, Tsingy de Bemaraha, Madagascar, CASENT9008848. 24. Epigynum, ventral. 25. Vulva, left half, dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.11513185" figureDoi-1="http://doi.org/10.5281/zenodo.11513187" httpUri-0="https://zenodo.org/record/11513185/files/figure.png" httpUri-1="https://zenodo.org/record/11513187/files/figure.png" pageId="8" pageNumber="337">Figs. 23-25</figureCitation>
) broad, quadrangle greater than 2.50, copulatory opening rims transverse, posterior margin of epigynum concave, vulva complex with large anterior lobe and slender ducts, vulva width less than 1.50 times length (
<figureCitation id="13262A0B4B60FFB32A9C92C0E64BF7D3" box="[984,1066,674,697]" captionStart="FIGURES" captionStartId="18.[155,167,1684,1703]" captionTargetBox="[224,1059,1157,1665]" captionTargetId="figure-120@18.[215,1081,1140,1672]" captionTargetPageId="18" captionText="FIGURES 24, 25. Vytfutia halandrefana new species, female genitalia drawings, Tsingy de Bemaraha, Madagascar, CASENT9008848. 24. Epigynum, ventral. 25. Vulva, left half, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513187" httpUri="https://zenodo.org/record/11513187/files/figure.png" pageId="8" pageNumber="337">Fig. 25</figureCitation>
); thoracic fovea oval, length less than 0.08 carapace length (
<figureCitation id="13262A0B4B60FFB32BCE92A0E0B8F7B3" box="[650,729,706,729]" captionStart="FIGURES" captionStartId="18.[155,167,1051,1070]" captionTargetBox="[216,1080,216,1035]" captionTargetId="figure-19@18.[216,1080,216,1035]" captionTargetPageId="18" captionText="FIGURES 20–23. Vytfutia halandrefana new species, female, Berenty, Madagascar, CASENT9012513, habitus photos. 20. Dorsal. 21. Ventral. 22. Left metatarsus-tarsus IV and apex of abdomen with spinnerets, ventral. 23. Epigynal region, dorsal. Scale bars 20–22 = 0.5 mm, 23 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.11513185" httpUri="https://zenodo.org/record/11513185/files/figure.png" pageId="8" pageNumber="337">Fig. 20</figureCitation>
); smallest
<taxonomicName id="4C1D4D0D4B60FFB32A0D92A0E1FAF7B3" authorityName="Deeleman-Reinhold" authorityYear="1986" box="[841,923,706,729]" class="Arachnida" family="Phyxelididae" genus="Vytfutia" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="337" phylum="Arthropoda" rank="genus">
<emphasis id="B969EA9C4B60FFB32A0D92A0E1FAF7B3" box="[841,923,706,729]" italics="true" pageId="8" pageNumber="337">Vytfutia</emphasis>
</taxonomicName>
, total length of females 2.12–2.42; male unknown.
</paragraph>
<paragraph id="8BA2368E4B60FFB329EF9360E078F0F6" blockId="8.[123,1174,219,1725]" pageId="8" pageNumber="337">
<emphasis id="B969EA9C4B60FFB329EF9360E28CF673" bold="true" box="[171,237,770,793]" pageId="8" pageNumber="337">Male:</emphasis>
Unknown.
<emphasis id="B969EA9C4B60FFB328359360E3A5F673" bold="true" box="[369,452,770,793]" pageId="8" pageNumber="337">Female</emphasis>
(
<typeStatus id="54A6882C4B60FFB328919360E056F673" box="[469,567,770,793]" pageId="8" pageNumber="337" type="holotype">holotype</typeStatus>
, CASENT9021513): Total length 2.12. Markings as in
<figureCitation id="13262A0B4B60FFB3293F9340E29FF653" box="[123,254,802,825]" captionStart="FIGURES" captionStartId="18.[155,167,1051,1070]" captionTargetBox="[216,1080,216,1035]" captionTargetId="figure-19@18.[216,1080,216,1035]" captionTargetPageId="18" captionText="FIGURES 20–23. Vytfutia halandrefana new species, female, Berenty, Madagascar, CASENT9012513, habitus photos. 20. Dorsal. 21. Ventral. 22. Left metatarsus-tarsus IV and apex of abdomen with spinnerets, ventral. 23. Epigynal region, dorsal. Scale bars 20–22 = 0.5 mm, 23 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.11513185" httpUri="https://zenodo.org/record/11513185/files/figure.png" pageId="8" pageNumber="337">Figs. 20, 21</figureCitation>
, carapace and chelicerae dark brown, labium and palpal coxae yellow-brown, venter and legs yellow-white; abdomen grey-white with dorsal brown cardiac marks broken into anterior and median parts, dorsum and sides with intermixed, scattered brown spots and white guanine deposits, venter plain yellow white. Carapace 0.95 long, 0.80 wide, 0.36 high; clypeus 0.04 high, height 0.80
<collectionCode id="ED0CAE4B4B60FFB329B893C1E346F6D0" box="[252,295,931,954]" country="Australia" name="Australian Museum" pageId="8" pageNumber="337" type="Museum">AM</collectionCode>
diameter; ocular area 0.18 long, 0.46 wide, OAL 1.07 times OQL; ratio of eyes
<collectionCode id="ED0CAE4B4B60FFB3293F93A1E2CBF6B0" box="[123,170,963,986]" country="Australia" name="Australian Museum" pageId="8" pageNumber="337" type="Museum">AM</collectionCode>
:AL:
<collectionCode id="ED0CAE4B4B60FFB3299393A1E363F6B0" box="[215,258,963,986]" country="USA" name="Pratt Museum" pageId="8" pageNumber="337">PM</collectionCode>
:
<collectionCode id="ED0CAE4B4B60FFB3284193A1E349F6B0" box="[261,296,963,986]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="8" pageNumber="337" type="Museum">PL</collectionCode>
, 1.00:1.11:1.11:1.22, diameter of
<collectionCode id="ED0CAE4B4B60FFB32BD393A1E0DFF6B0" box="[663,702,963,986]" country="USA" name="Pratt Museum" pageId="8" pageNumber="337">PM</collectionCode>
0.06; AM-AM 0.89
<collectionCode id="ED0CAE4B4B60FFB32AE593A1E1ADF6B0" box="[929,972,963,986]" country="Australia" name="Australian Museum" pageId="8" pageNumber="337" type="Museum">AM</collectionCode>
diameter, AM-AL 1.22
<collectionCode id="ED0CAE4B4B60FFB329EA9381E2BDF690" box="[174,220,995,1018]" country="Australia" name="Australian Museum" pageId="8" pageNumber="337" type="Museum">AM</collectionCode>
;
<collectionCode id="ED0CAE4B4B60FFB329A39381E35CF690" box="[231,317,995,1018]" pageId="8" pageNumber="337">PM-PM</collectionCode>
and PM-PL 1.60
<collectionCode id="ED0CAE4B4B60FFB328BE9381E045F690" box="[506,548,995,1018]" country="USA" name="Pratt Museum" pageId="8" pageNumber="337">PM</collectionCode>
; AL-PL 0.18 times
<collectionCode id="ED0CAE4B4B60FFB32BB79381E174F690" box="[755,789,995,1018]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="8" pageNumber="337" type="Museum">PL</collectionCode>
. Chelicerae 0.41 long; sternum 0.56 long, 0.50 wide; labium 0.18 long, 0.21 wide; palpal coxae 0.27 long, 0.15 wide. Femur I length 1.01 times carapace width. Spination greatly reduced compared to other
<taxonomicName id="4C1D4D0D4B60FFB32A9B9441E6F3F150" box="[991,1170,1059,1082]" pageId="8" pageNumber="337">
<emphasis id="B969EA9C4B60FFB32A9B9441E650F150" box="[991,1073,1059,1082]" italics="true" pageId="8" pageNumber="337">Vytfutia</emphasis>
species
</taxonomicName>
: Female: palp: femur d0-1-0, tibia d1-0-2, p0-0-1, tarsus p0-1-1, v1-1-2-2-1; leg I: femur d0-1-1-0, p0-0-0-1, tibia r0-1-1-0, metatarsus v2-0-0; leg II: femur d1-1-0, p0-0-1, tibia p1-1-0-2, r0-1-0, metatarsus v2-0-0-0; leg
<collectionCode id="ED0CAE4B4B60FFB328A794E6E060F1F1" box="[483,513,1156,1179]" pageId="8" pageNumber="337">III</collectionCode>
: femur d10-0-0; leg
<collectionCode id="ED0CAE4B4B60FFB32BAC94E6E167F1F1" box="[744,774,1156,1179]" pageId="8" pageNumber="337">IV</collectionCode>
: femur d10-0-0. Leg measurements: I: 0.81 + 0.34 + 0.72 + 0.62 + 0.56 = [3.05]; II: 0.75 + 0.31 + 0.56 + 0.50 + 0.31 = [2.43];
<collectionCode id="ED0CAE4B4B60FFB32D3794C6E6F0F1D1" box="[1139,1169,1188,1211]" pageId="8" pageNumber="337">III</collectionCode>
: 0.72 + 0.28 + 0.44 + 0.47 + 0.31 = [2.22];
<collectionCode id="ED0CAE4B4B60FFB32B1C94A6E017F1B1" box="[600,630,1220,1243]" pageId="8" pageNumber="337">IV</collectionCode>
: 0.81 + 0.34 + 0.59 + 0.56 + 0.28 = [2.58]; palp: 0.37 + 0.12 + 0.19 + na + 0.31 = [0.99]. Calamistrum as in
<figureCitation id="13262A0B4B60FFB32A4E9486E102F191" box="[778,867,1252,1275]" captionStart="FIGURES" captionStartId="18.[155,167,1051,1070]" captionTargetBox="[216,1080,216,1035]" captionTargetId="figure-19@18.[216,1080,216,1035]" captionTargetPageId="18" captionText="FIGURES 20–23. Vytfutia halandrefana new species, female, Berenty, Madagascar, CASENT9012513, habitus photos. 20. Dorsal. 21. Ventral. 22. Left metatarsus-tarsus IV and apex of abdomen with spinnerets, ventral. 23. Epigynal region, dorsal. Scale bars 20–22 = 0.5 mm, 23 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.11513185" httpUri="https://zenodo.org/record/11513185/files/figure.png" pageId="8" pageNumber="337">Figs. 22</figureCitation>
,
<figureCitation id="13262A0B4B60FFB32A2B9486E1D7F191" box="[879,950,1252,1275]" captionStart="FIGURES" captionStartId="24.[155,167,765,784]" captionTargetBox="[120,1173,216,746]" captionTargetId="figure-19@24.[120,1176,216,747]" captionTargetPageId="24" captionText="FIGURES 50–55. Vytfutia halandrefana new species, Berenty, CASENT9012420, female legs and palps, SEM. 50. Metatarsus IV, retrolateral, showing calamistrum. 51. Metatarsus IV, retrolateral, showing calamistral setae. 52. Tarsus IV, apical, showing STC and ITC. 53. Palpal femur, anterolateral, showing thorns (arrow). 54. Palpal femur, anterolateral thorn, close up. 55. Tarsus IV, retrolateral showing STC and ITC. Scale bars: 50, 300 µm; 51, 20 µm, 52, 55, 50 µm, 53, 200 µm, 54, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.11513205" httpUri="https://zenodo.org/record/11513205/files/figure.png" pageId="8" pageNumber="337">50, 51</figureCitation>
; palpal femur thorns as in
<figureCitation id="13262A0B4B60FFB329FD9566E321F071" box="[185,320,1284,1307]" captionStart="FIGURES" captionStartId="24.[155,167,765,784]" captionTargetBox="[120,1173,216,746]" captionTargetId="figure-19@24.[120,1176,216,747]" captionTargetPageId="24" captionText="FIGURES 50–55. Vytfutia halandrefana new species, Berenty, CASENT9012420, female legs and palps, SEM. 50. Metatarsus IV, retrolateral, showing calamistrum. 51. Metatarsus IV, retrolateral, showing calamistral setae. 52. Tarsus IV, apical, showing STC and ITC. 53. Palpal femur, anterolateral, showing thorns (arrow). 54. Palpal femur, anterolateral thorn, close up. 55. Tarsus IV, retrolateral showing STC and ITC. Scale bars: 50, 300 µm; 51, 20 µm, 52, 55, 50 µm, 53, 200 µm, 54, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.11513205" httpUri="https://zenodo.org/record/11513205/files/figure.png" pageId="8" pageNumber="337">Figs. 53, 54</figureCitation>
; tarsal claws
<collectionCode id="ED0CAE4B4B60FFB3289C9566E395F071" box="[472,500,1284,1307]" pageId="8" pageNumber="337">IV</collectionCode>
as in
<figureCitation id="13262A0B4B60FFB32B789566E0A2F071" box="[572,707,1284,1307]" captionStart="FIGURES" captionStartId="24.[155,167,765,784]" captionTargetBox="[120,1173,216,746]" captionTargetId="figure-19@24.[120,1176,216,747]" captionTargetPageId="24" captionText="FIGURES 50–55. Vytfutia halandrefana new species, Berenty, CASENT9012420, female legs and palps, SEM. 50. Metatarsus IV, retrolateral, showing calamistrum. 51. Metatarsus IV, retrolateral, showing calamistral setae. 52. Tarsus IV, apical, showing STC and ITC. 53. Palpal femur, anterolateral, showing thorns (arrow). 54. Palpal femur, anterolateral thorn, close up. 55. Tarsus IV, retrolateral showing STC and ITC. Scale bars: 50, 300 µm; 51, 20 µm, 52, 55, 50 µm, 53, 200 µm, 54, 30 µm." figureDoi="http://doi.org/10.5281/zenodo.11513205" httpUri="https://zenodo.org/record/11513205/files/figure.png" pageId="8" pageNumber="337">Figs. 52, 55</figureCitation>
. Epigynum as in
<figureCitation id="13262A0B4B60FFB32AC39566E182F071" box="[903,995,1284,1307]" captionStart="FIGURES" captionStartId="18.[155,167,1051,1070]" captionTargetBox="[216,1080,216,1035]" captionTargetId="figure-19@18.[216,1080,216,1035]" captionTargetPageId="18" captionText="FIGURES 20–23. Vytfutia halandrefana new species, female, Berenty, Madagascar, CASENT9012513, habitus photos. 20. Dorsal. 21. Ventral. 22. Left metatarsus-tarsus IV and apex of abdomen with spinnerets, ventral. 23. Epigynal region, dorsal. Scale bars 20–22 = 0.5 mm, 23 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.11513185" httpUri="https://zenodo.org/record/11513185/files/figure.png" pageId="8" pageNumber="337">Figs. 23</figureCitation>
,
<figureCitation id="13262A0B4B60FFB32AB59566E66FF071" box="[1009,1038,1284,1307]" captionStart="FIGURES" captionStartId="18.[155,167,1684,1703]" captionTargetBox="[224,1059,1157,1665]" captionTargetId="figure-120@18.[215,1081,1140,1672]" captionTargetPageId="18" captionText="FIGURES 24, 25. Vytfutia halandrefana new species, female genitalia drawings, Tsingy de Bemaraha, Madagascar, CASENT9008848. 24. Epigynum, ventral. 25. Vulva, left half, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513187" httpUri="https://zenodo.org/record/11513187/files/figure.png" pageId="8" pageNumber="337">24</figureCitation>
, vulva as in
<figureCitation id="13262A0B4B60FFB3293F9546E2ADF051" box="[123,204,1316,1339]" captionStart="FIGURES" captionStartId="18.[155,167,1684,1703]" captionTargetBox="[224,1059,1157,1665]" captionTargetId="figure-120@18.[215,1081,1140,1672]" captionTargetPageId="18" captionText="FIGURES 24, 25. Vytfutia halandrefana new species, female genitalia drawings, Tsingy de Bemaraha, Madagascar, CASENT9008848. 24. Epigynum, ventral. 25. Vulva, left half, dorsal." figureDoi="http://doi.org/10.5281/zenodo.11513187" httpUri="https://zenodo.org/record/11513187/files/figure.png" pageId="8" pageNumber="337">Fig. 25</figureCitation>
. Variation (N=2): Total length 2.12–2.42; carapace length 1.19–1.25 times width, height 0.45–0.56 width; PER 1.12–1.13 times
<collectionCode id="ED0CAE4B4B60FFB32B629526E03FF031" box="[550,606,1348,1371]" pageId="8" pageNumber="337">AER</collectionCode>
, 2.50–2.81 times OAL; OAL 1.04–1.07 times OQL; clypeal height 0.60–0.80 times
<collectionCode id="ED0CAE4B4B60FFB328899507E399F016" box="[461,504,1381,1404]" country="Australia" name="Australian Museum" pageId="8" pageNumber="337" type="Museum">AM</collectionCode>
diameter; cheliceral length 10.80–14.00 times clypeal height; sternum length 1.19–1.26 times width.
</paragraph>
<paragraph id="8BA2368E4B60FFB329EF95C7E1B3F0B6" blockId="8.[123,1174,219,1725]" pageId="8" pageNumber="337">
<emphasis id="B969EA9C4B60FFB329EF95C7E308F0D6" bold="true" box="[171,361,1445,1468]" pageId="8" pageNumber="337">Natural History:</emphasis>
Collection records from beating and pitfall trapping suggest that this species occurs on vegetation and on the ground in gallery forest and tropical dry forest.
</paragraph>
</subSubSection>
<subSubSection id="C30765054B60FFB329EF9587E63AF096" box="[171,1115,1509,1532]" pageId="8" pageNumber="337" type="distribution">
<paragraph id="8BA2368E4B60FFB329EF9587E63AF096" blockId="8.[123,1174,219,1725]" box="[171,1115,1509,1532]" pageId="8" pageNumber="337">
<emphasis id="B969EA9C4B60FFB329EF9587E35EF096" bold="true" box="[171,319,1509,1532]" pageId="8" pageNumber="337">Distribution:</emphasis>
Known from central and western
<collectingRegion id="49D9F86C4B60FFB32BEB9587E102F096" box="[687,867,1509,1532]" country="Madagascar" name="Toliara" pageId="8" pageNumber="337">Toliara Province</collectingRegion>
,
<collectingCountry id="F30A761E4B60FFB32A2A9587E18EF096" box="[878,1007,1509,1532]" name="Madagascar" pageId="8" pageNumber="337">Madagascar</collectingCountry>
(
<figureCitation id="13262A0B4B60FFB32ABA9587E62EF096" box="[1022,1103,1509,1532]" captionStart="FIGURE" captionStartId="26.[155,167,1071,1090]" captionTargetBox="[120,1176,216,1056]" captionTargetId="figure-19@26.[120,1176,216,1056]" captionTargetPageId="26" captionText="FIGURE 62. Vytfutia spp. Distributions on map of Eurasia and Australasia. Filled circle. Vytfutia bedel Deeleman-Reinhold, 1986, Holotype male from Gunung Leuser, border of National Park at Bohorok in primary rainforest, N. Sumatra, Indonesia. Triangle. Vytfutia labalaba new species, Holotype male from the wall of a building at 1550m elevation on Mt. Kinabalu, Sabah-Borneo, Malaysia. Diamond. Vytfutia halandrefana new species, Madagascar. Star. Vytfutia pallens Deeleman-Reinhold, 1989, male paratype from Niah Cave, Sarawak, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.11513211" httpUri="https://zenodo.org/record/11513211/files/figure.png" pageId="8" pageNumber="337">Fig. 62</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30765054B60FFB229EF9667E331F479" lastPageId="9" lastPageNumber="338" pageId="8" pageNumber="337" type="materials_examined">
<paragraph id="8BA2368E4B60FFB229EF9667E331F479" blockId="8.[123,1174,219,1725]" lastBlockId="9.[123,1173,220,275]" lastPageId="9" lastPageNumber="338" pageId="8" pageNumber="337">
<emphasis id="B969EA9C4B60FFB329EF9667E3F8F376" bold="true" box="[171,409,1541,1564]" pageId="8" pageNumber="337">Material Examined:</emphasis>
<materialsCitation id="3B753CD34B60FFB328EC9667E1C6F317" collectingDate="2002-02-03" collectingDateMax="2002-02-08" collectingDateMin="2002-02-03" collectionCode="CASENT" country="Madagascar" county="Reserve Prive Berenty" elevation="35" latitude="-24.956945" location="Mandrare River" longLatPrecision="21" longitude="46.27139" municipality="Foret de Bealoka" pageId="8" pageNumber="337" specimenCode="CASENT9021513" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<collectingCountry id="F30A761E4B60FFB328EC9667E00BF376" box="[424,618,1541,1564]" name="Madagascar" pageId="8" pageNumber="337">MADAGASCAR</collectingCountry>
:
<collectingRegion id="49D9F86C4B60FFB32B3F9667E16EF376" box="[635,783,1541,1564]" country="Madagascar" name="Toliara" pageId="8" pageNumber="337">Toliara Prov.</collectingRegion>
,
<collectingCounty id="62C34E024B60FFB32A669667E642F376" box="[802,1059,1541,1564]" pageId="8" pageNumber="337">Réserve Privé Berenty</collectingCounty>
,
<collectingMunicipality id="6BC6ACF44B60FFB32D719667E2B4F356" pageId="8" pageNumber="337">Forêt de Bealoka</collectingMunicipality>
,
<location id="8EC260554B60FFB329AC9647E3F5F356" LSID="urn:lsid:plazi:treatment:03B487984B60FFB2293F90B9E331F479:8EC260554B60FFB329AC9647E3F5F356" box="[232,404,1573,1596]" country="Madagascar" county="Reserve Prive Berenty" latitude="-24.956945" longLatPrecision="21" longitude="46.27139" municipality="Foret de Bealoka" name="Mandrare River" pageId="8" pageNumber="337" stateProvince="Toliara">Mandraré River</location>
,
<locationDeviation id="9A102FAC4B60FFB328EC9647E165F356" box="[424,772,1573,1596]" location="Amboasary" pageId="8" pageNumber="337">
<quantity id="4CE59B6B4B60FFB328EC9647E061F356" box="[424,512,1573,1596]" metricMagnitude="4" metricUnit="m" metricValue="1.46" pageId="8" pageNumber="337" unit="km" value="14.6">14.6 km</quantity>
329° NNW Amboasary
</locationDeviation>
, elev.
<quantity id="4CE59B6B4B60FFB32A0C9647E1E0F356" box="[840,897,1573,1596]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="337" unit="m" value="35.0">
<elevation id="0030D1BD4B60FFB32A0C9647E1E0F356" box="[840,897,1573,1596]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="337" unit="m" value="35.0">35 m</elevation>
</quantity>
,
<geoCoordinate id="EE2950494B60FFB32AC99647E666F356" box="[909,1031,1573,1596]" degrees="24" direction="south" minutes="57" orientation="latitude" pageId="8" pageNumber="337" precision="15" seconds="25" value="-24.956945">24°57ʹ25ʺS</geoCoordinate>
,
<geoCoordinate id="EE2950494B60FFB32D509647E6F0F356" box="[1044,1169,1573,1596]" degrees="46" direction="east" minutes="16" orientation="longitude" pageId="8" pageNumber="337" precision="15" seconds="17" value="46.27139">46°16ʹ17ʺE</geoCoordinate>
, beating low vegetation - gallery
<location id="8EC260554B60FFB328A19624E04BF337" LSID="urn:lsid:plazi:treatment:03B487984B60FFB2293F90B9E331F479:8EC260554B60FFB328A19624E04BF337" box="[485,554,1606,1629]" country="Madagascar" county="Reserve Prive Berenty" latitude="-24.956945" longLatPrecision="21" longitude="46.27139" municipality="Foret de Bealoka" name="Forest" pageId="8" pageNumber="337" stateProvince="Toliara">Forest</location>
,
<date id="FFA3104E4B60FFB32B7C9624E16DF337" box="[568,780,1606,1629]" pageId="8" pageNumber="337" value="2002-02-03" valueMax="2002-02-08" valueMin="2002-02-03">
<collectingDate id="EFE7E9A64B60FFB32B7C9624E16DF337" box="[568,780,1606,1629]" pageId="8" pageNumber="337" value="2002-02-03" valueMax="2002-02-08" valueMin="2002-02-03">3–8 February 2002</collectingDate>
</date>
, Fisher-Griswold Arthropod Survey Team, collection code BLF5320 (
<emphasis id="B969EA9C4B60FFB32B429604E009F317" box="[518,616,1638,1661]" italics="true" pageId="8" pageNumber="337">
<typeStatus id="54A6882C4B60FFB32B429604E009F317" box="[518,616,1638,1661]" pageId="8" pageNumber="337" type="holotype">Holotype</typeStatus>
</emphasis>
Female
<specimenCode id="DBBB9EF54B60FFB32B929604E1C1F317" box="[726,928,1638,1661]" collectionCode="CASENT" pageId="8" pageNumber="337">CASENT9021513</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B753CD34B60FFB32AFF9604E32DF3F7" collectingDate="2002-02-03" collectingDateMax="2002-02-08" collectingDateMin="2002-02-03" collectionCode="CASENT" country="Madagascar" county="Reserve Prive Berenty" elevation="35" latitude="-24.956945" location="Mandrare River" longLatPrecision="21" longitude="46.27139" municipality="Foret de Bealoka" pageId="8" pageNumber="337" specimenCode="CASENT9012420" specimenCount="1" stateProvince="Toliara Prov." typeStatus="holotype">
same data (Female
<specimenCode id="DBBB9EF54B60FFB3293F96E4E324F3F7" box="[123,325,1670,1693]" collectionCode="CASENT" pageId="8" pageNumber="337">CASENT9012420</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B753CD34B60FFB2281A96E4E32DF479" collectingDate="2001-11-06" collectingDateMax="2001-11-10" collectingDateMin="2001-11-06" collectionCode="CASENT" country="Madagascar" county="Parc National Tsingy de Bemaraha" elevation="50" lastPageId="9" lastPageNumber="338" latitude="-19.141945" location="Tombeau Vazimba" longLatPrecision="21" longitude="44.828056" municipality="Bekopaka" pageId="8" pageNumber="337" specimenCode="CASENT9008848" specimenCount="1" stateProvince="Toliara Prov." typeStatus="paratype">
<collectingCounty id="62C34E024B60FFB3281A96E4E090F3F7" box="[350,753,1670,1693]" pageId="8" pageNumber="337">Parc National Tsingy de Bemaraha</collectingCounty>
,
<quantity id="4CE59B6B4B60FFB32A4796E4E135F3F7" box="[771,852,1670,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="337" unit="km" value="3.4">3.4 km</quantity>
<geoCoordinate id="EE2950494B60FFB32A2696E4E1C4F3F7" box="[866,933,1670,1693]" degrees="93" direction="east" orientation="longitude" pageId="8" pageNumber="337" precision="55555" value="93.0">93° E</geoCoordinate>
<collectingMunicipality id="6BC6ACF44B60FFB32AF096E4E640F3F7" box="[948,1057,1670,1693]" pageId="8" pageNumber="337">Bekopaka</collectingMunicipality>
,
<location id="8EC260554B60FFB32D7796E4E2BAF3D7" LSID="urn:lsid:plazi:treatment:03B487984B60FFB2293F90B9E331F479:8EC260554B60FFB32D7796E4E2BAF3D7" country="Madagascar" county="Parc National Tsingy de Bemaraha" latitude="-19.141945" longLatPrecision="21" longitude="44.828056" municipality="Bekopaka" name="Tombeau Vazimba" pageId="8" pageNumber="337" stateProvince="Toliara Prov.">Tombeau Vazimba</location>
,
<geoCoordinate id="EE2950494B60FFB329AE96C4E336F3D7" box="[234,343,1702,1725]" degrees="19" direction="south" minutes="8" orientation="latitude" pageId="8" pageNumber="337" precision="15" seconds="31" value="-19.141945">19°8ʹ31ʺS</geoCoordinate>
,
<geoCoordinate id="EE2950494B60FFB3282396C4E385F3D7" box="[359,484,1702,1725]" degrees="44" direction="east" minutes="49" orientation="longitude" pageId="8" pageNumber="337" precision="15" seconds="41" value="44.828056">44°49ʹ41ʺE</geoCoordinate>
, elev.
<quantity id="4CE59B6B4B60FFB32B7496C4E000F3D7" box="[560,609,1702,1725]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="337" unit="m" value="50.0">
<elevation id="0030D1BD4B60FFB32B7496C4E000F3D7" box="[560,609,1702,1725]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="337" unit="m" value="50.0">50m</elevation>
</quantity>
,
<collectingMethod id="525C4E994B60FFB32B3596C4E085F3D7" box="[625,740,1702,1725]" pageId="8" pageNumber="337">pitfall trap</collectingMethod>
in tropical dry forest,
<date id="FFA3104E4B60FFB22AAD96C4E2D5F599" lastPageId="9" lastPageNumber="338" pageId="8" pageNumber="337" value="2001-11-06" valueMax="2001-11-10" valueMin="2001-11-06">
<collectingDate id="EFE7E9A64B60FFB22AAD96C4E2D5F599" lastPageId="9" lastPageNumber="338" pageId="8" pageNumber="337" value="2001-11-06" valueMax="2001-11-10" valueMin="2001-11-06">6-10 November 2001</collectingDate>
</date>
, Fisher-Griswold Arthropod Survey Team, collection code 4230 (
<emphasis id="B969EA9C4B61FFB22A9490BEE655F599" box="[976,1076,220,243]" italics="true" pageId="9" pageNumber="338">
<typeStatus id="54A6882C4B61FFB22A9490BEE655F599" box="[976,1076,220,243]" pageId="9" pageNumber="338" type="paratype">Paratype</typeStatus>
</emphasis>
female,
<specimenCode id="DBBB9EF54B61FFB2293F909EE324F479" box="[123,325,252,275]" collectionCode="CASENT" pageId="9" pageNumber="338">CASENT9008848</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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