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<document id="672690EB563B411F986546AC21D49CAA" ID-CLB-Dataset="168932" ID-DOI="10.5852/ejt.2022.844.1949" ID-GBIF-Dataset="3a08058c-14f8-4232-ad02-65ed5b28380f" ID-ISSN="2118-9773" ID-Zenodo-Dep="7222155" ID-ZooBank="193EDD91-B24D-455C-B8AA-8133586A00A1" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666126129383" checkinUser="felipe" docAuthor="Grzelak, Katarzyna &amp; Sørensen, Martin V." docDate="2022" docId="03B487885B3D2A08FDDF3FAF6638FC81" docLanguage="en" docName="ejt-844_1-108.pdf" docOrigin="European Journal of Taxonomy 844" docSource="http://dx.doi.org/10.5852/ejt.2022.844.1949" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Echinoderes landersi Grzelak &amp; Sørensen 2022, sp. nov." docType="treatment" docUuid="3F3C572F-E5CE-4645-AA8F-9472E698B2F4" docUuidSource="ZooBank" docVersion="12" lastPageNumber="25" masterDocId="FF8DFFF05B2C2A10FFB63A55654FFFB8" masterDocTitle="Echinoderes (Kinorhyncha: Cyclorhagida) from the Hikurangi Margin, New Zealand" masterLastPageNumber="108" masterPageNumber="1" pageNumber="18" updateTime="1699407836451" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2E3CF541E469B9CD42B37CAECDA2FAAB">Echinoderes (Kinorhyncha: Cyclorhagida) from the Hikurangi Margin, New Zealand</mods:title>
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<mods:namePart id="4506C5CE42E12D75927C7F5156598D65">Grzelak, Katarzyna</mods:namePart>
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<mods:affiliation id="80CCB52FD4D2584A784B757CCCBC1313">Polish Academy of Sciences, Institute of Oceanology, Sopot, Poland. &amp; Natural History Museum of Denmark, University of Copenhagen, DK- 2100 Copenhagen, Denmark.</mods:affiliation>
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<mods:namePart id="616B442F3C6B07E4C376018585603779">Sørensen, Martin V.</mods:namePart>
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<mods:affiliation id="7920A78F52C78B39C0CA47FED7074243">Natural History Museum of Denmark, University of Copenhagen, DK- 2100 Copenhagen, Denmark.</mods:affiliation>
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<treatment id="03B487885B3D2A08FDDF3FAF6638FC81" ID-DOI="http://doi.org/10.5281/zenodo.7225516" ID-GBIF-Taxon="203115730" ID-Zenodo-Dep="7225516" LSID="urn:lsid:zoobank.org:act:3F3C572F-E5CE-4645-AA8F-9472E698B2F4" httpUri="http://treatment.plazi.org/id/03B487885B3D2A08FDDF3FAF6638FC81" lastPageId="24" lastPageNumber="25" pageId="17" pageNumber="18">
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<heading id="D0EA81F25B3D2A01FDDF3FAF6684F9AE" box="[617,971,1530,1558]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
<taxonomicName id="4C1D4D1D5B3D2A01FDDF3FAF6626F9AD" ID-CoL="02504ba7-0733-4be4-a1a8-3e337087f730" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[617,873,1530,1557]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="18" phylum="Kinorhyncha" rank="species" species="landersi" status="sp. nov.">
<emphasis id="B969EA8C5B3D2A01FDDF3FAF6626F9AD" bold="true" box="[617,873,1530,1557]" italics="true" pageId="17" pageNumber="18">Echinoderes landersi</emphasis>
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<taxonomicNameLabel id="A25A57F75B3D2A01FCC73FA96684F9AE" box="[881,971,1532,1558]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BA2369E5B3D2A01FD163C1D66DBF9DB" blockId="17.[371,1217,1530,1635]" box="[672,916,1608,1635]" pageId="17" pageNumber="18">
<figureCitation id="13262A1B5B3D2A01FD163C1D6647F9DB" box="[672,776,1608,1635]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="19.[189,232,1963,1989]" captionStartId-1="20.[189,232,1891,1917]" captionStartId-2="22.[189,232,1822,1848]" captionTargetBox-0="[189,1399,265,1950]" captionTargetBox-1="[189,1399,1200,1848]" captionTargetBox-2="[199,1389,267,1786]" captionTargetId-0="figure-25@19.[189,1399,265,1950]" captionTargetId-1="figure-57@20.[189,1399,1199,1849]" captionTargetId-2="figure-105@22.[189,1399,265,1796]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="22" captionText-0="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." captionText-1="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." captionText-2="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.7222167" figureDoi-1="http://doi.org/10.5281/zenodo.7222169" figureDoi-2="http://doi.org/10.5281/zenodo.7222171" httpUri-0="https://zenodo.org/record/7222167/files/figure.png" httpUri-1="https://zenodo.org/record/7222169/files/figure.png" httpUri-2="https://zenodo.org/record/7222171/files/figure.png" pageId="17" pageNumber="18">Figs 57</figureCitation>
;
<tableCitation id="C69F03255B3D2A01FCA23C1D66DBF9DB" box="[788,916,1608,1635]" captionStart-0="Table 4" captionStart-1="Table 5" captionStartId-0="20.[189,258,265,291]" captionStartId-1="21.[189,258,265,291]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="Table 4. Measurements from light microscopy of Echinoderes landersi sp. nov. (in µm) from the Hikurangi Margin." captionText-1="Table 5. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes landersi sp. nov." pageId="17" pageNumber="18">Tables 45</tableCitation>
</paragraph>
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<heading id="D0EA81F25B3D2A01FF0B3CD96479F91E" bold="true" box="[189,310,1676,1702]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B969EA8C5B3D2A01FF0B3CD96479F91E" bold="true" box="[189,310,1676,1702]" pageId="17" pageNumber="18">Diagnosis</emphasis>
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<taxonomicName id="4C1D4D1D5B3D2A01FF0B3CEB6400F960" authorityName="Claparede" authorityYear="1863" box="[189,335,1726,1752]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="18" phylum="Kinorhyncha" rank="genus">
<emphasis id="B969EA8C5B3D2A01FF0B3CEB6400F960" box="[189,335,1726,1752]" italics="true" pageId="17" pageNumber="18">Echinoderes</emphasis>
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with spines in middorsal position on segments 4 and 6, and in lateroventral positions on segments 6 to 9. Tubes present in subdorsal, laterodorsal, sublateral and ventrolateral positions on segment 2, lateroventral positions on segment 5, subdorsal, midlateral and lateral accessory positions on segment 8, and midlateral positions on segments 9 and 10.
</paragraph>
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<heading id="D0EA81F25B3D2A01FF0B3D23640BF828" bold="true" box="[189,324,1910,1936]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B969EA8C5B3D2A01FF0B3D23640BF828" bold="true" box="[189,324,1910,1936]" pageId="17" pageNumber="18">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2369E5B3D2A01FF0B3DFD6481F850" blockId="17.[189,1399,1960,2025]" pageId="17" pageNumber="18">The species is named after Dr Stephen C. Landers in recognition of his contributions to kinorhynch taxonomy and ecology.</paragraph>
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<emphasis id="B969EA8C5B3E2A02FF0B3B5D64E5FE9A" bold="true" box="[189,426,264,290]" pageId="18" pageNumber="19">Material examined</emphasis>
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TAN1004 Voyage;
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TAN1004 Voyage;
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<emphasis id="B969EA8C5B3E2A02FF5C382064AEFD37" bold="true" box="[234,481,629,655]" pageId="18" pageNumber="19">Additional material</emphasis>
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, stn TAN1004/44;
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; NIWA TAN1004 Voyage;
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; personal reference collection of MVS. Mounted for SEM
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; NIWA TAN1004 Voyage; soft sediment; personal reference collection of MVS. Mounted for SEM
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.
</paragraph>
</subSubSection>
<subSubSection id="C30765155B3E2A07FF0B391F6019FD59" lastPageId="23" lastPageNumber="24" pageId="18" pageNumber="19" type="description">
<paragraph id="8BA2369E5B3E2A02FF0B391F6401FCDC" blockId="18.[189,334,842,868]" box="[189,334,842,868]" pageId="18" pageNumber="19">
<heading id="D0EA81F25B3E2A02FF0B391F6401FCDC" bold="true" box="[189,334,842,868]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B969EA8C5B3E2A02FF0B391F6401FCDC" bold="true" box="[189,334,842,868]" pageId="18" pageNumber="19">Description</emphasis>
</heading>
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<paragraph id="8BA2369E5B3E2A02FF0B392E6185FBBA" blockId="18.[188,1399,890,1026]" pageId="18" pageNumber="19">
GENERAL. Adults with head, neck and eleven trunk segments (
<figureCitation id="13262A1B5B3E2A02FC1E392F6159FC2D" box="[936,1046,890,917]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="19.[189,232,1963,1989]" captionStartId-1="20.[189,232,1891,1917]" captionStartId-2="22.[189,232,1822,1848]" captionTargetBox-0="[189,1399,265,1950]" captionTargetBox-1="[189,1399,1200,1848]" captionTargetBox-2="[199,1389,267,1786]" captionTargetId-0="figure-25@19.[189,1399,265,1950]" captionTargetId-1="figure-57@20.[189,1399,1199,1849]" captionTargetId-2="figure-105@22.[189,1399,265,1796]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="22" captionText-0="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." captionText-1="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." captionText-2="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.7222167" figureDoi-1="http://doi.org/10.5281/zenodo.7222169" figureDoi-2="http://doi.org/10.5281/zenodo.7222171" httpUri-0="https://zenodo.org/record/7222167/files/figure.png" httpUri-1="https://zenodo.org/record/7222169/files/figure.png" httpUri-2="https://zenodo.org/record/7222171/files/figure.png" pageId="18" pageNumber="19">Figs 57</figureCitation>
). Overview of measurements and dimensions in
<tableCitation id="C69F03255B3E2A02FE2E39CA64BEFC01" box="[408,497,927,953]" captionStart="Table 4" captionStartId="20.[189,258,265,291]" captionTargetPageId="20" captionText="Table 4. Measurements from light microscopy of Echinoderes landersi sp. nov. (in µm) from the Hikurangi Margin." pageId="18" pageNumber="19">Table 4</tableCitation>
. Distribution of cuticular structures, i.e., sensory spots, glandular cell outlets, spines and tubes, summarized in
<tableCitation id="C69F03255B3E2A02FDE5399667FFFC66" box="[595,688,963,990]" captionStart="Table 5" captionStartId="21.[189,258,265,291]" captionTargetPageId="21" captionText="Table 5. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes landersi sp. nov." pageId="18" pageNumber="19">Table 5</tableCitation>
. No details regarding scalid arrangement and morphology could be provided, because introverts of all specimens mounted for SEM fully retracted.
</paragraph>
<paragraph id="8BA2369E5B3E2A02FF0B3E7E647FFB36" blockId="18.[189,1401,1067,1166]" pageId="18" pageNumber="19">
NECK. With 16 placids. Midventral placid broadest, 11 µm in width and 16 µm in length, whereas all others narrower, measuring 7 µm in width at their bases (
<figureCitation id="13262A1B5B3E2A02FCC13E1A668EFBD2" box="[887,961,1103,1130]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="18" pageNumber="19">Fig. 5</figureCitation>
). Trichoscalid plates well developed (
<figureCitation id="13262A1B5B3E2A02FF703E21646DFB36" box="[198,290,1140,1166]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="18" pageNumber="19">Fig. 6B</figureCitation>
).
</paragraph>
<paragraph id="8BA2369E5B3E2A02FF0B3EE2606EFA3F" blockId="18.[187,1401,1207,1415]" pageId="18" pageNumber="19">
SEGMENT 1. Consists of complete cuticular ring. Sensory spots located on anterior half of segment, in subdorsal, laterodorsal and ventromedial positions; sensory spots on this and following segment with micropapillae surrounding central pore and long marginal hair (
<figureCitation id="13262A1B5B3E2A02FC033F556175FAA3" box="[949,1082,1280,1307]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="18" pageNumber="19">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B3E2A02FBFE3F556121FAA2" box="[1096,1134,1280,1306]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="18" pageNumber="19">7C</figureCitation>
). Glandular cell outlet
<typeStatus id="54A6883C5B3E2A02FF093F7065BDFA87" box="[191,242,1317,1343]" pageId="18" pageNumber="19">type</typeStatus>
1 present in middorsal and ventrolateral positions. Cuticular hairs relatively long, distributed evenly around segment. Posterior segment margin almost straight, forming pectinate fringe with very short, sawtooth-like fringe tips along dorsal margin and with slightly longer tips along ventral margin.
</paragraph>
<paragraph id="8BA2369E5B3E2A02FF0B3FE5607EF957" blockId="18.[187,1400,1456,1775]" pageId="18" pageNumber="19">
SEGMENT 2. Consists of complete cuticular ring, with tubes located in subdorsal, laterodorsal, sublateral and ventrolateral positions (
<figureCitation id="13262A1B5B3E2A02FDBF3F8067C1FA48" box="[521,654,1493,1520]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="18" pageNumber="19">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B3E2A02FD2A3F8067AAFA57" box="[668,741,1493,1519]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="18" pageNumber="19">6AB</figureCitation>
,
<figureCitation id="13262A1B5B3E2A02FD453F806671FA57" box="[755,830,1493,1519]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="18" pageNumber="19">7CD</figureCitation>
); in
<specimenCount id="9D1BFD175B3E2A02FCC53F806159FA57" box="[883,1046,1493,1519]" pageId="18" pageNumber="19" type="generic">one specimen</specimenCount>
right subdorsal tube missing. Sensory spots present in middorsal, laterodorsal and ventromedial positions. Glandular cell outlets
<typeStatus id="54A6883C5B3E2A02FA843FAF602AF9AC" box="[1330,1381,1530,1556]" pageId="18" pageNumber="19">type</typeStatus>
1 not observed. This structure better visible in LM than in SEM but none of LM specimens orientated in way that allowed detailed examination of segments, especially its ventral side; therefore, for this and following nine segments presence of glandular cell outlets
<typeStatus id="54A6883C5B3E2A02FC043C3D66AAF93A" box="[946,997,1640,1666]" pageId="18" pageNumber="19">type</typeStatus>
<quantity id="4CE59B7B5B3E2A02FC583C326153F939" box="[1006,1052,1639,1665]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="18" pageNumber="19" unit="in" value="1.0">1 in</quantity>
ventral positions can neither be confirmed nor rejected. Pachycyclus of anterior segment margin of regular thickness. Secondary pectinate fringe present near anterior segment margin of this and following segments, but usually covered by preceding segment. Cuticular hairs and pectinate fringe tips as on preceding segment.
</paragraph>
<paragraph id="8BA2369E5B3E2A02FF0B3D4D6111F851" blockId="18.[187,1399,1815,2025]" pageId="18" pageNumber="19">
SEGMENT 3. Present segment, and eight remaining ones, consist of one tergal and two sternal plates (
<figureCitation id="13262A1B5B3E2A02FF733D696402F8EF" box="[197,333,1852,1879]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="18" pageNumber="19">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B3E2A02FEE83D6964C9F8EE" box="[350,390,1852,1878]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="18" pageNumber="19">7D</figureCitation>
). Segment with subdorsal and midlateral sensory spots. On this and following six segments, cuticular hairs arranged in three or four rows across tergal plate, except for hairless laterodorsal areas; paraventral areas devoid of hairs on this and following seven segments (
<figureCitation id="13262A1B5B3E2A02FB143DD061BFF818" box="[1186,1264,1925,1952]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="18" pageNumber="19">Fig. 5</figureCitation>
). Posterior segment margin straight, terminating in pectinate fringe with longer and more slender fringe tips along dorsal margin than on preceding segments, otherwise as on preceding segment.
</paragraph>
<caption id="DF6266165B3F2A03FF0B3DFE67E8F850" ID-DOI="http://doi.org/10.5281/zenodo.7222167" ID-Zenodo-Dep="7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="19" pageNumber="20" startId="19.[189,232,1963,1989]" targetBox="[189,1399,265,1950]" targetPageId="19">
<paragraph id="8BA2369E5B3F2A03FF0B3DFE67E8F850" blockId="19.[189,1399,1963,2025]" pageId="19" pageNumber="20">
<emphasis id="B969EA8C5B3F2A03FF0B3DFE6442F87D" bold="true" box="[189,269,1963,1989]" pageId="19" pageNumber="20">Fig. 5.</emphasis>
Line art illustrations of
<taxonomicName id="4C1D4D1D5B3F2A03FD8E3DFE667BF87D" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[568,820,1963,1989]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="19" pageNumber="20" phylum="Kinorhyncha" rank="species" species="landersi" status="sp. nov.">
<emphasis id="B969EA8C5B3F2A03FD8E3DFE667BF87D" box="[568,820,1963,1989]" italics="true" pageId="19" pageNumber="20">Echinoderes landersi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B3F2A03FC893DF966D3F87E" box="[831,924,1964,1990]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B969EA8C5B3F2A03FC103DFE66F4F87D" bold="true" box="[934,955,1963,1989]" pageId="19" pageNumber="20">A</emphasis>
. ♀, dorsal view.
<emphasis id="B969EA8C5B3F2A03FB393DFE61ECF87D" bold="true" box="[1167,1187,1963,1989]" pageId="19" pageNumber="20">B</emphasis>
. ♀, ventral view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF6266165B382A04FF0B3B5C64D6FEFE" pageId="20" pageNumber="21" startId="20.[189,258,265,291]" targetBox="[388,1252,363,1180]" targetIsTable="true" targetPageId="20">
<paragraph id="8BA2369E5B382A04FF0B3B5C64D6FEFE" blockId="20.[189,1399,265,326]" pageId="20" pageNumber="21">
<emphasis id="B969EA8C5B382A04FF0B3B5C646BFE9B" bold="true" box="[189,292,265,291]" pageId="20" pageNumber="21">Table 4.</emphasis>
Measurements from light microscopy of
<taxonomicName id="4C1D4D1D5B382A04FC9C3B5C6167FE9B" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[810,1064,265,291]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="20" pageNumber="21" phylum="Kinorhyncha" rank="species" species="landersi" status="sp. nov.">
<emphasis id="B969EA8C5B382A04FC9C3B5C6167FE9B" box="[810,1064,265,291]" italics="true" pageId="20" pageNumber="21">Echinoderes landersi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B382A04FB833B5C61DAFE9B" box="[1077,1173,265,291]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
(in µm) from the Hikurangi Margin.
</paragraph>
</caption>
<paragraph id="8BA2369E5B382A04FE323B296134FB24" pageId="20" pageNumber="21">
<table id="F91DC43E5B38D5EFFE323B3E61ABFB24" box="[388,1252,363,1180]" gridcols="3" gridrows="22" pageId="20" pageNumber="21">
<tr id="352D34DC5B38D5EFFE323B3E61ABFE10" box="[388,1252,363,424]" gridrow="0" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323B3E674AFE10" box="[388,517,363,424]" gridcol="0" gridrow="0" pageId="20" pageNumber="21">
<emphasis id="B969EA8C5B382A04FE323B29674AFE2E" bold="true" box="[388,517,380,406]" pageId="20" pageNumber="21">Character</emphasis>
</th>
<th id="76FC5DA05B38D5EFFD113B3E66C6FE10" box="[679,905,363,424]" gridcol="1" gridrow="0" pageId="20" pageNumber="21">
<emphasis id="B969EA8C5B382A04FD693B3E6616FE1F" bold="true" pageId="20" pageNumber="21">Holotype NIWA-159402</emphasis>
(♀)
</th>
<th id="76FC5DA05B38D5EFFC433B3E61ABFE10" box="[1013,1252,363,424]" gridcol="2" gridrow="0" pageId="20" pageNumber="21">
<emphasis id="B969EA8C5B382A04FB823B3E61FBFE1F" bold="true" pageId="20" pageNumber="21">Paratype NHMD-916627</emphasis>
(♀)
</th>
</tr>
<tr id="352D34DC5B38D5EFFE323BE461ABFE74" box="[388,1252,433,460]" gridrow="1" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323BE4674AFE74" box="[388,517,433,460]" gridcol="0" gridrow="1" pageId="20" pageNumber="21">TL</th>
<td id="76FC5DA05B38D5EFFD113BE466C6FE74" box="[679,905,433,460]" gridcol="1" gridrow="1" pageId="20" pageNumber="21">333</td>
<td id="76FC5DA05B38D5EFFC433BE461ABFE74" box="[1013,1252,433,460]" gridcol="2" gridrow="1" pageId="20" pageNumber="21">351</td>
</tr>
<tr id="352D34DC5B38D5EFFE323B8061ABFE57" box="[388,1252,469,495]" gridrow="2" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323B80674AFE57" box="[388,517,469,495]" gridcol="0" gridrow="2" pageId="20" pageNumber="21">S1</th>
<td id="76FC5DA05B38D5EFFD113B8066C6FE57" box="[679,905,469,495]" gridcol="1" gridrow="2" pageId="20" pageNumber="21">27</td>
<td id="76FC5DA05B38D5EFFC433B8061ABFE57" box="[1013,1252,469,495]" gridcol="2" gridrow="2" pageId="20" pageNumber="21">24</td>
</tr>
<tr id="352D34DC5B38D5EFFE323BAD61ABFDAA" box="[388,1252,504,530]" gridrow="3" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323BAD674AFDAA" box="[388,517,504,530]" gridcol="0" gridrow="3" pageId="20" pageNumber="21">S2</th>
<td id="76FC5DA05B38D5EFFD113BAD66C6FDAA" box="[679,905,504,530]" gridcol="1" gridrow="3" pageId="20" pageNumber="21">34</td>
<td id="76FC5DA05B38D5EFFC433BAD61ABFDAA" box="[1013,1252,504,530]" gridcol="2" gridrow="3" pageId="20" pageNumber="21">21</td>
</tr>
<tr id="352D34DC5B38D5EFFE32384961ABFD8E" box="[388,1252,540,566]" gridrow="4" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323849674AFD8E" box="[388,517,540,566]" gridcol="0" gridrow="4" pageId="20" pageNumber="21">S3</th>
<td id="76FC5DA05B38D5EFFD11384966C6FD8E" box="[679,905,540,566]" gridcol="1" gridrow="4" pageId="20" pageNumber="21">40</td>
<td id="76FC5DA05B38D5EFFC43384961ABFD8E" box="[1013,1252,540,566]" gridcol="2" gridrow="4" pageId="20" pageNumber="21">37</td>
</tr>
<tr id="352D34DC5B38D5EFFE32386A61ABFDE1" box="[388,1252,575,601]" gridrow="5" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE32386A674AFDE1" box="[388,517,575,601]" gridcol="0" gridrow="5" pageId="20" pageNumber="21">S4</th>
<td id="76FC5DA05B38D5EFFD11386A66C6FDE1" box="[679,905,575,601]" gridcol="1" gridrow="5" pageId="20" pageNumber="21">42</td>
<td id="76FC5DA05B38D5EFFC43386A61ABFDE1" box="[1013,1252,575,601]" gridcol="2" gridrow="5" pageId="20" pageNumber="21">38</td>
</tr>
<tr id="352D34DC5B38D5EFFE32383761ABFDC5" box="[388,1252,610,637]" gridrow="6" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323837674AFDC5" box="[388,517,610,637]" gridcol="0" gridrow="6" pageId="20" pageNumber="21">S5</th>
<td id="76FC5DA05B38D5EFFD11383766C6FDC5" box="[679,905,610,637]" gridcol="1" gridrow="6" pageId="20" pageNumber="21">47</td>
<td id="76FC5DA05B38D5EFFC43383761ABFDC5" box="[1013,1252,610,637]" gridcol="2" gridrow="6" pageId="20" pageNumber="21">42</td>
</tr>
<tr id="352D34DC5B38D5EFFE3238D361ABFD19" box="[388,1252,646,673]" gridrow="7" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE3238D3674AFD19" box="[388,517,646,673]" gridcol="0" gridrow="7" pageId="20" pageNumber="21">S6</th>
<td id="76FC5DA05B38D5EFFD1138D366C6FD19" box="[679,905,646,673]" gridcol="1" gridrow="7" pageId="20" pageNumber="21">52</td>
<td id="76FC5DA05B38D5EFFC4338D361ABFD19" box="[1013,1252,646,673]" gridcol="2" gridrow="7" pageId="20" pageNumber="21">49</td>
</tr>
<tr id="352D34DC5B38D5EFFE3238FC61ABFD7C" box="[388,1252,681,708]" gridrow="8" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE3238FC674AFD7C" box="[388,517,681,708]" gridcol="0" gridrow="8" pageId="20" pageNumber="21">S7</th>
<td id="76FC5DA05B38D5EFFD1138FC66C6FD7C" box="[679,905,681,708]" gridcol="1" gridrow="8" pageId="20" pageNumber="21">54</td>
<td id="76FC5DA05B38D5EFFC4338FC61ABFD7C" box="[1013,1252,681,708]" gridcol="2" gridrow="8" pageId="20" pageNumber="21">50</td>
</tr>
<tr id="352D34DC5B38D5EFFE32389861ABFD5F" box="[388,1252,717,743]" gridrow="9" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323898674AFD5F" box="[388,517,717,743]" gridcol="0" gridrow="9" pageId="20" pageNumber="21">S8</th>
<td id="76FC5DA05B38D5EFFD11389866C6FD5F" box="[679,905,717,743]" gridcol="1" gridrow="9" pageId="20" pageNumber="21">49</td>
<td id="76FC5DA05B38D5EFFC43389861ABFD5F" box="[1013,1252,717,743]" gridcol="2" gridrow="9" pageId="20" pageNumber="21">44</td>
</tr>
<tr id="352D34DC5B38D5EFFE3238A561ABFCB2" box="[388,1252,752,778]" gridrow="10" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE3238A5674AFCB2" box="[388,517,752,778]" gridcol="0" gridrow="10" pageId="20" pageNumber="21">S9</th>
<td id="76FC5DA05B38D5EFFD1138A566C6FCB2" box="[679,905,752,778]" gridcol="1" gridrow="10" pageId="20" pageNumber="21">44</td>
<td id="76FC5DA05B38D5EFFC4338A561ABFCB2" box="[1013,1252,752,778]" gridcol="2" gridrow="10" pageId="20" pageNumber="21">40</td>
</tr>
<tr id="352D34DC5B38D5EFFE32394161ABFC96" box="[388,1252,788,814]" gridrow="11" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323941674AFC96" box="[388,517,788,814]" gridcol="0" gridrow="11" pageId="20" pageNumber="21">S10</th>
<td id="76FC5DA05B38D5EFFD11394166C6FC96" box="[679,905,788,814]" gridcol="1" gridrow="11" pageId="20" pageNumber="21">43</td>
<td id="76FC5DA05B38D5EFFC43394161ABFC96" box="[1013,1252,788,814]" gridcol="2" gridrow="11" pageId="20" pageNumber="21">40</td>
</tr>
<tr id="352D34DC5B38D5EFFE32396861ABFCEF" box="[388,1252,829,855]" gridrow="12" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323968674AFCEF" box="[388,517,829,855]" gridcol="0" gridrow="12" pageId="20" pageNumber="21">S11</th>
<td id="76FC5DA05B38D5EFFD11396866C6FCEF" box="[679,905,829,855]" gridcol="1" gridrow="12" pageId="20" pageNumber="21">42</td>
<td id="76FC5DA05B38D5EFFC43396861ABFCEF" box="[1013,1252,829,855]" gridcol="2" gridrow="12" pageId="20" pageNumber="21">39</td>
</tr>
<tr id="352D34DC5B38D5EFFE32393261ABFC39" box="[388,1252,871,897]" gridrow="13" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323932674AFC39" box="[388,517,871,897]" gridcol="0" gridrow="13" pageId="20" pageNumber="21">MD4 (ac)</th>
<td id="76FC5DA05B38D5EFFD11393266C6FC39" box="[679,905,871,897]" gridcol="1" gridrow="13" pageId="20" pageNumber="21">31</td>
<td id="76FC5DA05B38D5EFFC43393261ABFC39" box="[1013,1252,871,897]" gridcol="2" gridrow="13" pageId="20" pageNumber="21">24</td>
</tr>
<tr id="352D34DC5B38D5EFFE3239DF61ABFC1C" box="[388,1252,906,932]" gridrow="14" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE3239DF674AFC1C" box="[388,517,906,932]" gridcol="0" gridrow="14" pageId="20" pageNumber="21">MD6 (ac)</th>
<td id="76FC5DA05B38D5EFFD1139DF66C6FC1C" box="[679,905,906,932]" gridcol="1" gridrow="14" pageId="20" pageNumber="21">33</td>
<td id="76FC5DA05B38D5EFFC4339DF61ABFC1C" box="[1013,1252,906,932]" gridcol="2" gridrow="14" pageId="20" pageNumber="21">30</td>
</tr>
<tr id="352D34DC5B38D5EFFE3239F861ABFC70" box="[388,1252,941,968]" gridrow="15" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE3239F8674AFC70" box="[388,517,941,968]" gridcol="0" gridrow="15" pageId="20" pageNumber="21">LV6 (ac)</th>
<td id="76FC5DA05B38D5EFFD1139F866C6FC70" box="[679,905,941,968]" gridcol="1" gridrow="15" pageId="20" pageNumber="21">28</td>
<td id="76FC5DA05B38D5EFFC4339F861ABFC70" box="[1013,1252,941,968]" gridcol="2" gridrow="15" pageId="20" pageNumber="21">24</td>
</tr>
<tr id="352D34DC5B38D5EFFE32398461ABFC54" box="[388,1252,977,1004]" gridrow="16" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323984674AFC54" box="[388,517,977,1004]" gridcol="0" gridrow="16" pageId="20" pageNumber="21">LV7 (ac)</th>
<td id="76FC5DA05B38D5EFFD11398466C6FC54" box="[679,905,977,1004]" gridcol="1" gridrow="16" pageId="20" pageNumber="21">29</td>
<td id="76FC5DA05B38D5EFFC43398461ABFC54" box="[1013,1252,977,1004]" gridcol="2" gridrow="16" pageId="20" pageNumber="21">25</td>
</tr>
<tr id="352D34DC5B38D5EFFE3239A161ABFBB6" box="[388,1252,1012,1038]" gridrow="17" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE3239A1674AFBB6" box="[388,517,1012,1038]" gridcol="0" gridrow="17" pageId="20" pageNumber="21">LV8 (ac)</th>
<td id="76FC5DA05B38D5EFFD1139A166C6FBB6" box="[679,905,1012,1038]" gridcol="1" gridrow="17" pageId="20" pageNumber="21">34</td>
<td id="76FC5DA05B38D5EFFC4339A161ABFBB6" box="[1013,1252,1012,1038]" gridcol="2" gridrow="17" pageId="20" pageNumber="21">29</td>
</tr>
<tr id="352D34DC5B38D5EFFE323E4D61ABFB8A" box="[388,1252,1048,1074]" gridrow="18" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323E4D674AFB8A" box="[388,517,1048,1074]" gridcol="0" gridrow="18" pageId="20" pageNumber="21">LV9 (ac)</th>
<td id="76FC5DA05B38D5EFFD113E4D66C6FB8A" box="[679,905,1048,1074]" gridcol="1" gridrow="18" pageId="20" pageNumber="21">38</td>
<td id="76FC5DA05B38D5EFFC433E4D61ABFB8A" box="[1013,1252,1048,1074]" gridcol="2" gridrow="18" pageId="20" pageNumber="21">33</td>
</tr>
<tr id="352D34DC5B38D5EFFE323E6E61ABFBED" box="[388,1252,1083,1109]" gridrow="19" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323E6E674AFBED" box="[388,517,1083,1109]" gridcol="0" gridrow="19" pageId="20" pageNumber="21">LTS</th>
<td id="76FC5DA05B38D5EFFD113E6E66C6FBED" box="[679,905,1083,1109]" gridcol="1" gridrow="19" pageId="20" pageNumber="21">198</td>
<td id="76FC5DA05B38D5EFFC433E6E61ABFBED" box="[1013,1252,1083,1109]" gridcol="2" gridrow="19" pageId="20" pageNumber="21">173</td>
</tr>
<tr id="352D34DC5B38D5EFFE323E0A61ABFBC2" box="[388,1252,1119,1146]" gridrow="20" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323E0A674AFBC2" box="[388,517,1119,1146]" gridcol="0" gridrow="20" pageId="20" pageNumber="21">LTS/TL</th>
<td id="76FC5DA05B38D5EFFD113E0A66C6FBC2" box="[679,905,1119,1146]" gridcol="1" gridrow="20" pageId="20" pageNumber="21">59.6%</td>
<td id="76FC5DA05B38D5EFFC433E0A61ABFBC2" box="[1013,1252,1119,1146]" gridcol="2" gridrow="20" pageId="20" pageNumber="21">49.3%</td>
</tr>
<tr id="352D34DC5B38D5EFFE323ED761ABFB24" box="[388,1252,1154,1180]" gridrow="21" pageId="20" pageNumber="21">
<th id="76FC5DA05B38D5EFFE323ED7674AFB24" box="[388,517,1154,1180]" gridcol="0" gridrow="21" pageId="20" pageNumber="21">LTAS</th>
<td id="76FC5DA05B38D5EFFD113ED766C6FB24" box="[679,905,1154,1180]" gridcol="1" gridrow="21" pageId="20" pageNumber="21">70</td>
<td id="76FC5DA05B38D5EFFC433ED761ABFB24" box="[1013,1252,1154,1180]" gridcol="2" gridrow="21" pageId="20" pageNumber="21">60</td>
</tr>
</table>
</paragraph>
<caption id="DF6266165B382A04FF0B3D366489F85F" ID-DOI="http://doi.org/10.5281/zenodo.7222169" ID-Zenodo-Dep="7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1891,1917]" targetBox="[189,1399,1200,1848]" targetPageId="20">
<paragraph id="8BA2369E5B382A04FF0B3D366489F85F" blockId="20.[189,1399,1891,2023]" pageId="20" pageNumber="21">
<emphasis id="B969EA8C5B382A04FF0B3D36645EF8C5" bold="true" box="[189,273,1891,1917]" pageId="20" pageNumber="21">Fig. 6.</emphasis>
Light micrographs showing overview and details of
<taxonomicName id="4C1D4D1D5B382A04FC043D3661FDF8C5" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[946,1202,1891,1917]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="20" pageNumber="21" phylum="Kinorhyncha" rank="species" species="landersi" status="sp. nov.">
<emphasis id="B969EA8C5B382A04FC043D3661FDF8C5" box="[946,1202,1891,1917]" italics="true" pageId="20" pageNumber="21">Echinoderes landersi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B382A04FB763D31606EF8C6" box="[1216,1313,1892,1918]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B969EA8C5B382A04FA993D36600BF8C5" bold="true" box="[1327,1348,1891,1917]" pageId="20" pageNumber="21">A</emphasis>
. ♀, paratype (NHMD-916627).
<emphasis id="B969EA8C5B382A04FDBE3DD36770F818" bold="true" box="[520,575,1926,1952]" pageId="20" pageNumber="21">BD</emphasis>
. ♀, holotype (NIWA-159402).
<emphasis id="B969EA8C5B382A04FC053DD36687F818" bold="true" box="[947,968,1926,1952]" pageId="20" pageNumber="21">A</emphasis>
. Lateral overview.
<emphasis id="B969EA8C5B382A04FB1B3DD3618FF818" bold="true" box="[1197,1216,1926,1952]" pageId="20" pageNumber="21">B</emphasis>
. Segments 1 to 3, lateral view.
<emphasis id="B969EA8C5B382A04FEC33DFC64C6F87B" bold="true" box="[373,393,1961,1987]" pageId="20" pageNumber="21">C</emphasis>
. Segments 7 to 11, lateral view.
<emphasis id="B969EA8C5B382A04FCA63DFF666BF87C" bold="true" box="[784,804,1962,1988]" pageId="20" pageNumber="21">D</emphasis>
. Segments 4 to 7, lateral view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF6266165B392A05FF0B3B5C665CFEFF" pageId="21" pageNumber="22" startId="21.[189,258,265,291]" targetBox="[198,1376,391,845]" targetIsTable="true" targetPageId="21">
<paragraph id="8BA2369E5B392A05FF0B3B5C665CFEFF" blockId="21.[189,1398,265,327]" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FF0B3B5C646CFE9B" bold="true" box="[189,291,265,291]" pageId="21" pageNumber="22">Table 5.</emphasis>
Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in
<taxonomicName id="4C1D4D1D5B392A05FE0E3B7967FEFEFE" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[440,689,300,326]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="21" pageNumber="22" phylum="Kinorhyncha" rank="species" species="landersi" status="sp. nov.">
<emphasis id="B969EA8C5B392A05FE0E3B7967FEFEFE" box="[440,689,300,326]" italics="true" pageId="21" pageNumber="22">Echinoderes landersi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B392A05FD0E3B78665CFEFF" box="[696,787,301,327]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BA2369E5B392A05FF703BD261DDFCF4" pageId="21" pageNumber="22">
<table id="F91DC43E5B39D5EFFF703BD2602FFCF5" box="[198,1376,391,845]" gridcols="11" gridrows="12" pageId="21" pageNumber="22">
<tr id="352D34DC5B39D5EFFF703BD2602FFE19" box="[198,1376,391,417]" gridrow="0" pageId="21" pageNumber="22">
<th id="76FC5DA05B39D5EFFF703BD264D6FE19" box="[198,409,391,417]" gridcol="0" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FF703BD264D6FE19" bold="true" box="[198,409,391,417]" pageId="21" pageNumber="22">Position segment</emphasis>
</th>
<th id="76FC5DA05B39D5EFFE043BD264A4FE19" box="[434,491,391,417]" gridcol="1" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FE003BD264A8FE19" bold="true" box="[438,487,391,417]" pageId="21" pageNumber="22">MD</emphasis>
</th>
<th id="76FC5DA05B39D5EFFDA63BD26727FE19" box="[528,616,391,417]" gridcol="2" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FD9F3BD2671FFE19" bold="true" box="[553,592,391,417]" pageId="21" pageNumber="22">PD</emphasis>
</th>
<th id="76FC5DA05B39D5EFFD3F3BD267AEFE19" box="[649,737,391,417]" gridcol="3" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FD143BD26788FE19" bold="true" box="[674,711,391,417]" pageId="21" pageNumber="22">SD</emphasis>
</th>
<th id="76FC5DA05B39D5EFFD413BD26663FE19" box="[759,812,391,417]" gridcol="4" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FCB73BD26665FE19" bold="true" box="[769,810,391,417]" pageId="21" pageNumber="22">LD</emphasis>
</th>
<th id="76FC5DA05B39D5EFFCE43BD266CEFE19" box="[850,897,391,417]" gridcol="5" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FCE43BD266CEFE19" bold="true" box="[850,897,391,417]" pageId="21" pageNumber="22">ML</emphasis>
</th>
<th id="76FC5DA05B39D5EFFC0F3BD26692FE19" box="[953,989,391,417]" gridcol="6" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FC0F3BD26692FE19" bold="true" box="[953,989,391,417]" pageId="21" pageNumber="22">SL</emphasis>
</th>
<th id="76FC5DA05B39D5EFFBB53BD2611AFE19" box="[1027,1109,391,417]" gridcol="7" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FBAA3BD2610AFE19" bold="true" box="[1052,1093,391,417]" pageId="21" pageNumber="22">LA</emphasis>
</th>
<th id="76FC5DA05B39D5EFFBC73BD261D8FE19" box="[1137,1175,391,417]" gridcol="8" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FBC73BD261D8FE19" bold="true" box="[1137,1175,391,417]" pageId="21" pageNumber="22">LV</emphasis>
</th>
<th id="76FC5DA05B39D5EFFB723BD261B2FE19" box="[1220,1277,391,417]" gridcol="9" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FB7B3BD261B9FE19" bold="true" box="[1229,1270,391,417]" pageId="21" pageNumber="22">VL</emphasis>
</th>
<th id="76FC5DA05B39D5EFFA993BD2602FFE19" box="[1327,1376,391,417]" gridcol="10" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B969EA8C5B392A05FA993BD2602FFE19" bold="true" box="[1327,1376,391,417]" pageId="21" pageNumber="22">VM</emphasis>
</th>
</tr>
<tr id="352D34DC5B39D5EFFF703BEC602FFE6B" box="[198,1376,441,467]" gridrow="1" pageId="21" pageNumber="22" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76FC5DA05B39D5EFFF703BEC64D6FE6B" box="[198,409,441,467]" gridcol="0" gridrow="1" pageId="21" pageNumber="22">1</th>
<td id="76FC5DA05B39D5EFFE043BEC64A4FE6B" box="[434,491,441,467]" gridcol="1" gridrow="1" pageId="21" pageNumber="22">gco1</td>
<td id="76FC5DA05B39D5EFFD3F3BEC67AEFE6B" box="[649,737,441,467]" gridcol="3" gridrow="1" pageId="21" pageNumber="22">ss</td>
<td id="76FC5DA05B39D5EFFD413BEC6663FE6B" box="[759,812,441,467]" gridcol="4" gridrow="1" pageId="21" pageNumber="22">ss</td>
<td id="76FC5DA05B39D5EFFB723BEC61B2FE6B" box="[1220,1277,441,467]" gridcol="9" gridrow="1" pageId="21" pageNumber="22">gco1</td>
<td id="76FC5DA05B39D5EFFA993BEC602FFE6B" box="[1327,1376,441,467]" gridcol="10" gridrow="1" pageId="21" pageNumber="22">ss</td>
</tr>
<tr id="352D34DC5B39D5EFFF703B8B602FFE41" box="[198,1376,478,505]" gridrow="2" pageId="21" pageNumber="22" rowspan-2="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
<th id="76FC5DA05B39D5EFFF703B8B64D6FE41" box="[198,409,478,505]" gridcol="0" gridrow="2" pageId="21" pageNumber="22">2</th>
<td id="76FC5DA05B39D5EFFE043B8B64A4FE41" box="[434,491,478,505]" gridcol="1" gridrow="2" pageId="21" pageNumber="22">ss</td>
<td id="76FC5DA05B39D5EFFD3F3B8B67AEFE41" box="[649,737,478,505]" gridcol="3" gridrow="2" pageId="21" pageNumber="22">tu</td>
<td id="76FC5DA05B39D5EFFD413B8B6663FE41" box="[759,812,478,505]" gridcol="4" gridrow="2" pageId="21" pageNumber="22">tu,ss</td>
<td id="76FC5DA05B39D5EFFC0F3B8B6692FE41" box="[953,989,478,505]" gridcol="6" gridrow="2" pageId="21" pageNumber="22">tu</td>
<td id="76FC5DA05B39D5EFFB723B8B61B2FE41" box="[1220,1277,478,505]" gridcol="9" gridrow="2" pageId="21" pageNumber="22">tu</td>
<td id="76FC5DA05B39D5EFFA993B8B602FFE41" box="[1327,1376,478,505]" gridcol="10" gridrow="2" pageId="21" pageNumber="22">ss</td>
</tr>
<tr id="352D34DC5B39D5EFFF703851602FFDA7" box="[198,1376,516,543]" gridrow="3" pageId="21" pageNumber="22" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76FC5DA05B39D5EFFF70385164D6FDA7" box="[198,409,516,543]" gridcol="0" gridrow="3" pageId="21" pageNumber="22">3</th>
<td id="76FC5DA05B39D5EFFD3F385167AEFDA7" box="[649,737,516,543]" gridcol="3" gridrow="3" pageId="21" pageNumber="22">ss</td>
<td id="76FC5DA05B39D5EFFCE4385166CEFDA7" box="[850,897,516,543]" gridcol="5" gridrow="3" pageId="21" pageNumber="22">ss</td>
</tr>
<tr id="352D34DC5B39D5EFFF70387F602FFDFE" box="[198,1376,554,582]" gridrow="4" pageId="21" pageNumber="22" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76FC5DA05B39D5EFFF70387F64D6FDFE" box="[198,409,554,582]" gridcol="0" gridrow="4" pageId="21" pageNumber="22">4</th>
<td id="76FC5DA05B39D5EFFE04387F64A4FDFE" box="[434,491,554,582]" gridcol="1" gridrow="4" pageId="21" pageNumber="22">ac</td>
<td id="76FC5DA05B39D5EFFDA6387F6727FDFE" box="[528,616,554,582]" gridcol="2" gridrow="4" pageId="21" pageNumber="22">gco1</td>
</tr>
<tr id="352D34DC5B39D5EFFF70381A602FFDD3" box="[198,1376,591,619]" gridrow="5" pageId="21" pageNumber="22" rowspan-1="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="76FC5DA05B39D5EFFF70381A64D6FDD3" box="[198,409,591,619]" gridcol="0" gridrow="5" pageId="21" pageNumber="22">5</th>
<td id="76FC5DA05B39D5EFFDA6381A6727FDD3" box="[528,616,591,619]" gridcol="2" gridrow="5" pageId="21" pageNumber="22">gco1</td>
<td id="76FC5DA05B39D5EFFD3F381A67AEFDD3" box="[649,737,591,619]" gridcol="3" gridrow="5" pageId="21" pageNumber="22">ss</td>
<td id="76FC5DA05B39D5EFFCE4381A66CEFDD3" box="[850,897,591,619]" gridcol="5" gridrow="5" pageId="21" pageNumber="22">ss</td>
<td id="76FC5DA05B39D5EFFBC7381A61D8FDD3" box="[1137,1175,591,619]" gridcol="8" gridrow="5" pageId="21" pageNumber="22">tu</td>
<td id="76FC5DA05B39D5EFFA99381A602FFDD3" box="[1327,1376,591,619]" gridcol="10" gridrow="5" pageId="21" pageNumber="22">ss</td>
</tr>
<tr id="352D34DC5B39D5EFFF703820602FFD28" box="[198,1376,629,656]" gridrow="6" pageId="21" pageNumber="22" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="76FC5DA05B39D5EFFF70382064D6FD28" box="[198,409,629,656]" gridcol="0" gridrow="6" pageId="21" pageNumber="22">6</th>
<td id="76FC5DA05B39D5EFFE04382064A4FD28" box="[434,491,629,656]" gridcol="1" gridrow="6" pageId="21" pageNumber="22">ac</td>
<td id="76FC5DA05B39D5EFFDA638206727FD28" box="[528,616,629,656]" gridcol="2" gridrow="6" pageId="21" pageNumber="22">gco1,ss</td>
<td id="76FC5DA05B39D5EFFD3F382067AEFD28" box="[649,737,629,656]" gridcol="3" gridrow="6" pageId="21" pageNumber="22">ss</td>
<td id="76FC5DA05B39D5EFFCE4382066CEFD28" box="[850,897,629,656]" gridcol="5" gridrow="6" pageId="21" pageNumber="22">ss</td>
<td id="76FC5DA05B39D5EFFBC7382061D8FD28" box="[1137,1175,629,656]" gridcol="8" gridrow="6" pageId="21" pageNumber="22">ac</td>
<td id="76FC5DA05B39D5EFFA993820602FFD28" box="[1327,1376,629,656]" gridcol="10" gridrow="6" pageId="21" pageNumber="22">ss</td>
</tr>
<tr id="352D34DC5B39D5EFFF7038CE602FFD0E" box="[198,1376,667,694]" gridrow="7" pageId="21" pageNumber="22" rowspan-1="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="76FC5DA05B39D5EFFF7038CE64D6FD0E" box="[198,409,667,694]" gridcol="0" gridrow="7" pageId="21" pageNumber="22">7</th>
<td id="76FC5DA05B39D5EFFDA638CE6727FD0E" box="[528,616,667,694]" gridcol="2" gridrow="7" pageId="21" pageNumber="22">gco1</td>
<td id="76FC5DA05B39D5EFFD3F38CE67AEFD0E" box="[649,737,667,694]" gridcol="3" gridrow="7" pageId="21" pageNumber="22">ss</td>
<td id="76FC5DA05B39D5EFFCE438CE66CEFD0E" box="[850,897,667,694]" gridcol="5" gridrow="7" pageId="21" pageNumber="22">ss</td>
<td id="76FC5DA05B39D5EFFBC738CE61D8FD0E" box="[1137,1175,667,694]" gridcol="8" gridrow="7" pageId="21" pageNumber="22">ac</td>
<td id="76FC5DA05B39D5EFFA9938CE602FFD0E" box="[1327,1376,667,694]" gridcol="10" gridrow="7" pageId="21" pageNumber="22">ss</td>
</tr>
<tr id="352D34DC5B39D5EFFF703894602FFD64" box="[198,1376,705,732]" gridrow="8" pageId="21" pageNumber="22" rowspan-1="1" rowspan-10="1" rowspan-4="1" rowspan-6="1" rowspan-9="1">
<th id="76FC5DA05B39D5EFFF70389464D6FD64" box="[198,409,705,732]" gridcol="0" gridrow="8" pageId="21" pageNumber="22">8</th>
<td id="76FC5DA05B39D5EFFDA638946727FD64" box="[528,616,705,732]" gridcol="2" gridrow="8" pageId="21" pageNumber="22">gco1</td>
<td id="76FC5DA05B39D5EFFD3F389467AEFD64" box="[649,737,705,732]" gridcol="3" gridrow="8" pageId="21" pageNumber="22">ss,tu</td>
<td id="76FC5DA05B39D5EFFCE4389466CEFD64" box="[850,897,705,732]" gridcol="5" gridrow="8" pageId="21" pageNumber="22">tu</td>
<td id="76FC5DA05B39D5EFFBB53894611AFD64" box="[1027,1109,705,732]" gridcol="7" gridrow="8" pageId="21" pageNumber="22">tu</td>
<td id="76FC5DA05B39D5EFFBC7389461D8FD64" box="[1137,1175,705,732]" gridcol="8" gridrow="8" pageId="21" pageNumber="22">ac</td>
</tr>
<tr id="352D34DC5B39D5EFFF7038B2602FFCB9" box="[198,1376,743,769]" gridrow="9" pageId="21" pageNumber="22" rowspan-1="1" rowspan-10="1" rowspan-4="1" rowspan-6="1">
<th id="76FC5DA05B39D5EFFF7038B264D6FCB9" box="[198,409,743,769]" gridcol="0" gridrow="9" pageId="21" pageNumber="22">9</th>
<td id="76FC5DA05B39D5EFFDA638B26727FCB9" box="[528,616,743,769]" gridcol="2" gridrow="9" pageId="21" pageNumber="22">gco1</td>
<td id="76FC5DA05B39D5EFFD3F38B267AEFCB9" box="[649,737,743,769]" gridcol="3" gridrow="9" pageId="21" pageNumber="22">ss,ss</td>
<td id="76FC5DA05B39D5EFFCE438B266CEFCB9" box="[850,897,743,769]" gridcol="5" gridrow="9" pageId="21" pageNumber="22">tu</td>
<td id="76FC5DA05B39D5EFFBB538B2611AFCB9" box="[1027,1109,743,769]" gridcol="7" gridrow="9" pageId="21" pageNumber="22">si</td>
<td id="76FC5DA05B39D5EFFBC738B261D8FCB9" box="[1137,1175,743,769]" gridcol="8" gridrow="9" pageId="21" pageNumber="22">ac</td>
<td id="76FC5DA05B39D5EFFB7238B261B2FCB9" box="[1220,1277,743,769]" gridcol="9" gridrow="9" pageId="21" pageNumber="22">ss</td>
</tr>
<tr id="352D34DC5B39D5EFFF703958602FFC9F" box="[198,1376,781,807]" gridrow="10" pageId="21" pageNumber="22" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76FC5DA05B39D5EFFF70395864D6FC9F" box="[198,409,781,807]" gridcol="0" gridrow="10" pageId="21" pageNumber="22">10</th>
<td id="76FC5DA05B39D5EFFE04395864A4FC9F" box="[434,491,781,807]" gridcol="1" gridrow="10" pageId="21" pageNumber="22">gco1</td>
<td id="76FC5DA05B39D5EFFD3F395867AEFC9F" box="[649,737,781,807]" gridcol="3" gridrow="10" pageId="21" pageNumber="22">ss</td>
<td id="76FC5DA05B39D5EFFCE4395866CEFC9F" box="[850,897,781,807]" gridcol="5" gridrow="10" pageId="21" pageNumber="22">tu</td>
<td id="76FC5DA05B39D5EFFB72395861B2FC9F" box="[1220,1277,781,807]" gridcol="9" gridrow="10" pageId="21" pageNumber="22">ss</td>
</tr>
<tr id="352D34DC5B39D5EFFF703967602FFCF5" box="[198,1376,818,845]" gridrow="11" pageId="21" pageNumber="22" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-9="1">
<th id="76FC5DA05B39D5EFFF70396764D6FCF5" box="[198,409,818,845]" gridcol="0" gridrow="11" pageId="21" pageNumber="22">11</th>
<td id="76FC5DA05B39D5EFFD3F396767AEFCF5" box="[649,737,818,845]" gridcol="3" gridrow="11" pageId="21" pageNumber="22">ss,gco1</td>
<td id="76FC5DA05B39D5EFFBB53967611AFCF5" box="[1027,1109,818,845]" gridcol="7" gridrow="11" pageId="21" pageNumber="22">ltas(♀)</td>
<td id="76FC5DA05B39D5EFFBC7396761D8FCF5" box="[1137,1175,818,845]" gridcol="8" gridrow="11" pageId="21" pageNumber="22">lts</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA2369E5B392A05FF0B39D264EDFBB6" blockId="21.[189,1399,902,1038]" pageId="21" pageNumber="22">
SEGMENT 4. With spine in middorsal position. Spine relatively short (27 µm), only slightly exceeding beyond posterior segment margin (
<figureCitation id="13262A1B5B392A05FDD239FE6786FC7E" box="[612,713,939,966]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="21" pageNumber="22">Figs 5A</figureCitation>
,
<figureCitation id="13262A1B5B392A05FD6F39FE6668FC7D" box="[729,807,939,965]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="21" pageNumber="22">6A, D</figureCitation>
,
<figureCitation id="13262A1B5B392A05FC8039FE66CDFC7D" box="[822,898,939,965]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="21" pageNumber="22">7A, E</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B392A05FB1F39FE6193FC7D" box="[1193,1244,939,965]" pageId="21" pageNumber="22">type</typeStatus>
1 present in paradorsal positions. No other traits observed. Cuticular hairs and secondary pectinate fringe as on preceding segment.
</paragraph>
<paragraph id="8BA2369E5B392A05FF0B3E6366C9FB06" blockId="21.[189,1399,1078,1214]" pageId="21" pageNumber="22">
SEGMENT 5. With tubes in lateroventral positions (
<figureCitation id="13262A1B5B392A05FCAF3E636633FBE9" box="[793,892,1078,1105]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="21" pageNumber="22">Figs 5B</figureCitation>
,
<figureCitation id="13262A1B5B392A05FC3B3E6366FBFBE8" box="[909,948,1078,1104]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="21" pageNumber="22">7D</figureCitation>
). Sensory spots present in subdorsal, midlateral and ventromedial positions (
<figureCitation id="13262A1B5B392A05FD223E0F6654FBCD" box="[660,795,1114,1141]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="21" pageNumber="22">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B392A05FC9C3E0F661FFBCC" box="[810,848,1114,1140]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="21" pageNumber="22">6D</figureCitation>
,
<figureCitation id="13262A1B5B392A05FCD63E0F66E6FBCC" box="[864,937,1114,1140]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="21" pageNumber="22">7DE</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B392A05FB7A3E0E61B0FBCD" box="[1228,1279,1115,1141]" pageId="21" pageNumber="22">type</typeStatus>
1 present in paradorsal positions. Tips of pectinate fringe of posterior segment margin slightly longer than on preceding segment. Cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BA2369E5B392A05FF0B3EB36196FA29" blockId="21.[188,1401,1253,1425]" pageId="21" pageNumber="22">
SEGMENT 6. With spines in middorsal and lateroventral positions (
<figureCitation id="13262A1B5B392A05FC7B3EB0611EFAB8" box="[973,1105,1253,1280]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="21" pageNumber="22">Fig. 5AB</figureCitation>
). Middorsal spine, as on segment 4, relatively short (31 µm), only slightly exceeding beyond posterior segment margin (
<figureCitation id="13262A1B5B392A05FAA53F5F659BFAF1" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="21" pageNumber="22">Figs 6A, D</figureCitation>
,
<figureCitation id="13262A1B5B392A05FF513F7B6479FAF0" box="[231,310,1326,1352]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="21" pageNumber="22">7A, E</figureCitation>
). Sensory spots present in paradorsal, subdorsal, midlateral and ventromedial positions (
<figureCitation id="13262A1B5B392A05FF733F076403FAD5" box="[197,332,1362,1389]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="21" pageNumber="22">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B392A05FEEC3F0764CFFAD4" box="[346,384,1362,1388]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="21" pageNumber="22">6D</figureCitation>
,
<figureCitation id="13262A1B5B392A05FE393F066496FAD5" box="[399,473,1363,1389]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="21" pageNumber="22">7DE</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B392A05FD4C3F066662FAD5" box="[762,813,1363,1389]" pageId="21" pageNumber="22">type</typeStatus>
1 present in paradorsal positions (
<figureCitation id="13262A1B5B392A05FB783F07607CFAD5" box="[1230,1331,1362,1389]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="21" pageNumber="22">Figs 5A</figureCitation>
,
<figureCitation id="13262A1B5B392A05FAF73F076026FAD4" box="[1345,1385,1362,1388]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="21" pageNumber="22">6D</figureCitation>
). Pectinate fringe of posterior segment margin and cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BA2369E5B392A05FF0B3FEF67F0F9F8" blockId="21.[189,1399,1465,1601]" pageId="21" pageNumber="22">
SEGMENT 7. With spines in lateroventral positions, and sensory spots in subdorsal, midlateral and ventromedial positions (
<figureCitation id="13262A1B5B392A05FE573F8B6726FA41" box="[481,617,1502,1529]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="21" pageNumber="22">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B392A05FDCC3F8B678CFA40" box="[634,707,1502,1528]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="21" pageNumber="22">6CD</figureCitation>
,
<figureCitation id="13262A1B5B392A05FD653F8B67B8FA40" box="[723,759,1502,1528]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="21" pageNumber="22">7E</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B392A05FB973F8B611BFA40" box="[1057,1108,1502,1528]" pageId="21" pageNumber="22">type</typeStatus>
1 present in paradorsal positions. Tips of pectinate fringe of posterior segment margin slightly longer than on preceding segments. Segment otherwise as segment 6.
</paragraph>
<paragraph id="8BA2369E5B392A05FF0B3C3C6769F949" blockId="21.[187,1399,1641,1777]" pageId="21" pageNumber="22">
SEGMENT 8. With spines in lateroventral positions, and tubes in subdorsal, midlateral and lateral accessory positions (
<figureCitation id="13262A1B5B392A05FE8E3CD8648EF910" box="[312,449,1677,1704]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="21" pageNumber="22">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B392A05FE663CD864BAF91F" box="[464,501,1677,1703]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="21" pageNumber="22">6C</figureCitation>
,
<figureCitation id="13262A1B5B392A05FDB33CDB671EF910" box="[517,593,1678,1704]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="21" pageNumber="22">7GH</figureCitation>
). Sensory spots present in subdorsal positions only. Glandular cell outlets
<typeStatus id="54A6883C5B392A05FEA43CE7640AF974" box="[274,325,1714,1740]" pageId="21" pageNumber="22">type</typeStatus>
1 present in paradorsal positions. Pectinate fringe of posterior segment margin and cuticular hairs as on preceding segment.
</paragraph>
<paragraph id="8BA2369E5B392A05FF0B3D4C64A9F851" blockId="21.[187,1398,1816,2025]" pageId="21" pageNumber="22">
SEGMENT 9. With spines in lateroventral positions and tubes in midlateral positions (
<figureCitation id="13262A1B5B392A05FB003D4C6072F88B" box="[1206,1341,1816,1843]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="21" pageNumber="22">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B392A05FAFB3D4C603EF88B" box="[1357,1393,1817,1843]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="21" pageNumber="22">6C</figureCitation>
,
<figureCitation id="13262A1B5B392A05FF0B3D686445F8EF" box="[189,266,1853,1879]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="21" pageNumber="22">7GH</figureCitation>
). In
<specimenCount id="9D1BFD175B392A05FE8A3D6B6493F8EF" box="[316,476,1853,1880]" pageId="21" pageNumber="22" type="generic">one specimen</specimenCount>
additional tube on right side of segment located in subdorsal position (
<figureCitation id="13262A1B5B392A05FAA13D686027F8EF" box="[1303,1384,1853,1879]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="21" pageNumber="22">Fig. 7I</figureCitation>
). Two pairs of sensory spots located in subdorsal positions and one ventrolateral pair (
<figureCitation id="13262A1B5B392A05FB183D34607CF8C4" box="[1198,1331,1889,1916]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="21" pageNumber="22">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B392A05FAF73D346027F8C3" box="[1345,1384,1889,1915]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="21" pageNumber="22">7H</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B392A05FE0B3DD364BFF818" box="[445,496,1926,1952]" pageId="21" pageNumber="22">type</typeStatus>
1 present in paradorsal positions. Small, rounded sieve plates located in lateral accessory positions (
<figureCitation id="13262A1B5B392A05FE193DFF6743F87C" box="[431,524,1962,1988]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="21" pageNumber="22">Fig. 6C</figureCitation>
). Cuticular hair covering and pectinate fringe of posterior segment margin as on preceding segment.
</paragraph>
<caption id="DF6266165B3A2A06FF0B3D4B6674F850" ID-DOI="http://doi.org/10.5281/zenodo.7222171" ID-Zenodo-Dep="7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="22" pageNumber="23" startId="22.[189,232,1822,1848]" targetBox="[199,1389,267,1786]" targetPageId="22">
<paragraph id="8BA2369E5B3A2A06FF0B3D4B6674F850" blockId="22.[188,1399,1822,2025]" pageId="22" pageNumber="23">
<emphasis id="B969EA8C5B3A2A06FF0B3D4B6444F880" bold="true" box="[189,267,1822,1848]" pageId="22" pageNumber="23">Fig. 7.</emphasis>
Scanning electron micrographs showing overviews and details of
<taxonomicName id="4C1D4D1D5B3A2A06FBAD3D4B605BF880" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1051,1300,1822,1848]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="22" pageNumber="23" phylum="Kinorhyncha" rank="species" species="landersi" status="sp. nov.">
<emphasis id="B969EA8C5B3A2A06FBAD3D4B605BF880" box="[1051,1300,1822,1848]" italics="true" pageId="22" pageNumber="23">Echinoderes landersi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B3A2A06FAAA3D4A6038F881" box="[1308,1399,1823,1849]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B969EA8C5B3A2A06FF0B3D14659DF8E3" bold="true" box="[189,210,1857,1883]" pageId="22" pageNumber="23">A</emphasis>
. Dorsal overview.
<emphasis id="B969EA8C5B3A2A06FE053D146489F8E3" bold="true" box="[435,454,1857,1883]" pageId="22" pageNumber="23">B</emphasis>
. Lateral overview.
<emphasis id="B969EA8C5B3A2A06FD1D3D14678FF8E3" bold="true" box="[683,704,1857,1883]" pageId="22" pageNumber="23">C</emphasis>
. Segments 1 to 3, lateral view.
<emphasis id="B969EA8C5B3A2A06FB823D146106F8E3" bold="true" box="[1076,1097,1857,1883]" pageId="22" pageNumber="23">D</emphasis>
. Segments 2 to 6, ventral view.
<emphasis id="B969EA8C5B3A2A06FEB43D306459F8C7" bold="true" box="[258,278,1893,1919]" pageId="22" pageNumber="23">E</emphasis>
. Segments 4 to 7, dorsolateral view.
<emphasis id="B969EA8C5B3A2A06FD773D30679CF8C7" bold="true" box="[705,723,1893,1919]" pageId="22" pageNumber="23">F</emphasis>
. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view.
<emphasis id="B969EA8C5B3A2A06FEE13DD26423F819" bold="true" box="[343,364,1927,1953]" pageId="22" pageNumber="23">G</emphasis>
. Segments 8 to 10, lateral view.
<emphasis id="B969EA8C5B3A2A06FD4E3DDD6642F81A" bold="true" box="[760,781,1928,1954]" pageId="22" pageNumber="23">H</emphasis>
. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10.
<emphasis id="B969EA8C5B3A2A06FD0B3DFE6787F87D" bold="true" box="[701,712,1963,1989]" pageId="22" pageNumber="23">I</emphasis>
. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<paragraph id="8BA2369E5B3B2A07FF0B3B5C66F7FE4A" blockId="23.[189,1401,265,499]" pageId="23" pageNumber="24">
SEGMENT 10. With minute midlateral tubes located near posterior segment margin (
<figureCitation id="13262A1B5B3B2A07FB2A3B5C604FFE9C" box="[1180,1280,265,292]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="23" pageNumber="24">Figs 5A</figureCitation>
,
<figureCitation id="13262A1B5B3B2A07FAB83B5C607CFE9B" box="[1294,1331,265,291]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="23" pageNumber="24">6C</figureCitation>
,
<figureCitation id="13262A1B5B3B2A07FAF73B5C6026FE9B" box="[1345,1385,265,291]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="23" pageNumber="24">7H</figureCitation>
). Sensory spots present in subdorsal and ventrolateral positions (
<figureCitation id="13262A1B5B3B2A07FC113B7E6164FEFE" box="[935,1067,299,326]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="23" pageNumber="24">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B3B2A07FB8E3B79612FFEFE" box="[1080,1120,300,326]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="23" pageNumber="24">7H</figureCitation>
); in
<specimenCount id="9D1BFD175B3B2A07FB233B796079FEFD" box="[1173,1334,299,326]" pageId="23" pageNumber="24" type="generic">one specimen</specimenCount>
extra subdorsal sensory spot present on left side of segment (
<figureCitation id="13262A1B5B3B2A07FCF13B1B66EAFED0" box="[839,933,334,360]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="23" pageNumber="24">Fig. 7H</figureCitation>
). Glandular cell outlet
<typeStatus id="54A6883C5B3B2A07FB053B1A61A9FED1" box="[1203,1254,335,361]" pageId="23" pageNumber="24">type</typeStatus>
1 present in middorsal position. Cuticular hairs less dense on dorsal side than on preceding segment. Central part of tergal plate devoid of hairs. Hairs on sternal plates shorter than on preceding segments. Posterior segment margin of tergal plate straight, without fringe tips; margins of sternal plates concave, reaching posterior margin of terminal segment, with very short fringe tips.
</paragraph>
<paragraph id="8BA2369E5B3B2A07FF0B384C6019FD59" blockId="23.[187,1400,537,737]" pageId="23" pageNumber="24">
SEGMENT 11. With lateral terminal spines (
<figureCitation id="13262A1B5B3B2A07FD2D384C6653FD8C" box="[667,796,537,564]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="23" pageNumber="24">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B3B2A07FC90384C6600FD8B" box="[806,847,537,563]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="23" pageNumber="24">6A</figureCitation>
). Females with lateral terminal accessory spines (
<figureCitation id="13262A1B5B3B2A07FF7538696409FDEE" box="[195,326,571,598]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="23" pageNumber="24">Figs 5AB</figureCitation>
,
<figureCitation id="13262A1B5B3B2A07FEE438696436FDEE" box="[338,377,572,598]" captionStart="Fig" captionStartId="20.[189,232,1891,1917]" captionTargetBox="[189,1399,1200,1848]" captionTargetId="figure-57@20.[189,1399,1199,1849]" captionTargetPageId="20" captionText="Fig. 6. Light micrographs showing overview and details of Echinoderes landersi sp. nov. A. ♀, paratype (NHMD-916627). BD. ♀, holotype (NIWA-159402). A. Lateral overview. B. Segments 1 to 3, lateral view. C. Segments 7 to 11, lateral view. D. Segments 4 to 7, lateral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222169" httpUri="https://zenodo.org/record/7222169/files/figure.png" pageId="23" pageNumber="24">6A</figureCitation>
,
<figureCitation id="13262A1B5B3B2A07FE32386964E4FDEE" box="[388,427,572,598]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="23" pageNumber="24">7H</figureCitation>
); male conditions unknown. Sensory spots and glandular cell outlets
<typeStatus id="54A6883C5B3B2A07FB673869604BFDEE" box="[1233,1284,572,598]" pageId="23" pageNumber="24">type</typeStatus>
1 present in subdorsal positions; in
<specimenCount id="9D1BFD175B3B2A07FE5C380A67C4FDC0" box="[490,651,606,633]" pageId="23" pageNumber="24" type="generic">one specimen</specimenCount>
sensory spot on right side of segment missing. Segment devoid of cuticular hairs, but with very short cuticular hair-like structures covering paradorsal area and very short fringes covering margins of tergal and sternal plates. Tergal extensions triangular, with elongate and pointed tips (
<figureCitation id="13262A1B5B3B2A07FE3C389364A4FD59" box="[394,491,710,737]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="23" pageNumber="24">Figs 5A</figureCitation>
,
<figureCitation id="13262A1B5B3B2A07FE4E38936750FD58" box="[504,543,710,736]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="23" pageNumber="24">7H</figureCitation>
). Sternal extensions do not extend beyond tergal extensions (
<figureCitation id="13262A1B5B3B2A07FB5A38936007FD59" box="[1260,1352,710,737]" captionStart="Fig" captionStartId="19.[189,232,1963,1989]" captionTargetBox="[189,1399,265,1950]" captionTargetId="figure-25@19.[189,1399,265,1950]" captionTargetPageId="19" captionText="Fig. 5. Line art illustrations of Echinoderes landersi sp. nov. A. ♀, dorsal view. B. ♀, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222167" httpUri="https://zenodo.org/record/7222167/files/figure.png" pageId="23" pageNumber="24">Fig. 5B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30765155B3B2A07FF0B3953664AFCC9" pageId="23" pageNumber="24" type="distribution">
<paragraph id="8BA2369E5B3B2A07FF0B39536419FC98" blockId="23.[189,342,774,800]" box="[189,342,774,800]" pageId="23" pageNumber="24">
<heading id="D0EA81F25B3B2A07FF0B39536419FC98" bold="true" box="[189,342,774,800]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B969EA8C5B3B2A07FF0B39536419FC98" bold="true" box="[189,342,774,800]" pageId="23" pageNumber="24">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2369E5B3B2A07FF0B3960664AFCC9" blockId="23.[188,1399,821,881]" pageId="23" pageNumber="24">
Hikurangi slope and Pahaua Canyon,
<quantity id="4CE59B7B5B3B2A07FDC23960664AFCF7" box="[628,773,821,847]" metricMagnitude="2" metricUnit="m" metricValue="9.245000000000001" metricValueMax="11.209999999999999" metricValueMin="7.279999999999999" pageId="23" pageNumber="24" unit="m" value="924.5" valueMax="1121.0" valueMin="728.0">7281121 m</quantity>
b.s.l. See
<figureCitation id="13262A1B5B3B2A07FCCD396066F0FCF7" box="[891,959,821,847]" captionStart="Fig" captionStartId="2.[189,232,1961,1987]" captionTargetBox="[189,1399,1136,1922]" captionTargetId="figure-333@2.[189,1399,1136,1923]" captionTargetPageId="2" captionText="Fig. 1. Map showing the sampling stations. Colour codes indicate slopes (red), canyons (green), and seamounts (yellow). Inset shows New Zealand with a frame marking the sampling area." figureDoi="http://doi.org/10.5281/zenodo.7222159" httpUri="https://zenodo.org/record/7222159/files/figure.png" pageId="23" pageNumber="24">Fig. 1</figureCitation>
for a geographic overview of stations and
<tableCitation id="C69F03255B3B2A07FF5B3902640BFCC9" box="[237,324,855,881]" captionStart="Table 1" captionStartId="3.[185,254,549,577]" captionTargetBox="[192,1910,658,1734]" captionTargetPageId="3" captionText="Table 1 (continued on next four pages). Coordinates and basic data on sampling stations, and identities, type status and catalogue numbers of species of Echinoderes Claparède, 1863 from each station." httpUri="http://table.plazi.org/id/DF6266165B2F2A13FF0F38706644FDDC" pageId="23" pageNumber="24" tableUuid="DF6266165B2F2A13FF0F38706644FDDC">Table 1</tableCitation>
for station and specimen information.
</paragraph>
</subSubSection>
<subSubSection id="C30765155B3B2A08FF0B39CD6638FC81" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="discussion">
<paragraph id="8BA2369E5B3B2A07FF0B39CD6605FC0A" blockId="23.[189,842,919,946]" box="[189,842,919,946]" pageId="23" pageNumber="24">
<heading id="D0EA81F25B3B2A07FF0B39CD6605FC0A" bold="true" box="[189,842,919,946]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B969EA8C5B3B2A07FF0B39CD6605FC0A" bold="true" box="[189,842,919,946]" pageId="23" pageNumber="24">
Taxonomic remarks on
<taxonomicName id="4C1D4D1D5B3B2A07FE5339C267A9FC0A" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[485,742,919,946]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="landersi" status="sp. nov.">
<emphasis id="B969EA8C5B3B2A07FE5339C267A9FC0A" bold="true" box="[485,742,919,946]" italics="true" pageId="23" pageNumber="24">Echinoderes landersi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B3B2A07FD5B39CD6605FC0A" box="[749,842,920,946]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2369E5B3B2A07FF0B399367F1FAC6" blockId="23.[189,1400,966,1407]" pageId="23" pageNumber="24">
<taxonomicName id="4C1D4D1D5B3B2A07FF0B399364F6FC58" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[189,441,966,992]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="landersi" status="sp. nov.">
<emphasis id="B969EA8C5B3B2A07FF0B399364F6FC58" box="[189,441,966,992]" italics="true" pageId="23" pageNumber="24">Echinoderes landersi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B3B2A07FE723993676EFC58" box="[452,545,966,992]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
is easily distinguished from all other congeners by its combination of spines and tubes. The spine pattern on its dorsal side, with middorsal spines on segments 4 and 6, is already an uncommon feature within
<taxonomicName id="4C1D4D1D5B3B2A07FD173E5E667CFB9D" authorityName="Claparede" authorityYear="1863" box="[673,819,1035,1061]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="genus">
<emphasis id="B969EA8C5B3B2A07FD173E5E667CFB9D" box="[673,819,1035,1061]" italics="true" pageId="23" pageNumber="24">Echinoderes</emphasis>
</taxonomicName>
. This trait itself is shared only with
<taxonomicName id="4C1D4D1D5B3B2A07FB453E59643AFBF0" authority="Sorensen, 2014" authorityName="Sorensen" authorityYear="2014" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="astridae">
<emphasis id="B969EA8C5B3B2A07FB453E596039FB9D" box="[1267,1398,1035,1061]" italics="true" pageId="23" pageNumber="24">E. astridae</emphasis>
Sørensen, 2014
</taxonomicName>
,
<taxonomicName id="4C1D4D1D5B3B2A07FEC93E7B6784FBF0" authority="Higgins, 1982" authorityName="Higgins" authorityYear="1982" box="[383,715,1069,1096]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="bispinosus">
<emphasis id="B969EA8C5B3B2A07FEC93E7B6753FBFF" box="[383,540,1069,1095]" italics="true" pageId="23" pageNumber="24">E. bispinosus</emphasis>
<bibRefCitation id="EF8C4B6F5B3B2A07FD953E786784FBF0" author="Higgins R. P." box="[547,715,1069,1096]" pageId="23" pageNumber="24" pagination="305 - 316" refId="ref50573" refString="Higgins R. P. 1982. Three new species of Kinorhyncha from Bermuda. Transactions of the American Microscopical Society 101: 305 - 316. https: // doi. org / 10.2307 / 3225748" type="journal article" year="1982">Higgins, 1982</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1D4D1D5B3B2A07FD613E7B6118FBF0" authority="Yamasaki et al., 2020" authorityName="Yamasaki" authorityYear="2020" box="[727,1111,1069,1096]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="uozumii">
<emphasis id="B969EA8C5B3B2A07FD613E7B661BFBFF" box="[727,852,1070,1095]" italics="true" pageId="23" pageNumber="24">E. uozumii</emphasis>
Yamasaki
<emphasis id="B969EA8C5B3B2A07FC643E7B614AFBFF" box="[978,1029,1069,1095]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
., 2020
</taxonomicName>
and
<taxonomicName id="4C1D4D1D5B3B2A07FB383E7B6059FBFF" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1166,1302,1069,1095]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="dalzottoi" status="sp. nov.">
<emphasis id="B969EA8C5B3B2A07FB383E7B6059FBFF" box="[1166,1302,1069,1095]" italics="true" pageId="23" pageNumber="24">E. dalzottoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B3B2A07FAAB3E7B6038FBF0" box="[1309,1399,1070,1096]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
described below in the present study (
<bibRefCitation id="EF8C4B6F5B3B2A07FD223E056673FBD2" author="Higgins R. P." box="[660,828,1104,1130]" pageId="23" pageNumber="24" pagination="305 - 316" refId="ref50573" refString="Higgins R. P. 1982. Three new species of Kinorhyncha from Bermuda. Transactions of the American Microscopical Society 101: 305 - 316. https: // doi. org / 10.2307 / 3225748" type="journal article" year="1982">Higgins 1982</bibRefCitation>
; Sørensen 2014;
<bibRefCitation id="EF8C4B6F5B3B2A07FBA43E056064FBD2" author="Yamasaki H. &amp; Fujimoto S. &amp; Tanaka H." box="[1042,1323,1104,1130]" pageId="23" pageNumber="24" pagination="537 - 558" refId="ref54257" refString="Yamasaki H., Fujimoto S. &amp; Tanaka H. 2020 b. Three new meiobenthic species from a submarine cave in Japan: Echinoderes gama, E. kajiharai, and E. uozumii (Cyclorhagida: Kinorhyncha). Journal of the Marine Biological Association of the United Kingdom 100 (4): 537 - 558. https: // doi. org / 10.1017 / S 0025315420000429" type="journal article" year="2020">
Yamasaki
<emphasis id="B969EA8C5B3B2A07FB273E04619EFBD2" box="[1169,1233,1104,1130]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2020b
</bibRefCitation>
). This character in combination with its tubes on segment 2 makes
<taxonomicName id="4C1D4D1D5B3B2A07FC213E266155FB35" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[919,1050,1139,1165]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="landersi">
<emphasis id="B969EA8C5B3B2A07FC213E266155FB35" box="[919,1050,1139,1165]" italics="true" pageId="23" pageNumber="24">E. landersi</emphasis>
</taxonomicName>
unique among its congeners. The abovementioned species possess either three pairs of tubes as in
<taxonomicName id="4C1D4D1D5B3B2A07FC5A3EC3613BFB17" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1004,1140,1173,1199]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="dalzottoi">
<emphasis id="B969EA8C5B3B2A07FC5A3EC3613BFB17" box="[1004,1140,1173,1199]" italics="true" pageId="23" pageNumber="24">E. dalzottoi</emphasis>
</taxonomicName>
or only one pair as in
<taxonomicName id="4C1D4D1D5B3B2A07FF083EEC6470FB6A" authorityName="Sorensen" authorityYear="2014" box="[190,319,1208,1234]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="astridae">
<emphasis id="B969EA8C5B3B2A07FF083EEC6470FB6A" box="[190,319,1208,1234]" italics="true" pageId="23" pageNumber="24">E. astridae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D1D5B3B2A07FEF93EEC64A3FB6A" authorityName="Higgins" authorityYear="1982" box="[335,492,1208,1234]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="bispinosus">
<emphasis id="B969EA8C5B3B2A07FEF93EEC64A3FB6A" box="[335,492,1208,1234]" italics="true" pageId="23" pageNumber="24">E. bispinosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D1D5B3B2A07FD9F3EEC67E9FB6A" box="[553,678,1209,1234]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="uozumii">
<emphasis id="B969EA8C5B3B2A07FD9F3EEC67E9FB6A" box="[553,678,1209,1234]" italics="true" pageId="23" pageNumber="24">E. uozumii</emphasis>
</taxonomicName>
. Therefore, having four pairs of tubes on segment 2, located in subdorsal, laterodorsal, sublateral and ventrolateral positions, is uncommon and found exclusively in
<taxonomicName id="4C1D4D1D5B3B2A07FF693EAB642BFAAF" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[223,356,1277,1303]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="landersi">
<emphasis id="B969EA8C5B3B2A07FF693EAB642BFAAF" box="[223,356,1277,1303]" italics="true" pageId="23" pageNumber="24">E. landersi</emphasis>
</taxonomicName>
. The three latter species furthermore differ from
<taxonomicName id="4C1D4D1D5B3B2A07FC703EAB6105FAAF" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[966,1098,1277,1303]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="landersi">
<emphasis id="B969EA8C5B3B2A07FC703EAB6105FAAF" box="[966,1098,1277,1303]" italics="true" pageId="23" pageNumber="24">E. landersi</emphasis>
</taxonomicName>
by being equipped with glandular cell outlets
<typeStatus id="54A6883C5B3B2A07FE083F7564BEFA82" box="[446,497,1312,1338]" pageId="23" pageNumber="24">type</typeStatus>
2; such outlets are absent in
<taxonomicName id="4C1D4D1D5B3B2A07FCFA3F756682FA81" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[844,973,1311,1337]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="landersi">
<emphasis id="B969EA8C5B3B2A07FCFA3F756682FA81" box="[844,973,1311,1337]" italics="true" pageId="23" pageNumber="24">E. landersi</emphasis>
</taxonomicName>
. The regular hair covering found in
<taxonomicName id="4C1D4D1D5B3B2A07FF083F166470FAE4" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[190,319,1346,1372]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="landersi">
<emphasis id="B969EA8C5B3B2A07FF083F166470FAE4" box="[190,319,1346,1372]" italics="true" pageId="23" pageNumber="24">E. landersi</emphasis>
</taxonomicName>
also distinguishes it from
<taxonomicName id="4C1D4D1D5B3B2A07FDC23F1667B3FAE4" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[628,764,1346,1372]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="dalzottoi">
<emphasis id="B969EA8C5B3B2A07FDC23F1667B3FAE4" box="[628,764,1346,1372]" italics="true" pageId="23" pageNumber="24">E. dalzottoi</emphasis>
</taxonomicName>
, which is characterized by having a trunk cuticle with perforation sites only but no cuticular hairs.
</paragraph>
<paragraph id="8BA2369E5B3B2A07FF0B3FF36483F850" blockId="23.[188,1400,1445,2025]" pageId="23" pageNumber="24">
In addition,
<taxonomicName id="4C1D4D1D5B3B2A07FEE73FF3649CFA07" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[337,467,1445,1471]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="landersi" status="sp. nov.">
<emphasis id="B969EA8C5B3B2A07FEE73FF3649CFA07" box="[337,467,1445,1471]" italics="true" pageId="23" pageNumber="24">E. landersi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B3B2A07FE6D3FF36778FA78" box="[475,567,1446,1472]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
can easily be distinguished from these species by other traits, among which the composition of segment 8 with tubes in subdorsal, midlateral and lateral accessory positions is the most important, as these are not present in any other species of
<taxonomicName id="4C1D4D1D5B3B2A07FC493FBF61C0F9BC" authorityName="Claparede" authorityYear="1863" box="[1023,1167,1514,1540]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="genus">
<emphasis id="B969EA8C5B3B2A07FC493FBF61C0F9BC" box="[1023,1167,1514,1540]" italics="true" pageId="23" pageNumber="24">Echinoderes</emphasis>
</taxonomicName>
. Numerous species have various combinations of tubes on this particular segment, but no other species has three pairs of tubes. There are several species with two pairs of tubes, in laterodorsal and lateral accessory positions as observed in, e.g.,
<taxonomicName id="4C1D4D1D5B3B2A07FE0F3C066652F9D4" authority="Higgins, 1983" authorityName="Higgins" authorityYear="1983" box="[441,797,1618,1644]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="abbreviatus">
<emphasis id="B969EA8C5B3B2A07FE0F3C066729F9D4" box="[441,614,1618,1644]" italics="true" pageId="23" pageNumber="24">E. abbreviatus</emphasis>
<bibRefCitation id="EF8C4B6F5B3B2A07FDC73C076652F9D4" author="Higgins R. P." box="[625,797,1618,1644]" pageId="23" pageNumber="24" pagination="1 - 131" refId="ref50610" refString="Higgins R. P. 1983. The Atlantic barrier reef ecosystem at Carrie Bow Cay, Belize, II: Kinorhyncha. Smithsonian Contributions to Marine Science 18: 1 - 131. https: // doi. org / 10.5479 / si. 01960768.18.1" type="journal article" year="1983">Higgins, 1983</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1D4D1D5B3B2A07FC9A3C0661D4F9D4" authority="Pardos et al., 2016" authorityName="Pardos" authorityYear="2016" box="[812,1179,1618,1645]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="belenae">
<emphasis id="B969EA8C5B3B2A07FC9A3C0666E4F9D4" box="[812,939,1618,1644]" italics="true" pageId="23" pageNumber="24">E. belenae</emphasis>
Pardos
<emphasis id="B969EA8C5B3B2A07FBA63C066108F9D4" box="[1040,1095,1618,1644]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
., 2016
</taxonomicName>
or
<taxonomicName id="4C1D4D1D5B3B2A07FB7E3C066434F937" authority="Sorensen, 2006" authorityName="Sorensen" authorityYear="2006" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="intermedius">
<emphasis id="B969EA8C5B3B2A07FB7E3C066038F9D4" box="[1224,1399,1618,1644]" italics="true" pageId="23" pageNumber="24">E. intermedius</emphasis>
<bibRefCitation id="EF8C4B6F5B3B2A07FF0B3C206434F937" author="Sorensen M. V." box="[189,379,1653,1679]" pageId="23" pageNumber="24" pagination="51 - 77" refId="ref52724" refString="Sorensen M. V. 2006. New kinorhynchs from Panama, with a discussion of some phylogenetically significant cuticular structures. Meiofauna Marina 15: 51 - 77." type="journal article" year="2006">Sørensen, 2006</bibRefCitation>
</taxonomicName>
, or in subdorsal and lateral accessory positions as found in
<taxonomicName id="4C1D4D1D5B3B2A07FBDB3C23644EF909" authority="(Zelinka, 1928)" baseAuthorityName="Zelinka" baseAuthorityYear="1928" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="capitatus">
<emphasis id="B969EA8C5B3B2A07FBDB3C2361B0F937" box="[1133,1279,1654,1679]" italics="true" pageId="23" pageNumber="24">E. capitatus</emphasis>
(
<bibRefCitation id="EF8C4B6F5B3B2A07FAA23C2065B6F909" author="Zelinka C." pageId="23" pageNumber="24" refId="ref54329" refString="Zelinka C. 1928. Monographie der Echinodera. Verlag Wilhelm Engelmann, Leipzig. Manuscript received: 18 May 2022" type="book" year="1928">Zelinka, 1928</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C1D4D1D5B3B2A07FEF63CCD67A9F909" authority="GaOrdonez et al., 2008" authorityName="GaOrdonez" authorityYear="2008" box="[320,742,1687,1714]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="isabelae">
<emphasis id="B969EA8C5B3B2A07FEF63CCD6489F909" box="[320,454,1687,1713]" italics="true" pageId="23" pageNumber="24">E. isabelae</emphasis>
<bibRefCitation id="EF8C4B6F5B3B2A07FE663CC267A9F909" author="GaOrdonez D. &amp; Pardos F. &amp; Benito J." box="[464,742,1687,1714]" pageId="23" pageNumber="24" pagination="95 - 111" refId="ref50172" refString="GaOrdonez D., Pardos F. &amp; Benito J. 2008. Three new Echinoderes (Kinorhyncha, Cyclorhagida) from North Spain, with new evolutionary aspects in the genus. Zoologischer Anzeiger 247: 95 - 111. https: // doi. org / 10.1016 / j. jcz. 2007.07.001" type="journal article" year="2008">
<superScript id="7C689BD65B3B2A07FE663CC2671DF909" attach="both" box="[464,594,1687,1713]" fontSize="6" pageId="23" pageNumber="24">GaOrdóñez</superScript>
<emphasis id="B969EA8C5B3B2A07FDEA3CCD67D5F909" box="[604,666,1687,1713]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
, 2008
</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF8C4B6F5B3B2A07FD4E3CC266D3F909" author="Zelinka C." box="[760,924,1687,1713]" pageId="23" pageNumber="24" refId="ref54329" refString="Zelinka C. 1928. Monographie der Echinodera. Verlag Wilhelm Engelmann, Leipzig. Manuscript received: 18 May 2022" type="book" year="1928">Zelinka 1928</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B3B2A07FC1D3CC2611FF909" author="Higgins R. P." box="[939,1104,1687,1713]" pageId="23" pageNumber="24" pagination="1 - 131" refId="ref50610" refString="Higgins R. P. 1983. The Atlantic barrier reef ecosystem at Carrie Bow Cay, Belize, II: Kinorhyncha. Smithsonian Contributions to Marine Science 18: 1 - 131. https: // doi. org / 10.5479 / si. 01960768.18.1" type="journal article" year="1983">Higgins 1983</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B3B2A07FBD63CC2603DF909" author="GaOrdonez D. &amp; Pardos F. &amp; Benito J." box="[1120,1394,1687,1714]" pageId="23" pageNumber="24" pagination="95 - 111" refId="ref50172" refString="GaOrdonez D., Pardos F. &amp; Benito J. 2008. Three new Echinoderes (Kinorhyncha, Cyclorhagida) from North Spain, with new evolutionary aspects in the genus. Zoologischer Anzeiger 247: 95 - 111. https: // doi. org / 10.1016 / j. jcz. 2007.07.001" type="journal article" year="2008">
<superScript id="7C689BD65B3B2A07FBD63CC261AEF909" attach="both" box="[1120,1249,1687,1713]" fontSize="6" pageId="23" pageNumber="24">GaOrdóñez</superScript>
<emphasis id="B969EA8C5B3B2A07FB5A3CCD606DF909" box="[1260,1314,1687,1713]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
. 2008
</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B3B2A07FF0B3CEF643EF96C" author="Sorensen M. V." box="[189,369,1722,1748]" pageId="23" pageNumber="24" pagination="51 - 77" refId="ref52724" refString="Sorensen M. V. 2006. New kinorhynchs from Panama, with a discussion of some phylogenetically significant cuticular structures. Meiofauna Marina 15: 51 - 77." type="journal article" year="2006">Sørensen 2006</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B3B2A07FEC93CEF6720F96C" author="Pardos F. &amp; Herranz M. &amp; Sanchez N." box="[383,623,1722,1748]" pageId="23" pageNumber="24" pagination="26 - 47" refId="ref52161" refString="Pardos F., Herranz M. &amp; Sanchez N. 2016 b. Two sides of a coin: The Phylum Kinorhyncha in Panama. II) Pacific Panama. Zoologischer Anzeiger 265: 26 - 47. https: // doi. org / 10.1016 / j. jcz. 2016.06.006" type="journal article" year="2016">
Pardos
<emphasis id="B969EA8C5B3B2A07FE6F3CEE6758F96C" box="[473,535,1722,1748]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2016b
</bibRefCitation>
); nevertheless, the most common tube pattern is the presence of only one pair of tubes on segment 8, typically in a sublateral or lateral accessory position. Furthermore,
<taxonomicName id="4C1D4D1D5B3B2A07FF0B3D556471F8A1" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[189,318,1791,1817]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="landersi">
<emphasis id="B969EA8C5B3B2A07FF0B3D556471F8A1" box="[189,318,1791,1817]" italics="true" pageId="23" pageNumber="24">E. landersi</emphasis>
</taxonomicName>
possesses midlateral tubes on segment 9, which is another relatively rare trait, shared with only four species, i.e.,
<taxonomicName id="4C1D4D1D5B3B2A07FE7E3D7766D8F884" authority="Higgins &amp; Rao, 1979" authorityName="Higgins &amp; Rao" authorityYear="1979" box="[456,919,1825,1852]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="andamanensis">
<emphasis id="B969EA8C5B3B2A07FE7E3D7767DEF883" box="[456,657,1825,1851]" italics="true" pageId="23" pageNumber="24">E. andamanensis</emphasis>
<bibRefCitation id="EF8C4B6F5B3B2A07FD2E3D7466D8F884" author="Higgins R. P. &amp; Rao G. C." box="[664,919,1825,1852]" pageId="23" pageNumber="24" pagination="75 - 85" refId="ref50699" refString="Higgins R. P. &amp; Rao G. C. 1979. Kinorhynchs from the Andaman Islands. Zoological Journal of the Linnean Society 67: 75 - 85." type="journal article" year="1979">Higgins &amp; Rao, 1979</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1D4D1D5B3B2A07FC153D776144F883" box="[931,1035,1826,1851]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="annae">
<emphasis id="B969EA8C5B3B2A07FC153D776144F883" box="[931,1035,1826,1851]" italics="true" pageId="23" pageNumber="24">E. annae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D1D5B3B2A07FBAD3D7765B7F8E6" authority="Higgins, 1967" authorityName="Higgins" authorityYear="1967" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="newcaledoniensis">
<emphasis id="B969EA8C5B3B2A07FBAD3D776044F883" box="[1051,1291,1825,1851]" italics="true" pageId="23" pageNumber="24">E. newcaledoniensis</emphasis>
<bibRefCitation id="EF8C4B6F5B3B2A07FAA43D7465B7F8E6" author="Higgins R. P." pageId="23" pageNumber="24" pagination="75 - 90" refId="ref50500" refString="Higgins R. P. 1967. The Kinorhyncha of New-Caledonia. Expedition francaise sur les Recifs coralliens de la Nouvelle-Caledonie 2: 75 - 90. Fondation Singer-Polignac, Paris." type="journal article" year="1967">Higgins, 1967</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1D4D1D5B3B2A07FE803D10649FF8E6" authorityName="Yamasaki" authorityYear="2016" box="[310,464,1860,1886]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="serratulus">
<emphasis id="B969EA8C5B3B2A07FE803D10649FF8E6" box="[310,464,1860,1886]" italics="true" pageId="23" pageNumber="24">E. serratulus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8C4B6F5B3B2A07FE693D1167CAF8E6" author="Higgins R. P." box="[479,645,1860,1886]" pageId="23" pageNumber="24" pagination="75 - 90" refId="ref50500" refString="Higgins R. P. 1967. The Kinorhyncha of New-Caledonia. Expedition francaise sur les Recifs coralliens de la Nouvelle-Caledonie 2: 75 - 90. Fondation Singer-Polignac, Paris." type="journal article" year="1967">Higgins 1967</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B3B2A07FD253D1166C0F8E6" author="Higgins R. P. &amp; Rao G. C." box="[659,911,1860,1887]" pageId="23" pageNumber="24" pagination="75 - 85" refId="ref50699" refString="Higgins R. P. &amp; Rao G. C. 1979. Kinorhynchs from the Andaman Islands. Zoological Journal of the Linnean Society 67: 75 - 85." type="journal article" year="1979">Higgins &amp; Rao 1979</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B3B2A07FC2B3D1161ECF8E6" author="Sorensen M. V. &amp; Gasiorowski L. &amp; Randso P. V. &amp; Sanchez N. &amp; Neves R. C." box="[925,1187,1860,1887]" pageId="23" pageNumber="24" pagination="3 - 27" refId="ref53116" refString="Sorensen M. V., Gasiorowski L., Randso P. V., Sanchez N. &amp; Neves R. C. 2016 a. First report of kinorhynchs from Singapore, with the description of three new species. Raffles Bulletin of Zoology 64: 3 - 27." type="journal article" year="2016">
Sørensen
<emphasis id="B969EA8C5B3B2A07FBA53D106108F8E6" box="[1043,1095,1860,1886]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
. 2016a
</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B3B2A07FB193D116024F8E6" author="Yamasaki H." box="[1199,1387,1860,1886]" pageId="23" pageNumber="24" pagination="32" refId="ref53664" refString="Yamasaki H. 2016. Two new Echinoderes species (Echinoderidae, Cyclorhagida, Kinorhyncha) from Nha Trang, Vietnam. Zoological Studies 55: e 32. https: // doi. org / 10.6620 / ZS. 2016.55 - 32" type="journal article" year="2016">Yamasaki 2016</bibRefCitation>
). Nevertheless, none of these species can in any way be confused with
<taxonomicName id="4C1D4D1D5B3B2A07FC613D326119F839" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[983,1110,1895,1921]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="23" pageNumber="24" phylum="Kinorhyncha" rank="species" species="landersi">
<emphasis id="B969EA8C5B3B2A07FC613D326119F839" box="[983,1110,1895,1921]" italics="true" pageId="23" pageNumber="24">E. landersi</emphasis>
</taxonomicName>
due to their significantly different middorsal spine patterns. Hence, besides its uncommon middorsal spine pattern with spines on segments 4 and 6 only, the new species is very easily recognized by the nature of its cuticular structures of segments 2, 8 and 9.
</paragraph>
<paragraph id="8BA2369E5B342A08FF0B3B5C6638FC81" blockId="24.[188,1399,265,825]" pageId="24" pageNumber="25">
One noteworthy observation in
<taxonomicName id="4C1D4D1D5B342A08FDFF3B5F679FFE9B" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[585,720,265,291]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="24" pageNumber="25" phylum="Kinorhyncha" rank="species" species="landersi" status="sp. nov.">
<emphasis id="B969EA8C5B342A08FDFF3B5F679FFE9B" box="[585,720,265,291]" italics="true" pageId="24" pageNumber="25">E. landersi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B342A08FD6B3B5F6671FE9C" box="[733,830,266,292]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
is the inconsistent tube pattern found in
<specimenCount id="9D1BFD175B342A08FAFA3B5F6463FEFE" pageId="24" pageNumber="25" type="generic">one specimen</specimenCount>
(
<figureCitation id="13262A1B5B342A08FE8D3B7964E6FEFF" box="[315,425,300,327]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="24" pageNumber="25">Fig. 7F, I</figureCitation>
). However, in contrast to recent observations of intraspecific variation in species of
<taxonomicName id="4C1D4D1D5B342A08FF0B3B056400FED2" authorityName="Claparede" authorityYear="1863" box="[189,335,336,362]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="24" pageNumber="25" phylum="Kinorhyncha" rank="genus">
<emphasis id="B969EA8C5B342A08FF0B3B056400FED2" box="[189,335,336,362]" italics="true" pageId="24" pageNumber="25">Echinoderes</emphasis>
</taxonomicName>
where pairs of tubes may be present or absent, it was only one tube missing from the pair in
<taxonomicName id="4C1D4D1D5B342A08FF0A3B206472FE36" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[188,317,372,398]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="24" pageNumber="25" phylum="Kinorhyncha" rank="species" species="landersi">
<emphasis id="B969EA8C5B342A08FF0A3B206472FE36" box="[188,317,372,398]" italics="true" pageId="24" pageNumber="25">E. landersi</emphasis>
</taxonomicName>
. Variation in tube patterns, regarding their presence or absence, has been documented, e.g., in
<taxonomicName id="4C1D4D1D5B342A08FF6B3BCD6783FE0A" authority="Grzelak &amp; Sorensen, 2017" authorityName="Grzelak &amp; Sorensen" authorityYear="2017" box="[221,716,407,434]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="24" pageNumber="25" phylum="Kinorhyncha" rank="species" species="daenerysae">
<emphasis id="B969EA8C5B342A08FF6B3BCD64C9FE09" box="[221,390,407,433]" italics="true" pageId="24" pageNumber="25">E. daenerysae</emphasis>
Grzelak &amp; Sørensen,
<quantity id="4CE59B7B5B342A08FD273BCD6783FE0A" box="[657,716,408,434]" metricMagnitude="1" metricUnit="m" metricValue="5.12318" pageId="24" pageNumber="25" unit="in" value="2017.0">2017</quantity>
</taxonomicName>
in
<bibRefCitation id="EF8C4B6F5B342A08FD403BC26170FE0A" author="Grzelak K. &amp; Sorensen M. V." box="[758,1087,407,434]" pageId="24" pageNumber="25" pagination="113 - 147" refId="ref50257" refString="Grzelak K. &amp; Sorensen M. V. 2018. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Spitsbergen, with additional information about known Arctic species. Marine Biology Research 14: 113 - 147. https: // doi. org / 10.1080 / 17451000.2017.1367096" type="journal article" year="2018">Grzelak &amp; Sørensen (2018)</bibRefCitation>
for ventrolateral tubes on segment 2, in
<taxonomicName id="4C1D4D1D5B342A08FED23BE96666FE6D" authority="Higgins &amp; Kristensen, 1988" authorityName="Higgins &amp; Kristensen" authorityYear="1988" box="[356,809,443,469]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="24" pageNumber="25" phylum="Kinorhyncha" rank="species" species="eximus">
<emphasis id="B969EA8C5B342A08FED23BE96499FE6D" box="[356,470,444,469]" italics="true" pageId="24" pageNumber="25">E. eximus</emphasis>
<bibRefCitation id="EF8C4B6F5B342A08FE683BEE6666FE6D" author="Higgins R. P. &amp; Kristensen R. M." box="[478,809,443,469]" pageId="24" pageNumber="25" pagination="1 - 56" refId="ref50656" refString="Higgins R. P. &amp; Kristensen R. M. 1988. Kinorhyncha from Disko Island, West Greenland. Smithsonian Contributions to Zoology 458: 1 - 56. https: // doi. org / 10.5479 / si. 00810282.458" type="journal article" year="1988">Higgins &amp; Kristensen, 1988</bibRefCitation>
</taxonomicName>
for sublateral tubes on segment 9, in
<taxonomicName id="4C1D4D1D5B342A08FB513BE96038FE6D" authorityName="Karling" authorityYear="1954" box="[1255,1399,443,469]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="24" pageNumber="25" phylum="Kinorhyncha" rank="species" species="levanderi">
<emphasis id="B969EA8C5B342A08FB513BE96038FE6D" box="[1255,1399,443,469]" italics="true" pageId="24" pageNumber="25">E. levanderi</emphasis>
</taxonomicName>
for subdorsal tubes on segment 2, and in
<taxonomicName id="4C1D4D1D5B342A08FD763B8A6663FE40" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[704,812,478,504]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="24" pageNumber="25" phylum="Kinorhyncha" rank="species" species="frodoi" status="sp. nov.">
<emphasis id="B969EA8C5B342A08FD763B8A6663FE40" box="[704,812,478,504]" italics="true" pageId="24" pageNumber="25">E. frodoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B342A08FC813B8A66D9FE41" box="[823,918,479,505]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
(present study) for midlateral tubes on segment 1 (
<bibRefCitation id="EF8C4B6F5B342A08FEFA385767C3FDA4" author="Grzelak K. &amp; Sorensen M. V." box="[332,652,514,540]" pageId="24" pageNumber="25" pagination="113 - 147" refId="ref50257" refString="Grzelak K. &amp; Sorensen M. V. 2018. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Spitsbergen, with additional information about known Arctic species. Marine Biology Research 14: 113 - 147. https: // doi. org / 10.1080 / 17451000.2017.1367096" type="journal article" year="2018">Grzelak &amp; Sørensen 2018</bibRefCitation>
; Sørensen 2018). Among specimens of
<taxonomicName id="4C1D4D1D5B342A08FBC2385661B6FDA4" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1140,1273,514,540]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="24" pageNumber="25" phylum="Kinorhyncha" rank="species" species="landersi">
<emphasis id="B969EA8C5B342A08FBC2385661B6FDA4" box="[1140,1273,514,540]" italics="true" pageId="24" pageNumber="25">E. landersi</emphasis>
</taxonomicName>
, however,
<specimenCount id="9D1BFD175B342A08FF0B3873642FFDF8" box="[189,352,550,576]" pageId="24" pageNumber="25" type="generic">one specimen</specimenCount>
lacks the subdorsal tube on the right side on segment 2 (
<figureCitation id="13262A1B5B342A08FBAC38736139FDF8" box="[1050,1142,550,576]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="24" pageNumber="25">Fig. 7F</figureCitation>
), but has a subdorsal tube on the right side of segment 9 (
<figureCitation id="13262A1B5B342A08FD30381C6793FDDC" box="[646,732,585,612]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="24" pageNumber="25">Fig. 7I</figureCitation>
). Tubes in subdorsal positions on segment 9 were not observed in other specimens. A similar inconsistency was described by
<bibRefCitation id="EF8C4B6F5B342A08FBA438386039FD3F" author="Yamasaki H. &amp; Dal Zotto M." box="[1042,1398,621,648]" pageId="24" pageNumber="25" pagination="189 - 205" refId="ref53781" refString="Yamasaki H. &amp; Dal Zotto M. 2019. Investigation of echinoderid kinorhynchs described 90 years ago: redescription of Echinoderes capitatus (Zelinka, 1928) and Echinoderes ferrugineus Zelinka, 1928. Zoologischer Anzeiger 282: 189 - 205. https: // doi. org / 10.1016 / j. jcz. 2019.05.013" type="journal article" year="2019">Yamasaki &amp; Dal Zotto (2019)</bibRefCitation>
for specimens of
<taxonomicName id="4C1D4D1D5B342A08FE3C38C46759FD12" baseAuthorityName="Zelinka" baseAuthorityYear="1928" box="[394,534,657,682]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="24" pageNumber="25" phylum="Kinorhyncha" rank="species" species="capitatus">
<emphasis id="B969EA8C5B342A08FE3C38C46759FD12" box="[394,534,657,682]" italics="true" pageId="24" pageNumber="25">E. capitatus</emphasis>
</taxonomicName>
that lack a ventrolateral tube on one side on segment 2 (
<bibRefCitation id="EF8C4B6F5B342A08FB0738C5640BFD76" author="Yamasaki H. &amp; Dal Zotto M." pageId="24" pageNumber="25" pagination="189 - 205" refId="ref53781" refString="Yamasaki H. &amp; Dal Zotto M. 2019. Investigation of echinoderid kinorhynchs described 90 years ago: redescription of Echinoderes capitatus (Zelinka, 1928) and Echinoderes ferrugineus Zelinka, 1928. Zoologischer Anzeiger 282: 189 - 205. https: // doi. org / 10.1016 / j. jcz. 2019.05.013" type="journal article" year="2019">Yamasaki &amp; Dal Zotto 2019</bibRefCitation>
). Moreover, we observed that
<specimenCount id="9D1BFD175B342A08FD1138E06607FD76" box="[679,840,692,719]" pageId="24" pageNumber="25" type="generic">one specimen</specimenCount>
lacks a subdorsal sensory spot on the right side of segment 10, and that this lacking sensory spot seems to be relocated to segment 9 (
<figureCitation id="13262A1B5B342A08FB63388D6079FD4A" box="[1237,1334,728,754]" captionStart="Fig" captionStartId="22.[189,232,1822,1848]" captionTargetBox="[199,1389,267,1786]" captionTargetId="figure-105@22.[189,1399,265,1796]" captionTargetPageId="22" captionText="Fig. 7. Scanning electron micrographs showing overviews and details of Echinoderes landersi sp. nov. A. Dorsal overview. B. Lateral overview. C. Segments 1 to 3, lateral view. D. Segments 2 to 6, ventral view. E. Segments 4 to 7, dorsolateral view. F. Segments 1 to 3 of female with inconsistent tube pattern, dorsal view. G. Segments 8 to 10, lateral view. H. Segments 8 to 11, dorsal view, with inset showing close-up of midlateral tube on segment 10. I. Segments 8 to 9 of female with inconsistent tube pattern, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222171" httpUri="https://zenodo.org/record/7222171/files/figure.png" pageId="24" pageNumber="25">Fig. 7H</figureCitation>
). For now, we cannot explain the reason for this observed variation, but it seems most likely that it is due to individual abnormalities, rather than intraspecific variation.
</paragraph>
</subSubSection>
</treatment>
</document>