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<document id="80F262A58BC331CB88D0C36E0EA8B201" ID-DOI="10.5252/geodiversitas2023v45a21" ID-ISSN="1638-9395" ID-Zenodo-Dep="10144332" ID-ZooBank="urn:lsid:zoobank.org:pub:395206B2-2C0A-44E4-9F81-4C871D9A39EF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700154359418" checkinUser="plazi" docAuthor="Albesa, Joaquín &amp; Neubauer, Thomas A." docDate="2023" docId="03B087F2BB26FFEFFF5CFB068BD4FEA8" docLanguage="en" docName="geodiversitas.2023.45.21.pdf" docOrigin="Geodiversitas 45 (21)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a21.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Euglandininae Baker 1941" docType="treatment" docVersion="2" lastPageNumber="648" masterDocId="FF89FF8ABB21FFE8FFB2FFD28816FFF9" masterDocTitle="Shifting shell morphology in a Late Miocene-Pliocene land snail species lineage (Gastropoda: Stylommatophora: Spiraxidae), with the description of a new species" masterLastPageNumber="657" masterPageNumber="643" pageNumber="648" updateTime="1700154790388" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="973DC81328DA2FACB652FC0F67142E25">Shifting shell morphology in a Late Miocene-Pliocene land snail species lineage (Gastropoda: Stylommatophora: Spiraxidae), with the description of a new species</mods:title>
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<mods:namePart id="3FC45A435F6DB78DBE473477B07C9056">Albesa, Joaquín</mods:namePart>
<mods:affiliation id="959807CE79482E18477D619B0D2816DE">Departamento de Botánica y Geología, Universitat de València, 46100 Burjassot (Spain) and Museu Valencià dHistoria Natural, P. O. Box 8460, 46018 Alginet (Spain) joaquin. albesa @ uv. es</mods:affiliation>
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<mods:namePart id="4F86BE43D86164FEF57D48AD6F7B1C47">Neubauer, Thomas A.</mods:namePart>
<mods:affiliation id="3B6DB542DC7B57EEFBAF0BD215156AC1">SNSB - Bavarian State Collection for Palaeontology and Geology, Richard-Wagner-Strasse 10, 80333 Munich (Germany) and Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands)</mods:affiliation>
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<subSubSection id="C303656FBB26FFEFFF5CFB068A8BFB17" box="[238,669,1236,1262]" pageId="7" pageNumber="648" type="nomenclature">
<paragraph id="8BA636E4BB26FFEFFF5CFB068A8BFB17" blockId="7.[238,669,1236,1262]" box="[238,669,1236,1262]" pageId="7" pageNumber="648">
<heading id="D0EE8188BB26FFEFFF5CFB068A8BFB17" box="[238,669,1236,1262]" centered="true" fontSize="9" level="2" pageId="7" pageNumber="648" reason="3">
Subfamily
<taxonomicName id="4C194D67BB26FFEFFED0FB068A8BFB17" ID-CoL="7NXMQ" authority="Baker, 1941" authorityName="Baker" authorityYear="1941" box="[354,669,1236,1262]" class="Gastropoda" family="Spiraxidae" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="subFamily" subFamily="Euglandininae">
<smallCapsWord id="8D40A038BB26FFEFFED0FB068A02FB15" baselines="1255,1255" box="[354,532,1236,1262]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Euglandininae" pageId="7" pageNumber="648">EUGLANDININAE</smallCapsWord>
<bibRefCitation id="EF884B15BB26FFEFFDA9FB068A8BFB17" author="BAKER H. B." box="[539,669,1236,1262]" pageId="7" pageNumber="648" pagination="51 - 61" refId="ref9723" refString="BAKER H. B. 1941. - Outline of American Oleacininae and new species from Mexico. The Nautilus 55: 51 - 61. https: // www. biodiversitylibrary. org / page / 8519954" type="journal article" year="1941">Baker, 1941</bibRefCitation>
</taxonomicName>
</heading>
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<subSubSection id="C303656FBB26FFEFFF36FAC68BD4FEA8" pageId="7" pageNumber="648" type="discussion">
<paragraph id="8BA636E4BB26FFEFFF36FAC688FEFAD5" blockId="7.[130,776,1300,2028]" box="[132,232,1300,1326]" pageId="7" pageNumber="648">
<smallCapsWord id="8D40A038BB26FFEFFF36FAC688FEFAD5" baselines="1319,1319" box="[132,232,1300,1326]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="7" pageNumber="648">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8BA636E4BB26FFEFFF36FAE18909F895" blockId="7.[130,776,1300,2028]" pageId="7" pageNumber="648">
The genera
<taxonomicName id="4C194D67BB26FFEFFEB8FAE189BDFAB4" authorityName="Wenz" authorityYear="1914" box="[266,427,1331,1357]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="genus">
<emphasis id="B96DEAF6BB26FFEFFEB8FAE189BDFAB4" box="[266,427,1331,1357]" italics="true" pageId="7" pageNumber="648">Palaeoglandina</emphasis>
</taxonomicName>
,
<taxonomicName id="4C194D67BB26FFEFFE09FAE18A58FAB4" authorityName="Wenz" authorityYear="1914" box="[443,590,1331,1357]" class="Gastropoda" family="Oleacinidae" genus="Pseudoleacina" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="genus">
<emphasis id="B96DEAF6BB26FFEFFE09FAE18A58FAB4" box="[443,590,1331,1357]" italics="true" pageId="7" pageNumber="648">Pseudoleacina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D67BB26FFEFFD3EFAE188A2FA94" authorityName="Pfeffer" authorityYear="1930" class="Gastropoda" family="Oleacinidae" genus="Trachyglandina" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="genus">
<emphasis id="B96DEAF6BB26FFEFFD3EFAE188A2FA94" italics="true" pageId="7" pageNumber="648">Trachyglandina</emphasis>
</taxonomicName>
are traditionally included in
<taxonomicName id="4C194D67BB26FFEFFE40FA818A64FA94" authorityName="H. Adams &amp; A. Adams" authorityYear="1855" box="[498,626,1363,1389]" class="Gastropoda" family="Oleacinidae" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="family">Oleacinidae</taxonomicName>
(
<bibRefCitation id="EF884B15BB26FFEFFD31FA868B16FA94" author="WENZ W." box="[643,768,1363,1390]" pageId="7" pageNumber="648" pagination="737 - 1068" refId="ref13770" refString="WENZ W. 1923. - Fossilium Catalogus I: Animalia. Gastropoda extramarina tertiaria. Pars 20. W. Junk, Berlin: 737 - 1068. https: // www. biodiversitylibrary. org / page / 40596679" type="book chapter" year="1923">Wenz 1923</bibRefCitation>
;
<bibRefCitation id="EF884B15BB26FFEFFF36FAA18927FA74" author="NORDSIECK H." box="[132,305,1395,1421]" pageId="7" pageNumber="648" pagination="153 - 185" refId="ref12749" refString="NORDSIECK H. 2014. - Annotated check-list of the genera of fossil land snails (Gastropoda: Stylommatophora) of western and central Europe (Cretaceous - Pliocene), with description of new taxa. Archiv fur Molluskenkunde 143 (2): 153 - 185. https: // doi. org / 10.1127 / arch. moll / 143 / 2014 / 153 - 185" type="journal article" year="2014">Nordsieck 2014</bibRefCitation>
;
<bibRefCitation id="EF884B15BB26FFEFFE8DFAA18A9CFA74" author="HARZHAUSER M. &amp; NEUBAUER T. A." box="[319,650,1395,1421]" pageId="7" pageNumber="648" pagination="103610" refId="ref11406" refString="HARZHAUSER M. &amp; NEUBAUER T. A. 2021. - A review of the land snail faunas of the European Cenozoic - composition, diversity and turnovers. Earth-Science Reviews 217: 103610. https: // doi. org / 10.1016 / j. earscirev. 2021.103610" type="journal article" year="2021">Harzhauser &amp; Neubauer 2021</bibRefCitation>
). However, this systematic placement is based on an outdated concept that unites true oleacinids and European species presently considered to belong in
<taxonomicName id="4C194D67BB26FFEFFE3DFA0189E8FA14" authorityName="Baker" authorityYear="1939" box="[399,510,1491,1517]" class="Gastropoda" family="Spiraxidae" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="family">Spiraxidae</taxonomicName>
(compare e.g.
<bibRefCitation id="EF884B15BB26FFEFFD12FA0188AAF9F4" author="SCHILEYKO A. A." pageId="7" pageNumber="648" pagination="731 - 880" refId="ref13265" refString="SCHILEYKO A. A. 2000. - Treatise on Recent terrestrial pulmonate molluscs. Part 6. Rhytididae, Chlamydephoridae, Systrophiidae, Haplotrematidae, Streptaxidae, Spiraxidae, Oleacinidae, Testacellidae. Ruthenica, Supplement 2: 731 - 880." type="journal article" year="2000">Schileyko 2000</bibRefCitation>
vs
<bibRefCitation id="EF884B15BB26FFEFFF52FA2189ECF9F4" author="HAUSDORF B. &amp; SOLVERY S." box="[224,506,1523,1549]" pageId="7" pageNumber="648" pagination="307 - 310" refId="ref11654" refString="HAUSDORF B. &amp; SOLVERY S. 2021. - Introduction of the Predatory Land Snail Species Poiretia delesserti (Spiraxidae) in France. Malacologia 63 (2): 307 - 310. https: // doi. org / 10.4002 / 040.063.0209" type="journal article" year="2021">Hausdorf &amp; Solvery 2021</bibRefCitation>
).
<bibRefCitation id="EF884B15BB26FFEFFDA2FA218AC6F9F4" author="NORDSIECK H." box="[528,720,1523,1549]" pageId="7" pageNumber="648" pagination="153 - 185" refId="ref12749" refString="NORDSIECK H. 2014. - Annotated check-list of the genera of fossil land snails (Gastropoda: Stylommatophora) of western and central Europe (Cretaceous - Pliocene), with description of new taxa. Archiv fur Molluskenkunde 143 (2): 153 - 185. https: // doi. org / 10.1127 / arch. moll / 143 / 2014 / 153 - 185" type="journal article" year="2014">Nordsieck (2014)</bibRefCitation>
considered the genera
<taxonomicName id="4C194D67BB26FFEFFEF7F9C089F7F9D5" authorityName="Wenz" authorityYear="1914" box="[325,481,1554,1580]" class="Gastropoda" family="Oleacinidae" genus="Palaeoglandina" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="genus">
<emphasis id="B96DEAF6BB26FFEFFEF7F9C089F7F9D5" box="[325,481,1554,1580]" italics="true" pageId="7" pageNumber="648">Palaeoglandina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D67BB26FFEFFDA3F9C08A88F9D5" authorityName="Wenz" authorityYear="1914" box="[529,670,1554,1580]" class="Gastropoda" family="Oleacinidae" genus="Pseudoleacina" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="genus">
<emphasis id="B96DEAF6BB26FFEFFDA3F9C08A88F9D5" box="[529,670,1554,1580]" italics="true" pageId="7" pageNumber="648">Pseudoleacina</emphasis>
</taxonomicName>
to belong in
<taxonomicName id="4C194D67BB26FFEFFF13F9E08957F9B5" authorityName="Baker" authorityYear="1941" box="[161,321,1586,1612]" class="Gastropoda" family="Spiraxidae" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="subFamily" subFamily="Euglandininae">Euglandininae</taxonomicName>
, which is today ranked as a subfamily of
<taxonomicName id="4C194D67BB26FFEFFF34F98088E3F995" authorityName="Baker" authorityYear="1939" box="[134,245,1618,1644]" class="Gastropoda" family="Spiraxidae" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="family">Spiraxidae</taxonomicName>
(see also
<bibRefCitation id="EF884B15BB26FFEFFEE4F98089E7F995" author="SALVADOR R. B." box="[342,497,1618,1644]" pageId="7" pageNumber="648" pagination="157 - 171" refId="ref13217" refString="SALVADOR R. B. 2013. - The fossil pulmonate snails of Sandelzhausen (Early / Middle Miocene, Germany): Succineidae, Testacelloidea and Helicoidea. Zootaxa 3721: 157 - 171. https: // doi. org / 10.11646 / zootaxa. 3721.2.3" type="journal article" year="2013">
<collectingCountry id="F30E7674BB26FFEFFEE4F98089A4F995" box="[342,434,1618,1644]" name="El Salvador" pageId="7" pageNumber="648">Salvador</collectingCountry>
2013
</bibRefCitation>
). We follow that concept, which is congruent with the similarity of the fossil species and modern representatives of the subfamily, such as
<taxonomicName id="4C194D67BB26FFEFFD3CF9408B11F955" authorityName="Crosse &amp; P. Fischer" authorityYear="1870" box="[654,775,1682,1708]" class="Gastropoda" family="Spiraxidae" genus="Euglandina" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="genus">
<emphasis id="B96DEAF6BB26FFEFFD3CF9408B11F955" box="[654,775,1682,1708]" italics="true" pageId="7" pageNumber="648">Euglandina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D67BB26FFEFFF03F96189ACF935" authority="(Schileyko 2000)" baseAuthorityName="Schileyko" baseAuthorityYear="2000" box="[177,442,1714,1740]" class="Gastropoda" family="Spiraxidae" genus="Poiretia" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="genus">
<emphasis id="B96DEAF6BB26FFEFFF03F9618916F935" box="[177,256,1715,1740]" italics="true" pageId="7" pageNumber="648">Poiretia</emphasis>
(
<bibRefCitation id="EF884B15BB26FFEFFEBCF96089A5F935" author="SCHILEYKO A. A." box="[270,435,1714,1740]" pageId="7" pageNumber="648" pagination="731 - 880" refId="ref13265" refString="SCHILEYKO A. A. 2000. - Treatise on Recent terrestrial pulmonate molluscs. Part 6. Rhytididae, Chlamydephoridae, Systrophiidae, Haplotrematidae, Streptaxidae, Spiraxidae, Oleacinidae, Testacellidae. Ruthenica, Supplement 2: 731 - 880." type="journal article" year="2000">Schileyko 2000</bibRefCitation>
)
</taxonomicName>
. The monotypic genus
<taxonomicName id="4C194D67BB26FFEFFD1FF96088C2F915" authorityName="Pfeffer" authorityYear="1930" class="Gastropoda" family="Oleacinidae" genus="Trachyglandina" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="genus">
<emphasis id="B96DEAF6BB26FFEFFD1FF96088C2F915" italics="true" pageId="7" pageNumber="648">Trachyglandina</emphasis>
</taxonomicName>
is here only tentatively referred to the
<taxonomicName id="4C194D67BB26FFEFFDD7F9008B15F915" authorityName="Baker" authorityYear="1941" box="[613,771,1746,1772]" class="Gastropoda" family="Spiraxidae" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="subFamily" subFamily="Euglandininae">Euglandininae</taxonomicName>
, since it is based solely on a fragment; until further material of the
<typeStatus id="54A28846BB26FFEFFF78F8C088EEF8D5" box="[202,248,1810,1836]" pageId="7" pageNumber="648">type</typeStatus>
species,
<taxonomicName id="4C194D67BB26FFEFFEE1F8C08AFAF8D5" authority="Pfeffer, 1930" authorityName="Pfeffer" authorityYear="1930" box="[339,748,1810,1836]" class="Gastropoda" family="Oleacinidae" genus="Trachyglandina" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="species" species="dietrichii">
<emphasis id="B96DEAF6BB26FFEFFEE1F8C08A4EF8D5" box="[339,600,1810,1836]" italics="true" pageId="7" pageNumber="648">Trachyglandina dietrichii</emphasis>
<bibRefCitation id="EF884B15BB26FFEFFDEDF8C08AFAF8D5" author="PFEFFER G." box="[607,748,1810,1836]" pageId="7" pageNumber="648" pagination="153 - 380" refId="ref12818" refString="PFEFFER G. 1930. - Zur Kenntnis tertiarer Landschnecken. Geologisch-Palaontologische Abhandlungen, neue Folge 17 (3): 153 - 380." type="journal article" year="1930">Pfeffer, 1930</bibRefCitation>
</taxonomicName>
, is discovered both genus and
<typeStatus id="54A28846BB26FFEFFE11F8E089C7F8B5" box="[419,465,1842,1868]" pageId="7" pageNumber="648">type</typeStatus>
species should be considered
<emphasis id="B96DEAF6BB26FFEFFF31F8808900F892" box="[131,278,1873,1899]" italics="true" pageId="7" pageNumber="648">nomina dubia</emphasis>
.
</paragraph>
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The revised placement of the three genera is also supported by the geographic distribution of modern
<taxonomicName id="4C194D67BB26FFEFFDECF8438B16F852" authorityName="Baker" authorityYear="1941" box="[606,768,1937,1963]" class="Gastropoda" family="Spiraxidae" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="subFamily" subFamily="Euglandininae">Euglandininae</taxonomicName>
, some of which occur in the Mediterranean region and the Caucasus (e.g.
<bibRefCitation id="EF884B15BB26FFEFFE96F80389DBF812" author="SCHILEYKO A. A." box="[292,461,2001,2027]" pageId="7" pageNumber="648" pagination="731 - 880" refId="ref13265" refString="SCHILEYKO A. A. 2000. - Treatise on Recent terrestrial pulmonate molluscs. Part 6. Rhytididae, Chlamydephoridae, Systrophiidae, Haplotrematidae, Streptaxidae, Spiraxidae, Oleacinidae, Testacellidae. Ruthenica, Supplement 2: 731 - 880." type="journal article" year="2000">Schileyko 2000</bibRefCitation>
;
<bibRefCitation id="EF884B15BB26FFEFFE69F8038ADEF812" author="WELTER-SCHULTES F." box="[475,712,2001,2027]" pageId="7" pageNumber="648" refId="ref13689" refString="WELTER-SCHULTES F. 2012. - European Non-Marine Molluscs, a Guide for Species Identification. Planet Poster Editions, Gottingen, 760 p." type="book" year="2012">Welter-Schultes 2012</bibRefCitation>
;
<bibRefCitation id="EF884B15BB26FFEFFD67F8038BA4FF0B" author="HELWERDA R. A." box="[725,946,215,2027]" pageId="7" pageNumber="648" pagination="35 - 48" refId="ref11699" refString="HELWERDA R. A. 2015. - Predatory Poiretia (Stylommatophora, Oleacinidae) snails: histology and observations. Vita Malacologica 13: 35 - 48. https: // repository. naturalis. nl / pub / 588032" type="journal article" year="2015">Helwerda 2015</bibRefCitation>
), while true
<taxonomicName id="4C194D67BB26FFEFFBF4FF058CDCFF08" authorityName="H. Adams &amp; A. Adams" authorityYear="1855" box="[1094,1226,215,241]" class="Gastropoda" family="Oleacinidae" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="family">Oleacinidae</taxonomicName>
are restricted to the Americas (
<bibRefCitation id="EF884B15BB26FFEFFC17FF258C45FEE8" author="SCHILEYKO A. A." box="[933,1107,247,273]" pageId="7" pageNumber="648" pagination="731 - 880" refId="ref13265" refString="SCHILEYKO A. A. 2000. - Treatise on Recent terrestrial pulmonate molluscs. Part 6. Rhytididae, Chlamydephoridae, Systrophiidae, Haplotrematidae, Streptaxidae, Spiraxidae, Oleacinidae, Testacellidae. Ruthenica, Supplement 2: 731 - 880." type="journal article" year="2000">Schileyko 2000</bibRefCitation>
;
<bibRefCitation id="EF884B15BB26FFEFFBD7FF258D75FEE8" author="BARKER G. M. &amp; EFFORD M. G." box="[1125,1379,247,273]" pageId="7" pageNumber="648" pagination="279 - 404" refId="ref9827" refString="BARKER G. M. &amp; EFFORD M. G. 2004. - Predatory gastropods as natural enemies of terrestrial gastropods and other invertebrates, in BARKER G. M. (ed.), Natural Enemies of Terrestrial Molluscs. Cabi, Wallingford: 279 - 404. https: // doi. org / 10.1079 / 9780851993195.0279" type="book chapter" year="2004">Barker &amp; Efford 2004</bibRefCitation>
) (note that some of these authors included the relevant genera in
<taxonomicName id="4C194D67BB26FFEFFC9FFEE58BA0FEA8" authorityName="H. Adams &amp; A. Adams" authorityYear="1855" box="[813,950,311,337]" class="Gastropoda" family="Oleacinidae" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="648" phylum="Mollusca" rank="family">Oleacinidae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>