treatments-xml/data/03/B0/87/03B087E55B19FFEEFF79F7EBFCB5FC5E.xml
2024-06-21 12:22:17 +02:00

195 lines
24 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="8C74A1069F7382F9048EBBFAD2B46C37" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B19FFEEFF79F7EBFCB5FC5E" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena ramuquintana Perkins 2011, new species" docType="treatment" docVersion="6" lastPageNumber="126" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="125" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="E7B9DA604CC7F886B4275F48533A4829" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EE42362C6B4504500FCC206B0444D0A8">
<mods:title id="BDF2C5565F955272DAE01E60A2A49656">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
</mods:titleInfo>
<mods:name id="76EE400BAD66D1502F6AC716C468027D" type="personal">
<mods:role id="1C19B6E6FB0C0494AC9B5C5005C5919A">
<mods:roleTerm id="5ABBB4BE3E8E8B01A9CEF7C80C82378C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CCCF9EFDFBA7CFBF703BB681E9349D05">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="70CA585E13AE9FE0D8B20D6D4AB36A4C">text</mods:typeOfResource>
<mods:relatedItem id="E0DEE8682970E53E2E22ABC34D8B9F69" type="host">
<mods:titleInfo id="4B13302B045717F5E0B9B9A56E650E3C">
<mods:title id="4AA982ECAF7D102268E1B3FF4B9A6727">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8A9F4FDB4A81C19DB474A0C06F6EED4C">
<mods:date id="8860E804A33913DA9BD6D8C451B2D379">2011</mods:date>
<mods:detail id="E4E6D205B8A0782927E02F241CBE7750" type="pubDate">
<mods:number id="71D80B4538891908E4C3CFBA87042C45">2011-07-08</mods:number>
</mods:detail>
<mods:detail id="8F6F0CB8E3107D331952C90D3FAE0C72" type="volume">
<mods:number id="7F2967637FBC41A37D56D7F8020F5171">2944</mods:number>
</mods:detail>
<mods:detail id="069D5332B9D31DBFF4DEB5B960D40BA8" type="issue">
<mods:number id="1CEE98D9CA57BD51E9ABDB42BF2DAABF">1</mods:number>
</mods:detail>
<mods:extent id="64B0D5BF4FA41C1B172D66F5BB7AAE32" unit="page">
<mods:start id="E344373D13B1F70F00B594258423012A">1</mods:start>
<mods:end id="BDFC70B3CB5F978AD4C5FE0A2C9EF919">417</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="C47FDE2A0A64B51A33ABB9FB1DD9AAC6">
<mods:url id="37547B94C0B86AAA2ACF2F1A2A8A4DB3">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
</mods:location>
<mods:classification id="888496DD9791441BB95F73D4A514B226">journal article</mods:classification>
<mods:identifier id="CBA8E95F6D4F39A65E44E3D112016228" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
<mods:identifier id="29ED136B00F906968288933DB74EE248" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="F5D74AF69710A9260E973E8655FDAB00" type="Zenodo-Dep">5283420</mods:identifier>
</mods:mods>
<treatment id="03B087E55B19FFEEFF79F7EBFCB5FC5E" ID-DOI="http://doi.org/10.5281/zenodo.5291787" ID-Zenodo-Dep="5291787" LSID="urn:lsid:plazi:treatment:03B087E55B19FFEEFF79F7EBFCB5FC5E" httpUri="http://treatment.plazi.org/id/03B087E55B19FFEEFF79F7EBFCB5FC5E" lastPageId="125" lastPageNumber="126" pageId="124" pageNumber="125">
<subSubSection id="C30365785B19FFEFFF79F7EBFDD5FA76" box="[151,611,1478,1504]" pageId="124" pageNumber="125" type="nomenclature">
<paragraph id="8BA636F35B19FFEFFF79F7EBFDD5FA76" blockId="124.[151,611,1478,1537]" box="[151,611,1478,1504]" pageId="124" pageNumber="125">
<heading id="D0EE819F5B19FFEFFF79F7EBFDD5FA76" bold="true" box="[151,611,1478,1504]" fontSize="11" level="1" pageId="124" pageNumber="125" reason="1">
<emphasis id="B96DEAE15B19FFEFFF79F7EBFDD5FA76" bold="true" box="[151,611,1478,1504]" pageId="124" pageNumber="125">
<taxonomicName id="4C194D705B19FFEFFF79F7EBFE72FA76" ID-CoL="3N3GG" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,452,1478,1504]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="ramuquintana" status="sp. nov.">
<emphasis id="B96DEAE15B19FFEFFF79F7EBFE72FA76" bold="true" box="[151,452,1478,1504]" italics="true" pageId="124" pageNumber="125">Hydraena ramuquintana</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B19FFEFFE3DF7EBFDD5FA76" box="[467,611,1478,1504]" pageId="124" pageNumber="125" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B19FFEFFF79F7C4FE34F997" box="[151,386,1513,1537]" pageId="124" pageNumber="125" type="description">
<paragraph id="8BA636F35B19FFEFFF79F7C4FE34F997" blockId="124.[151,611,1478,1537]" box="[151,386,1513,1537]" pageId="124" pageNumber="125">
(
<figureCitation id="13222A765B19FFEFFF71F7C4FEBAF997" box="[159,268,1513,1537]" captionStart="FIGURE 230" captionStartId="317.[151,250,1753,1776]" captionTargetBox="[208,1388,225,1713]" captionTargetId="figure-26@317.[186,1401,193,1732]" captionTargetPageId="317" captionText="FIGURE 230. Hydraena ramuquintana, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283862" httpUri="https://zenodo.org/record/5283862/files/figure.png" pageId="124" pageNumber="125">Figs. 230</figureCitation>
,
<figureCitation id="13222A765B19FFEFFEF8F7C4FEF5F997" box="[278,323,1513,1537]" captionStart="FIGURES 232233" captionStartId="319.[151,264,1650,1673]" captionTargetBox="[317,1288,216,1619]" captionTargetId="figure-29@319.[264,1323,193,1629]" captionTargetPageId="319" captionText="FIGURES 232233. Aedeagi of holotypes. 232. H. ramuquintana. 233. H. buquintana." figureDoi="http://doi.org/10.5281/zenodo.5283868" httpUri="https://zenodo.org/record/5283868/files/figure.png" pageId="124" pageNumber="125">232</figureCitation>
,
<figureCitation id="13222A765B19FFEFFEA0F7C4FECFF997" box="[334,377,1513,1537]" captionStart="FIGURES 555558" captionStartId="414.[151,264,1650,1673]" captionTargetBox="[165,1421,226,1573]" captionTargetId="figure-23@414.[165,1421,193,1629]" captionTargetPageId="414" captionText="FIGURES 555558. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5295737" httpUri="https://zenodo.org/record/5295737/files/figure.png" pageId="124" pageNumber="125">558</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B19FFEFFF79F401FB03F9F1" pageId="124" pageNumber="125" type="materials_examined">
<paragraph id="8BA636F35B19FFEFFF79F401FB03F9F1" blockId="124.[151,1436,1579,2036]" pageId="124" pageNumber="125">
Type Material.
<materialsCitation id="3B713CAE5B19FFEFFEBEF401FD19F9F1" collectingDate="1991-06-21" collectionCode="MCZ" collectorName="D. Larson" elevation="140" latitude="-5.633333" location="Ramu Valley" longLatPrecision="1305" longitude="145.36667" pageId="124" pageNumber="125" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<typeStatus id="54A288515B19FFEFFEBEF401FE0BF9D2" box="[336,445,1580,1604]" pageId="124" pageNumber="125" type="holotype">
<emphasis id="B96DEAE15B19FFEFFEBEF401FE0BF9D2" bold="true" box="[336,445,1580,1604]" pageId="124" pageNumber="125">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B19FFEFFDC8F401FCB2F9D2" bold="true" box="[550,772,1579,1604]" pageId="124" pageNumber="125">
<collectingRegion id="49DDF8115B19FFEFFDC8F401FD49F9D2" box="[550,767,1579,1604]" country="Papua New Guinea" name="Madang" pageId="124" pageNumber="125">Madang Province</collectingRegion>
:
</emphasis>
<location id="8EC660285B19FFEFFCFDF401FC11F9D2" LSID="urn:lsid:plazi:treatment:03B087E55B19FFEEFF79F7EBFCB5FC5E:8EC660285B19FFEFFCFDF401FC11F9D2" box="[787,935,1580,1604]" latitude="-5.633333" longLatPrecision="1305" longitude="145.36667" name="Ramu Valley" pageId="124" pageNumber="125" stateProvince="Madang">Ramu Valley</location>
,
<locationDeviation id="9A142FD15B19FFEFFC5DF401FBCCF9D2" box="[947,1146,1580,1604]" location="Brahman" pageId="124" pageNumber="125">
<quantity id="4CE19B165B19FFEFFC5DF401FC5BF9D2" box="[947,1005,1580,1604]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="124" pageNumber="125" unit="km" value="6.0">6 km</quantity>
N Brahman
</locationDeviation>
, [GE est.]
<quantity id="4CE19B165B19FFEFFB1AF401FAC0F9D5" box="[1268,1398,1580,1604]" metricMagnitude="2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="124" pageNumber="125" unit="m" value="140.0" valueMax="150.0" valueMin="130.0">
<elevation id="0034D1C05B19FFEFFB1AF401FAC0F9D5" box="[1268,1398,1580,1604]" metricMagnitude="2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="124" pageNumber="125" unit="m" value="140.0" valueMax="150.0" valueMin="130.0">130150 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B19FFEFFA6CF401FF66F9F1" degrees="5" direction="south" minutes="38" orientation="latitude" pageId="124" pageNumber="125" precision="925" value="-5.633333">5° 38' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B19FFEFFF32F462FEE2F9F1" box="[220,340,1615,1639]" degrees="145" direction="east" minutes="22" orientation="longitude" pageId="124" pageNumber="125" precision="925" value="145.36667">145° 22' E</geoCoordinate>
,
<date id="FFA710335B19FFEFFE8EF462FE6FF9F1" box="[352,473,1615,1639]" pageId="124" pageNumber="125" value="1991-06-21">
<collectingDate id="EFE3E9DB5B19FFEFFE8EF462FE6FF9F1" box="[352,473,1615,1639]" pageId="124" pageNumber="125" value="1991-06-21">21 vi 1991</collectingDate>
</date>
,
<collectorName id="26EC53255B19FFEFFE0BF47DFDE0F9F1" box="[485,598,1616,1639]" pageId="124" pageNumber="125">D. Larson</collectorName>
(
<collectionCode id="ED08AE365B19FFEFFD8AF462FD11F9F1" box="[612,679,1615,1639]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="124" pageNumber="125" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B19FFEFFD54F47DFB07F9F1" box="[698,1201,1615,1640]" collectingDate="1991-06-21" collectionCode="MCZ" collectorName="D. Larson" elevation="140" latitude="-5.633333" location="Ramu Valley" longLatPrecision="1305" longitude="145.36667" pageId="124" pageNumber="125" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<emphasis id="B96DEAE15B19FFEFFD54F47DFC84F9FE" bold="true" box="[698,818,1616,1640]" pageId="124" pageNumber="125">
<typeStatus id="54A288515B19FFEFFD54F47DFC84F9FE" box="[698,818,1616,1640]" pageId="124" pageNumber="125" type="paratype">Paratypes</typeStatus>
</emphasis>
: Same data as holotype (2
<collectionCode id="ED08AE365B19FFEFFB89F462FB1FF9F1" box="[1127,1193,1615,1639]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="124" pageNumber="125" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B19FFEFFF29F45EFCD4F8A9" pageId="124" pageNumber="125" type="diagnosis">
<paragraph id="8BA636F35B19FFEFFF29F45EFCD4F8A9" blockId="124.[151,1436,1579,2036]" pageId="124" pageNumber="125">
<emphasis id="B96DEAE15B19FFEFFF29F45EFE63F91A" bold="true" box="[199,469,1651,1676]" pageId="124" pageNumber="125">Differential Diagnosis.</emphasis>
Similar to
<taxonomicName id="4C194D705B19FFEFFDB7F458FD57F91A" authorityName="Perkins" authorityYear="2011" box="[601,737,1652,1676]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="quintana">
<emphasis id="B96DEAE15B19FFEFFDB7F458FDDBF91A" box="[601,621,1653,1676]" italics="true" pageId="124" pageNumber="125">H</emphasis>
.
<emphasis id="B96DEAE15B19FFEFFD90F458FD57F91A" box="[638,737,1653,1676]" italics="true" pageId="124" pageNumber="125">quintana</emphasis>
</taxonomicName>
in dorsal habitus, small size (ca.
<quantity id="4CE19B165B19FFEFFB89F459FB64F91A" box="[1127,1234,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="124" pageNumber="125" unit="mm" value="1.21">1.21 mm</quantity>
), and subserial to random discal elytral punctures (
<figureCitation id="13222A765B19FFEFFDE0F4BAFDCDF939" box="[526,635,1687,1711]" captionStart="FIGURE 227" captionStartId="315.[151,250,1753,1776]" captionTargetBox="[256,1331,214,1724]" captionTargetId="figure-26@315.[240,1347,193,1732]" captionTargetPageId="315" captionText="FIGURE 227. Hydraena quintana, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283860" httpUri="https://zenodo.org/record/5283860/files/figure.png" pageId="124" pageNumber="125">Figs. 227</figureCitation>
,
<figureCitation id="13222A765B19FFEFFD66F4BAFD00F939" box="[648,694,1687,1711]" captionStart="FIGURE 230" captionStartId="317.[151,250,1753,1776]" captionTargetBox="[208,1388,225,1713]" captionTargetId="figure-26@317.[186,1401,193,1732]" captionTargetPageId="317" captionText="FIGURE 230. Hydraena ramuquintana, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283862" httpUri="https://zenodo.org/record/5283862/files/figure.png" pageId="124" pageNumber="125">230</figureCitation>
); differing therefrom ventrally in plaque ratios,
<taxonomicName id="4C194D705B19FFEFFB37F4B4FA2AF939" authorityName="Perkins" authorityYear="2011" box="[1241,1436,1688,1712]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="ramuquintana">
<emphasis id="B96DEAE15B19FFEFFB37F4B4FB5BF926" box="[1241,1261,1689,1712]" italics="true" pageId="124" pageNumber="125">H</emphasis>
.
<emphasis id="B96DEAE15B19FFEFFB13F4B5FA2AF939" box="[1277,1436,1688,1711]" italics="true" pageId="124" pageNumber="125">ramuquintana</emphasis>
</taxonomicName>
having the plaques less convergent anteriorly and P2 wider (ratios ca. 3/1.5/5/4 vs. 2/2/73). Males of the two species are immediately differentiated by the highly modified abdominal ventrites of
<taxonomicName id="4C194D705B19FFEFFBA5F4CCFB65F961" authorityName="Perkins" authorityYear="2011" box="[1099,1235,1760,1784]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="quintana">
<emphasis id="B96DEAE15B19FFEFFBA5F4CCFBE9F96E" box="[1099,1119,1761,1784]" italics="true" pageId="124" pageNumber="125">H</emphasis>
.
<emphasis id="B96DEAE15B19FFEFFB9EF4CDFB65F961" box="[1136,1235,1760,1783]" italics="true" pageId="124" pageNumber="125">quintana</emphasis>
</taxonomicName>
(SEM
<figureCitation id="13222A765B19FFEFFAC4F4F2FF75F88A" captionStart="FIGURES 352357" captionStartId="363.[151,264,1650,1673]" captionTargetBox="[181,1409,197,1625]" captionTargetId="figure-21@363.[177,1410,193,1629]" captionTargetPageId="363" captionText="FIGURES 352357. Scanning electron micrographs of H. quintana." figureDoi="http://doi.org/10.5281/zenodo.5284005" httpUri="https://zenodo.org/record/5284005/files/figure.png" pageId="124" pageNumber="125">Figs. 356, 357</figureCitation>
,
<figureCitation id="13222A765B19FFEFFF21F529FF4BF88A" box="[207,253,1796,1820]" captionStart="FIGURES 358363" captionStartId="364.[151,264,1650,1673]" captionTargetBox="[185,1402,197,1621]" captionTargetId="figure-29@364.[182,1405,193,1629]" captionTargetPageId="364" captionText="FIGURES 358363. Scanning electron micrographs of H. quintana (358361) and H. otiarca (362363)." figureDoi="http://doi.org/10.5281/zenodo.5284007" httpUri="https://zenodo.org/record/5284007/files/figure.png" pageId="124" pageNumber="125">361</figureCitation>
). The male genitalia of all three species in the
<emphasis id="B96DEAE15B19FFEFFCE8F529FCC6F88A" box="[774,880,1796,1820]" italics="true" pageId="124" pageNumber="125">Quintana</emphasis>
group distinctively differ from one another in the shapes of the processes of the distal piece (
<figureCitation id="13222A765B19FFEFFD96F50AFD55F8A9" box="[632,739,1831,1855]" captionStart="FIGURES 228229" captionStartId="316.[151,264,1648,1671]" captionText="FIGURES 228229. Aedeagi of holotypes. 228. H. apexa. 229. H. quintana." pageId="124" pageNumber="125">Figs. 229</figureCitation>
,
<figureCitation id="13222A765B19FFEFFD01F50AFCE3F8A9" box="[751,853,1831,1855]" captionStart="FIGURES 232233" captionStartId="319.[151,264,1650,1673]" captionTargetBox="[317,1288,216,1619]" captionTargetId="figure-29@319.[264,1323,193,1629]" captionTargetPageId="319" captionText="FIGURES 232233. Aedeagi of holotypes. 232. H. ramuquintana. 233. H. buquintana." figureDoi="http://doi.org/10.5281/zenodo.5283868" httpUri="https://zenodo.org/record/5283868/files/figure.png" pageId="124" pageNumber="125">232, 233</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B19FFEEFF29F566FA94FCCA" lastPageId="125" lastPageNumber="126" pageId="124" pageNumber="125" type="description">
<paragraph id="8BA636F35B19FFEFFF29F566FDF5F862" blockId="124.[151,1436,1579,2036]" pageId="124" pageNumber="125">
<emphasis id="B96DEAE15B19FFEFFF29F566FEEEF8F2" bold="true" box="[199,344,1867,1892]" inLineHeading="true" pageId="124" pageNumber="125" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B19FFEFFE42F561FDB8F8F2" box="[428,526,1868,1892]" pageId="124" pageNumber="125" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.21/0.50; head 0.22/0.30; pronotum 0.33/0.41, PA 0.32, PB 0.37; elytra 0.71/0.50. Dorsum of head dark brown to piceous, lateral areas of clypeus slightly lighter; pronotum testaceous in front of and behind brown to dark brown, diffusely margined, fascia, ratios of color bands, as measured in midline, ca. 5/11/5; elytra brown; legs light brown; maxillary palpi testaceous, tip not darker. Dorsum shining.
</paragraph>
<paragraph id="8BA636F35B18FFEEFF29F2B5FB33FE59" blockId="125.[151,1437,151,968]" pageId="125" pageNumber="126">Frons punctures ca. 1xef near eyes, smaller and sparser medially; interstices shining, 15xpd. Clypeus effacedly microreticulate laterally, very finely sparsely punctulate and shining medially. Mentum effacedly microreticulate, very sparsely very finely punctulate, weakly shining. Postmentum microreticulate in shallow median depression, otherwise finely sparsely punctate. Genae very slightly raised, shining, with shallow, obsolete median impression, without posterior ridge. Pronotum transverse, ca. median 3/4 of anterior margin slightly arcuate to posterior, sides weakly arcuate, slightly emarginate between midlength and posterior angle; punctures on disc very fine and very sparse, slightly smaller than largest frons punctures, interstices shining, ca. 68xpd, punctures anteriorly and posteriorly slightly larger than those on disc; PF1 and PF4 absent; PF2 very shallow, completely shallowly confluent in midline to form U-shaped impression; PF3 moderately deep, wide.</paragraph>
<paragraph id="8BA636F35B18FFEEFF29F3F1FB60FDC9" blockId="125.[151,1437,151,968]" pageId="125" pageNumber="126">Elytra widest at about midlength; summit of posterior declivity at or very near midlength; lateral explanate margins narrow; on basal 1/3 punctures subserial to random, ca. 1xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 13xpd. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming moderately deep angle with one another.</paragraph>
<paragraph id="8BA636F35B18FFEEFF29F041FA94FCCA" blockId="125.[151,1437,151,968]" pageId="125" pageNumber="126">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 3/1.5/5/4. P1 laminate; median carina sinuate in profile, slightly raised behind coxae. P2 length/width ca. 5/3, sides very slightly diverging toward blunt, raised apex. Plaques slightly rounded, at sides of median depression, anteriorly tapering and slightly convergent. Metaventrite between mesocoxae very weakly raised in midline. AIS width at straight posterior margin slightly wider than P2. All legs of moderate length. Profemur (male) without tubercle on ventral margin; protibia straight, very slightly widened on medial margin at about midlength. Meso- and metatibia straight, slender. Abdominal apex very slightly asymmetrical; last tergite (male) with very small apicomedian notch. Aedeagus as illustrated (
<figureCitation id="13222A765B18FFEEFB5DF169FAA0FCCA" box="[1203,1302,836,860]" captionStart="FIGURES 232233" captionStartId="319.[151,264,1650,1673]" captionTargetBox="[317,1288,216,1619]" captionTargetId="figure-29@319.[264,1323,193,1629]" captionTargetPageId="319" captionText="FIGURES 232233. Aedeagi of holotypes. 232. H. ramuquintana. 233. H. buquintana." figureDoi="http://doi.org/10.5281/zenodo.5283868" httpUri="https://zenodo.org/record/5283868/files/figure.png" pageId="125" pageNumber="126">Fig. 232</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B18FFEEFF29F145FAB4FCE9" box="[199,1282,871,896]" pageId="125" pageNumber="126" type="etymology">
<paragraph id="8BA636F35B18FFEEFF29F145FAB4FCE9" blockId="125.[151,1437,151,968]" box="[199,1282,871,896]" pageId="125" pageNumber="126">
<emphasis id="B96DEAE15B18FFEEFF29F145FEFBFC16" bold="true" box="[199,333,872,896]" inLineHeading="true" pageId="125" pageNumber="126" reason="1">Etymology.</emphasis>
Named in reference to the geographical distribution and relationship to
<taxonomicName id="4C194D705B18FFEEFB9BF144FB4CFCE9" authorityName="Perkins" authorityYear="2011" box="[1141,1274,872,896]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="quintana">
<emphasis id="B96DEAE15B18FFEEFB9BF144FB3FFC16" box="[1141,1161,873,896]" italics="true" pageId="125" pageNumber="126">H</emphasis>
.
<emphasis id="B96DEAE15B18FFEEFB79F145FB4CFCE9" box="[1175,1274,872,895]" italics="true" pageId="125" pageNumber="126">quintana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B18FFEEFF29F1A6FCB5FC5E" pageId="125" pageNumber="126" type="distribution">
<paragraph id="8BA636F35B18FFEEFF29F1A6FCB5FC5E" blockId="125.[151,1437,151,968]" pageId="125" pageNumber="126">
<emphasis id="B96DEAE15B18FFEEFF29F1A6FED6FC32" bold="true" box="[199,352,907,932]" inLineHeading="true" pageId="125" pageNumber="126" reason="1">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54A288515B18FFEEFD35F1A1FCBDFC32" box="[731,779,908,932]" pageId="125" pageNumber="126">type</typeStatus>
locality near Brahman in the Ramu Valley; this is in the eastern arm of Area 3; elevation
<quantity id="4CE19B165B18FFEEFDE8F182FD33FC51" box="[518,645,943,967]" metricMagnitude="2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="125" pageNumber="126" unit="m" value="140.0" valueMax="150.0" valueMin="130.0">130150 m</quantity>
(
<figureCitation id="13222A765B18FFEEFD7AF182FD45FC51" box="[660,755,943,967]" captionStart="FIGURES 555558" captionStartId="414.[151,264,1650,1673]" captionTargetBox="[165,1421,226,1573]" captionTargetId="figure-23@414.[165,1421,193,1629]" captionTargetPageId="414" captionText="FIGURES 555558. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5295737" httpUri="https://zenodo.org/record/5295737/files/figure.png" pageId="125" pageNumber="126">Fig. 558</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>