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<document id="299C1A509D7F0A9A854E5AEE000B9F56" ID-CLB-Dataset="45364" ID-DOI="10.11646/zootaxa.3826.1.3" ID-GBIF-Dataset="6f4b1535-6c6b-4446-aa26-b1ba83ad9126" ID-ISSN="1175-5326" ID-Zenodo-Dep="228537" ID-ZooBank="53549D58-8F38-47B5-879F-0245E900C131" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460444968431" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Métais, Grégoire, Bartolini, Annachiara, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H., Marivaux, Laurent &amp; Welcomme, Jean-Loup" docDate="2014" docId="03A087E00E4EFFB5FF7FFC3CFA5EF8EE" docLanguage="en" docName="zt03826p138.pdf" docOrigin="Zootaxa 3826 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyriopsis Merle, Pacaud, Métais, Bartolini, Lashari, Brohi, Solangi, Marivaux &amp; Welcomme, 2014, gen. nov." docType="treatment" docUuid="EBE6EA53-FF6B-4026-850A-8F3DC63C07FE" docUuidSource="ZooBank" docVersion="10" lastPageNumber="108" masterDocId="FF99FF980E49FFB2FFE8FFEEFFDAFFAB" masterDocTitle="Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography" masterLastPageNumber="138" masterPageNumber="101" pageNumber="108" updateTime="1698333283847" updateUser="ExternalLinkService">
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<mods:title id="525B27FE1E369D159C5D692E74DE9DC6">Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography</mods:title>
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<mods:namePart id="0F22008B261917AF6373F06C30EC9AB1">Merle, Didier</mods:namePart>
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<mods:namePart id="3ECB9A675F06A1754516A3002C7DDD99">Bartolini, Annachiara</mods:namePart>
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<mods:namePart id="7C1363FDCA0900A35E62A7A0D41BF9D0">Brohi, Imdad A.</mods:namePart>
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<treatment id="03A087E00E4EFFB5FF7FFC3CFA5EF8EE" ID-DOI="http://doi.org/10.5281/zenodo.6137806" ID-GBIF-Taxon="119509276" ID-Zenodo-Dep="6137806" LSID="urn:lsid:zoobank.org:act:EBE6EA53-FF6B-4026-850A-8F3DC63C07FE" httpUri="http://treatment.plazi.org/id/03A087E00E4EFFB5FF7FFC3CFA5EF8EE" lastPageNumber="108" pageId="7" pageNumber="108">
<subSubSection id="C313657D0E4EFFB5FF7FFC3CFE0BFC47" box="[151,465,978,1004]" pageId="7" pageNumber="108" type="nomenclature">
<paragraph id="8BB636F60E4EFFB5FF7FFC3CFE0BFC47" blockId="7.[151,465,978,1004]" box="[151,465,978,1004]" pageId="7" pageNumber="108">
<heading id="D0FE819A0E4EFFB5FF7FFC3CFE0BFC47" bold="true" box="[151,465,978,1004]" fontSize="11" level="1" pageId="7" pageNumber="108" reason="1">
<emphasis id="B97DEAE40E4EFFB5FF7FFC3CFE0BFC47" bold="true" box="[151,465,978,1004]" pageId="7" pageNumber="108">
Genus
<taxonomicName id="4C094D750E4EFFB5FF06FC3CFE81FC47" box="[238,347,978,1004]" class="Gastropoda" family="Volutidae" genus="Lyriopsis" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E4EFFB5FF06FC3CFE81FC47" bold="true" box="[238,347,978,1004]" italics="true" pageId="7" pageNumber="108">Lyriopsis</emphasis>
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<taxonomicNameLabel id="A24E579F0E4EFFB5FE8BFC3CFE0BFC47" box="[355,465,978,1004]" pageId="7" pageNumber="108" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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</subSubSection>
<subSubSection id="C313657D0E4EFFB5FF7FFBF7FB08FB84" box="[151,1234,1049,1071]" pageId="7" pageNumber="108" type="materials_examined">
<paragraph id="8BB636F60E4EFFB5FF7FFBF7FB08FB84" blockId="7.[151,1234,1049,1071]" box="[151,1234,1049,1071]" pageId="7" pageNumber="108">
<emphasis id="B97DEAE40E4EFFB5FF7FFBF7FEF8FB84" bold="true" box="[151,290,1049,1071]" pageId="7" pageNumber="108">
<typeStatus id="54B288540E4EFFB5FF7FFBF7FF16FB84" box="[151,204,1049,1071]" pageId="7" pageNumber="108">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C094D750E4EFFB5FEC1FBF4FD4BFB84" authority="Vredenburg, 1923" authorityName="Vredenburg" authorityYear="1923" box="[297,657,1049,1071]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="cossmanni">
<emphasis id="B97DEAE40E4EFFB5FEC1FBF4FE0BFB84" box="[297,465,1050,1071]" italics="true" pageId="7" pageNumber="108">Lyria cossmanni</emphasis>
<bibRefCitation id="EF984B070E4EFFB5FE3FFBF7FD4BFB84" author="Vredenburg" box="[471,657,1049,1071]" pageId="7" pageNumber="108" refString="Vredenburg, E. W. (1923) Indian Tertiary Gastropoda, IV. Olividae, Harpidae, Marginellidae, Volutidae and Mitridae, with comparative diagnosis of new species. Records of the Geological Survey of India, 54, 243 - 276." type="journal article" year="1923">Vredenburg, 1923</bibRefCitation>
</taxonomicName>
. Early Eocene from Lakhra Formation (Sindh,
<collectingCountry id="F31E76660E4EFFB5FB85FBF7FB12FB84" box="[1133,1224,1049,1071]" name="Pakistan" pageId="7" pageNumber="108">Pakistan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C313657D0E4EFFB5FF7FFBB2FAD7FBDE" box="[151,1293,1116,1141]" pageId="7" pageNumber="108" type="etymology">
<paragraph id="8BB636F60E4EFFB5FF7FFBB2FAD7FBDE" blockId="7.[151,1437,1116,1861]" box="[151,1293,1116,1141]" pageId="7" pageNumber="108">
<emphasis id="B97DEAE40E4EFFB5FF7FFBB2FEC4FBDE" bold="true" box="[151,286,1116,1141]" pageId="7" pageNumber="108">Etymology.</emphasis>
Combination of
<taxonomicName id="4C094D750E4EFFB5FE34FBB3FDCDFBDF" box="[476,535,1117,1140]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E4EFFB5FE34FBB3FDCDFBDF" box="[476,535,1117,1140]" italics="true" pageId="7" pageNumber="108">Lyria</emphasis>
</taxonomicName>
with the suffix
<emphasis id="B97DEAE40E4EFFB5FD20FBB3FCDBFBDF" box="[712,769,1117,1140]" italics="true" pageId="7" pageNumber="108">opsis</emphasis>
(resembling in appearance). Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C313657D0E4EFFB5FF2FFB91FBADFA83" pageId="7" pageNumber="108" type="diagnosis">
<paragraph id="8BB636F60E4EFFB5FF2FFB91FD98FB4B" blockId="7.[151,1437,1116,1861]" pageId="7" pageNumber="108">
<emphasis id="B97DEAE40E4EFFB5FF2FFB91FE9BFB33" bold="true" box="[199,321,1151,1176]" pageId="7" pageNumber="108">Diagnosis.</emphasis>
A ventricose
<emphasis id="B97DEAE40E4EFFB5FE34FB6EFDECFB3C" box="[476,566,1152,1175]" italics="true" pageId="7" pageNumber="108">
<taxonomicName id="4C094D750E4EFFB5FE0CFB6EFDF4FB3C" box="[484,558,1152,1175]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">Lyria</taxonomicName>
</emphasis>
shape with costae slightly opisthocline, rather flexuous in their posterior part, not corresponding from whorl to whorl. Whorls rather depressed, slightly convex, terraced. Four oblique, strong columellar folds. Protoconch bulbous.
</paragraph>
<paragraph id="8BB636F60E4EFFB5FF2FFB02FBADFA83" blockId="7.[151,1437,1116,1861]" pageId="7" pageNumber="108">
<emphasis id="B97DEAE40E4EFFB5FF2FFB02FE53FAAE" bold="true" box="[199,393,1260,1285]" pageId="7" pageNumber="108">Included species</emphasis>
.
<taxonomicName id="4C094D750E4EFFB5FE7FFB03FCFAFAAF" authority="Vredenburg, 1923" authorityName="Vredenburg" authorityYear="1923" box="[407,800,1260,1285]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="cossmanni">
<emphasis id="B97DEAE40E4EFFB5FE7FFB03FD8AFAAF" box="[407,592,1261,1284]" italics="true" pageId="7" pageNumber="108">Lyria cossmanni</emphasis>
<bibRefCitation id="EF984B070E4EFFB5FDBFFB02FCFAFAAF" author="Vredenburg" box="[599,800,1260,1285]" pageId="7" pageNumber="108" refString="Vredenburg, E. W. (1923) Indian Tertiary Gastropoda, IV. Olividae, Harpidae, Marginellidae, Volutidae and Mitridae, with comparative diagnosis of new species. Records of the Geological Survey of India, 54, 243 - 276." type="journal article" year="1923">Vredenburg, 1923</bibRefCitation>
</taxonomicName>
(Lakhra Formation, Sindh Province,
<collectingCountry id="F31E76660E4EFFB5FB2AFB02FAF9FAAE" box="[1218,1315,1260,1285]" name="Pakistan" pageId="7" pageNumber="108">Pakistan</collectingCountry>
) and
<taxonomicName id="4C094D750E4EFFB5FA88FB03FE78FA83" authority="Cox, 1930" authorityName="Cox" authorityYear="1930" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="samanaensis">
<emphasis id="B97DEAE40E4EFFB5FA88FB03FEFCFA83" italics="true" pageId="7" pageNumber="108">Lyria samanaensis</emphasis>
<bibRefCitation id="EF984B070E4EFFB5FEC5FAFEFE78FA83" author="Cox" box="[301,418,1296,1320]" pageId="7" pageNumber="108" refString="Cox, L. R. (1930) The fossil fauna of the Samana Range and some neighbouring areas. Part 8. The mollusc of the Hangu Shales. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 15, 129 - 222." type="journal article" year="1930">Cox, 1930</bibRefCitation>
</taxonomicName>
(Hangu Formation, Paleocene, Khyber Pakhtunkhwa,
<collectingCountry id="F31E76660E4EFFB5FBEFFAE1FBB1FA83" box="[1031,1131,1295,1320]" name="Pakistan" pageId="7" pageNumber="108">Pakistan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C313657D0E4EFFB5FF2FFADAFA5EF8EE" pageId="7" pageNumber="108" type="discussion">
<paragraph id="8BB636F60E4EFFB5FF2FFADAFA5EF8EE" blockId="7.[151,1437,1116,1861]" pageId="7" pageNumber="108">
<emphasis id="B97DEAE40E4EFFB5FF2FFADAFE96FAE6" bold="true" box="[199,332,1332,1357]" pageId="7" pageNumber="108">Discussion.</emphasis>
<bibRefCitation id="EF984B070E4EFFB5FEBCFADAFD44FAE7" author="Cossmann" box="[340,670,1332,1357]" pageId="7" pageNumber="108" refString="Cossmann, M. &amp; Pissarro, G. (1909) The Mollusca of the Ranikot Series. Part 1, Cephalopoda and Gastropoda. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 3 (1), 1 - 83." type="journal article" year="1909">Cossmann &amp; Pissarro (1909)</bibRefCitation>
,
<bibRefCitation id="EF984B070E4EFFB5FD44FADAFC58FAE7" author="Vredenburg" box="[684,898,1332,1357]" pageId="7" pageNumber="108" refString="Vredenburg, E. W. (1923) Indian Tertiary Gastropoda, IV. Olividae, Harpidae, Marginellidae, Volutidae and Mitridae, with comparative diagnosis of new species. Records of the Geological Survey of India, 54, 243 - 276." type="journal article" year="1923">Vredenburg (1923)</bibRefCitation>
and
<bibRefCitation id="EF984B070E4EFFB5FC53FADAFBE7FAE7" author="Cox" box="[955,1085,1332,1356]" pageId="7" pageNumber="108" refString="Cox, L. R. (1930) The fossil fauna of the Samana Range and some neighbouring areas. Part 8. The mollusc of the Hangu Shales. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 15, 129 - 222." type="journal article" year="1930">Cox (1930)</bibRefCitation>
attributed these two species to the genus
<taxonomicName id="4C094D750E4EFFB5FEE0FAB7FE99FADB" box="[264,323,1369,1392]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E4EFFB5FEE0FAB7FE99FADB" box="[264,323,1369,1392]" italics="true" pageId="7" pageNumber="108">Lyria</emphasis>
</taxonomicName>
and compared them to three Eocene species from the Paris Basin:
<taxonomicName id="4C094D750E4EFFB5FBD8FAB7FF00FA3F" authority="Lamarck, 1803" authorityName="Lamarck" authorityYear="1803" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="harpula" subGenus="Lyria">
<emphasis id="B97DEAE40E4EFFB5FBD8FAB7FAFBFADB" box="[1072,1313,1367,1392]" italics="true" pageId="7" pageNumber="108">Lyria (Lyria) harpula</emphasis>
(
<bibRefCitation id="EF984B070E4EFFB5FAD8FAB9FF08FA3F" author="Lamarck" pageId="7" pageNumber="108" refString="Lamarck, J. - B. (1803) Memoires sur les fossiles des environs de Paris comprenant la determination des especes qui appartiennent aux animaux marins sans vertebres, et dont la plupart sont figures dans la collection des velins du Museum. Annales du Museum d'Histoire naturelle, 1, 474 - 479." type="journal article" year="1803">Lamarck, 1803</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C094D750E4EFFB5FF00FA93FD7AFA3E" authority="d'Orbigny, 1850" authorityName="d'Orbigny" authorityYear="1850" box="[232,672,1404,1429]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="subturgidula" subGenus="Lyria">
<emphasis id="B97DEAE40E4EFFB5FF00FA93FE13FA3E" box="[232,457,1404,1429]" italics="true" pageId="7" pageNumber="108">L. (L.) subturgidula</emphasis>
(dOrbigny, 1850)
</taxonomicName>
(=
<taxonomicName id="4C094D750E4EFFB5FD24FA92FB9EFA3E" authority="Deshayes, 1835" authorityName="Deshayes" authorityYear="1835" box="[716,1092,1404,1429]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="turgidula">
<emphasis id="B97DEAE40E4EFFB5FD24FA92FC5EFA3E" box="[716,900,1404,1429]" italics="true" pageId="7" pageNumber="108">Voluta turgidula</emphasis>
Deshayes, 1835
</taxonomicName>
, not of Brocchi, 1814) and
<taxonomicName id="4C094D750E4EFFB5FA6DFA93FDF4FA13" authority="Morlet, 1888" authorityName="Morlet" authorityYear="1888" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="coroni" subGenus="Pseudolyria">
<emphasis id="B97DEAE40E4EFFB5FA6DFA93FE5AFA13" italics="true" pageId="7" pageNumber="108">L. (Pseudolyria) coroni</emphasis>
(Morlet, 1888)
</taxonomicName>
. According to these authors, strong but narrowly rounded costae and a biconic shape are shared with the Eocene
<taxonomicName id="4C094D750E4EFFB5FDF2FA2BFD8CFA77" box="[538,598,1477,1500]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E4EFFB5FDF2FA2BFD8CFA77" box="[538,598,1477,1500]" italics="true" pageId="7" pageNumber="108">Lyria</emphasis>
</taxonomicName>
species of the Paris Basin and even with many other Cenozoic species of
<taxonomicName id="4C094D750E4EFFB5FF7FFA07FF08F9AB" box="[151,210,1513,1536]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E4EFFB5FF7FFA07FF08F9AB" box="[151,210,1513,1536]" italics="true" pageId="7" pageNumber="108">Lyria</emphasis>
</taxonomicName>
. Thus,
<taxonomicName id="4C094D750E4EFFB5FEDBFA07FE40F9AB" box="[307,410,1513,1536]" class="Gastropoda" family="Volutidae" genus="Lyriopsis" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E4EFFB5FEDBFA07FE40F9AB" box="[307,410,1513,1536]" italics="true" pageId="7" pageNumber="108">Lyriopsis</emphasis>
</taxonomicName>
is placed in the tribe
<taxonomicName id="4C094D750E4EFFB5FD58FA09FD25F9AB" box="[688,767,1511,1536]" class="Gastropoda" family="Volutidae" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="tribe" tribe="Lyriini">Lyriini</taxonomicName>
, but four characters distinguish members of
<taxonomicName id="4C094D750E4EFFB5FAF4FA07FA8DF9AB" box="[1308,1367,1513,1536]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E4EFFB5FAF4FA07FA8DF9AB" box="[1308,1367,1513,1536]" italics="true" pageId="7" pageNumber="108">Lyria</emphasis>
</taxonomicName>
from
<taxonomicName id="4C094D750E4EFFB5FF7FF9E3FF24F98F" box="[151,254,1549,1572]" class="Gastropoda" family="Volutidae" genus="Lyriopsis" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E4EFFB5FF7FF9E3FF24F98F" box="[151,254,1549,1572]" italics="true" pageId="7" pageNumber="108">Lyriopsis</emphasis>
</taxonomicName>
: the morphology of the costae, the construction of the columellar folds, the shape of the whorl, and the protoconch. In
<emphasis id="B97DEAE40E4EFFB5FEA2F9DFFE60F9EC" box="[330,442,1584,1608]" italics="true" pageId="7" pageNumber="108">
L. (
<taxonomicName id="4C094D750E4EFFB5FE9CF9DEFE68F9EC" box="[372,434,1584,1607]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">Lyria</taxonomicName>
)
</emphasis>
, the posterior ends of the costae are straight, whereas they are obviously sinuous in
<taxonomicName id="4C094D750E4EFFB5FF7FF9BBFF24F9C7" box="[151,254,1621,1644]" class="Gastropoda" family="Volutidae" genus="Lyriopsis" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E4EFFB5FF7FF9BBFF24F9C7" box="[151,254,1621,1644]" italics="true" pageId="7" pageNumber="108">Lyriopsis</emphasis>
</taxonomicName>
. Regarding the subgenus
<taxonomicName id="4C094D750E4EFFB5FDC8F9BAFC9BF9C7" authority="Martin, 1931" authorityName="Martin" authorityYear="1931" box="[544,833,1620,1645]" class="Gastropoda" family="Volutidae" genus="Pseudolyria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E4EFFB5FDC8F9BAFD72F9C6" box="[544,680,1620,1645]" italics="true" pageId="7" pageNumber="108">Pseudolyria</emphasis>
<bibRefCitation id="EF984B070E4EFFB5FD46F9BAFC9BF9C7" author="Martin" box="[686,833,1620,1645]" pageId="7" pageNumber="108" refString="Martin, K. (1931) Mollusken aus dem Obereocan von Nanggulan. Dienst van den Mijnbouw in Nederlandsch-Indie, Wetenschappelijke Mededeelingen, 18, 1 - 56." type="journal article" year="1931">Martin, 1931</bibRefCitation>
</taxonomicName>
[
<typeStatus id="54B288540E4EFFB5FCA6F9BBFC5BF9C6" box="[846,897,1621,1645]" pageId="7" pageNumber="108">type</typeStatus>
species
<taxonomicName id="4C094D750E4EFFB5FC08F9BAFAA3F9C7" authority="Martin, 1931" authorityName="Martin" authorityYear="1931" box="[992,1401,1620,1645]" class="Gastropoda" family="Volutidae" genus="Pseudolyria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="species" species="ventricola">
<emphasis id="B97DEAE40E4EFFB5FC08F9BAFB3AF9C6" box="[992,1248,1620,1645]" italics="true" pageId="7" pageNumber="108">Pseudolyria ventricola</emphasis>
<bibRefCitation id="EF984B070E4EFFB5FB0FF9BAFAA3F9C7" author="Martin" box="[1255,1401,1620,1645]" pageId="7" pageNumber="108" refString="Martin, K. (1931) Mollusken aus dem Obereocan von Nanggulan. Dienst van den Mijnbouw in Nederlandsch-Indie, Wetenschappelijke Mededeelingen, 18, 1 - 56." type="journal article" year="1931">Martin, 1931</bibRefCitation>
</taxonomicName>
by monotypy], the costae are more sinuous than in
<emphasis id="B97DEAE40E4EFFB5FD27F997FCE5F924" box="[719,831,1656,1680]" italics="true" pageId="7" pageNumber="108">
L. (
<taxonomicName id="4C094D750E4EFFB5FD11F996FCEDF924" box="[761,823,1656,1679]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">Lyria</taxonomicName>
)
</emphasis>
, but the sinuosity is more anteriorly placed than in
<taxonomicName id="4C094D750E4EFFB5FF7FF973FF24F91F" box="[151,254,1693,1716]" class="Gastropoda" family="Volutidae" genus="Lyriopsis" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E4EFFB5FF7FF973FF24F91F" box="[151,254,1693,1716]" italics="true" pageId="7" pageNumber="108">Lyriopsis</emphasis>
</taxonomicName>
. Moreover, the outer lip of
<emphasis id="B97DEAE40E4EFFB5FDDBF973FD30F91E" box="[563,746,1692,1717]" italics="true" pageId="7" pageNumber="108">
L. (
<taxonomicName id="4C094D750E4EFFB5FDB1F972FD38F91E" box="[601,738,1692,1717]" class="Gastropoda" family="Volutidae" genus="Pseudolyria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">Pseudolyria</taxonomicName>
)
</emphasis>
is crenulated, whereas it is smooth in
<taxonomicName id="4C094D750E4EFFB5FB7DF973FB26F91F" box="[1173,1276,1693,1716]" class="Gastropoda" family="Volutidae" genus="Lyriopsis" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E4EFFB5FB7DF973FB26F91F" box="[1173,1276,1693,1716]" italics="true" pageId="7" pageNumber="108">Lyriopsis</emphasis>
</taxonomicName>
. In
<emphasis id="B97DEAE40E4EFFB5FAC1F973FA4FF918" box="[1321,1429,1692,1716]" italics="true" pageId="7" pageNumber="108">
L. (
<taxonomicName id="4C094D750E4EFFB5FAA6F972FA56F918" box="[1358,1420,1692,1715]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">Lyria</taxonomicName>
)
</emphasis>
, the inner lip of most species displays two or three strong basal folds and several weaker folds adapically, whereas only four strong folds have been observed in the species referred to
<taxonomicName id="4C094D750E4EFFB5FC47F90BFBCCF957" box="[943,1046,1765,1788]" class="Gastropoda" family="Volutidae" genus="Lyriopsis" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E4EFFB5FC47F90BFBCCF957" box="[943,1046,1765,1788]" italics="true" pageId="7" pageNumber="108">Lyriopsis</emphasis>
</taxonomicName>
. The shape of the whorl is rather depressed, slightly convex and terraced in
<taxonomicName id="4C094D750E4EFFB5FD67F8E7FD2CF88B" box="[655,758,1801,1824]" class="Gastropoda" family="Volutidae" genus="Lyriopsis" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E4EFFB5FD67F8E7FD2CF88B" box="[655,758,1801,1824]" italics="true" pageId="7" pageNumber="108">Lyriopsis</emphasis>
</taxonomicName>
species, whereas the whorls are strongly convex in most species of
<taxonomicName id="4C094D750E4EFFB5FEE5F8C3FE44F8E8" box="[269,414,1836,1860]" class="Gastropoda" family="Volutidae" genus="Lyria" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus">
<emphasis id="B97DEAE40E4EFFB5FEE5F8C3FE44F8E8" box="[269,414,1836,1860]" italics="true" pageId="7" pageNumber="108">Lyria (Lyria)</emphasis>
</taxonomicName>
. For these reasons, the placement of
<taxonomicName id="4C094D750E4EFFB5FCD6F8C3FC7FF8EF" box="[830,933,1837,1860]" class="Gastropoda" family="Volutidae" genus="Lyriopsis" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B97DEAE40E4EFFB5FCD6F8C3FC7FF8EF" box="[830,933,1837,1860]" italics="true" pageId="7" pageNumber="108">Lyriopsis</emphasis>
</taxonomicName>
in the
<taxonomicName id="4C094D750E4EFFB5FC1AF8C2FBE4F8EE" box="[1010,1086,1836,1861]" class="Gastropoda" family="Volutidae" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="108" phylum="Mollusca" rank="tribe" tribe="Lyriini">Lyriini</taxonomicName>
is provisional and uncertain.
</paragraph>
</subSubSection>
</treatment>
</document>