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<document id="BEBB651D09D0EDA7F303C79A2D346A01" ID-DOI="10.11646/zootaxa.4038.1.1" ID-GBIF-Dataset="066e84f1-206b-40cb-a615-2126696b813d" ID-ISSN="1175-5326" ID-Zenodo-Dep="289876" ID-ZooBank="1B62B78C-AA59-4417-A4FC-1CC9CED745E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461316934850" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Bordoni, Arnaldo" docDate="2015" docId="039D87F5552DFFD8FF4AFB18FCE6BD81" docLanguage="en" docName="zt04038p094.pdf" docOrigin="Zootaxa 4038 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Thyreocephalus pilosus Roth 1851" docType="treatment" docVersion="7" lastPageNumber="26" masterDocId="FFA4FF8D553BFFC1FFDDFFA6FFFFBE25" masterDocTitle="Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="23" updateTime="1698618126966" updateUser="plazi">
<mods:mods id="6FB8271EA9B1424DEA094C96D5273C9E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="800753392B6BC72F60013EBCF990ECD4">
<mods:title id="0B2361DFB7C127C5F3FAFE97B8D056E6">Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name id="A162F05D53410A57B925A249327B434D" type="personal">
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</mods:role>
<mods:namePart id="C6825A0416ED57B4E1A455BDB7D4287F">Janák, Jiří</mods:namePart>
</mods:name>
<mods:name id="8F426D105EC454A03FCA22DFA4DC92B8" type="personal">
<mods:role id="5D0C05C21940CB7791252908781176E3">
<mods:roleTerm id="273D80B043A2A90744F00A4D45108B8F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6C12C334E0F70063B211484377FD0614">Bordoni, Arnaldo</mods:namePart>
</mods:name>
<mods:typeOfResource id="373A0F0C5F118DCC6C5CC9819121A3D7">text</mods:typeOfResource>
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<mods:title id="650F018FFBAAD2882A64EEBC93A0315A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="348DE79A00094590E3E685CD4763B3BA">
<mods:date id="9207056BD66B4BBFA5F0977EEC50BE20">2015</mods:date>
<mods:detail id="FD0009572C4BF5021993754DABDA06B4" type="volume">
<mods:number id="C6716BEEABDE9424FCAE890B98FFC4B4">4038</mods:number>
</mods:detail>
<mods:detail id="EC5267975B2B8E031518F919DD29B201" type="issue">
<mods:number id="A07E2324D9DABBE9570D4D88C6072E7A">1</mods:number>
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<mods:start id="30190CBDC1B111372FB82B6B8023BD0C">1</mods:start>
<mods:end id="67D79A97B118908D0F5701BBC47CF2F0">94</mods:end>
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<mods:classification id="CBF03778C49F9D45E84F3626BD845AF2">journal article</mods:classification>
<mods:identifier id="31363D7E15E2D2A4D095C4415767497F" type="DOI">10.11646/zootaxa.4038.1.1</mods:identifier>
<mods:identifier id="88FC7BE269BF4D6D33CA146C15B654A5" type="GBIF-Dataset">066e84f1-206b-40cb-a615-2126696b813d</mods:identifier>
<mods:identifier id="AC129B07680183D8A6C10F47F806527B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D9771D8D3A9ED1492B9B2D23856112D2" type="Zenodo-Dep">289876</mods:identifier>
<mods:identifier id="245521D9B14C0CF3F2CA38DEECD82C82" type="ZooBank">1B62B78C-AA59-4417-A4FC-1CC9CED745E0</mods:identifier>
</mods:mods>
<treatment id="039D87F5552DFFD8FF4AFB18FCE6BD81" ID-DOI="http://doi.org/10.5281/zenodo.6101621" ID-GBIF-Taxon="127693945" ID-Zenodo-Dep="6101621" LSID="urn:lsid:plazi:treatment:039D87F5552DFFD8FF4AFB18FCE6BD81" httpUri="http://treatment.plazi.org/id/039D87F5552DFFD8FF4AFB18FCE6BD81" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23">
<subSubSection id="C32E6568552DFFD7FF4AFB18FEA7BADF" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8B8B36E3552DFFD7FF4AFB18FDA9BAFD" blockId="22.[151,598,1214,1274]" box="[151,598,1214,1240]" pageId="22" pageNumber="23">
<heading id="D0C3818F552DFFD7FF4AFB18FDA9BAFD" bold="true" box="[151,598,1214,1240]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<taxonomicName id="4C344D60552DFFD7FF4AFB18FDA9BAFD" ID-CoL="56RY3" authority="Roth, 1851" authorityName="Roth" authorityYear="1851" box="[151,598,1214,1240]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B940EAF1552DFFD7FF4AFB18FDA9BAFD" bold="true" box="[151,598,1214,1240]" pageId="22" pageNumber="23">
<emphasis id="B940EAF1552DFFD7FF4AFB18FE4ABAFD" bold="true" box="[151,437,1214,1240]" italics="true" pageId="22" pageNumber="23">Thyreocephalus pilosus</emphasis>
(
<bibRefCitation id="EFA54B12552DFFD7FE19FB18FDB1BAFD" author="Roth" box="[452,590,1214,1240]" pageId="22" pageNumber="23" refString="Roth, J. R. (1851) Diagnosen neuer Coleoptera aus Abyssinien. Archiv fur Naturgeschichte, 17, 115 - 133." type="journal article" year="1851">Roth, 1851</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8B36E3552DFFD7FF4AFB47FEA7BADF" blockId="22.[151,598,1214,1274]" box="[151,344,1249,1274]" pageId="22" pageNumber="23">
(
<figureCitation id="130F2A66552DFFD7FF42FB47FF03BADF" box="[159,252,1249,1274]" captionStart="FIGURE 29" captionStartId="16.[151,250,1593,1615]" captionTargetBox="[162,1433,201,1578]" captionTargetId="figure@16.[151,1436,193,1581]" captionTargetPageId="16" captionText="FIGURE 29. Distribution of species in Thyreocephalus pilosus group (● = T. africanus (Bernhauer), Δ = T. brunneiventris (Tottenham), ▲ = T. pilosus (Roth), ☆ T. pseudoafricanus sp. nov., ○ = T. secretus (Bernhauer), ★ = T. tsingidianus sp. nov.)." httpUri="https://zenodo.org/record/289884/files/figure.png" pageId="22" pageNumber="23">Figs. 29</figureCitation>
,
<figureCitation id="130F2A66552DFFD7FED5FB47FEAFBADF" box="[264,336,1249,1274]" captionStart="FIGURES 45 – 50" captionStartId="24.[151,264,1942,1964]" captionTargetBox="[237,1348,208,1910]" captionTargetId="figure@24.[222,1365,193,1921]" captionTargetPageId="24" captionText="FIGURES 45 – 50. Thyreocephalus pilosus (Roth); 45, head and pronotum; 46, labrum; 47, male tergite 10; 48, male sternite 9; 49, aedeagus; 50, paramere." httpUri="https://zenodo.org/record/289888/files/figure.png" pageId="22" pageNumber="23">45–50</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32E6568552DFFD7FF4AFA8FFDA1BBBE" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="8B8B36E3552DFFD7FF4AFA8FFDF6BB1A" blockId="22.[151,1436,1321,1435]" box="[151,521,1321,1343]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB2411CD552DFFD7FF4AFA8FFDF6BB1A" box="[151,521,1321,1343]" pageId="22" pageNumber="23">
<taxonomicName id="4C344D60552DFFD7FF4AFA8FFDFABB1A" ID-CoL="5C68T" authority="Roth 1851: 118" authorityName="Roth" authorityPageNumber="118" authorityYear="1851" box="[151,517,1321,1343]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B940EAF1552DFFD7FF4AFA8FFEA0BB1A" box="[151,351,1321,1343]" italics="true" pageId="22" pageNumber="23">Xantholinus pilosus</emphasis>
<treatmentCitation id="0A9510F2552DFFD7FEB8FA8FFDFABB1A" author="Roth" box="[357,517,1321,1343]" page="118" pageId="22" pageNumber="23" year="1851">
<bibRefCitation id="EFA54B12552DFFD7FEB8FA8FFE2CBB1A" author="Roth" box="[357,467,1321,1343]" pageId="22" pageNumber="23" refString="Roth, J. R. (1851) Diagnosen neuer Coleoptera aus Abyssinien. Archiv fur Naturgeschichte, 17, 115 - 133." type="journal article" year="1851">Roth 1851</bibRefCitation>
: 118
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8B36E3552DFFD7FF4AFAEEFE67BB59" blockId="22.[151,1436,1321,1435]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB2411CD552DFFD7FF4AFAEEFE67BB59" pageId="22" pageNumber="23">
<emphasis id="B940EAF1552DFFD7FF4AFAEEFEBDBB7B" box="[151,322,1352,1374]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="4C344D60552DFFD7FF4AFAEEFEC3BB7B" box="[151,316,1352,1374]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pilosus">Eulissus pilosus</taxonomicName>
;
</emphasis>
<treatmentCitation id="0A9510F2552DFFD7FE94FAEEFDCFBB7B" author="Eppelsheim" box="[329,560,1352,1374]" page="208" pageId="22" pageNumber="23" year="1895">
<bibRefCitation id="EFA54B12552DFFD7FE94FAEEFE02BB7B" author="Eppelsheim" box="[329,509,1352,1374]" pageId="22" pageNumber="23" refString="Eppelsheim, E. (1895) Beitrag zur Staphylinenfauna West-Afrika's. Zweites Stuck. Deutsche Entomologische Zeitschrift, 1895 (1), 113 - 141. http: // dx. doi. org / 10.1002 / mmnd. 48018950125" type="journal article" year="1895">Eppelsheim 1895</bibRefCitation>
: 208
</treatmentCitation>
;
<treatmentCitation id="0A9510F2552DFFD7FDE6FAEEFCDEBB7B" author="Bernhauer" box="[571,801,1352,1374]" page="478" pageId="22" pageNumber="23" year="1912">
<bibRefCitation id="EFA54B12552DFFD7FDE6FAEEFD12BB7B" author="Bernhauer" box="[571,749,1352,1374]" pageId="22" pageNumber="23" refString="Bernhauer, M. (1912 b) Neue Staphyliniden aus Zentral- und Deutsch-Ostafrika. In: Schubotz, H., Wissenschaftliche Ergebnisse der deutschen Zentral-Afrika-Expedition 1907 - 1908 unter Fuhrung Adolf Friedrichs, Herzog zu Mecklenburg. Zoologie III. Klinkhardt &amp; Biermann, Leipzig, pp. 469 - 486." type="book chapter" year="1912" yearSuffix="b">Bernhauer 1912b</bibRefCitation>
: 478
</treatmentCitation>
;
<treatmentCitation id="0A9510F2552DFFD7FCF6FAEEFB81BB7B" author="Bernhauer" box="[811,1150,1352,1374]" page="311" pageId="22" pageNumber="23" year="1914">
<bibRefCitation id="EFA54B12552DFFD7FCF6FAEEFBB2BB7B" author="Bernhauer" box="[811,1101,1352,1374]" pageId="22" pageNumber="23" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Junk, W. &amp; Schenkling, S. (Eds), Coleopterorum Catalogus, 5 (57). Junk, Berlin, pp. 289 - 408." type="journal article" year="1914">Bernhauer &amp; Schubert 1914</bibRefCitation>
: 311
</treatmentCitation>
;
<treatmentCitation id="0A9510F2552DFFD7FB54FAEEFA9ABB7B" author="Tottenham" box="[1161,1381,1352,1374]" page="224" pageId="22" pageNumber="23" year="1956">
<bibRefCitation id="EFA54B12552DFFD7FB54FAEEFACDBB7B" author="Tottenham" box="[1161,1330,1352,1374]" pageId="22" pageNumber="23" refString="Tottenham, C. E. (1956) Contributions a l'etude de la faune entomologique du Ruanda-Urundi (Mission P. Basilewsky 1953). LXXXVII. Coleoptera: Staphylinidae: Steninae, Xantholinae, Staphylinidae, Tachyporinae and Pygosteninae). Annales du Musee Royal du Congo Belge, Tervuren (Ser. in 8 o), Sciences Zoologiques, 51, 221 - 332." type="journal article" year="1956">Tottenham 1956</bibRefCitation>
: 224
</treatmentCitation>
, 255;
<treatmentCitation id="0A9510F2552DFFD7FF1AFAC1FE6ABB59" author="Herman" box="[199,405,1382,1404]" page="3628" pageId="22" pageNumber="23" year="2001">
<bibRefCitation id="EFA54B12552DFFD7FF1AFAC1FEAABB59" author="Herman" box="[199,341,1382,1404]" pageId="22" pageNumber="23" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">Herman 2001</bibRefCitation>
: 3628
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8B36E3552DFFD7FF4AFA23FDA1BBBE" blockId="22.[151,1436,1321,1435]" box="[151,606,1413,1435]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB2411CD552DFFD7FF4AFA23FDA1BBBE" box="[151,606,1413,1435]" pageId="22" pageNumber="23">
<emphasis id="B940EAF1552DFFD7FF4AFA23FE71BBBE" box="[151,398,1413,1435]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="4C344D60552DFFD7FF4AFA23FE77BBBE" ID-CoL="56RY3" box="[151,392,1413,1435]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pilosus">Thyreocephalus pilosus</taxonomicName>
;
</emphasis>
<treatmentCitation id="0A9510F2552DFFD7FE49FA23FDA6BBBE" author="Cameron" box="[404,601,1413,1435]" page="33" pageId="22" pageNumber="23" year="1950">
<bibRefCitation id="EFA54B12552DFFD7FE49FA23FDCCBBBE" author="Cameron" box="[404,563,1413,1435]" pageId="22" pageNumber="23" refString="Cameron, M. (1950) Staphylinidae (Coleoptera Polyphaga). Explorations du Parc National Albert. Mission G. F. de Witte (1933 - 1935), 59, 1 - 85." type="journal article" year="1950">
<collectingCountry id="F3237673552DFFD7FE49FA23FE0CBBBE" box="[404,499,1413,1435]" name="Cameroon" pageId="22" pageNumber="23">Cameron</collectingCountry>
, 1950
</bibRefCitation>
: 33
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32E6568552DFFD8FF4AFA64FBCFBEF0" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8B8B36E3552DFFD7FF4AFA64FE5CBBFE" blockId="22.[151,1437,1474,2034]" box="[151,419,1474,1499]" pageId="22" pageNumber="23">
<emphasis id="B940EAF1552DFFD7FF4AFA64FECABBFE" bold="true" box="[151,309,1474,1499]" pageId="22" pageNumber="23">
<typeStatus id="548F8841552DFFD7FF4AFA64FF2DBBFE" box="[151,210,1474,1499]" pageId="22" pageNumber="23">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3237673552DFFD7FEE1FA64FE60BBFE" box="[316,415,1474,1499]" name="Ethiopia" pageId="22" pageNumber="23">Ethiopia</collectingCountry>
.
</paragraph>
<paragraph id="8B8B36E3552DFFD7FF1AFA40FC34B84E" blockId="22.[151,1437,1474,2034]" pageId="22" pageNumber="23">
<emphasis id="B940EAF1552DFFD7FF1AFA40FD7FBBDA" bold="true" box="[199,640,1510,1535]" pageId="22" pageNumber="23">
<typeStatus id="548F8841552DFFD7FF1AFA40FEFDBBDA" box="[199,258,1510,1535]" pageId="22" pageNumber="23">Type</typeStatus>
material examined.
<typeStatus id="548F8841552DFFD7FDD7FA40FD7FBBDA" box="[522,640,1510,1535]" pageId="22" pageNumber="23" type="lectotype">Lectotype</typeStatus>
</emphasis>
(by present designation) ♂:
<collectingCountry id="F3237673552DFFD7FC3CFA40FB92BBDA" box="[993,1133,1510,1535]" name="Ethiopia" pageId="22" pageNumber="23">
<emphasis id="B940EAF1552DFFD7FC3CFA40FB92BBDA" bold="true" box="[993,1133,1510,1535]" pageId="22" pageNumber="23">ETHIOPIA</emphasis>
</collectingCountry>
: “
<collectingCountry id="F3237673552DFFD7FB50FA40FAFBBBDA" box="[1165,1284,1510,1535]" name="Ethiopia" pageId="22" pageNumber="23">Abyssinia</collectingCountry>
”, “
<typeStatus id="548F8841552DFFD7FAF7FA41FA9BBBDA" box="[1322,1380,1511,1535]" pageId="22" pageNumber="23">Type</typeStatus>
von
<taxonomicName id="4C344D60552DFFD7FF4AF9ACFE3DB806" authority="Roth" authorityName="Roth" box="[151,450,1546,1571]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B940EAF1552DFFD7FF4AF9ACFE85B806" box="[151,378,1546,1571]" italics="true" pageId="22" pageNumber="23">Xantholinus pilosus</emphasis>
Roth
</taxonomicName>
”, “
<taxonomicName id="4C344D60552DFFD7FE37F9ACFCAEB806" authority="Roth" authorityName="Roth" box="[490,849,1546,1571]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B940EAF1552DFFD7FE37F9ACFD07B806" box="[490,760,1546,1571]" italics="true" pageId="22" pageNumber="23">Thyreocephalus pilosus</emphasis>
(Roth)
</taxonomicName>
, J. Janák det. 1992”. The specimen is chosen as
<typeStatus id="548F8841552DFFD7FF4AF988FEFDB862" box="[151,258,1582,1607]" pageId="22" pageNumber="23" type="lectotype">lectotype</typeStatus>
. The labels “
<typeStatus id="548F8841552DFFD7FE47F989FDE6B862" box="[410,537,1583,1607]" pageId="22" pageNumber="23" type="lectotype">Lectotypus</typeStatus>
<taxonomicName id="4C344D60552DFFD7FDFEF988FBE3B862" authority="Roth, Janak &amp; Bordoni" authorityName="Roth, Janak &amp; Bordoni" box="[547,1052,1582,1607]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B940EAF1552DFFD7FDFEF988FCFCB862" box="[547,771,1582,1607]" italics="true" pageId="22" pageNumber="23">Xantholinus pilosus</emphasis>
Roth, Janák &amp; Bordoni
</taxonomicName>
des. 2014” and “
<taxonomicName id="4C344D60552DFFD7FB37F988FEC6B84E" authority="Roth" authorityName="Roth" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B940EAF1552DFFD7FB37F988FF19B84E" italics="true" pageId="22" pageNumber="23">Thyreocephalus pilosus</emphasis>
(Roth)
</taxonomicName>
, Janák &amp; Bordoni det. 2014” were added to this specimen.
</paragraph>
<paragraph id="8B8B36E3552DFFD7FF1AF9D0FEA0B9D7" blockId="22.[151,1437,1474,2034]" pageId="22" pageNumber="23">
<emphasis id="B940EAF1552DFFD7FF1AF9D0FD1AB8AA" bold="true" box="[199,741,1654,1679]" pageId="22" pageNumber="23">
Additional material examined.
<collectingCountry id="F3237673552DFFD7FDE1F9D0FD1AB8AA" box="[572,741,1654,1679]" name="Cameroon" pageId="22" pageNumber="23">CAMEROON</collectingCountry>
</emphasis>
: 1 ♀: Joko, Colin (
<collectionCode id="ED25AE26552DFFD7FC63F9D1FC00B8AA" box="[958,1023,1655,1679]" pageId="22" pageNumber="23">JJRC</collectionCode>
).
<collectingCountry id="F3237673552DFFD7FBCFF9D0FEFEB896" name="Democratic Republic of the Congo" pageId="22" pageNumber="23">
<emphasis id="B940EAF1552DFFD7FBCFF9D0FEFEB896" bold="true" pageId="22" pageNumber="23">DEMOCRATIC REPUBLIC OF CONGO</emphasis>
</collectingCountry>
: 3 spec.:
<collectingCountry id="F3237673552DFFD7FEB7F93CFE57B896" box="[362,424,1690,1715]" name="Democratic Republic of the Congo" pageId="22" pageNumber="23">Zaire</collectingCountry>
, Umg. Lwiro,
<date id="FF8A1023552DFFD7FD94F93CFD5DB896" box="[585,674,1690,1715]" pageId="22" pageNumber="23" value="1985-03">iii.1985</date>
, W. Banyw'ne (
<collectionCode id="ED25AE26552DFFD7FC92F93CFC60B897" LSID="urn:lsid:biocol.org:col:34840" box="[847,927,1690,1714]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="22" pageNumber="23">SMNS</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FC77F93CFC12B897" box="[938,1005,1690,1714]" pageId="22" pageNumber="23">ABFI</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FC2AF93CFBD5B897" LSID="urn:lsid:biocol.org:col:33211" box="[1015,1066,1690,1714]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="22" pageNumber="23">DEI</collectionCode>
); 1 spec.: Kivu, Masisi (
<collectionCode id="ED25AE26552DFFD7FAEAF93CFA70B897" box="[1335,1423,1690,1714]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="22" pageNumber="23">IRSNB</collectionCode>
); 1 ♀: Lulua, Kapanga,
<date id="FF8A1023552DFFD7FE45F91BFE1EB8F3" box="[408,481,1725,1750]" pageId="22" pageNumber="23" value="1933-01">i.1933</date>
, G. F. Overlaet (
<collectionCode id="ED25AE26552DFFD7FD43F918FD08B8F3" box="[670,759,1726,1750]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="22" pageNumber="23">MRAC</collectionCode>
).
<emphasis id="B940EAF1552DFFD7FCD6F91BFC6BB8F3" bold="true" box="[779,916,1725,1750]" pageId="22" pageNumber="23">
<collectingCountry id="F3237673552DFFD7FCD6F91BFC70B8F3" box="[779,911,1725,1750]" name="Eritrea" pageId="22" pageNumber="23">ERITREA</collectingCountry>
:
</emphasis>
4 spec.: Adua, xi (
<collectionCode id="ED25AE26552DFFD7FBADF918FB31B8F3" box="[1136,1230,1726,1750]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="23">NHMW</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FB06F918FAD0B8F3" box="[1243,1327,1726,1750]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="22" pageNumber="23">MFNB</collectionCode>
); 1 spec.: Adua, vi (
<collectionCode id="ED25AE26552DFFD7FED5F947FEA1B8DC" box="[264,350,1761,1785]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="22" pageNumber="23">MFNB</collectionCode>
); 16 spec.: Adua (
<collectionCode id="ED25AE26552DFFD7FDF6F947FD7FB8DC" LSID="urn:lsid:biocol.org:col:34795" box="[555,640,1761,1785]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="22" pageNumber="23">FMNH</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FD51F944FD21B8DC" box="[652,734,1762,1785]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="22" pageNumber="23">MFNB</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FD36F944FCB8B8DC" box="[747,839,1762,1785]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="23">NHMW</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FC8EF944FC5BB8DC" box="[851,932,1762,1785]" pageId="22" pageNumber="23">NMHP</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FC6DF947FC0CB8DC" box="[944,1011,1761,1785]" pageId="22" pageNumber="23">ABFI</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FC23F947FBC1B8DC" box="[1022,1086,1761,1785]" pageId="22" pageNumber="23">JJRC</collectionCode>
); 3 spec.:
<collectingCountry id="F3237673552DFFD7FB6CF947FAEEB8DF" box="[1201,1297,1761,1786]" name="Eritrea" pageId="22" pageNumber="23">Erythrée</collectingCountry>
(
<collectionCode id="ED25AE26552DFFD7FAFCF947FA86B8DC" box="[1313,1401,1761,1785]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="22" pageNumber="23">IRSNB</collectionCode>
); 1 spec.: Asmara (
<collectionCode id="ED25AE26552DFFD7FE8DF8A3FE59B938" box="[336,422,1797,1821]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="22" pageNumber="23">MFNB</collectionCode>
).
<collectingCountry id="F3237673552DFFD7FE60F8A2FDB6B938" box="[445,585,1796,1821]" name="Ethiopia" pageId="22" pageNumber="23">
<emphasis id="B940EAF1552DFFD7FE60F8A2FDB6B938" bold="true" box="[445,585,1796,1821]" pageId="22" pageNumber="23">ETHIOPIA</emphasis>
</collectingCountry>
: 16 spec.: Abyssinien (
<collectionCode id="ED25AE26552DFFD7FCBAF8A3FC44B938" box="[871,955,1797,1821]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="22" pageNumber="23">MFNB</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FC16F8A3FC03B939" LSID="urn:lsid:biocol.org:col:33211" box="[971,1020,1797,1820]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="22" pageNumber="23">DEI</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FBD3F8A3FBAEB938" box="[1038,1105,1797,1821]" pageId="22" pageNumber="23">ABFI</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FBBCF8A3FB49B938" box="[1121,1206,1797,1821]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="22" pageNumber="23">IRSNB</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FB1AF8A3FAD7B938" box="[1223,1320,1797,1821]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="23">NHMW</collectionCode>
); 1 spec.:
<collectingCountry id="F3237673552DFFD7FF4AF881FEF5B965" box="[151,266,1831,1856]" name="Ethiopia" pageId="22" pageNumber="23">Abyssinia</collectingCountry>
, Dr. Hofmann (
<collectionCode id="ED25AE26552DFFD7FE15F88EFDD4B965" box="[456,555,1832,1856]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="23">NHMW</collectionCode>
); 1 spec.: Lac Daka,
<date id="FF8A1023552DFFD7FCEDF881FC54B965" box="[816,939,1831,1856]" pageId="22" pageNumber="23" value="1914-03-14">14.iii.1914</date>
(
<collectionCode id="ED25AE26552DFFD7FC62F88EFBFFB965" box="[959,1024,1832,1856]" pageId="22" pageNumber="23">JJRC</collectionCode>
); 1 spec.:
<collectingCountry id="F3237673552DFFD7FB5DF881FB0FB965" box="[1152,1264,1831,1856]" name="Ethiopia" pageId="22" pageNumber="23">Abyssinie</collectingCountry>
cent.,
<collectingCountry id="F3237673552DFFD7FA9AF88EFA63B91A" box="[1351,1436,1832,1855]" name="Morocco" pageId="22" pageNumber="23">Maraco</collectingCountry>
(
<collectionCode id="ED25AE26552DFFD7FF42F8EAFF1FB941" box="[159,224,1868,1892]" pageId="22" pageNumber="23">JJRC</collectionCode>
); 1 spec.: Arussi Galla, Ganale Gudda,
<date id="FF8A1023552DFFD7FD4BF8EDFCF7B941" box="[662,776,1867,1892]" pageId="22" pageNumber="23" value="1893-03" valueMax="1893-05-31" valueMin="1893-03-00">iii–v.1893</date>
,V. Bottego (
<collectionCode id="ED25AE26552DFFD7FC4EF8EAFC02B941" box="[915,1021,1868,1892]" pageId="22" pageNumber="23">MCSNG</collectionCode>
); 1 spec.: Illubabor Prov., Umgebung Bedele,
<date id="FF8A1023552DFFD7FF2EF8C8FEB7B9A2" box="[243,328,1902,1927]" pageId="22" pageNumber="23" value="1994-04">iv.1994</date>
, Werner (
<collectionCode id="ED25AE26552DFFD7FE6BF8C9FE15B9A2" LSID="urn:lsid:biocol.org:col:33211" box="[438,490,1903,1927]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="22" pageNumber="23">DEI</collectionCode>
); 1 spec.: Grambela, Illubabor,
<date id="FF8A1023552DFFD7FC91F8C8FC51B9A2" box="[844,942,1902,1927]" pageId="22" pageNumber="23" value="2001-05-05">5.v.2001</date>
, Werner &amp; Sudre (
<collectionCode id="ED25AE26552DFFD7FB5FF8C9FB49B9A2" LSID="urn:lsid:biocol.org:col:33211" box="[1154,1206,1903,1927]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="22" pageNumber="23">DEI</collectionCode>
);
<specimenCount id="9D32FD6A552DFFD7FB17F8C9FB0BB9A2" box="[1226,1268,1903,1927]" pageId="22" pageNumber="23" type="generic">1 ♂</specimenCount>
: Jimma, Kaffa prov.,
<date id="FF8A1023552DFFD7FF03F834FEC2B98E" box="[222,317,1938,1963]" pageId="22" pageNumber="23" value="1971-07">vii.1971</date>
, G. de Rougemont (
<collectionCode id="ED25AE26552DFFD7FDC0F835FD8EB98E" box="[541,625,1939,1963]" pageId="22" pageNumber="23">GROB</collectionCode>
).
<collectingCountry id="F3237673552DFFD7FD5EF834FD01B98E" box="[643,766,1938,1963]" name="Uganda" pageId="22" pageNumber="23">
<emphasis id="B940EAF1552DFFD7FD5EF834FD01B98E" bold="true" box="[643,766,1938,1963]" pageId="22" pageNumber="23">UGANDA</emphasis>
</collectingCountry>
: 1 spec.: without additional data (
<collectionCode id="ED25AE26552DFFD7FBA3F835FB2BB98E" box="[1150,1236,1939,1963]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="22" pageNumber="23">NMPC</collectionCode>
); 1 spec.:
<collectingCountry id="F3237673552DFFD7FA98F834FA63B98E" box="[1349,1436,1938,1963]" name="Uganda" pageId="22" pageNumber="23">Uganda</collectingCountry>
Central,
<collectingCountry id="F3237673552DFFD7FF25F813FEE1B9EB" box="[248,286,1973,1998]" name="Switzerland" pageId="22" pageNumber="23">Ch</collectingCountry>
.,
<date id="FF8A1023552DFFD7FEF2F813FE69B9EB" box="[303,406,1973,1998]" pageId="22" pageNumber="23" value="1909-01" valueMax="1909-02-31" valueMin="1909-01-00">i–ii.1909</date>
, Alluaud (
<collectionCode id="ED25AE26552DFFD7FDD0F810FD9CB9EB" box="[525,611,1974,1998]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="22" pageNumber="23">NMPC</collectionCode>
); 3 spec.: Victoria Nyanza, Arcipelago di Sesse, Bugala, 1908, E. Bayon (
<collectionCode id="ED25AE26552DFFD7FF43F87CFEFCB9D7" box="[158,259,2010,2034]" pageId="22" pageNumber="23">MCSNG</collectionCode>
,
<collectionCode id="ED25AE26552DFFD7FED3F87CFEACB9D7" box="[270,339,2010,2034]" pageId="22" pageNumber="23">ABFI</collectionCode>
).
</paragraph>
<caption id="DF4B666B552CFFD6FF4AF800FE74B9FE" httpUri="https://zenodo.org/record/289887/files/figure.png" pageId="23" pageNumber="24" targetBox="[195,1376,206,1912]" targetPageId="23">
<paragraph id="8B8B36E3552CFFD6FF4AF800FE74B9FE" blockId="23.[151,1435,1958,2011]" pageId="23" pageNumber="24">
<emphasis id="B940EAF1552CFFD6FF4AF800FEB3B999" bold="true" box="[151,332,1958,1980]" pageId="23" pageNumber="24">FIGURES 40–44</emphasis>
.
<taxonomicName id="4C344D60552CFFD6FE84F800FD58B999" box="[345,679,1958,1980]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="pseudoafricanus" status="sp. nov.">
<emphasis id="B940EAF1552CFFD6FE84F800FD58B999" box="[345,679,1958,1980]" italics="true" pageId="23" pageNumber="24">Thyreocephalus pseudoafricanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273578A552CFFD6FD70F800FD02B999" box="[685,765,1958,1980]" pageId="23" pageNumber="24" rank="species">
<emphasis id="B940EAF1552CFFD6FD70F800FD02B999" bold="true" box="[685,765,1958,1980]" pageId="23" pageNumber="24">sp. nov.</emphasis>
</taxonomicNameLabel>
; 40, head and pronotum; 41, labrum; 42, male tergite 10; 43, male sternite 9; 44, aedeagus.
</paragraph>
</caption>
<caption id="DF4B666B5523FFD9FF4AF830FE4DB9EE" httpUri="https://zenodo.org/record/289888/files/figure.png" pageId="24" pageNumber="25" targetBox="[237,1348,208,1910]" targetPageId="24">
<paragraph id="8B8B36E35523FFD9FF4AF830FE4DB9EE" blockId="24.[151,1436,1942,1995]" pageId="24" pageNumber="25">
<emphasis id="B940EAF15523FFD9FF4AF830FEB2B989" bold="true" box="[151,333,1942,1964]" pageId="24" pageNumber="25">FIGURES 45–50</emphasis>
.
<taxonomicName id="4C344D605523FFD9FE84F830FD6CB989" authority="Roth" authorityName="Roth" box="[345,659,1942,1964]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B940EAF15523FFD9FE84F830FDB7B989" box="[345,584,1942,1964]" italics="true" pageId="24" pageNumber="25">Thyreocephalus pilosus</emphasis>
(Roth)
</taxonomicName>
; 45, head and pronotum; 46, labrum; 47, male tergite 10; 48, male sternite 9; 49, aedeagus; 50, paramere.
</paragraph>
</caption>
<paragraph id="8B8B36E35522FFD8FF1AFF31FBCFBEF0" blockId="25.[151,1437,151,933]" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FF1AFF31FB9FBE95" bold="true" box="[199,1120,151,176]" pageId="25" pageNumber="26">
Cited material (not examined).
<collectingCountry id="F32376735522FFD8FD92FF31FBA4BE95" box="[591,1115,151,176]" name="Democratic Republic of the Congo" pageId="25" pageNumber="26">DEMOCRATIC REPUBLIC OF CONGO</collectingCountry>
:
</emphasis>
Parc Nat. Albert-Rutshuru (
<bibRefCitation id="EFA54B125522FFD8FF43FF1AFEAEBEF0" author="Cameron" box="[158,337,188,213]" pageId="25" pageNumber="26" refString="Cameron, M. (1950) Staphylinidae (Coleoptera Polyphaga). Explorations du Parc National Albert. Mission G. F. de Witte (1933 - 1935), 59, 1 - 85." type="journal article" year="1950">
<collectingCountry id="F32376735522FFD8FF43FF1AFEF6BEF1" box="[158,265,188,212]" name="Cameroon" pageId="25" pageNumber="26">Cameron</collectingCountry>
, 1950
</bibRefCitation>
).
<collectingCountry id="F32376735522FFD8FEB9FF1AFE1EBEF0" box="[356,481,188,213]" name="Rwanda" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FEB9FF1AFE1EBEF0" bold="true" box="[356,481,188,213]" pageId="25" pageNumber="26">RWANDA</emphasis>
</collectingCountry>
: terr. Nyanza, Gitarama,
<quantity id="4CCC9B065522FFD8FD20FF1AFCADBEF1" box="[765,850,188,213]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="25" pageNumber="26" unit="m" value="1850.0">1850 m</quantity>
(
<bibRefCitation id="EFA54B125522FFD8FCBDFF1AFBDBBEF0" author="Tottenham" box="[864,1060,188,213]" pageId="25" pageNumber="26" refString="Tottenham, C. E. (1956) Contributions a l'etude de la faune entomologique du Ruanda-Urundi (Mission P. Basilewsky 1953). LXXXVII. Coleoptera: Staphylinidae: Steninae, Xantholinae, Staphylinidae, Tachyporinae and Pygosteninae). Annales du Musee Royal du Congo Belge, Tervuren (Ser. in 8 o), Sciences Zoologiques, 51, 221 - 332." type="journal article" year="1956">Tottenham, 1956</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E65685522FFD8FF1AFF79FD03BCD5" pageId="25" pageNumber="26" type="description">
<paragraph id="8B8B36E35522FFD8FF1AFF79FA7EBFD0" blockId="25.[151,1437,151,933]" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FF1AFF79FE8BBEDD" bold="true" box="[199,372,223,248]" pageId="25" pageNumber="26">Redescription.</emphasis>
Body length
<quantity id="4CCC9B065522FFD8FDD6FF46FD78BED2" box="[523,647,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="25" pageNumber="26" unit="mm" value="16.0" valueMax="18.0" valueMin="14.0">14–18 mm</quantity>
; length from anterior margin of head to posterior margin of elytra:
<quantity id="4CCC9B065522FFD8FAAFFF46FF39BF39" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="25" pageNumber="26" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
. Head and pronotum black, elytra yellowish, with scutellum and adjacent area infuscate, almost black; metasternum black, abdomen yellowish with tergites 3 and 4 almost completely brown black; first visible sternite at least partly black; antennae brown dark; legs brown black. Shiny, very robust and large. Head and pronotum with very fine micro-punctation. Head and pronotum and related punctation as in
<figureCitation id="130F2A665522FFD8FBDCFEC9FBAABFAD" box="[1025,1109,367,392]" captionStart="FIGURES 45 – 50" captionStartId="24.[151,264,1942,1964]" captionTargetBox="[237,1348,208,1910]" captionTargetId="figure@24.[222,1365,193,1921]" captionTargetPageId="24" captionText="FIGURES 45 – 50. Thyreocephalus pilosus (Roth); 45, head and pronotum; 46, labrum; 47, male tergite 10; 48, male sternite 9; 49, aedeagus; 50, paramere." httpUri="https://zenodo.org/record/289888/files/figure.png" pageId="25" pageNumber="26">Fig. 45</figureCitation>
; labrum as in
<figureCitation id="130F2A665522FFD8FB25FEC9FAB4BFAD" box="[1272,1355,367,392]" captionStart="FIGURES 45 – 50" captionStartId="24.[151,264,1942,1964]" captionTargetBox="[237,1348,208,1910]" captionTargetId="figure@24.[222,1365,193,1921]" captionTargetPageId="24" captionText="FIGURES 45 – 50. Thyreocephalus pilosus (Roth); 45, head and pronotum; 46, labrum; 47, male tergite 10; 48, male sternite 9; 49, aedeagus; 50, paramere." httpUri="https://zenodo.org/record/289888/files/figure.png" pageId="25" pageNumber="26">Fig. 46</figureCitation>
. Elytra sub-quadrate, as long and wide as pronotum, with slightly rounded sides and less marked humeral angles. Surface with very sparse punctation, arranged in three series, one near suture, one median and one lateral. Abdomen with transverse micro-striation and very fine and sparse punctation, arranged in some series, more dense on the sides.
</paragraph>
<paragraph id="8B8B36E35522FFD8FF1AFE59FBA2BC45" blockId="25.[151,1437,151,933]" pageId="25" pageNumber="26">
<collectingCountry id="F32376735522FFD8FF1AFE59FEFBBC3D" box="[199,260,511,536]" name="Malta" pageId="25" pageNumber="26">Male</collectingCountry>
. Lateral parts of temples with series of moderately coarse punctures. Tergite 10 and sternite 9 of male genital segment as in
<figureCitation id="130F2A665522FFD8FE41FD82FDD3BC19" box="[412,556,548,573]" captionStart="FIGURES 45 – 50" captionStartId="24.[151,264,1942,1964]" captionTargetBox="[237,1348,208,1910]" captionTargetId="figure@24.[222,1365,193,1921]" captionTargetPageId="24" captionText="FIGURES 45 – 50. Thyreocephalus pilosus (Roth); 45, head and pronotum; 46, labrum; 47, male tergite 10; 48, male sternite 9; 49, aedeagus; 50, paramere." httpUri="https://zenodo.org/record/289888/files/figure.png" pageId="25" pageNumber="26">Figs. 47, 48</figureCitation>
. Aedeagus (
<figureCitation id="130F2A665522FFD8FD63FD82FCBDBC18" box="[702,834,548,573]" captionStart="FIGURES 45 – 50" captionStartId="24.[151,264,1942,1964]" captionTargetBox="[237,1348,208,1910]" captionTargetId="figure@24.[222,1365,193,1921]" captionTargetPageId="24" captionText="FIGURES 45 – 50. Thyreocephalus pilosus (Roth); 45, head and pronotum; 46, labrum; 47, male tergite 10; 48, male sternite 9; 49, aedeagus; 50, paramere." httpUri="https://zenodo.org/record/289888/files/figure.png" pageId="25" pageNumber="26">Fig. 49, 50</figureCitation>
) elongate,
<quantity id="4CCC9B065522FFD8FC1AFD82FBDDBC19" box="[967,1058,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="25" pageNumber="26" unit="mm" value="1.7">1.7 mm</quantity>
long, with dilated median lobe; parameres symmetrical; inner sac very narrow, tube-like, covered with small spinulae.
</paragraph>
<paragraph id="8B8B36E35522FFD8FF1AFDCAFBBCBCA1" blockId="25.[151,1437,151,933]" box="[199,1091,620,645]" pageId="25" pageNumber="26">Female. Lateral parts of temples impunctate or at most with few fine punctures.</paragraph>
<paragraph id="8B8B36E35522FFD8FF1AFD29FD03BCD5" blockId="25.[151,1437,151,933]" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FF1AFD29FE2EBC8D" bold="true" box="[199,465,655,680]" pageId="25" pageNumber="26">Differential diagnosis.</emphasis>
<taxonomicName id="4C344D605522FFD8FE07FD29FD1ABC8D" box="[474,741,655,680]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B940EAF15522FFD8FE07FD29FD1ABC8D" box="[474,741,655,680]" italics="true" pageId="25" pageNumber="26">Thyreocephalus pilosus</emphasis>
</taxonomicName>
differs from the most similar
<taxonomicName id="4C344D605522FFD8FB95FD37FF27BCE8" authority="Tottenham, 1956" authorityName="Tottenham" authorityYear="1956" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="brunneiventris">
<emphasis id="B940EAF15522FFD8FB95FD37FAF5BC8D" box="[1096,1290,655,680]" italics="true" pageId="25" pageNumber="26">T. brunneiventris</emphasis>
(
<bibRefCitation id="EFA54B125522FFD8FAC1FD29FF2FBCE8" author="Tottenham" pageId="25" pageNumber="26" refString="Tottenham, C. E. (1956) Contributions a l'etude de la faune entomologique du Ruanda-Urundi (Mission P. Basilewsky 1953). LXXXVII. Coleoptera: Staphylinidae: Steninae, Xantholinae, Staphylinidae, Tachyporinae and Pygosteninae). Annales du Musee Royal du Congo Belge, Tervuren (Ser. in 8 o), Sciences Zoologiques, 51, 221 - 332." type="journal article" year="1956">Tottenham, 1956</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C344D605522FFD8FECCFD13FD9DBCE8" authority="Tottenham, 1956" authorityName="Tottenham" authorityYear="1956" box="[273,610,692,717]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="secretus">
<emphasis id="B940EAF15522FFD8FECCFD13FE76BCE9" box="[273,393,693,716]" italics="true" pageId="25" pageNumber="26">T. secretus</emphasis>
(
<bibRefCitation id="EFA54B125522FFD8FE45FD12FDA5BCE8" author="Tottenham" box="[408,602,692,717]" pageId="25" pageNumber="26" refString="Tottenham, C. E. (1956) Contributions a l'etude de la faune entomologique du Ruanda-Urundi (Mission P. Basilewsky 1953). LXXXVII. Coleoptera: Staphylinidae: Steninae, Xantholinae, Staphylinidae, Tachyporinae and Pygosteninae). Annales du Musee Royal du Congo Belge, Tervuren (Ser. in 8 o), Sciences Zoologiques, 51, 221 - 332." type="journal article" year="1956">Tottenham, 1956</bibRefCitation>
)
</taxonomicName>
in the abdomen with at least partly black first visible sternite and by the apically broadly rounded median lobe of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C32E65685522FFD8FF1AFD5AFB6EBD1D" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8B8B36E35522FFD8FF1AFD5AFB6EBD1D" blockId="25.[151,1437,151,933]" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FF1AFD5AFE9FBD30" bold="true" box="[199,352,764,789]" pageId="25" pageNumber="26">Distribution.</emphasis>
The species is distributed in
<collectingCountry id="F32376735522FFD8FD69FD5AFCD3BD31" box="[692,812,764,788]" name="Cameroon" pageId="25" pageNumber="26">Cameroon</collectingCountry>
,
<collectingCountry id="F32376735522FFD8FCE4FD5AFB5EBD31" box="[825,1185,764,789]" name="Democratic Republic of the Congo" pageId="25" pageNumber="26">Democratic Republic of Congo</collectingCountry>
,
<collectingCountry id="F32376735522FFD8FB72FD5AFB01BD30" box="[1199,1278,764,789]" name="Eritrea" pageId="25" pageNumber="26">Eritrea</collectingCountry>
,
<collectingCountry id="F32376735522FFD8FAD6FD5AFA95BD30" box="[1291,1386,764,789]" name="Ethiopia" pageId="25" pageNumber="26">Ethiopia</collectingCountry>
and
<collectingCountry id="F32376735522FFD8FF4AFCB9FF11BD1D" box="[151,238,799,824]" name="Uganda" pageId="25" pageNumber="26">Uganda</collectingCountry>
(
<figureCitation id="130F2A665522FFD8FF20FCB9FEADBD1D" box="[253,338,799,824]" captionStart="FIGURE 29" captionStartId="16.[151,250,1593,1615]" captionTargetBox="[162,1433,201,1578]" captionTargetId="figure@16.[151,1436,193,1581]" captionTargetPageId="16" captionText="FIGURE 29. Distribution of species in Thyreocephalus pilosus group (● = T. africanus (Bernhauer), Δ = T. brunneiventris (Tottenham), ▲ = T. pilosus (Roth), ☆ T. pseudoafricanus sp. nov., ○ = T. secretus (Bernhauer), ★ = T. tsingidianus sp. nov.)." httpUri="https://zenodo.org/record/289884/files/figure.png" pageId="25" pageNumber="26">Fig. 29</figureCitation>
). The identification of specimens cited from
<collectingCountry id="F32376735522FFD8FC95FCB9FC5BBD1D" box="[840,932,799,824]" name="Rwanda" pageId="25" pageNumber="26">Rwanda</collectingCountry>
has to be confirmed.
</paragraph>
</subSubSection>
<subSubSection id="C32E65685522FFD8FF1AFCE2FCE6BD81" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8B8B36E35522FFD8FF1AFCE2FCE6BD81" blockId="25.[151,1437,151,933]" pageId="25" pageNumber="26">
<emphasis id="B940EAF15522FFD8FF1AFCE2FEC4BD78" bold="true" box="[199,315,836,861]" pageId="25" pageNumber="26">Remarks.</emphasis>
<collectingCountry id="F32376735522FFD8FE99FCE2FE9CBD79" box="[324,355,836,860]" name="American Samoa" pageId="25" pageNumber="26">As</collectingCountry>
the superior line of pronotal hypomeron is turning downwards well before middle, joining or almost joining inferior line next to front margin of procoxae and continuing onto anterior margin of pronotum, the species was transferred from
<taxonomicName id="4C344D605522FFD8FE02FC2AFDC3BD80" box="[479,572,908,933]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF15522FFD8FE02FC2AFDC3BD80" box="[479,572,908,933]" italics="true" pageId="25" pageNumber="26">Eulissus</emphasis>
</taxonomicName>
to
<taxonomicName id="4C344D605522FFD8FDBDFC2AFCEDBD80" box="[608,786,908,933]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF15522FFD8FDBDFC2AFCEDBD80" box="[608,786,908,933]" italics="true" pageId="25" pageNumber="26">Thyreocephalus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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