291 lines
31 KiB
XML
291 lines
31 KiB
XML
<document id="BE49ED61BD781ECD9D22161F873129ED" ID-CLB-Dataset="3285" ID-DOI="10.11646/zootaxa.5072.1.1" ID-GBIF-Dataset="d74cb174-4e73-4b24-8240-071f098aa4d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5728873" ID-ZooBank="9CCBE63C-F8B9-46A7-A6EB-F7FCCC781C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637912784011" checkinUser="plazi" docAuthor="Lee, Soo Bin & Nam, Gi-Soo" docDate="2021" docId="039D87BAFFFCFFE5FF65F9FFFE8366CD" docLanguage="en" docName="zootaxa.5072.1.1.pdf" docOrigin="Zootaxa 5072 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calosoma kimi Lee & Nam 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="5" masterDocId="FFA4FFC2FFFEFFE1FFF2FFF9FFC26144" masterDocTitle="A new species of fossil Calosoma (Coleoptera: Carabidae) from the Geumgwangdong Formation (Early Miocene), South Korea" masterLastPageNumber="11" masterPageNumber="1" pageNumber="3" updateTime="1699470320573" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="528E9B9041B7C717CCACBB74AC2C4DD2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B4051C48072263725C689EF19587AC7C">
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<mods:title id="4CEC5005AC194C6690EAF885BCD2199E">A new species of fossil Calosoma (Coleoptera: Carabidae) from the Geumgwangdong Formation (Early Miocene), South Korea</mods:title>
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</mods:titleInfo>
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<mods:name id="E5B50DC8C50D128D436C860EF1787265" type="personal">
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<mods:role id="98B46C229B140E5C63CD4D97282F23E1">
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<mods:roleTerm id="0D59BDD61406EA361CEFE9FDE3E1747B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="030676F920FE5831E172CA744FE82E93">Lee, Soo Bin</mods:namePart>
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<mods:nameIdentifier id="71BC30AA184F283B8952A2A36FEC4189" type="ORCID">0000-0003-2112-3897</mods:nameIdentifier>
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<mods:affiliation id="4D23EABC303F1E8892ED865B5DC08DB5">Department of Science Education, Gongju National University of Education 27 Ungjinro, Gongjusi, Chungcheonnamdo, Souh Korea. dinos 20000 @ naver. com; https: // orcid. org / 0000 - 0003 - 2112 - 3897</mods:affiliation>
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<mods:nameIdentifier id="44AE7BDBE8143D6D8C62D11669F85726" type="email">dinos20000@naver.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="B1DEDA298D4FB11CA46A06C01829CA21" type="personal">
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<mods:role id="43E48746331E2EE3640650865C0693BB">
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<mods:roleTerm id="DE28F43C626164D99CAC159D30CBD008">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8D7622D480C90A2CA8D27540A0AA76BA">Nam, Gi-Soo</mods:namePart>
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<mods:affiliation id="6DC9233B6DE83B6B36F3145F9622FE28">Department of Science Education, Gongju National University of Education 27 Ungjinro, Gongjusi, Chungcheonnamdo, Souh Korea. dinos 20000 @ naver. com; https: // orcid. org / 0000 - 0003 - 2112 - 3897</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="D9094E0C1D47117EF97FC6B85D842413">text</mods:typeOfResource>
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<mods:titleInfo id="8699CE7D0F1884CDB246A1B746375FD0">
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<mods:title id="217BB315617AFB0599185DE1B768F2F0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="33E171FBADD72CE17120EF8D50E5F6EE">
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<mods:date id="D789AAE6F8962B8ABC69DEE5FCBCEA80">2021</mods:date>
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<mods:detail id="C97799ADB33D0D5867E45F64C5354A67" type="pubDate">
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<mods:number id="3B49B2F2CC991B266D2E1A2E69CC7F6F">2021-11-26</mods:number>
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</mods:detail>
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<mods:detail id="3079EAF8DF6FE2A3BD55CEA34ED4F170" type="volume">
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<mods:number id="181E9A11F917818A6B2B5828444CA7E7">5072</mods:number>
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</mods:detail>
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<mods:detail id="21EC4331898F50EF7E0ACD5DFC82A7EC" type="issue">
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<mods:number id="236BBA86EED313055037FA3FAF87AE99">1</mods:number>
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<mods:extent id="C0B84B2CB592E90ADE3B6BE3549DFE13" unit="page">
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<mods:start id="F243845B1993A243F6D7A1D80F10B2E4">1</mods:start>
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<mods:end id="EF4E47A188546836CD77BE5AE8347E4A">11</mods:end>
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</mods:extent>
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<mods:classification id="FB3D8B0CE8BE677C0B453BD5B473B1A4">journal article</mods:classification>
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<mods:identifier id="91E6035A32F9D7F31D3BC3730D9D3E32" type="CLB-Dataset">3285</mods:identifier>
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<mods:identifier id="F56F938373BAB21427DB3DB2A0F1ABBC" type="DOI">10.11646/zootaxa.5072.1.1</mods:identifier>
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<mods:identifier id="2E73F041AE8E72BDBB6B9B2FAB8C0FEB" type="GBIF-Dataset">d74cb174-4e73-4b24-8240-071f098aa4d1</mods:identifier>
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<mods:identifier id="25835F6B330C89B61DDCA1366437BF55" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9B29DC3BE0E772BCC1B33AB73DB6C88D" type="Zenodo-Dep">5728873</mods:identifier>
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<mods:identifier id="6A0132DA874203DE4BA86BB5612248C8" type="ZooBank">9CCBE63C-F8B9-46A7-A6EB-F7FCCC781C33</mods:identifier>
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</mods:mods>
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<treatment id="039D87BAFFFCFFE5FF65F9FFFE8366CD" ID-DOI="http://doi.org/10.5281/zenodo.5729731" ID-GBIF-Taxon="190868546" ID-Zenodo-Dep="5729731" LSID="urn:lsid:plazi:treatment:039D87BAFFFCFFE5FF65F9FFFE8366CD" httpUri="http://treatment.plazi.org/id/039D87BAFFFCFFE5FF65F9FFFE8366CD" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
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<subSubSection id="C32E6527FFFCFFE3FF65F9FFFD346764" box="[151,758,1542,1569]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8B8B36ACFFFCFFE3FF65F9FFFD346764" blockId="2.[151,758,1542,1569]" box="[151,758,1542,1569]" pageId="2" pageNumber="3">
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<heading id="D0C381C0FFFCFFE3FF65F9FFFD346764" bold="true" box="[151,758,1542,1569]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="B940EABEFFFCFFE3FF65F9FFFD346764" bold="true" box="[151,758,1542,1569]" pageId="2" pageNumber="3">
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Species
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<taxonomicName id="4C344D2FFFFCFFE3FF08F9FFFD536764" authority="Lee & Nam, 2021" authorityName="Lee & Nam" authorityYear="2021" box="[250,657,1542,1569]" class="Insecta" family="Carabidae" genus="Calosoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="kimi" status="sp. nov.">
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<emphasis id="B940EABEFFFCFFE3FF08F9FFFE726765" bold="true" box="[250,432,1542,1569]" italics="true" pageId="2" pageNumber="3">Calosoma kimi</emphasis>
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Lee & Nam, 2021
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</taxonomicName>
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<taxonomicNameLabel id="A27357C5FFFCFFE3FD6AF9FFFD346764" box="[664,758,1542,1568]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6527FFFCFFE3FF65F9B7FDEF67F4" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="8B8B36ACFFFCFFE3FF65F9B7FC8D672C" blockId="2.[151,1436,1614,1929]" box="[151,847,1614,1641]" pageId="2" pageNumber="3">
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<emphasis id="B940EABEFFFCFFE3FF65F9B7FF3D672C" bold="true" box="[151,255,1614,1640]" pageId="2" pageNumber="3">Material</emphasis>
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.
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<materialsCitation id="3B5C3CF1FFFCFFE3FEFFF9B7FDEC672C" ID-GBIF-Occurrence="3412172301" box="[269,558,1614,1640]" pageId="2" pageNumber="3" specimenCount="2" typeStatus="holotype">
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<typeStatus id="548F880EFFFCFFE3FEFFF9B7FEB7672C" box="[269,373,1614,1640]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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GNUE-311001,
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</materialsCitation>
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<materialsCitation id="3B5C3CF1FFFCFFE3FDC7F9B6FC8E672C" ID-GBIF-Occurrence="3412172302" box="[565,844,1614,1641]" pageId="2" pageNumber="3" specimenCount="2" typeStatus="paratype">
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<typeStatus id="548F880EFFFCFFE3FDC7F9B6FD55672D" box="[565,663,1615,1641]" pageId="2" pageNumber="3" type="paratype">Paratype</typeStatus>
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GNUE-311002
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8B36ACFFFCFFE3FF35F98BFB2C67C9" blockId="2.[151,1436,1614,1929]" box="[199,1262,1650,1677]" pageId="2" pageNumber="3">
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<emphasis id="B940EABEFFFCFFE3FF35F98BFEEB67C8" bold="true" box="[199,297,1650,1676]" pageId="2" pageNumber="3">Locality</emphasis>
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. Geumgwangri, Donghaemyeon, Namgu, Pohangsi, Gyeongsangbukdo,
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<collectingCountry id="F323763CFFFCFFE3FBA8F98BFB2867C9" box="[1114,1258,1650,1677]" name="South Korea" pageId="2" pageNumber="3">South Korea</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B8B36ACFFFCFFE3FF35F96FFDEF67F4" blockId="2.[151,1436,1614,1929]" box="[199,557,1686,1713]" pageId="2" pageNumber="3">
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<emphasis id="B940EABEFFFCFFE3FF35F96FFEB867F4" bold="true" box="[199,378,1686,1712]" pageId="2" pageNumber="3">Geological Age</emphasis>
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. Early Miocene
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6527FFFCFFE3FF35F943FA596791" box="[199,1435,1722,1749]" pageId="2" pageNumber="3" type="etymology">
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<paragraph id="8B8B36ACFFFCFFE3FF35F943FA596791" blockId="2.[151,1436,1614,1929]" box="[199,1435,1722,1749]" pageId="2" pageNumber="3">
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<emphasis id="B940EABEFFFCFFE3FF35F943FE8A6790" bold="true" box="[199,328,1722,1748]" pageId="2" pageNumber="3">Etymology</emphasis>
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. Honoring Professor Jong Heon Kim for his important work on fossils from Pohang,
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<collectingCountry id="F323763CFFFCFFE3FAF4F943FA556791" box="[1286,1431,1722,1749]" name="South Korea" pageId="2" pageNumber="3">South Korea</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6527FFFCFFE3FF35F927FDAA66CC" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="8B8B36ACFFFCFFE3FF35F927FDAA66CC" blockId="2.[151,1436,1614,1929]" pageId="2" pageNumber="3">
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<emphasis id="B940EABEFFFCFFE3FF35F927FEF867BC" bold="true" box="[199,314,1758,1784]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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. The 5
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<superScript id="7C419BE4FFFCFFE3FE74F927FE5167A8" attach="left" box="[390,403,1758,1772]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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interval connected to the 7
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<superScript id="7C419BE4FFFCFFE3FD4CF927FD0967A8" attach="left" box="[702,715,1758,1772]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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interval and the 6
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<superScript id="7C419BE4FFFCFFE3FC62F927FC5F67A8" attach="left" box="[912,925,1758,1772]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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interval surrounded at the apex of elytron; the 11
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<superScript id="7C419BE4FFFCFFE3FF40F8FBFF7D6654" attach="left" box="[178,191,1794,1808]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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and 13
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<superScript id="7C419BE4FFFCFFE3FEE3F8FBFEDC6654" attach="left" box="[273,286,1794,1808]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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intervals connected and surround the 12
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<superScript id="7C419BE4FFFCFFE3FD10F8FBFD2D6654" attach="left" box="[738,751,1794,1808]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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interval; the 10
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<superScript id="7C419BE4FFFCFFE3FC6CF8FBFC696654" attach="left" box="[926,939,1794,1808]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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and 14
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<superScript id="7C419BE4FFFCFFE3FC0EF8FBFBCB6654" attach="left" box="[1020,1033,1794,1808]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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intervals connected and surrounded the 11
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<superScript id="7C419BE4FFFCFFE3FF2DF8DFFF2E6670" attach="left" box="[223,236,1830,1844]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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and 13
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<superScript id="7C419BE4FFFCFFE3FEB6F8DFFE936670" attach="none" box="[324,337,1830,1844]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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; the 16
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<superScript id="7C419BE4FFFCFFE3FE59F8DFFE7A6670" attach="left" box="[427,440,1830,1844]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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to 18
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<superScript id="7C419BE4FFFCFFE3FE0CF8DFFDC96670" attach="left" box="[510,523,1830,1844]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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intervals smaller. Scales on the 13
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<superScript id="7C419BE4FFFCFFE3FC50F8DFFC6D6670" attach="left" box="[930,943,1830,1844]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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to 16
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<superScript id="7C419BE4FFFCFFE3FC07F8DFFBC06670" attach="left" box="[1013,1026,1830,1844]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
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interval small and deeply grooved, forming checkered patterns from the base to the middle part of the elytron and forming overlapping patterns from the middle part to the apex of the elytron.
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</paragraph>
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</subSubSection>
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<caption id="DF4B6624FFFDFFE2FF65FBA1FC066534" ID-DOI="http://doi.org/10.5281/zenodo.5728877" ID-Zenodo-Dep="5728877" box="[151,964,1112,1136]" httpUri="https://zenodo.org/record/5728877/files/figure.png" pageId="3" pageNumber="4" startId="3.[151,250,1112,1136]" targetBox="[152,1435,181,1088]" targetPageId="3">
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<paragraph id="8B8B36ACFFFDFFE2FF65FBA1FC066534" blockId="3.[151,964,1112,1136]" box="[151,964,1112,1136]" pageId="3" pageNumber="4">
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<emphasis id="B940EABEFFFDFFE2FF65FBA1FED16534" bold="true" box="[151,275,1112,1136]" pageId="3" pageNumber="4">FIGURE 2.</emphasis>
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Photograph of specimens.
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<emphasis id="B940EABEFFFDFFE2FDD4FBA1FD886534" bold="true" box="[550,586,1112,1136]" pageId="3" pageNumber="4">(a).</emphasis>
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GNUE-311001.
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<emphasis id="B940EABEFFFDFFE2FD05FBA1FCDE6534" bold="true" box="[759,796,1112,1136]" pageId="3" pageNumber="4">(b).</emphasis>
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GNUE-311002.
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</paragraph>
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</caption>
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<subSubSection id="C32E6527FFFDFFE5FF35FB67FE8366CD" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
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<paragraph id="8B8B36ACFFFDFFE2FF35FB67FAE66429" blockId="3.[151,1437,1182,2037]" pageId="3" pageNumber="4">
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<emphasis id="B940EABEFFFDFFE2FF35FB67FE9B65FC" bold="true" box="[199,345,1182,1208]" pageId="3" pageNumber="4">Description.</emphasis>
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GNUE-311001 is sedimented in shaly mudstone with a damaged leaf (
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<taxonomicName id="4C344D2FFFFDFFE2FB9FFB66FB7165FC" box="[1133,1203,1183,1208]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
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<emphasis id="B940EABEFFFDFFE2FB9FFB66FB7165FC" box="[1133,1203,1183,1208]" italics="true" pageId="3" pageNumber="4">Fagus</emphasis>
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</taxonomicName>
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?) (
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<figureCitation id="130F2A29FFFDFFE2FB25FB67FADE65FC" box="[1239,1308,1182,1208]" captionStart="FIGURE 2" captionStartId="3.[151,250,1112,1136]" captionTargetBox="[152,1435,181,1088]" captionTargetId="figure-19@3.[151,1435,181,1088]" captionTargetPageId="3" captionText="FIGURE 2. Photograph of specimens. (a). GNUE-311001. (b). GNUE-311002." figureDoi="http://doi.org/10.5281/zenodo.5728877" httpUri="https://zenodo.org/record/5728877/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
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). The head, half of the thorax, both damaged elytra, and parts of profemur and protibia and right mesotarsus are preserved. The total length of the specimen from the tip of the head to the apex of the elytra is
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<quantity id="4CCC9B49FFFDFFE2FBD7FB1FFB496445" box="[1061,1163,1254,1281]" metricMagnitude="-2" metricUnit="m" metricValue="3.07" pageId="3" pageNumber="4" unit="mm" value="30.7">30.7 mm</quantity>
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, and the width from the margin of the elytron to the outer margin of the pronotum is
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<quantity id="4CCC9B49FFFDFFE2FCCBFAF3FC5E6461" box="[825,924,1290,1317]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="3" pageNumber="4" unit="mm" value="12.4">12.4 mm</quantity>
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(
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<figureCitation id="130F2A29FFFDFFE2FC59FAF3FB826461" box="[939,1088,1290,1317]" captionStart="FIGURE 3" captionStartId="4.[151,250,1617,1641]" captionTargetBox="[151,1418,309,1594]" captionTargetId="figure-107@4.[151,1436,309,1594]" captionTargetPageId="4" captionText="FIGURE 3. Calosoma kimi sp. nov. holotype (GNUE-311001). (a). Photograph of the specimen. (b). Sketch of the specimen." figureDoi="http://doi.org/10.5281/zenodo.5728879" httpUri="https://zenodo.org/record/5728879/files/figure.png" pageId="3" pageNumber="4">Fig. 3a and b</figureCitation>
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). The shape of the fossil is flat, probably because of sedimentation. GNUE-311002 is in mold form. It was sedimented in the shale, only preserved with the right elytron (
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<figureCitation id="130F2A29FFFDFFE2FE66FAABFDE86428" box="[404,554,1362,1388]" captionStart="FIGURE 4" captionStartId="5.[151,250,1966,1990]" captionTargetBox="[157,1327,345,1942]" captionTargetId="figure-74@5.[155,1432,345,1942]" captionTargetPageId="5" captionText="FIGURE 4. Calosoma kimi sp. nov. paratype (GNUE-311002). (a). Photograph of the specimen. (b). Sketch of the specimen." figureDoi="http://doi.org/10.5281/zenodo.5728881" httpUri="https://zenodo.org/record/5728881/files/figure.png" pageId="3" pageNumber="4">Fig. 4a and b</figureCitation>
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. The length of the GNUE-311002 is
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<quantity id="4CCC9B49FFFDFFE2FC34FAABFBD56429" box="[966,1047,1362,1389]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="4" unit="mm" value="23.0">23 mm</quantity>
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, and the width is
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<quantity id="4CCC9B49FFFDFFE2FB2EFAABFAE26429" box="[1244,1312,1362,1389]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="4" unit="mm" value="6.0">6 mm</quantity>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8B8B36ACFFFDFFE2FF35FA8FFB286701" blockId="3.[151,1437,1182,2037]" pageId="3" pageNumber="4">
|
|
Head: Head is well preserved with loss of left maxillary palp and several antennae segments on the GNUE-311001. The left 1
|
|
<superScript id="7C419BE4FFFDFFE2FE97FA63FEB264EC" attach="left" box="[357,368,1434,1448]" fontSize="6" pageId="3" pageNumber="4">st</superScript>
|
|
and 3
|
|
<superScript id="7C419BE4FFFDFFE2FE47FA63FE0164EC" attach="left" box="[437,451,1434,1448]" fontSize="6" pageId="3" pageNumber="4">rd</superScript>
|
|
antennomeres and the right 2
|
|
<superScript id="7C419BE4FFFDFFE2FCE3FA63FCE364EC" attach="left" box="[785,801,1434,1448]" fontSize="6" pageId="3" pageNumber="4">nd</superScript>
|
|
and 3
|
|
<superScript id="7C419BE4FFFDFFE2FC94FA63FCB664EC" attach="left" box="[870,884,1434,1448]" fontSize="6" pageId="3" pageNumber="4">rd</superScript>
|
|
antennomeres are preserved. The right maxillary palp is well preserved. The length of the last and 2
|
|
<superScript id="7C419BE4FFFDFFE2FD4EFA47FD0E6488" attach="left" box="[700,716,1470,1484]" fontSize="6" pageId="3" pageNumber="4">nd</superScript>
|
|
maxillary palpomeres is approximately
|
|
<quantity id="4CCC9B49FFFDFFE2FB7AFA47FB09649D" box="[1160,1227,1470,1497]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="mm" value="1.0">1 mm</quantity>
|
|
. Both dentaries are poorly preserved, but it seems that the dentaries are protruded. The right 1
|
|
<superScript id="7C419BE4FFFDFFE2FC22FA1BFC1964B4" attach="left" box="[976,987,1506,1520]" fontSize="6" pageId="3" pageNumber="4">st</superScript>
|
|
and 2
|
|
<superScript id="7C419BE4FFFDFFE2FBEEFA1BFBEE64B4" attach="left" box="[1052,1068,1506,1520]" fontSize="6" pageId="3" pageNumber="4">nd</superScript>
|
|
labial palpomeres are preserved. Compound eyes are not preserved, but a trace of the compound eyes is poorly preserved on the middle part of the head. The shape of the head is triangular. The length of the head is
|
|
<quantity id="4CCC9B49FFFDFFE2FC77F9D3FC0A6701" box="[901,968,1578,1605]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="4" unit="mm" value="3.0">3 mm</quantity>
|
|
, and the width is
|
|
<quantity id="4CCC9B49FFFDFFE2FB7FF9D3FB246701" box="[1165,1254,1578,1605]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="4" unit="mm" value="4.7">4.7 mm</quantity>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8B8B36ACFFFDFFE2FF35F9B7FBDB67BD" blockId="3.[151,1437,1182,2037]" pageId="3" pageNumber="4">
|
|
Thorax: On the GNUE-311001, only a part of the right pronotum is preserved. The side of the pronotum is rounded and bordered in shape, which is a diagnostic feature of
|
|
<taxonomicName id="4C344D2FFFFDFFE2FC9CF98BFB1767C9" authority="(Hashmi et al. 2005)" baseAuthorityName="Hashmi" baseAuthorityYear="2005" box="[878,1237,1650,1677]" class="Insecta" family="Carabidae" genus="Calosoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B940EABEFFFDFFE2FC9CF98BFC2267C8" box="[878,992,1650,1676]" italics="true" pageId="3" pageNumber="4">Calosoma</emphasis>
|
|
(
|
|
<bibRefCitation id="EFA54B5DFFFDFFE2FC1DF98BFB0F67C9" author="Hashmi, S. N. A. & Kamaluddin, S. & Hussain, S. Z." box="[1007,1229,1650,1677]" pageId="3" pageNumber="4" pagination="259 - 272" refId="ref3877" refString="Hashmi, S. N. A., Kamaluddin, S. & Hussain, S. Z. (2005) Revision of the genus Calosoma Weber (Coleoptera: Carabidae) with two new species from Pakistan with their cladistic relationships. International Journal of Biology and Biotechnology, 2 (2), 259 - 272." type="journal article" year="2005">
|
|
Hashmi
|
|
<emphasis id="B940EABEFFFDFFE2FBBDF98AFB4367C8" box="[1103,1153,1650,1676]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
|
|
. 2005
|
|
</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
. The shape of the pronotum is rounded but slightly angled at the middle part of the lateral margin. The length of the pronotum is
|
|
<quantity id="4CCC9B49FFFDFFE2FAA7F96FFA5B67F5" box="[1365,1433,1686,1713]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="5.0">5 mm</quantity>
|
|
. The width of the right pronotum is
|
|
<quantity id="4CCC9B49FFFDFFE2FDDAF943FDAE6791" box="[552,620,1722,1749]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="4" unit="mm" value="4.0">4 mm</quantity>
|
|
. Although the pronotum is not completely preserved, on the basis of the width of the right pronotum, we assume the full width of the pronotum is
|
|
<quantity id="4CCC9B49FFFDFFE2FC22F927FBD667BD" box="[976,1044,1758,1785]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="4" unit="mm" value="8.0">8 mm</quantity>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8B8B36ACFFFDFFE5FF35F8FBFD566059" blockId="3.[151,1437,1182,2037]" lastBlockId="4.[151,1437,150,285]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
|
Elytron: On the GNUE-311001, both elytra are partly preserved. One big hole is grooved on the left elytron and from the middle to the apex of the medial part of the left elytron, and the outline of the left elytron is also partly not preserved. On the right elytron, only a part of the wing base is preserved. GNUE-311002 consists of partly preserved elytron (
|
|
<figureCitation id="130F2A29FFFDFFE2FF07F897FEF866CC" box="[245,314,1902,1928]" captionStart="FIGURE 4" captionStartId="5.[151,250,1966,1990]" captionTargetBox="[157,1327,345,1942]" captionTargetId="figure-74@5.[155,1432,345,1942]" captionTargetPageId="5" captionText="FIGURE 4. Calosoma kimi sp. nov. paratype (GNUE-311002). (a). Photograph of the specimen. (b). Sketch of the specimen." figureDoi="http://doi.org/10.5281/zenodo.5728881" httpUri="https://zenodo.org/record/5728881/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
|
|
). The shape of the elytron is quadrangle-like. The medial outlet line is thin. The total countable striae are 19 and intervals are 18. The 1st interval is small. The 4
|
|
<superScript id="7C419BE4FFFDFFE2FCEFF86BFCE866E4" attach="left" box="[797,810,1938,1952]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
|
to 8
|
|
<superScript id="7C419BE4FFFDFFE2FCA8F86BFCA566E4" attach="left" box="[858,871,1938,1952]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
|
intervals are large and become smaller from the 9
|
|
<superScript id="7C419BE4FFFDFFE2FA62F86BFA5F66E4" attach="none" box="[1424,1437,1938,1952]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
|
interval. The 16
|
|
<superScript id="7C419BE4FFFDFFE2FEBAF84FFE976680" attach="left" box="[328,341,1974,1988]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
|
to 18
|
|
<superScript id="7C419BE4FFFDFFE2FE66F84FFE636680" attach="left" box="[404,417,1974,1988]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
|
intervals are smallest and fainter than other intervals. Transversal lines are clearly showing from the 5
|
|
<superScript id="7C419BE4FFFDFFE2FEF9F823FEDA66AC" attach="left" box="[267,280,2010,2024]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
|
to 16
|
|
<superScript id="7C419BE4FFFDFFE2FEA5F823FEA666AC" attach="left" box="[343,356,2010,2024]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
|
intervals. Grooved dimples are shown on the 4
|
|
<superScript id="7C419BE4FFFDFFE2FC80F823FCBD66AC" attach="none" box="[882,895,2010,2024]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
|
, 8
|
|
<superScript id="7C419BE4FFFDFFE2FC69F823FC6A66AC" attach="none" box="[923,936,2010,2024]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
|
, and 12
|
|
<superScript id="7C419BE4FFFDFFE2FBF2F823FBCF66AC" attach="left" box="[1024,1037,2010,2024]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
|
intervals, creating a triploid pattern (
|
|
<bibRefCitation id="EFA54B5DFFFAFFE5FF6DFF6EFEF561F5" author="Jeannel, R." box="[159,311,151,177]" pageId="4" pageNumber="5" refId="ref4016" refString="Jeannel, R. (1940) Les Calosomes. Memoires du Musee National D'Histoire Naturelle. Editions du Musee, Paris, 255 pp." type="book" year="1940">Jeannel 1940</bibRefCitation>
|
|
). The 5
|
|
<superScript id="7C419BE4FFFAFFE5FE7AFF6FFE5761E0" attach="left" box="[392,405,150,164]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
and 7
|
|
<superScript id="7C419BE4FFFAFFE5FE2AFF6FFE2761E0" attach="left" box="[472,485,150,164]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
intervals are connected at the apex of the elytron and surround by the 6
|
|
<superScript id="7C419BE4FFFAFFE5FB0EFF6FFACB61E0" attach="left" box="[1276,1289,150,164]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
interval. The 11
|
|
<superScript id="7C419BE4FFFAFFE5FF40FF43FF7D618C" attach="left" box="[178,191,186,200]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
and 13
|
|
<superScript id="7C419BE4FFFAFFE5FEE2FF43FEDF618C" attach="left" box="[272,285,186,200]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
intervals are connected at the near margin of the elytron and surround the 12
|
|
<superScript id="7C419BE4FFFAFFE5FB9DFF43FBBE618C" attach="left" box="[1135,1148,186,200]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
interval. The 10
|
|
<superScript id="7C419BE4FFFAFFE5FAC0FF43FAFD618C" attach="left" box="[1330,1343,186,200]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
and 14
|
|
<superScript id="7C419BE4FFFAFFE5FA62FF43FA5F618C" attach="none" box="[1424,1437,186,200]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
intervals meet and surround the 11
|
|
<superScript id="7C419BE4FFFAFFE5FDD2FF27FDEF61A8" attach="left" box="[544,557,222,236]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
to 13
|
|
<superScript id="7C419BE4FFFAFFE5FD9DFF27FDBE61A8" attach="left" box="[623,636,222,236]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
intervals at the apex of the elytron. These interval connection patterns are clearly visible on GNUE-311002 (
|
|
<figureCitation id="130F2A29FFFAFFE5FDB0FEFAFD446059" box="[578,646,258,285]" captionStart="FIGURE 5" captionStartId="6.[151,250,1696,1720]" captionTargetBox="[155,1432,181,1672]" captionTargetId="figure-27@6.[155,1432,181,1672]" captionTargetPageId="6" captionText="FIGURE 5. The apex of the wing of C.kimi sp. nov. paratype (GNUE-311002). (a). Photograph of the 10th to 14th interval connection patterns. (b). Sketch of the 10th to 14th interval connection patterns. (c). Photograph of the 5th to 7th interval connection patterns. (d). Sketch of the 5th to 7th interval connection patterns." figureDoi="http://doi.org/10.5281/zenodo.5728883" httpUri="https://zenodo.org/record/5728883/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="DF4B6624FFFAFFE5FF65F9A8FA55672D" ID-DOI="http://doi.org/10.5281/zenodo.5728879" ID-Zenodo-Dep="5728879" box="[151,1431,1617,1641]" httpUri="https://zenodo.org/record/5728879/files/figure.png" pageId="4" pageNumber="5" startId="4.[151,250,1617,1641]" targetBox="[151,1418,309,1594]" targetPageId="4">
|
|
<paragraph id="8B8B36ACFFFAFFE5FF65F9A8FA55672D" blockId="4.[151,1431,1617,1641]" box="[151,1431,1617,1641]" pageId="4" pageNumber="5">
|
|
<emphasis id="B940EABEFFFAFFE5FF65F9A8FED1672D" bold="true" box="[151,275,1617,1641]" pageId="4" pageNumber="5">FIGURE 3.</emphasis>
|
|
<taxonomicName id="4C344D2FFFFAFFE5FEEBF9A8FE70672D" authority="Lee & Nam, 2021" authorityName="Lee & Nam" authorityYear="2021" box="[281,434,1617,1641]" class="Insecta" family="Carabidae" genus="Calosoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kimi" status="sp. nov.">
|
|
<emphasis id="B940EABEFFFAFFE5FEEBF9A8FE70672D" box="[281,434,1617,1641]" italics="true" pageId="4" pageNumber="5">Calosoma kimi</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="B940EABEFFFAFFE5FE4AF9A8FDCB672D" bold="true" box="[440,521,1617,1641]" pageId="4" pageNumber="5">
|
|
<taxonomicNameLabel id="A27357C5FFFAFFE5FE4AF9A8FDCB672D" box="[440,521,1617,1641]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
|
</emphasis>
|
|
holotype (GNUE-311001).
|
|
<emphasis id="B940EABEFFFAFFE5FCD4F9A8FC88672D" bold="true" box="[806,842,1617,1641]" pageId="4" pageNumber="5">(a).</emphasis>
|
|
Photograph of the specimen.
|
|
<emphasis id="B940EABEFFFAFFE5FB8BF9A8FB5C672D" bold="true" box="[1145,1182,1617,1641]" pageId="4" pageNumber="5">(b).</emphasis>
|
|
Sketch of the specimen.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8B8B36ACFFFAFFE5FF35F942FB14665A" blockId="4.[151,1436,1723,1929]" pageId="4" pageNumber="5">
|
|
The scales are deeply grooved from the 9
|
|
<superScript id="7C419BE4FFFAFFE5FD7DF942FD5E678D" attach="left" box="[655,668,1723,1737]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
to 15
|
|
<superScript id="7C419BE4FFFAFFE5FD28F942FD25678D" attach="left" box="[730,743,1723,1737]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
interval. The scales on the 17
|
|
<superScript id="7C419BE4FFFAFFE5FBDFF942FBF8678D" attach="left" box="[1069,1082,1723,1737]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
and 18
|
|
<superScript id="7C419BE4FFFAFFE5FB79F942FB5A678D" attach="left" box="[1163,1176,1723,1737]" fontSize="6" pageId="4" pageNumber="5">th</superScript>
|
|
intervals are extremely small. From the wing base to the middle of the elytron, the scales are arranged in a check pattern. However, from the middle to the margin of the elytron, the scales are arranged in an overlapping pattern (
|
|
<figureCitation id="130F2A29FFFAFFE5FB71F8FAFB05665A" box="[1155,1223,1795,1822]" captionStart="FIGURE 6" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[407,1180,181,1920]" captionTargetId="figure-19@7.[407,1180,181,1920]" captionTargetPageId="7" captionText="FIGURE 6. Scales on elytron of C. kimi sp. nov. Scales on the above represents a checkered patterns. Scales on the bottom represents an overlapping patterns." figureDoi="http://doi.org/10.5281/zenodo.5728885" httpUri="https://zenodo.org/record/5728885/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8B8B36ACFFFAFFE5FF35F8D1FE8366CD" blockId="4.[151,1436,1723,1929]" pageId="4" pageNumber="5">Leg: On GNUE-311001, part of the left profemur and protibia segments and the right leg tarsus (assumed mesotarsus) are preserved. The tarsus on the right leg is forward to the upside. We assume that it is broken and folded upward.</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |