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<document id="B76AD8622054E81CF96DA6CF33081792" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625623959389" checkinUser="felipe" docAuthor="Published, First" docDate="2007" docId="039D4153547C955236FE7D52FC5DFCA4" docLanguage="en" docName="zt01396p084.pdf" docOrigin="Zootaxa 1396" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Arrup areolatus" docType="treatment" docVersion="2" lastPageNumber="17" masterDocId="FFA4392B5473954236697820FFDBFFA8" masterDocTitle="The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha)" masterLastPageNumber="84" masterPageNumber="1" pageNumber="16" updateTime="1698992576941" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3849409B247BB38E0F889734BC65AB5F">The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha)</mods:title>
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<mods:date id="7BED52F5F1CC400ACB470B5D0509ABDA">2007</mods:date>
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<mods:number id="36A1125C59304325F32E1BAF6C4AFEAB">2007-01-22</mods:number>
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<treatment id="039D4153547C955236FE7D52FC5DFCA4" LSID="urn:lsid:plazi:treatment:039D4153547C955236FE7D52FC5DFCA4" httpUri="http://treatment.plazi.org/id/039D4153547C955236FE7D52FC5DFCA4" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection id="C32EA3CE547C954D36FE7D52FDE5FA24" box="[151,574,1394,1420]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8B8BF045547C954D36FE7D52FDE5FA24" blockId="15.[151,574,1394,1455]" box="[151,574,1394,1420]" pageId="15" pageNumber="16">
<heading id="D0C34729547C954D36FE7D52FDE5FA24" bold="true" box="[151,574,1394,1420]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName id="4C348BC6547C954D36FE7D52FDE5FA24" ID-CoL="GV68" authority="(Shinohara, 1957)" baseAuthorityName="Shinohara" baseAuthorityYear="1957" box="[151,574,1394,1420]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="areolatus">
<emphasis id="B9402C57547C954D36FE7D52FDE5FA24" bold="true" box="[151,574,1394,1420]" pageId="15" pageNumber="16">
<emphasis id="B9402C57547C954D36FE7D52FE8CFA24" bold="true" box="[151,343,1394,1420]" italics="true" pageId="15" pageNumber="16">Arrup areolatus</emphasis>
(
<bibRefCitation id="EFA58DB4547C954D370E7D52FDEDFA24" author="Shinohara, K." box="[359,566,1394,1420]" pageId="15" pageNumber="16" pagination="187 - 190" refId="ref54889" refString="Shinohara, K. (1957) Taxonomical and morphological study on Myriapoda: I: Two new species of Chilopoda. Zoological Magazine, Tokyo, 66 (4), 187 - 190." type="journal article" year="1957">Shinohara, 1957</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32EA3CE547C954D36FE7DB5FB32FA54" pageId="15" pageNumber="16" type="description">
<paragraph id="8B8BF045547C954D36FE7DB5FF3CFA07" blockId="15.[151,574,1394,1455]" box="[151,231,1429,1455]" pageId="15" pageNumber="16">
<figureCitation id="130FECC0547C954D36FE7DB5FF3CFA07" box="[151,231,1429,1455]" captionStart="FIGURES 69" captionStartId="11.[151,270,1889,1913]" captionTargetBox="[282,1281,564,1847]" captionTargetId="figure-183@11.[264,1323,554,1894]" captionTargetPageId="11" captionText="FIGURES 69. 67, Arrup cf. holstii, male from Mt. Kenashi, Honshu: 6, anterior part of head, ventral (maxillary complex removed); 7, forcipular segment and forcipules, dorsal (the poison calyx is indicated by solid lines, setae not drawn). Areolation is drawn only partially. 8, Arrup sauteri: forcipular segment and left forcipule, ventral (setae not drawn); redrawn from Silvestri (1919). 9, Arrup areolatus: maxillary complex, ventral; redrawn from Shinohara (1957)." pageId="15" pageNumber="16">Figs. 9</figureCitation>
</paragraph>
<paragraph id="8B8BF045547C954D36FE7DC5FB32FA54" blockId="15.[151,1257,1509,1628]" box="[151,1257,1509,1532]" pageId="15" pageNumber="16">
[1]
<taxonomicName id="4C348BC6547C954D36AE7DC5FDA2FA54" authority="Shinohara 1957" authorityName="Shinohara" authorityYear="1957" box="[199,633,1509,1532]" class="Chilopoda" family="Mecistocephalidae" genus="Nodocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="areolatus">
<emphasis id="B9402C57547C954D36AE7DC5FE11FA54" box="[199,458,1509,1532]" italics="true" pageId="15" pageNumber="16">Nodocephalus areolatus</emphasis>
<bibRefCitation id="EFA58DB4547C954D37B97DC5FDA2FA54" author="Shinohara, K." box="[464,633,1509,1532]" pageId="15" pageNumber="16" pagination="187 - 190" refId="ref54889" refString="Shinohara, K. (1957) Taxonomical and morphological study on Myriapoda: I: Two new species of Chilopoda. Zoological Magazine, Tokyo, 66 (4), 187 - 190." type="journal article" year="1957">Shinohara 1957</bibRefCitation>
</taxonomicName>
— Zool. Mag.
<collectingRegion id="49F03EA7547C954D35767DC5FCB8FA54" box="[799,867,1509,1532]" country="Japan" name="Tokyo" pageId="15" pageNumber="16">Tokyo</collectingRegion>
66: 187 (original description); fig. 1
</paragraph>
</subSubSection>
<subSubSection id="C32EA3CE547C954D36AE7E25FC97F9F4" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8B8BF045547C954D36AE7E25FBDFF9B4" blockId="15.[151,1257,1509,1628]" box="[199,1028,1541,1564]" pageId="15" pageNumber="16">
<taxonomicName id="4C348BC6547C954D36AE7E25FD59F9B4" authority=": Murakami 1993" authorityName="Murakami" authorityYear="1993" box="[199,642,1541,1564]" class="Chilopoda" family="Mecistocephalidae" genus="Nodocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="areolatus">
<emphasis id="B9402C57547C954D36AE7E25FE12F9B4" box="[199,457,1541,1564]" italics="true" pageId="15" pageNumber="16">Nodocephalus areolatus</emphasis>
:
<bibRefCitation id="EFA58DB4547C954D37B17E25FD59F9B4" author="Murakami, Y." box="[472,642,1541,1564]" pageId="15" pageNumber="16" pagination="95 - 107" refId="ref54511" refString="Murakami, Y. (1993) Arthropoda: Diplopoda, Pauropoda, Chilopoda, Symphyla. In: Environmental Agency Japan (Ed.), List of species of Animals in Japan: Invertebrates I. The Shizen-Kankyo-Kenkyu Center, Tokyo, 5, 95 - 107." type="journal article" year="1993">Murakami 1993</bibRefCitation>
</taxonomicName>
— List Species Anim.
<collectingCountry id="F323B0D5547C954D35147E25FC62F9B4" box="[893,953,1541,1564]" name="Japan" pageId="15" pageNumber="16">Japan</collectingCountry>
5: 105
</paragraph>
<paragraph id="8B8BF045547C954D36AE7E05FB61F994" blockId="15.[151,1257,1509,1628]" box="[199,1210,1573,1596]" pageId="15" pageNumber="16">
<taxonomicName id="4C348BC6547C954D36AE7E05FDA4F994" authority=": Shinohara 1999" authorityName="Shinohara" authorityYear="1999" box="[199,639,1573,1596]" class="Chilopoda" family="Mecistocephalidae" genus="Nodocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="areolatus">
<emphasis id="B9402C57547C954D36AE7E05FE11F994" box="[199,458,1573,1596]" italics="true" pageId="15" pageNumber="16">Nodocephalus areolatus</emphasis>
:
<bibRefCitation id="EFA58DB4547C954D37B17E05FDA4F994" author="Shinohara, K." box="[472,639,1573,1596]" pageId="15" pageNumber="16" pagination="693 - 709" refId="ref55374" refString="Shinohara, K. (1999) Chilopoda. In: Aoki, J. (Ed.), Pictorial keys to soil animals of Japan. Tokay University Press, Tokyo, 693 - 709." type="book chapter" year="1999">Shinohara 1999</bibRefCitation>
</taxonomicName>
— Pictorial Keys Soil Anim.
<collectingCountry id="F323B0D5547C954D35A87E05FBD9F994" box="[961,1026,1573,1596]" name="Japan" pageId="15" pageNumber="16">Japan</collectingCountry>
: 709; fig. 6 (703)
</paragraph>
<paragraph id="8B8BF045547C954D36AE7E65FC97F9F4" blockId="15.[151,1257,1509,1628]" box="[199,844,1605,1628]" pageId="15" pageNumber="16">
<taxonomicName id="4C348BC6547C954D36AE7E65FD9AF9F4" ID-CoL="GV68" authority=": Foddai et al. 2003" authorityName="Foddai" authorityYear="2003" box="[199,577,1605,1628]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="areolatus">
<emphasis id="B9402C57547C954D36AE7E65FEAAF9F4" box="[199,369,1605,1628]" italics="true" pageId="15" pageNumber="16">Arrup areolatus</emphasis>
:
<bibRefCitation id="EFA58DB4547C954D37167E65FD9AF9F4" author="Foddai, D. &amp; Bonato, L. &amp; Pereira, L. A. &amp; Minelli, A." box="[383,577,1605,1628]" pageId="15" pageNumber="16" refId="ref53487" refString="Foddai, D., Bonato, L., Pereira, L. A. &amp; Minelli, A., 2003 Phylogeny and systematics of the Arrupinae (Chilopoda Geophilomorpha Mecistocephalidae) with description of a new dwarfed species. Journal of Natural History, 37,1247 - 1267." type="book" year="2003">
Foddai
<emphasis id="B9402C57547C954D37B97E65FDDDF9F4" box="[464,518,1605,1628]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2003
</bibRefCitation>
</taxonomicName>
— J. Nat. Hist. 37: 1262
</paragraph>
</subSubSection>
<subSubSection id="C32EA3CE547C955236FE7EAEFC5DFCA4" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8B8BF045547C954D36FE7EAEFDD7F8B7" blockId="15.[151,1437,1677,2023]" pageId="15" pageNumber="16">
<emphasis id="B9402C57547C954D36FE7EAEFED5F90F" box="[151,270,1678,1703]" italics="true" pageId="15" pageNumber="16">Diagnosis</emphasis>
. An
<taxonomicName id="4C348BC6547C954D372F7EAEFE33F90F" box="[326,488,1677,1703]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9402C57547C954D372F7EAEFE57F90F" box="[326,396,1678,1703]" italics="true" pageId="15" pageNumber="16">Arrup</emphasis>
species
</taxonomicName>
with body length reaching about
<quantity id="4CCC5DA0547C954D35047EADFC71F90F" box="[877,938,1677,1703]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="cm" value="2.0">2 cm</quantity>
. Basal part of the medial projections of the first maxillae about 3 times as long as the distal lobe, with swollen internal margin. Distal article of the telopodite of the second maxillae without claw. Forcipular segment I with a well developed tooth, subtriangular, acute and projecting.
</paragraph>
<paragraph id="8B8BF045547C954D36AF7F0EFABFF8EF" blockId="15.[151,1437,1677,2023]" box="[198,1380,1837,1863]" pageId="15" pageNumber="16">
<emphasis id="B9402C57547C954D36AF7F0EFEB3F8EF" box="[198,360,1838,1863]" italics="true" pageId="15" pageNumber="16">Type material</emphasis>
.
<typeStatus id="548F4EE7547C954D371E7F0DFE32F8EF" box="[375,489,1837,1863]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
: female, about
<quantity id="4CCC5DA0547C954D34F37F0DFD36F8EF" box="[666,749,1837,1863]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="16" unit="mm" value="18.0">18 mm</quantity>
long, adult.
<typeStatus id="548F4EE7547C954D35E97F0DFC2CF8EF" box="[896,1015,1837,1863]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D323BCC547C954D326B7F0DFB59F8EF" box="[1026,1154,1837,1863]" pageId="15" pageNumber="16" type="juvenile">2 juveniles</specimenCount>
(
<bibRefCitation id="EFA58DB4547C954D32F87F0DFA8CF8EF" author="Shinohara, K." box="[1169,1367,1837,1863]" pageId="15" pageNumber="16" pagination="187 - 190" refId="ref54889" refString="Shinohara, K. (1957) Taxonomical and morphological study on Myriapoda: I: Two new species of Chilopoda. Zoological Magazine, Tokyo, 66 (4), 187 - 190." type="journal article" year="1957">Shinohara, 1957</bibRefCitation>
).
</paragraph>
<paragraph id="8B8BF045547C954D36AF7F76FC0FF8C7" blockId="15.[151,1437,1677,2023]" box="[198,980,1877,1903]" pageId="15" pageNumber="16">
<emphasis id="B9402C57547C954D36AF7F76FE80F8C7" box="[198,347,1878,1903]" italics="true" pageId="15" pageNumber="16">
<typeStatus id="548F4EE7547C954D36AF7F76FF26F8C7" box="[198,253,1878,1903]" pageId="15" pageNumber="16">Type</typeStatus>
locality
</emphasis>
. Okyô­daira (Chichibu), Honshu (
<bibRefCitation id="EFA58DB4547C954D35687F75FC1CF8C7" author="Shinohara, K." box="[769,967,1877,1903]" pageId="15" pageNumber="16" pagination="187 - 190" refId="ref54889" refString="Shinohara, K. (1957) Taxonomical and morphological study on Myriapoda: I: Two new species of Chilopoda. Zoological Magazine, Tokyo, 66 (4), 187 - 190." type="journal article" year="1957">Shinohara, 1957</bibRefCitation>
).
</paragraph>
<paragraph id="8B8BF045547C954D36AF7F5EFC0DF83F" blockId="15.[151,1437,1677,2023]" box="[198,982,1917,1943]" pageId="15" pageNumber="16">
<emphasis id="B9402C57547C954D36AF7F5EFDD2F83F" box="[198,521,1918,1943]" italics="true" pageId="15" pageNumber="16">
Depository of
<typeStatus id="548F4EE7547C954D37047F5EFE45F83F" box="[365,414,1918,1943]" pageId="15" pageNumber="16">type</typeStatus>
material
</emphasis>
. Coll. K. Shinohara (
<bibRefCitation id="EFA58DB4547C954D356A7F5DFC12F83F" author="Shinohara, K." box="[771,969,1917,1943]" pageId="15" pageNumber="16" pagination="187 - 190" refId="ref54889" refString="Shinohara, K. (1957) Taxonomical and morphological study on Myriapoda: I: Two new species of Chilopoda. Zoological Magazine, Tokyo, 66 (4), 187 - 190." type="journal article" year="1957">Shinohara, 1957</bibRefCitation>
).
</paragraph>
<paragraph id="8B8BF045547C954D36AF7F86FE21F817" blockId="15.[151,1437,1677,2023]" box="[198,506,1957,1983]" pageId="15" pageNumber="16">
<emphasis id="B9402C57547C954D36AF7F86FE7FF817" box="[198,420,1958,1983]" italics="true" pageId="15" pageNumber="16">Material examined</emphasis>
. None.
</paragraph>
<paragraph id="8B8BF045547C955236AF7FEEFD3CFDEA" blockId="15.[151,1437,1677,2023]" lastBlockId="16.[151,1437,152,780]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="B9402C57547C954D36AF7FEEFE8BF84F" box="[198,336,1998,2023]" italics="true" pageId="15" pageNumber="16">Description</emphasis>
. Body size of adults reaching at least about
<quantity id="4CCC5DA0547C954D35037FEDFC70F84F" box="[874,939,1997,2023]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="cm" value="2.0">2 cm</quantity>
. Body colour yellow, head and forcipular segment deep orange. Head about 1.5 times as long as wide. Clypeal ratio about 3.5. Clypeal setae: about 12 pairs, including one median pair anterior to the plagulae and a group on each side reaching the anterior corner. Labrum: anterior ala triangular, medial margin reduced to a vertex; posterior margin of side­pieces almost straight. First maxillae: anterior corners of coxosternum not projecting; each medial projection with 34 spines on the internal margin and 2 setae at mid­length; basal part very large and distally rounded, about 3 times as long as the distal lobe; distal lobe emerging abruptly from the basal part, with an acute angle; each telopodite with 2 setae at mid­length, distal lobe attenuated. Second maxillae: article I of the telopodite about 2.5 times as long as wide, article III slender, about 4.3 times as long as wide; claw absent. Forcipules: article I about 1.5 times as long as wide, distal tooth acute, and distinctly projecting outwards; article II without teeth; article III with a tubercle; tarsungulum with one distinct basal tooth. Last leg­bearing segment: sternum semicircular; about 14 pores on each coxopleuron.
</paragraph>
<paragraph id="8B8BF0455463955236AF7A70FDB3FDC2" blockId="16.[151,1437,152,780]" box="[198,616,592,618]" pageId="16" pageNumber="17">
<emphasis id="B9402C575463955236AF7A70FDB9FDC1" box="[198,610,592,617]" italics="true" pageId="16" pageNumber="17">Distribution in the considered area</emphasis>
.
</paragraph>
<paragraph id="8B8BF0455463955236AF7A5AFCCCFD3C" blockId="16.[151,1437,152,780]" box="[198,791,632,660]" pageId="16" pageNumber="17">
Honshu: Okyô­daira (Chichibu)
<superScript id="7C415D0D5463955234397A58FDB8FD2E" attach="left" box="[592,611,632,646]" fontSize="6" pageId="16" pageNumber="17">[1]</superScript>
(
<typeStatus id="548F4EE754639552341B7A5AFD7DFD3C" box="[626,678,634,660]" pageId="16" pageNumber="17">type</typeStatus>
locality).
</paragraph>
<paragraph id="8B8BF0455463955236AF7A83FCB9FD14" blockId="16.[151,1437,152,780]" box="[198,866,674,700]" pageId="16" pageNumber="17">
<emphasis id="B9402C575463955236AF7A83FE6DFD14" box="[198,438,675,700]" italics="true" pageId="16" pageNumber="17">General distribution</emphasis>
. Only known from the
<typeStatus id="548F4EE75463955234A37A82FD27FD14" box="[714,764,674,700]" pageId="16" pageNumber="17">type</typeStatus>
locality.
</paragraph>
<paragraph id="8B8BF0455463955236AF7AEBFC5DFCA4" blockId="16.[151,1437,152,780]" pageId="16" pageNumber="17">
<emphasis id="B9402C575463955236AF7AEBFEF6FD4C" box="[198,301,715,740]" italics="true" pageId="16" pageNumber="17">Remarks</emphasis>
. The unusual elongation of the distal article of the telopodite of the second maxillae, inferred from the original drawing, may represent a diagnostic character.
</paragraph>
</subSubSection>
</treatment>
</document>