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<document id="930DFF87AA1CC1ABB1C246FA25D95B73" ID="10.11646/zootaxa.4231.1.11SLASH10070" ID-CLB-Dataset="37061" ID-DOI="10.11646/zootaxa.4231.1.11" ID-GBIF-Dataset="93b6aba3-df53-42ac-8731-aeead58f58fb" ID-ISSN="1175-5326" ID-Zenodo-Dep="293607" ID-ZooBank="9444631E-01A8-451E-81F3-C9126D1CB602" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1486632309353" checkinUser="plazi" docAuthor="Lopes, Rogério B., Noll, Fernando B. &amp; Stange, Lionel A." docDate="2017" docId="039B0E59FFA10F32FF7AC1E0FC12FA0A" docLanguage="en" docName="zootaxa.4231.1.11SLASH10070.pdf" docOrigin="Zootaxa 4231 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zethus wileyi" docType="treatment" docVersion="10" lastPageNumber="138" masterDocId="FFA27621FFA00F33FFEDC217FFB3FF91" masterDocTitle="Zethus (Zethus) wileyi (Hymenoptera, Vespidae, Eumeninae), a new species with its own species-group" masterLastPageNumber="144" masterPageNumber="137" pageNumber="138" updateTime="1698436705272" updateUser="ExternalLinkService">
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<mods:title id="F678DF936F096E3941498B8B608BD137">Zethus (Zethus) wileyi (Hymenoptera, Vespidae, Eumeninae), a new species with its own species-group</mods:title>
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<mods:namePart id="18B3DBE04CBEBE43458D568D0EA5913A">Lopes, Rogério B.</mods:namePart>
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<mods:namePart id="E98D71E81E917E68377F489F0F6759D6">Noll, Fernando B.</mods:namePart>
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<mods:namePart id="2F3B706EF2D7CC74110CC263A41EB12F">Stange, Lionel A.</mods:namePart>
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<mods:date id="112CE75CF4FDA66609C8DDF97F57B5A6">2017</mods:date>
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<subSubSection id="C328ECC4FFA10F32FF7AC1E0FD6FFB83" box="[151,732,1015,1042]" pageId="1" pageNumber="138" type="nomenclature">
<paragraph id="8B8DBF4FFFA10F32FF7AC1E0FD6FFB83" blockId="1.[151,732,1015,1042]" box="[151,732,1015,1042]" pageId="1" pageNumber="138">
<heading id="D0C50823FFA10F32FF7AC1E0FD6FFB83" bold="true" box="[151,732,1015,1042]" fontSize="11" level="1" pageId="1" pageNumber="138" reason="1">
<emphasis id="B946635DFFA10F32FF7AC1E0FD6FFB83" bold="true" box="[151,732,1015,1042]" pageId="1" pageNumber="138">
<taxonomicName id="4C32C4CCFFA10F32FF7AC1E0FE84FB80" box="[151,311,1015,1041]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="wileyi">
<emphasis id="B946635DFFA10F32FF7AC1E0FE84FB80" bold="true" box="[151,311,1015,1041]" italics="true" pageId="1" pageNumber="138">Zethus wileyi</emphasis>
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species-group, new species-group
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<subSubSection id="C328ECC4FFA10F32FF7AC629FD54FB0E" pageId="1" pageNumber="138" type="diagnosis">
<paragraph id="8B8DBF4FFFA10F32FF7AC629FD17FBEA" blockId="1.[151,1436,1086,1435]" pageId="1" pageNumber="138">
<emphasis id="B946635DFFA10F32FF7AC629FEB9FBC6" bold="true" box="[151,266,1086,1111]" pageId="1" pageNumber="138">Diagnosis</emphasis>
. Stem of T2 longer than stem of T1; F11 of male falciform, long and sharp; submedian carina present; and complex male genitalia (described below).
</paragraph>
<paragraph id="8B8DBF4FFFA10F32FF2AC691FD54FB0E" blockId="1.[151,1436,1086,1435]" box="[199,743,1158,1183]" pageId="1" pageNumber="138">
<emphasis id="B946635DFFA10F32FF2AC691FE22FB0E" bold="true" box="[199,401,1158,1183]" pageId="1" pageNumber="138">Included species.</emphasis>
<taxonomicName id="4C32C4CCFFA10F32FE75C691FD30FB0E" authority="Stange" authorityName="Stange" box="[408,643,1158,1183]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="wileyi" status="sp. nov.">
<emphasis id="B946635DFFA10F32FE75C691FD9EFB0F" box="[408,557,1158,1182]" italics="true" pageId="1" pageNumber="138">Zethus wileyi</emphasis>
Stange
</taxonomicName>
,
<taxonomicNameLabel id="A275DE26FFA10F32FD60C691FD54FB0E" box="[653,743,1158,1183]" pageId="1" pageNumber="138" rank="species">
<emphasis id="B946635DFFA10F32FD60C691FD54FB0E" bold="true" box="[653,743,1158,1183]" pageId="1" pageNumber="138">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C328ECC4FFA10F32FF2AC6BDFB3FFABE" pageId="1" pageNumber="138" type="description">
<paragraph id="8B8DBF4FFFA10F32FF2AC6BDFB3FFABE" blockId="1.[151,1436,1086,1435]" pageId="1" pageNumber="138">
<emphasis id="B946635DFFA10F32FF2AC6BDFEE9FB52" bold="true" box="[199,346,1194,1219]" pageId="1" pageNumber="138">Systematics.</emphasis>
The squared anterior margin of S1 along with rounded aspect of the basal plate of the aedeagus are synapomorphies that support this species as sister group to the
<taxonomicName id="4C32C4CCFFA10F32FC16C6D8FB24FB77" box="[1019,1175,1230,1254]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="hilarianus">
<emphasis id="B946635DFFA10F32FC16C6D8FB24FB77" box="[1019,1175,1230,1254]" italics="true" pageId="1" pageNumber="138">Z. hilarianus</emphasis>
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species-group, while plesiomorphies such as the hirsute ventral surface of the fore femur of the male, slit-shaped propodeal aperture and convergent antero-lateral margin of S1 keep it from being part of the extant species-group.
</paragraph>
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<subSubSection id="C328ECC4FFA10F32FF2AC72DFC12FA0A" pageId="1" pageNumber="138" type="discussion">
<paragraph id="8B8DBF4FFFA10F32FF2AC72DFC12FA0A" blockId="1.[151,1436,1086,1435]" pageId="1" pageNumber="138">
<emphasis id="B946635DFFA10F32FF2AC72DFE88FAC2" bold="true" box="[199,315,1338,1363]" pageId="1" pageNumber="138">Remarks.</emphasis>
Following Bohart and Stanges (1965) key, the specimen (
<figureCitation id="1309A3CAFFA10F32FC23C72DFBBDFAC2" box="[974,1038,1338,1363]" captionStart="FIGURE 1" captionStartId="2.[151,250,690,712]" captionTargetBox="[151,1435,195,666]" captionTargetId="figure@2.[151,1435,195,666]" captionTargetPageId="2" captionText="FIGURE 1. Zethus wileyi sp. nov., ♂, holotype. a. Head; b. Habitus. Scale = 1. 0 mm" httpUri="https://zenodo.org/record/311435/files/figure.png" pageId="1" pageNumber="138">Fig. 1</figureCitation>
) runs to the
<taxonomicName id="4C32C4CCFFA10F32FB73C72CFF7CFAE6" authority="de Saussure, 1855" authorityName="de Saussure" authorityYear="1855" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="smithii">
<emphasis id="B946635DFFA10F32FB73C72CFABBFAC2" box="[1182,1288,1339,1363]" italics="true" pageId="1" pageNumber="138">Z. smithii</emphasis>
de
<bibRefCitation id="EFA3C2BEFFA10F32FADCC72CFF7CFAE6" author="Saussure" pageId="1" pageNumber="138" refString="Saussure, H. de (1855) Etudes sur la famille des vespides 3. La Monographie des Masariens et un supplement a la Monographie des Eumeniens. V. Masson, Paris &amp; J. Cherbuliez, Geneva, 352 pp" type="book" year="1855">Saussure, 1855</bibRefCitation>
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species-group, which is now part of the
<taxonomicName id="4C32C4CCFFA10F32FD7BC748FC4AFAE6" authority="de Saussure, 1855" authorityName="de Saussure" authorityYear="1855" box="[662,1017,1374,1399]" class="Insecta" family="Eumenidae" genus="Zethus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="hilarianus">
<emphasis id="B946635DFFA10F32FD7BC748FC94FAE7" box="[662,807,1374,1398]" italics="true" pageId="1" pageNumber="138">Z. hilarianus</emphasis>
de
<bibRefCitation id="EFA3C2BEFFA10F32FCBDC748FC4AFAE6" author="Saussure" box="[848,1017,1374,1399]" pageId="1" pageNumber="138" refString="Saussure, H. de (1855) Etudes sur la famille des vespides 3. La Monographie des Masariens et un supplement a la Monographie des Eumeniens. V. Masson, Paris &amp; J. Cherbuliez, Geneva, 352 pp" type="book" year="1855">Saussure, 1855</bibRefCitation>
</taxonomicName>
species-group, according to
<bibRefCitation id="EFA3C2BEFFA10F32FAD1C748FEBBFA0A" pageId="1" pageNumber="138" refString="Lopes, R. B., Kumagai, A. F., Stange, L. A. &amp; Hermes, M. G. (2015) Cladistic analysis and systematic revision of the Zethus smithii (de Saussure) species-group sensu Bohart &amp; Stange (Hymenoptera: Vespidae: Eumeninae). Invertebrate Systematics, 29, 421 - 443." type="journal article">
Lopes
<emphasis id="B946635DFFA10F32FA65C748FF07FA0A" italics="true" pageId="1" pageNumber="138">et al.</emphasis>
(2015)
</bibRefCitation>
. However, it does not agree with any of the known species.
</paragraph>
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