treatments-xml/data/03/99/76/03997670FFBFE6688ECBFC5F773A6B5A.xml
2024-06-21 12:22:17 +02:00

498 lines
63 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="E24EBEA8BD932C0BA9D5592C45311433" ID-DOI="10.5281/zenodo.195835" ID-GBIF-Dataset="5fe3da15-3dd3-495f-9c15-7f9d606315aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="195835" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460234920266" checkinUser="plazi" docAuthor="Emberson, Rowan M." docDate="2010" docId="03997670FFBFE6688ECBFC5F773A6B5A" docLanguage="en" docName="zt02501p053.pdf" docOrigin="Zootaxa 2501" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Longicheles Valle 1953" docType="treatment" docVersion="8" lastPageNumber="45" masterDocId="FFA00E08FFBBE6608E5CFFF873136F26" masterDocTitle="A reappraisal of some basal lineages of the family Macrochelidae, with the description of a new genus (Acarina: Mesostigmata)" masterLastPageNumber="53" masterPageNumber="37" pageNumber="41" updateTime="1698239325266" updateUser="plazi">
<mods:mods id="1706FFD86C466B5892A2592A0855F0B5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7F57BAA648A2329A0BD80F7B7E8266AB">
<mods:title id="DD4FA3D5665E37C09342FE927E2D8340">A reappraisal of some basal lineages of the family Macrochelidae, with the description of a new genus (Acarina: Mesostigmata)</mods:title>
</mods:titleInfo>
<mods:name id="9FDB02E4CC813F19CADAC2C0B9407BB0" type="personal">
<mods:role id="BC39E9B64A9DF4853A646A49BCA69500">
<mods:roleTerm id="106A166D33410F2B57E242A25CE55694">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1A842629B17DBC65D5AF8C27BC183B9">Emberson, Rowan M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CAC769347F4DEBFE07A2F6ADCA46DEC6">text</mods:typeOfResource>
<mods:relatedItem id="980E235C212E535C57323CC489A6C686" type="host">
<mods:titleInfo id="1EF587B3F82D8CC9D139916D54D854ED">
<mods:title id="59C127E8D6A9DDAE8C62C623ABE1C466">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="700FA3CF4137CE298A13FD0A5127872A">
<mods:date id="1E9EFFF4834017E07A104E838851AA70">2010</mods:date>
<mods:detail id="22006400BF323F690BCFEA6EED7CD578" type="volume">
<mods:number id="810625BC139BE6645DE4D515EACF72A4">2501</mods:number>
</mods:detail>
<mods:extent id="95000DC2F1EFC6A0F149ABE4C9BE5739" unit="page">
<mods:start id="972C933FEEC730F9C37B7267FC818C6B">37</mods:start>
<mods:end id="4BF26440A6DBDCC6D7A844478386B2DE">53</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1DBAF070CF19CF058460ED2941EED7B1">journal article</mods:classification>
<mods:identifier id="9A8FD44C2979229C508ED3AEAB6178BC" type="DOI">10.5281/zenodo.195835</mods:identifier>
<mods:identifier id="D0C5F328E38E855607A658FF74B7E3A9" type="GBIF-Dataset">5fe3da15-3dd3-495f-9c15-7f9d606315aa</mods:identifier>
<mods:identifier id="7867C694D6E04F82F252CA9CD84E11F9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3019E9E83E993649CE0A48568E0FA231" type="Zenodo-Dep">195835</mods:identifier>
</mods:mods>
<treatment id="03997670FFBFE6688ECBFC5F773A6B5A" ID-DOI="http://doi.org/10.5281/zenodo.6212246" ID-GBIF-Taxon="119387875" ID-Zenodo-Dep="6212246" LSID="urn:lsid:plazi:treatment:03997670FFBFE6688ECBFC5F773A6B5A" httpUri="http://treatment.plazi.org/id/03997670FFBFE6688ECBFC5F773A6B5A" lastPageId="8" lastPageNumber="45" pageId="4" pageNumber="41">
<subSubSection id="C32A94EDFFBFE6648ECBFC5F72AA6CE7" box="[151,441,935,961]" pageId="4" pageNumber="41" type="nomenclature">
<paragraph id="8B8FC766FFBFE6648ECBFC5F72AA6CE7" blockId="4.[151,441,935,961]" box="[151,441,935,961]" pageId="4" pageNumber="41">
<heading id="D0C7700AFFBFE6648ECBFC5F72AA6CE7" bold="true" box="[151,441,935,961]" fontSize="11" level="1" pageId="4" pageNumber="41" reason="1">
<taxonomicName id="4C30BCE5FFBFE6648ECBFC5F72AA6CE7" authority="Valle, 1953" authorityName="Valle" authorityYear="1953" box="[151,441,935,961]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBFE6648ECBFC5F72AA6CE7" bold="true" box="[151,441,935,961]" pageId="4" pageNumber="41">
<emphasis id="B9441B74FFBFE6648ECBFC5F72396CE7" bold="true" box="[151,298,935,961]" italics="true" pageId="4" pageNumber="41">Longicheles</emphasis>
<bibRefCitation id="EFA1BA97FFBFE6648F6DFC5F72AA6CE7" author="Valle" box="[305,441,935,961]" pageId="4" pageNumber="53" refString="Valle, A. (1953) Revisione di generi e sottogeneri Berlesiani di Acari. (Primo contributo). Redia, 38, 316 - 360 + 3 plates." type="book chapter" year="1953">Valle, 1953</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A94EDFFBFE6648ECBFC0870B86BDC" pageId="4" pageNumber="41" type="reference_group">
<paragraph id="8B8FC766FFBFE6648ECBFC08725A6B01" blockId="4.[151,1436,1008,1127]" pageId="4" pageNumber="41">
<treatmentCitationGroup id="AB20E048FFBFE6648ECBFC08725A6B01" pageId="4" pageNumber="41">
<taxonomicName id="4C30BCE5FFBFE6648ECBFC0871626B21" authority="Valle, 1953: 323" authorityName="Valle" authorityPageNumber="323" authorityYear="1953" box="[151,625,1008,1031]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Longicheles">
<emphasis id="B9441B74FFBFE6648ECBFC08721C6B21" box="[151,271,1008,1031]" italics="true" pageId="4" pageNumber="41">Geholaspis</emphasis>
(
<emphasis id="B9441B74FFBFE6648F7FFC0872B76B21" box="[291,420,1008,1031]" italics="true" pageId="4" pageNumber="41">Longicheles</emphasis>
)
<treatmentCitation id="0A91E177FFBFE6648FEBFC0871626B21" author="Valle" box="[439,625,1008,1031]" page="323" pageId="4" pageNumber="41" year="1953">
<bibRefCitation id="EFA1BA97FFBFE6648FEBFC08712B6B21" author="Valle" box="[439,568,1008,1031]" pageId="4" pageNumber="53" refString="Valle, A. (1953) Revisione di generi e sottogeneri Berlesiani di Acari. (Primo contributo). Redia, 38, 316 - 360 + 3 plates." type="book chapter" year="1953">Valle, 1953</bibRefCitation>
: 323
</treatmentCitation>
</taxonomicName>
, 343.
<typeStatus id="548B79C4FFBFE6648CE6FC0871FC6B21" box="[698,751,1008,1031]" pageId="4" pageNumber="41">Type</typeStatus>
species:
<taxonomicName id="4C30BCE5FFBFE6648D05FC0876066B21" authority="Berlese, 1904" authorityName="Berlese" authorityYear="1904" box="[857,1301,1008,1031]" class="Arachnida" family="Macrochelidae" genus="Holostaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="mandibularis">
<emphasis id="B9441B74FFBFE6648D05FC0877636B21" box="[857,1136,1008,1031]" italics="true" pageId="4" pageNumber="41">Holostaspis mandibularis</emphasis>
<bibRefCitation id="EFA1BA97FFBFE6648A27FC0876066B21" author="Berlese" box="[1147,1301,1008,1031]" pageId="4" pageNumber="51" refString="Berlese, A. (1904) Acari nuovi. Manipulus II. Redia, 1, 258 - 280." type="journal article" year="1904">Berlese, 1904</bibRefCitation>
</taxonomicName>
, by original designation.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8FC766FFBFE6648ECBFBC8714C6B41" blockId="4.[151,1436,1008,1127]" pageId="4" pageNumber="41">
<treatmentCitationGroup id="AB20E048FFBFE6648ECBFBC8714C6B41" pageId="4" pageNumber="41">
<taxonomicName id="4C30BCE5FFBFE6648ECBFBC872B46B61" authority="(Longicheles)" baseAuthorityName="Longicheles" box="[151,423,1072,1095]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBFE6648ECBFBC8721C6B61" box="[151,271,1072,1095]" italics="true" pageId="4" pageNumber="41">Geholaspis</emphasis>
(
<emphasis id="B9441B74FFBFE6648F42FBC8728C6B61" box="[286,415,1072,1095]" italics="true" pageId="4" pageNumber="41">Longicheles</emphasis>
)
</taxonomicName>
.—
<treatmentCitation id="0A91E177FFBFE6648F8CFBC8719A6B61" author="Krauss" box="[464,649,1072,1095]" page="36" pageId="4" pageNumber="41" year="1970">
<bibRefCitation id="EFA1BA97FFBFE6648F8CFBC871736B61" author="Krauss" box="[464,608,1072,1095]" pageId="4" pageNumber="52" refString="Krauss, W. (1970) Die europaischen Arten der Gattungen Macrocheles Latreille 1829 und Geholaspis Berlese 1918. (Eine systematische Studie aus dem Jahr 1960). Acarologie, Schriftenreihe fur Vergleichende Milbenkunde, 14, 1 - 43 + 20 plates." type="book chapter" year="1970">Krauss, 1970</bibRefCitation>
: 36
</treatmentCitation>
;
<treatmentCitation id="0A91E177FFBFE6648CCFFBC870D86B61" author="Hyatt" box="[659,971,1072,1095]" page="74" pageId="4" pageNumber="41" year="1988">
<bibRefCitation id="EFA1BA97FFBFE6648CCFFBC870B06B61" author="Hyatt" box="[659,931,1072,1095]" pageId="4" pageNumber="52" refString="Hyatt, K. H. &amp; Emberson, R. M. (1988) A review of the Macrochelidae (Acari: Mesostigmata) of the British Isles. Bulletin of the British Museum (Natural History), Zoology, 54, 63 - 125 + 6 plates." type="book chapter" year="1988">Hyatt &amp; Emberson, 1988</bibRefCitation>
: 74
</treatmentCitation>
;
<treatmentCitation id="0A91E177FFBFE6648D8AFBC8779F6B61" author="Masan" box="[982,1164,1072,1095]" page="55" pageId="4" pageNumber="41" year="2003">
<bibRefCitation id="EFA1BA97FFBFE6648D8AFBC877776B61" author="Masan" box="[982,1124,1072,1095]" pageId="4" pageNumber="53" refString="Masan, P. (2003) Macrochelid mites of Slovakia (Acari: Mesostigmata: Macrochelidae). Institute of Zoology, Slovak Academy of Sciences, Bratislava. 149 pp." type="book" year="2003">Mašán, 2003</bibRefCitation>
: 55
</treatmentCitation>
.
<taxonomicName id="4C30BCE5FFBFE6648ECBFBA8720B6B41" box="[151,280,1104,1127]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBFE6648ECBFBA8720B6B41" box="[151,280,1104,1127]" italics="true" pageId="4" pageNumber="41">Longicheles</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0A91E177FFBFE6648F1CFBA871486B41" author="Athias-Henriot" box="[320,603,1104,1127]" page="241" pageId="4" pageNumber="41" year="1968">
<bibRefCitation id="EFA1BA97FFBFE6648F1CFBA871356B41" author="Athias-Henriot" box="[320,550,1104,1127]" pageId="4" pageNumber="51" refString="Athias-Henriot, C. (1968) L'appareil d'insemination laelapoide (Acariens anactinotriches: Laelapoidea, P). Premieres observations. Possibilite d'emploi a des fins taxonomiques. Bulletin scientifique de Bourgogne, 25, 229 - 274." type="journal article" year="1968">Athias-Henriot, 1968</bibRefCitation>
: 241
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8FC766FFBFE6648ECBFB6D70B86BDC" blockId="4.[151,1437,1173,1983]" pageId="4" pageNumber="41">
Geholaspini with massively developed, elongate, multidentate chelicerae (
<figureCitation id="130BDBE3FFBFE6648A7BFB6D77606B89" box="[1063,1139,1173,1199]" captionStart="FIGURES 1 7. 1 3" captionStartId="5.[151,269,1907,1931]" captionTargetBox="[187,1412,188,1863]" captionTargetId="figure@5.[151,1436,159,1883]" captionTargetPageId="5" captionText="FIGURES 1 7. 1 3, Dissoloncha superbus (Hull), female. 1. Epistome; 2. Ventri-anal shield; 3. Sacculus complex. 4 5. Geholaspis longispinosus (Kramer), female. 4. Chelicera; 5. Epistome. 6. Longicheles hortorum (Berlese) comb. nov., female, chelicera with accessory teeth on external face. 7. Longicheles mandibularis (Berlese), female, epistome." httpUri="https://zenodo.org/record/195836/files/figure.png" pageId="4" pageNumber="41">Fig. 6</figureCitation>
); epistome with narrow, distally bifid median process (
<figureCitation id="130BDBE3FFBFE6648C58FB42715D6BF2" box="[516,590,1210,1236]" captionStart="FIGURES 1 7. 1 3" captionStartId="5.[151,269,1907,1931]" captionTargetBox="[187,1412,188,1863]" captionTargetId="figure@5.[151,1436,159,1883]" captionTargetPageId="5" captionText="FIGURES 1 7. 1 3, Dissoloncha superbus (Hull), female. 1. Epistome; 2. Ventri-anal shield; 3. Sacculus complex. 4 5. Geholaspis longispinosus (Kramer), female. 4. Chelicera; 5. Epistome. 6. Longicheles hortorum (Berlese) comb. nov., female, chelicera with accessory teeth on external face. 7. Longicheles mandibularis (Berlese), female, epistome." httpUri="https://zenodo.org/record/195836/files/figure.png" pageId="4" pageNumber="41">Fig. 7</figureCitation>
); dorsal setae j5 strongly displaced posteriorly, lying well posterior to setae j6 (
<figureCitation id="130BDBE3FFBFE6648EA3FB1872556BDC" box="[255,326,1248,1274]" captionStart="FIGURES 8 11. 8 9" captionStartId="6.[151,270,1681,1705]" captionTargetBox="[159,1420,186,1607]" captionTargetId="figure@6.[151,1436,140,1660]" captionTargetPageId="6" captionText="FIGURES 8 11. 8 9. Longicheles hortorum (Berlese), female. 8. Dorsum, showing posterior displacement of setae j 5; 9. Anterior dorsal region, showing position and characters of setae j 1, j 2 and z 1. 10. Longicheles longulus (Berlese) comb. nov., female, anterior dorsal region, showing position and characters of setae j 1, j 2 and z 1. 11. Reductholaspis analis (Hyatt &amp; Emberson) comb. nov., female, dorsal shield." httpUri="https://zenodo.org/record/195837/files/figure.png" pageId="4" pageNumber="41">Fig. 8</figureCitation>
); ventri-anal shield with five pairs of pre-anal setae.
</paragraph>
</subSubSection>
<subSubSection id="C32A94EDFFBFE6688E98FAFD773A6B5A" lastPageId="8" lastPageNumber="45" pageId="4" pageNumber="41" type="description">
<paragraph id="8B8FC766FFBFE6648E98FAFD71D1680C" blockId="4.[151,1437,1173,1983]" pageId="4" pageNumber="41">
The dorsal shield has 28 pairs of setae, which are mainly pilose or plumose distally, except for a variable number of aciculate setae in the j-J series and sometimes one to three setae in the z-Z series. Setae z
<date id="FF8EE1A6FFBFE6648B6FFAD2766A6A62" box="[1331,1401,1322,1348]" pageId="4" pageNumber="41">1 may</date>
be short and pilose, or aciculate to slightly spatulate, and longer than j1. Setae j5 are displaced posteriorly, so that they are mesad and well posterior to j6 (
<figureCitation id="130BDBE3FFBFE6648CCEFA8D71CC6AA9" box="[658,735,1397,1423]" captionStart="FIGURES 8 11. 8 9" captionStartId="6.[151,270,1681,1705]" captionTargetBox="[159,1420,186,1607]" captionTargetId="figure@6.[151,1436,140,1660]" captionTargetPageId="6" captionText="FIGURES 8 11. 8 9. Longicheles hortorum (Berlese), female. 8. Dorsum, showing posterior displacement of setae j 5; 9. Anterior dorsal region, showing position and characters of setae j 1, j 2 and z 1. 10. Longicheles longulus (Berlese) comb. nov., female, anterior dorsal region, showing position and characters of setae j 1, j 2 and z 1. 11. Reductholaspis analis (Hyatt &amp; Emberson) comb. nov., female, dorsal shield." httpUri="https://zenodo.org/record/195837/files/figure.png" pageId="4" pageNumber="41">Fig. 8</figureCitation>
). The ventral setae are mostly simple except toward the posterior lateral margins. The sternal shield has punctate sculpture, without a distinct pattern of lines, and the metasternal plates are free and have the usual seta and pore. The epigynal shield is subtriangular, with a truncate posterior margin and covers strongly developed lateral epigynal sclerites. The ventri-anal shield is usually longer than broad; it has five pairs of pre-anal setae; one or both pairs of the Zv series may be plumose, and the inguinal pores are near the anterior lateral angles. The cheliceral digits are elongate and multidentate, the fixed chela always has more than five teeth, often many more (
<figureCitation id="130BDBE3FFBFE6648A12F9AD77856949" box="[1102,1174,1621,1647]" captionStart="FIGURES 1 7. 1 3" captionStartId="5.[151,269,1907,1931]" captionTargetBox="[187,1412,188,1863]" captionTargetId="figure@5.[151,1436,159,1883]" captionTargetPageId="5" captionText="FIGURES 1 7. 1 3, Dissoloncha superbus (Hull), female. 1. Epistome; 2. Ventri-anal shield; 3. Sacculus complex. 4 5. Geholaspis longispinosus (Kramer), female. 4. Chelicera; 5. Epistome. 6. Longicheles hortorum (Berlese) comb. nov., female, chelicera with accessory teeth on external face. 7. Longicheles mandibularis (Berlese), female, epistome." httpUri="https://zenodo.org/record/195836/files/figure.png" pageId="4" pageNumber="41">Fig. 6</figureCitation>
), and may have a row of accessory teeth on the external face (
<taxonomicName id="4C30BCE5FFBFE6648C3FF983707C69B2" authority="Berlese" authorityName="Berlese" box="[611,879,1658,1684]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="hortorum">
<emphasis id="B9441B74FFBFE6648C3FF983716069B2" box="[611,627,1659,1684]" italics="true" pageId="4" pageNumber="41">L</emphasis>
.
<emphasis id="B9441B74FFBFE6648CDDF98371E369B2" box="[641,752,1659,1684]" italics="true" pageId="4" pageNumber="41">hortorum</emphasis>
(Berlese))
</taxonomicName>
, while the movable chela has at least eight teeth and often have a row of accessory teeth on the external face (
<figureCitation id="130BDBE3FFBFE6648DCCF95870CF699C" box="[912,988,1696,1722]" captionStart="FIGURES 1 7. 1 3" captionStartId="5.[151,269,1907,1931]" captionTargetBox="[187,1412,188,1863]" captionTargetId="figure@5.[151,1436,159,1883]" captionTargetPageId="5" captionText="FIGURES 1 7. 1 3, Dissoloncha superbus (Hull), female. 1. Epistome; 2. Ventri-anal shield; 3. Sacculus complex. 4 5. Geholaspis longispinosus (Kramer), female. 4. Chelicera; 5. Epistome. 6. Longicheles hortorum (Berlese) comb. nov., female, chelicera with accessory teeth on external face. 7. Longicheles mandibularis (Berlese), female, epistome." httpUri="https://zenodo.org/record/195836/files/figure.png" pageId="4" pageNumber="41">Fig. 6</figureCitation>
). The dentate anterior margin of the epistome is more or less transverse and has a narrow, bifid, median process, which may be distally dissected (
<figureCitation id="130BDBE3FFBFE6648EFCF91273FA6822" box="[160,233,1770,1796]" captionStart="FIGURES 1 7. 1 3" captionStartId="5.[151,269,1907,1931]" captionTargetBox="[187,1412,188,1863]" captionTargetId="figure@5.[151,1436,159,1883]" captionTargetPageId="5" captionText="FIGURES 1 7. 1 3, Dissoloncha superbus (Hull), female. 1. Epistome; 2. Ventri-anal shield; 3. Sacculus complex. 4 5. Geholaspis longispinosus (Kramer), female. 4. Chelicera; 5. Epistome. 6. Longicheles hortorum (Berlese) comb. nov., female, chelicera with accessory teeth on external face. 7. Longicheles mandibularis (Berlese), female, epistome." httpUri="https://zenodo.org/record/195836/files/figure.png" pageId="4" pageNumber="41">Fig. 7</figureCitation>
). Spermathecal structures are generally not apparent, but the elongate tubuli and rami of two species have been illustrated by
<bibRefCitation id="EFA1BA97FFBFE6648FE9F8E871AD680C" author="Athias-Henriot" box="[437,702,1808,1834]" pageId="4" pageNumber="51" refString="Athias-Henriot, C. (1968) L'appareil d'insemination laelapoide (Acariens anactinotriches: Laelapoidea, P). Premieres observations. Possibilite d'emploi a des fins taxonomiques. Bulletin scientifique de Bourgogne, 25, 229 - 274." type="journal article" year="1968">Athias-Henriot (1968)</bibRefCitation>
.
</paragraph>
<paragraph id="8B8FC766FFBFE6648E98F8CD729B6869" blockId="4.[151,1437,1173,1983]" box="[196,392,1845,1871]" pageId="4" pageNumber="41">Males unknown.</paragraph>
<paragraph id="8B8FC766FFBFE6648E98F8A2727B6899" blockId="4.[151,1437,1173,1983]" pageId="4" pageNumber="41">Protonymphs, where known, have a posterior extension of the hind margin of the podonotal shield into a median lobe bearing setae j5. This extension is probably associated with a posterior migration of the cheliceral retractor muscles.</paragraph>
<caption id="DF4F97EEFFBEE6658ECBF88B765E68EC" httpUri="https://zenodo.org/record/195836/files/figure.png" pageId="5" pageNumber="42" targetBox="[187,1412,188,1863]" targetPageId="5">
<paragraph id="8B8FC766FFBEE6658ECBF88B765E68EC" blockId="5.[151,1436,1907,1994]" pageId="5" pageNumber="42">
<emphasis id="B9441B74FFBEE6658ECBF88B725168AD" bold="true" box="[151,322,1907,1931]" pageId="5" pageNumber="42">FIGURES 17.</emphasis>
13,
<taxonomicName id="4C30BCE5FFBEE6658F21F88B71A068AC" authority="Hull" authorityName="Hull" box="[381,691,1907,1930]" class="Arachnida" family="Macrochelidae" genus="Dissoloncha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="superbus">
<emphasis id="B9441B74FFBEE6658F21F88B717A68AC" box="[381,617,1907,1930]" italics="true" pageId="5" pageNumber="42">Dissoloncha superbus</emphasis>
(Hull)
</taxonomicName>
, female. 1. Epistome; 2. Ventri-anal shield; 3. Sacculus complex. 45.
<taxonomicName id="4C30BCE5FFBEE6658ECBF86B7109688C" authority="Kramer" authorityName="Kramer" box="[151,538,1939,1962]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="longispinosus">
<emphasis id="B9441B74FFBEE6658ECBF86B72BF688C" box="[151,428,1939,1962]" italics="true" pageId="5" pageNumber="42">Geholaspis longispinosus</emphasis>
(Kramer)
</taxonomicName>
, female. 4. Chelicera; 5. Epistome. 6.
<taxonomicName id="4C30BCE5FFBEE6658DE0F86B7605688C" authority="Berlese" authorityName="Berlese" box="[956,1302,1939,1962]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="hortorum" status="comb. nov.">
<emphasis id="B9441B74FFBEE6658DE0F86B77BF688C" box="[956,1196,1939,1962]" italics="true" pageId="5" pageNumber="42">Longicheles hortorum</emphasis>
(Berlese)
</taxonomicName>
<taxonomicNameLabel id="A277A60FFFBEE6658B42F86B768B688C" box="[1310,1432,1939,1962]" pageId="5" pageNumber="42">comb. nov.</taxonomicNameLabel>
, female, chelicera with accessory teeth on external face. 7.
<taxonomicName id="4C30BCE5FFBEE6658D58F84B779668EC" authority="Berlese" authorityName="Berlese" box="[772,1157,1971,1994]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="mandibularis">
<emphasis id="B9441B74FFBEE6658D58F84B770968EC" box="[772,1050,1971,1994]" italics="true" pageId="5" pageNumber="42">Longicheles mandibularis</emphasis>
(Berlese)
</taxonomicName>
, female, epistome.
</paragraph>
</caption>
<caption id="DF4F97EEFFBDE6668ECBF969703B682E" httpUri="https://zenodo.org/record/195837/files/figure.png" pageId="6" pageNumber="43" targetBox="[159,1420,186,1607]" targetPageId="6">
<paragraph id="8B8FC766FFBDE6668ECBF969703B682E" blockId="6.[151,1437,1681,1801]" pageId="6" pageNumber="43">
<emphasis id="B9441B74FFBDE6668ECBF9697243698F" bold="true" box="[151,336,1681,1705]" pageId="6" pageNumber="43">FIGURES 811.</emphasis>
89.
<taxonomicName id="4C30BCE5FFBDE6668FD1F96A71F4698E" authority="Berlese" authorityName="Berlese" box="[397,743,1681,1705]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="hortorum">
<emphasis id="B9441B74FFBDE6668FD1F96A7169698F" box="[397,634,1682,1705]" italics="true" pageId="6" pageNumber="43">Longicheles hortorum</emphasis>
(Berlese)
</taxonomicName>
, female. 8. Dorsum, showing posterior displacement of setae j5; 9. Anterior dorsal region, showing position and characters of setae j1, j2 and z1. 10.
<taxonomicName id="4C30BCE5FFBDE6668A18F94A768E69EE" authority="Berlese" authorityName="Berlese" box="[1092,1437,1713,1737]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="longulus" status="comb. nov.">
<emphasis id="B9441B74FFBDE6668A18F94A762369EF" box="[1092,1328,1714,1737]" italics="true" pageId="6" pageNumber="43">Longicheles longulus</emphasis>
(Berlese)
</taxonomicName>
<taxonomicNameLabel id="A277A60FFFBDE6668ECBF929720169CE" box="[151,274,1745,1768]" pageId="6" pageNumber="43">comb. nov.</taxonomicNameLabel>
, female, anterior dorsal region, showing position and characters of setae j1, j2 and z1. 11.
<taxonomicName id="4C30BCE5FFBDE6668AAAF92A72A9682E" authority="Hyatt &amp; Emberson" authorityName="Hyatt &amp; Emberson" class="Arachnida" family="Macrochelidae" genus="Reductholaspis" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="analis" status="comb. nov.">
<emphasis id="B9441B74FFBDE6668AAAF92A73CB682F" italics="true" pageId="6" pageNumber="43">Reductholaspis analis</emphasis>
(Hyatt &amp; Emberson)
</taxonomicName>
<taxonomicNameLabel id="A277A60FFFBDE6668F9CF909712A682E" box="[448,569,1777,1800]" pageId="6" pageNumber="43">comb. nov.</taxonomicNameLabel>
, female, dorsal shield.
</paragraph>
</caption>
<paragraph id="8B8FC766FFBDE6678E9AF8C972C06EB4" blockId="6.[151,1437,1841,2017]" lastBlockId="7.[151,1437,152,402]" lastPageId="7" lastPageNumber="44" pageId="6" pageNumber="43">
<emphasis id="B9441B74FFBDE6668E9AF8C97207686D" bold="true" box="[198,276,1841,1867]" pageId="6" pageNumber="43">Notes.</emphasis>
<bibRefCitation id="EFA1BA97FFBDE6668F47F8CA72BE686A" author="Valle" box="[283,429,1842,1868]" pageId="6" pageNumber="53" refString="Valle, A. (1953) Revisione di generi e sottogeneri Berlesiani di Acari. (Primo contributo). Redia, 38, 316 - 360 + 3 plates." type="book chapter" year="1953">Valle (1953)</bibRefCitation>
originally distinguished
<taxonomicName id="4C30BCE5FFBDE6668C8CF8CA704D686D" box="[720,862,1842,1867]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBDE6668C8CF8CA704D686D" box="[720,862,1842,1867]" italics="true" pageId="6" pageNumber="43">Longicheles</emphasis>
</taxonomicName>
from other subgenera of
<taxonomicName id="4C30BCE5FFBDE6668ADAF8CA7619686D" box="[1158,1290,1842,1867]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBDE6668ADAF8CA7619686D" box="[1158,1290,1842,1867]" italics="true" pageId="6" pageNumber="43">Geholaspis</emphasis>
</taxonomicName>
on the basis of two main characters: the elongate nature of the cheliceral digits in
<taxonomicName id="4C30BCE5FFBDE6668A45F8AF77A16856" box="[1049,1202,1879,1904]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBDE6668A45F8AF77A16856" box="[1049,1202,1879,1904]" italics="true" pageId="6" pageNumber="43">Longicheles</emphasis>
</taxonomicName>
, and the posterior displacement of setae j5 (setae
<quantity id="4CC86A83FFBDE6668C58F884712C68B0" box="[516,575,1916,1942]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="6" pageNumber="43" unit="in" value="18.0">18 in</quantity>
his system). The length of the cheliceral digits was expressed as the ratio of dorsal shield length to the length of the movable chela. The posterior displacement of setae j5 was expressed as the ratio of dorsal shield length to the distance between setae z5 and j5 (setae 13 and
<quantity id="4CC86A83FFBDE6668B68F83F766268C7" box="[1332,1393,1991,2017]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="6" pageNumber="43" unit="in" value="18.0">18 in</quantity>
his system). Valles characters are still considered valid and important in defining the genus.
<bibRefCitation id="EFA1BA97FFBCE6678AE4FF6073FA6FF1" author="Evans" pageId="7" pageNumber="52" refString="Evans, G. O. &amp; Browning, E. (1956) British mites of the subfamily Macrochelinae Tragardh (Gamasina - Macrochelidae). Bulletin of the British Museum (Natural History), Zoology, 4, 1 - 56 + 4 plates." type="book chapter" year="1956">Evans &amp; Browning (1956)</bibRefCitation>
introduced an additional character based on the length of setae z1 (M
<quantity id="4CC86A83FFBCE6678A11FF45776B6FF1" box="[1101,1144,189,215]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="44" unit="in" value="1.0">1 in</quantity>
their system), in which
<taxonomicName id="4C30BCE5FFBCE6678ECBFF1A72366FDD" box="[151,293,226,251]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBCE6678ECBFF1A72366FDD" box="[151,293,226,251]" italics="true" pageId="7" pageNumber="44">Longicheles</emphasis>
</taxonomicName>
was thought to have short setae and
<emphasis id="B9441B74FFBCE6678C8EFF1A707B6FDD" box="[722,872,226,251]" italics="true" pageId="7" pageNumber="44">
<taxonomicName id="4C30BCE5FFBCE6678C8EFF1A70456FDD" box="[722,854,226,251]" class="Arachnida" family="Macrochelidae" genus="Geholaspis" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus" sensu="stricto" status="s">Geholaspis</taxonomicName>
<taxonomicNameLabel id="A277A60FFFBCE6678D01FF1A707B6FDD" box="[861,872,226,251]" pageId="7" pageNumber="44" sensu="stricto">s</taxonomicNameLabel>
</emphasis>
.
<emphasis id="B9441B74FFBCE6678D2BFF1A70876FDD" box="[887,916,226,251]" italics="true" pageId="7" pageNumber="44">str</emphasis>
. to have long, simple setae. This was shown by
<bibRefCitation id="EFA1BA97FFBCE6678EE1FEF072796E04" author="Balogh" box="[189,362,264,290]" pageId="7" pageNumber="51" refString="Balogh, J. (1958) Macrocheliden aus Bulgarien (Acarina, Mesostigmata). Acta entomologica Musei Nationalis Prague, 32, 247 - 256." type="journal article" year="1958">Balogh (1958)</bibRefCitation>
not to hold for all species of
<taxonomicName id="4C30BCE5FFBCE6678C91FEF0704F6E07" box="[717,860,264,289]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBCE6678C91FEF0704F6E07" box="[717,860,264,289]" italics="true" pageId="7" pageNumber="44">Longicheles</emphasis>
</taxonomicName>
. He described
<taxonomicName id="4C30BCE5FFBCE6678A4DFEF076316E04" authority="Balogh" authorityName="Balogh" box="[1041,1314,264,290]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="longisetosus">
<emphasis id="B9441B74FFBCE6678A4DFEF077326E07" box="[1041,1057,264,289]" italics="true" pageId="7" pageNumber="44">L</emphasis>
.
<emphasis id="B9441B74FFBCE6678A6DFEF077D76E07" box="[1073,1220,264,289]" italics="true" pageId="7" pageNumber="44">longisetosus</emphasis>
Balogh
</taxonomicName>
as having long, simple z1 setae, but with all the other characters of
<taxonomicName id="4C30BCE5FFBCE6678D15FED570CA6E60" box="[841,985,301,326]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBCE6678D15FED570CA6E60" box="[841,985,301,326]" italics="true" pageId="7" pageNumber="44">Longicheles</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFA1BA97FFBCE6678DB5FED5776D6E61" author="Valle" box="[1001,1150,301,327]" pageId="7" pageNumber="53" refString="Valle, A. (1963) Intorno ad una nuova specie del genere Geholaspis, Berl. (Acari, Macrochelidae). Bollettino delle sedute dell'Accademia Gioenia di Scienze naturali in Catania, 7, 419 - 427." type="journal article" year="1963">Valle (1963)</bibRefCitation>
and
<bibRefCitation id="EFA1BA97FFBCE6678AE6FED573F56E4A" author="Valle" pageId="7" pageNumber="53" refString="Valle, A., &amp; Mazzoleni, M. (1967) Nuove specie Mediterranee del genere Geholaspis Berl. e considerazioni intorno al sottogenere Longicheles Valle (Acari, Macrochelidae). Rendiconti Istituto Lombardo - accademia di Scienze e Lettere, 101 B, 385 - 403 + 5 plates." type="book chapter" year="1967">Valle &amp; Mazzoleni (1967)</bibRefCitation>
subsequently described four additional species of
<taxonomicName id="4C30BCE5FFBCE6678D60FEAA70D96E4D" box="[828,970,338,363]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFBCE6678D60FEAA70D96E4D" box="[828,970,338,363]" italics="true" pageId="7" pageNumber="44">Longicheles</emphasis>
</taxonomicName>
, from Mediterranean islands, which all have long, simple z1 setae.
</paragraph>
<caption id="DF4F97EEFFBCE6678ECBF86471CB68D5" httpUri="https://zenodo.org/record/195838/files/figure.png" pageId="7" pageNumber="44" targetBox="[202,1362,402,1947]" targetPageId="7">
<paragraph id="8B8FC766FFBCE6678ECBF86471CB68D5" blockId="7.[151,1436,1948,2036]" pageId="7" pageNumber="44">
<emphasis id="B9441B74FFBCE6678ECBF864724D6892" bold="true" box="[151,350,1948,1972]" pageId="7" pageNumber="44">FIGURES 1217.</emphasis>
1215.
<taxonomicName id="4C30BCE5FFBCE6678FEAF86470956895" authority="Hyatt &amp; Emberson" authorityName="Hyatt &amp; Emberson" box="[438,902,1948,1971]" class="Arachnida" family="Macrochelidae" genus="Reductholaspis" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="analis" status="comb. nov.">
<emphasis id="B9441B74FFBCE6678FEAF86471B06895" box="[438,675,1948,1971]" italics="true" pageId="7" pageNumber="44">Reductholaspis analis</emphasis>
(Hyatt &amp; Emberson)
</taxonomicName>
<taxonomicNameLabel id="A277A60FFFBCE6678DD2F86477146895" box="[910,1031,1948,1971]" pageId="7" pageNumber="44">comb. nov.</taxonomicNameLabel>
, female. 12. Chelicera; 13. Epistome; 14. Anal shield; 15. gnathosoma venter, showing position of hypostomal setae. 16.
<taxonomicName id="4C30BCE5FFBCE6678A68F844768B68F5" authority="Koch" authorityName="Koch" box="[1076,1432,1980,2003]" class="Arachnida" family="Macrochelidae" genus="Nothrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="B9441B74FFBCE6678A68F84477DC68F5" box="[1076,1231,1980,2003]" italics="true" pageId="7" pageNumber="44">Nothrholaspis</emphasis>
<emphasis id="B9441B74FFBCE6678A84F844765368F5" box="[1240,1344,1980,2003]" italics="true" pageId="7" pageNumber="44">carinatus</emphasis>
(Koch)
</taxonomicName>
, epistome. 17.
<taxonomicName id="4C30BCE5FFBCE6678F70F824717568D5" authority="Koch" authorityName="Koch" box="[300,614,2012,2035]" class="Arachnida" family="Macrochelidae" genus="Macrholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="opacus">
<emphasis id="B9441B74FFBCE6678F70F824710268D5" box="[300,529,2012,2035]" italics="true" pageId="7" pageNumber="44">Macrholaspis opacus</emphasis>
(Koch)
</taxonomicName>
, epistome.
</paragraph>
</caption>
<paragraph id="8B8FC766FFB3E6688E98FF6072B96EFA" blockId="8.[151,1437,152,1148]" pageId="8" pageNumber="45">
<bibRefCitation id="EFA1BA97FFB3E6688E98FF6071186F94" author="Valle" box="[196,523,152,178]" pageId="8" pageNumber="53" refString="Valle, A., &amp; Mazzoleni, M. (1967) Nuove specie Mediterranee del genere Geholaspis Berl. e considerazioni intorno al sottogenere Longicheles Valle (Acari, Macrochelidae). Rendiconti Istituto Lombardo - accademia di Scienze e Lettere, 101 B, 385 - 403 + 5 plates." type="book chapter" year="1967">Valle &amp; Mazzoleni (1967)</bibRefCitation>
divided the species of
<taxonomicName id="4C30BCE5FFB3E6688D72FF6070D76F97" box="[814,964,152,177]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB3E6688D72FF6070D76F97" box="[814,964,152,177]" italics="true" pageId="8" pageNumber="45">Longicheles</emphasis>
</taxonomicName>
into four groups based mainly on the dentition and relative length of the cheliceral digits, but also the pilosity of seta j2. However,
<bibRefCitation id="EFA1BA97FFB3E6688AAAFF4576886FF1" author="Masan" box="[1270,1435,189,215]" pageId="8" pageNumber="53" refString="Masan, P. (2003) Macrochelid mites of Slovakia (Acari: Mesostigmata: Macrochelidae). Institute of Zoology, Slovak Academy of Sciences, Bratislava. 149 pp." type="book" year="2003">Mašán (2003)</bibRefCitation>
has pointed out that in Central Europe there are two distinct groups of species within
<taxonomicName id="4C30BCE5FFB3E6688AC8FF1A76316FDD" box="[1172,1314,226,251]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB3E6688AC8FF1A76316FDD" box="[1172,1314,226,251]" italics="true" pageId="8" pageNumber="45">Longicheles</emphasis>
</taxonomicName>
, based on the length, pilosity, and position of the j1 and z1 setae. These groupings can be extended to all known species of
<taxonomicName id="4C30BCE5FFB3E6688EEAFED572576E60" box="[182,324,301,326]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB3E6688EEAFED572576E60" box="[182,324,301,326]" italics="true" pageId="8" pageNumber="45">Longicheles</emphasis>
</taxonomicName>
and are here referred to as the
<taxonomicName id="4C30BCE5FFB3E6688CF2FED570596E60" box="[686,842,301,326]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="mandibularis">
<emphasis id="B9441B74FFB3E6688CF2FED570596E60" box="[686,842,301,326]" italics="true" pageId="8" pageNumber="45">mandibularis</emphasis>
</taxonomicName>
species group, in which setae j1 are close together so that their bases are virtually contiguous and setae z1 are short and distally pilose (
<figureCitation id="130BDBE3FFB3E6688AE0FEAA763A6E4A" box="[1212,1321,338,364]" captionStart="FIGURES 8 11. 8 9" captionStartId="6.[151,270,1681,1705]" captionTargetBox="[159,1420,186,1607]" captionTargetId="figure@6.[151,1436,140,1660]" captionTargetPageId="6" captionText="FIGURES 8 11. 8 9. Longicheles hortorum (Berlese), female. 8. Dorsum, showing posterior displacement of setae j 5; 9. Anterior dorsal region, showing position and characters of setae j 1, j 2 and z 1. 10. Longicheles longulus (Berlese) comb. nov., female, anterior dorsal region, showing position and characters of setae j 1, j 2 and z 1. 11. Reductholaspis analis (Hyatt &amp; Emberson) comb. nov., female, dorsal shield." httpUri="https://zenodo.org/record/195837/files/figure.png" pageId="8" pageNumber="45">Fig. 8, 9</figureCitation>
), and the
<taxonomicName id="4C30BCE5FFB3E6688ECBFE8073EE6EB7" box="[151,253,376,401]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B9441B74FFB3E6688ECBFE8073EE6EB7" box="[151,253,376,401]" italics="true" pageId="8" pageNumber="45">longulus</emphasis>
</taxonomicName>
species group, in which setae j1 are distinctly separated and setae z1 are longer than setae j1 (
<figureCitation id="130BDBE3FFB3E6688B30FE8073AB6E91" captionStart="FIGURES 8 11. 8 9" captionStartId="6.[151,270,1681,1705]" captionTargetBox="[159,1420,186,1607]" captionTargetId="figure@6.[151,1436,140,1660]" captionTargetPageId="6" captionText="FIGURES 8 11. 8 9. Longicheles hortorum (Berlese), female. 8. Dorsum, showing posterior displacement of setae j 5; 9. Anterior dorsal region, showing position and characters of setae j 1, j 2 and z 1. 10. Longicheles longulus (Berlese) comb. nov., female, anterior dorsal region, showing position and characters of setae j 1, j 2 and z 1. 11. Reductholaspis analis (Hyatt &amp; Emberson) comb. nov., female, dorsal shield." httpUri="https://zenodo.org/record/195837/files/figure.png" pageId="8" pageNumber="45">Fig. 10</figureCitation>
). In the
<taxonomicName id="4C30BCE5FFB3E6688F47FE6572926E90" box="[283,385,413,438]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B9441B74FFB3E6688F47FE6572926E90" box="[283,385,413,438]" italics="true" pageId="8" pageNumber="45">longulus</emphasis>
</taxonomicName>
species group, setae z1 are usually aciculate or occasionally slightly spatulate/pectinate (
<taxonomicName id="4C30BCE5FFB3E6688EFDFE3A728D6EFA" authority="Balogh" authorityName="Balogh" box="[161,414,450,476]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="bulgaricus">
<emphasis id="B9441B74FFB3E6688EFDFE3A73A26EFD" box="[161,177,450,475]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688EE3FE3A722F6EFD" box="[191,316,450,475]" italics="true" pageId="8" pageNumber="45">bulgaricus</emphasis>
Balogh
</taxonomicName>
).
</paragraph>
<paragraph id="8B8FC766FFB3E6688E98FE1077816C21" blockId="8.[151,1437,152,1148]" pageId="8" pageNumber="45">
The most distinctive feature of
<taxonomicName id="4C30BCE5FFB3E6688C1CFE1071C26D27" box="[576,721,488,513]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB3E6688C1CFE1071C26D27" box="[576,721,488,513]" italics="true" pageId="8" pageNumber="45">Longicheles</emphasis>
</taxonomicName>
is the development of the elongate, multidentate cheliceral digits, which are unique in the family though somewhat similar to those of
<taxonomicName id="4C30BCE5FFB3E6688A77FDF5768F6D01" authority="Krantz. The" authorityName="Krantz. The" box="[1067,1436,525,551]" class="Arachnida" family="Macrochelidae" genus="Lordocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="rykei">
<emphasis id="B9441B74FFB3E6688A77FDF576106D00" box="[1067,1283,525,550]" italics="true" pageId="8" pageNumber="45">Lordocheles rykei</emphasis>
Krantz. The
</taxonomicName>
fixed chela is nearly four times as long (from the tip to the dorsal seta) as they are wide. This development may be associated with the posterior migration of dorsal setae j5 to a position well posterior to j6 (
<bibRefCitation id="EFA1BA97FFB3E6688B57FDA0769D6D54" author="Valle" box="[1291,1422,600,626]" pageId="8" pageNumber="53" refString="Valle, A. (1953) Revisione di generi e sottogeneri Berlesiani di Acari. (Primo contributo). Redia, 38, 316 - 360 + 3 plates." type="book chapter" year="1953">Valle 1953</bibRefCitation>
), and the posterior projection of the hind margin of the podonotal shield in protonymphs of, at least,
<taxonomicName id="4C30BCE5FFB3E6688BDFFD8572B66D9A" authority="Berlese" authorityName="Berlese" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="mandibularis">
<emphasis id="B9441B74FFB3E6688BDFFD8576806DB0" box="[1411,1427,637,662]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688ECBFD5A72206D9D" box="[151,307,674,699]" italics="true" pageId="8" pageNumber="45">mandibularis</emphasis>
(Berlese)
</taxonomicName>
and
<taxonomicName id="4C30BCE5FFB3E6688F83FD5A71CC6D9A" authority="Berlese" authorityName="Berlese" box="[479,735,674,700]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="hortorum">
<emphasis id="B9441B74FFB3E6688F83FD5A72FC6D9D" box="[479,495,674,699]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688FA2FD5A717E6D9D" box="[510,621,674,699]" italics="true" pageId="8" pageNumber="45">hortorum</emphasis>
(Berlese)
</taxonomicName>
(
<bibRefCitation id="EFA1BA97FFB3E6688CB3FD5A70D76D9A" author="Hirschmann" box="[751,964,674,700]" pageId="8" pageNumber="52" refString="Hirschmann, W. (1987) Chaetotaxie und porotaxie bei Geholaspis mandibularis (Berlese 1904) und Geholaspis longispinosus (Kramer 1876) (Parasitiformes: Trichopygidiina). Acarologia, 28, 107 - 115." type="journal article" year="1987">Hirschmann 1987</bibRefCitation>
,
<bibRefCitation id="EFA1BA97FFB3E6688D8DFD5A77E66D9A" author="Hyatt" box="[977,1269,674,700]" pageId="8" pageNumber="52" refString="Hyatt, K. H. &amp; Emberson, R. M. (1988) A review of the Macrochelidae (Acari: Mesostigmata) of the British Isles. Bulletin of the British Museum (Natural History), Zoology, 54, 63 - 125 + 6 plates." type="book chapter" year="1988">Hyatt &amp; Emberson 1988</bibRefCitation>
). This suite of unique apomorphic characters fully justifies the elevation of
<taxonomicName id="4C30BCE5FFB3E6688D24FD30771A6DC7" box="[888,1033,712,737]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB3E6688D24FD30771A6DC7" box="[888,1033,712,737]" italics="true" pageId="8" pageNumber="45">Longicheles</emphasis>
</taxonomicName>
to separate generic status as first suggested by
<bibRefCitation id="EFA1BA97FFB3E6688F6AFD15712D6C21" author="Athias-Henriot" box="[310,574,749,775]" pageId="8" pageNumber="51" refString="Athias-Henriot, C. (1968) L'appareil d'insemination laelapoide (Acariens anactinotriches: Laelapoidea, P). Premieres observations. Possibilite d'emploi a des fins taxonomiques. Bulletin scientifique de Bourgogne, 25, 229 - 274." type="journal article" year="1968">Athias-Henriot (1968)</bibRefCitation>
, but not generally accepted by subsequent authors.
</paragraph>
<paragraph id="8B8FC766FFB3E6688E98FCEA71506CE4" blockId="8.[151,1437,152,1148]" pageId="8" pageNumber="45">
Although known from North
<collectingCountry id="F32787F6FFB3E6688C6EFCEA718C6C0A" box="[562,671,786,812]" name="United States of America" pageId="8" pageNumber="45">America</collectingCountry>
,
<taxonomicName id="4C30BCE5FFB3E6688CF0FCEA70526C0D" box="[684,833,786,811]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B9441B74FFB3E6688CF0FCEA70526C0D" box="[684,833,786,811]" italics="true" pageId="8" pageNumber="45">Longicheles</emphasis>
</taxonomicName>
is mainly European in distribution, having been recorded from
<collectingCountry id="F32787F6FFB3E6688F1AFCC0728A6C74" box="[326,409,824,850]" name="Ireland" pageId="8" pageNumber="45">Ireland</collectingCountry>
to European
<collectingCountry id="F32787F6FFB3E6688C64FCC071956C74" box="[568,646,824,850]" name="Russia" pageId="8" pageNumber="45">Russia</collectingCountry>
(
<bibRefCitation id="EFA1BA97FFB3E6688CCAFCC070FB6C74" author="Bregetova" box="[662,1000,824,850]" pageId="8" pageNumber="51" refString="Bregetova, N. G. &amp; Koroleva, E. V. (1960) The macrochelid mites (Gamasoidea, Macrochelidae) in the USSR. Akademiia Nauk SSSR Zoologicheskii institut Parazitologicheskii Sbornik, 29, 32 - 154. [in Russian]." type="journal article" year="1960">Bregetova &amp; Koroleva 1960</bibRefCitation>
,
<bibRefCitation id="EFA1BA97FFB3E6688DA9FCC076096C74" author="Hyatt" box="[1013,1306,824,850]" pageId="8" pageNumber="52" refString="Hyatt, K. H. &amp; Emberson, R. M. (1988) A review of the Macrochelidae (Acari: Mesostigmata) of the British Isles. Bulletin of the British Museum (Natural History), Zoology, 54, 63 - 125 + 6 plates." type="book chapter" year="1988">Hyatt &amp; Emberson 1988</bibRefCitation>
), but again seems to be most diverse in Central and Southern Europe (
<bibRefCitation id="EFA1BA97FFB3E6688D1AFCA570DA6C51" author="Valle" box="[838,969,861,887]" pageId="8" pageNumber="53" refString="Valle, A. (1953) Revisione di generi e sottogeneri Berlesiani di Acari. (Primo contributo). Redia, 38, 316 - 360 + 3 plates." type="book chapter" year="1953">Valle 1953</bibRefCitation>
,
<bibRefCitation id="EFA1BA97FFB3E6688D89FCA577016C51" author="Valle" box="[981,1042,861,887]" pageId="8" pageNumber="53" refString="Valle, A. (1963) Intorno ad una nuova specie del genere Geholaspis, Berl. (Acari, Macrochelidae). Bollettino delle sedute dell'Accademia Gioenia di Scienze naturali in Catania, 7, 419 - 427." type="journal article" year="1963">1963</bibRefCitation>
,
<bibRefCitation id="EFA1BA97FFB3E6688A43FCA576526C51" author="Valle" box="[1055,1345,861,887]" pageId="8" pageNumber="53" refString="Valle, A., &amp; Mazzoleni, M. (1967) Nuove specie Mediterranee del genere Geholaspis Berl. e considerazioni intorno al sottogenere Longicheles Valle (Acari, Macrochelidae). Rendiconti Istituto Lombardo - accademia di Scienze e Lettere, 101 B, 385 - 403 + 5 plates." type="book chapter" year="1967">Valle &amp; Mazzoleni 1967</bibRefCitation>
,
<bibRefCitation id="EFA1BA97FFB3E6688B11FCA573CB6CBA" author="Masan" pageId="8" pageNumber="53" refString="Masan, P. (2003) Macrochelid mites of Slovakia (Acari: Mesostigmata: Macrochelidae). Institute of Zoology, Slovak Academy of Sciences, Bratislava. 149 pp." type="book" year="2003">Mašán 2003</bibRefCitation>
).
<taxonomicName id="4C30BCE5FFB3E6688EB0FC7A71366CBD" box="[236,549,898,923]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="mandibularis">
<emphasis id="B9441B74FFB3E6688EB0FC7A71366CBD" box="[236,549,898,923]" italics="true" pageId="8" pageNumber="45">Longicheles mandibularis</emphasis>
</taxonomicName>
has been introduced to
<collectingCountry id="F32787F6FFB3E6688D1AFC7A70A76CBA" box="[838,948,898,924]" name="Australia" pageId="8" pageNumber="45">Australia</collectingCountry>
(
<bibRefCitation id="EFA1BA97FFB3E6688D98FC7A77606CBA" author="Halliday" box="[964,1139,898,924]" pageId="8" pageNumber="52" refString="Halliday, R. B. (2001) Mesostigmatid mite fauna of Jenolan Caves, New South Wales (Acari: Mesostigmata). Australian Journal of Entomology, 40, 299 - 311." type="journal article" year="2001">Halliday 2001</bibRefCitation>
) and
<collectingCountry id="F32787F6FFB3E6688AEBFC7A764B6CBA" box="[1207,1368,898,924]" name="New Zealand" pageId="8" pageNumber="45">New Zealand</collectingCountry>
(new record), evidently by human agency.
</paragraph>
<paragraph id="8B8FC766FFB3E6688E98FC35773A6B5A" blockId="8.[151,1437,152,1148]" pageId="8" pageNumber="45">
<emphasis id="B9441B74FFB3E6688E98FC3572896CC1" bold="true" box="[196,410,973,999]" pageId="8" pageNumber="45">Included species:</emphasis>
<taxonomicName id="4C30BCE5FFB3E6688FFDFC3571366CC0" box="[417,549,973,999]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B9441B74FFB3E6688FFDFC3572A26CC0" box="[417,433,973,998]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688F9CFC3571366CC0" box="[448,549,973,998]" italics="true" pageId="8" pageNumber="45">longulus</emphasis>
</taxonomicName>
species group:
<taxonomicName id="4C30BCE5FFB3E6688C80FC35778C6CC1" authority="Valle &amp; Mazzoleni, 1967" authorityName="Valle &amp; Mazzoleni" authorityYear="1967" box="[732,1183,973,999]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="bianchii" status="comb. nov.">
<emphasis id="B9441B74FFB3E6688C80FC3571FF6CC0" box="[732,748,973,998]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688CA7FC3570486CC0" box="[763,859,973,998]" italics="true" pageId="8" pageNumber="45">bianchii</emphasis>
(
<bibRefCitation id="EFA1BA97FFB3E6688D36FC3577866CC1" author="Valle" box="[874,1173,973,999]" pageId="8" pageNumber="53" refString="Valle, A., &amp; Mazzoleni, M. (1967) Nuove specie Mediterranee del genere Geholaspis Berl. e considerazioni intorno al sottogenere Longicheles Valle (Acari, Macrochelidae). Rendiconti Istituto Lombardo - accademia di Scienze e Lettere, 101 B, 385 - 403 + 5 plates." type="book chapter" year="1967">Valle &amp; Mazzoleni, 1967</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB3E6688AF6FC35766B6CC1" box="[1194,1400,973,999]" pageId="8" pageNumber="45">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB3E6688BD8FC3572C66B2A" authority="Balogh, 1958" authorityName="Balogh" authorityYear="1958" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="bulgaricus">
<emphasis id="B9441B74FFB3E6688BD8FC3576876CC0" box="[1412,1428,973,998]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688ECBFC0A72056B2D" box="[151,278,1010,1035]" italics="true" pageId="8" pageNumber="45">bulgaricus</emphasis>
(
<bibRefCitation id="EFA1BA97FFB3E6688F7AFC0A72DF6B2A" author="Balogh" box="[294,460,1010,1036]" pageId="8" pageNumber="51" refString="Balogh, J. (1958) Macrocheliden aus Bulgarien (Acarina, Mesostigmata). Acta entomologica Musei Nationalis Prague, 32, 247 - 256." type="journal article" year="1958">Balogh, 1958</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFB3E6688FBFFC0A708B6B2A" authority="Valle &amp; Mazzoleni, 1967" authorityName="Valle &amp; Mazzoleni" authorityYear="1967" box="[483,920,1010,1036]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="ilvana" status="comb. nov.">
<emphasis id="B9441B74FFB3E6688FBFFC0A72E06B2D" box="[483,499,1010,1035]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688C5FFC0A715E6B2D" box="[515,589,1010,1035]" italics="true" pageId="8" pageNumber="45">ilvana</emphasis>
(
<bibRefCitation id="EFA1BA97FFB3E6688C01FC0A709C6B2A" author="Valle" box="[605,911,1010,1036]" pageId="8" pageNumber="53" refString="Valle, A., &amp; Mazzoleni, M. (1967) Nuove specie Mediterranee del genere Geholaspis Berl. e considerazioni intorno al sottogenere Longicheles Valle (Acari, Macrochelidae). Rendiconti Istituto Lombardo - accademia di Scienze e Lettere, 101 B, 385 - 403 + 5 plates." type="book chapter" year="1967">Valle &amp; Mazzoleni, 1967</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB3E6688DF8FC0A77656B2A" box="[932,1142,1010,1036]" pageId="8" pageNumber="45">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB3E6688ADFFC0A76846B2A" authority="Valle, 1963" authorityName="Valle" authorityYear="1963" box="[1155,1431,1010,1036]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="lagrecai" status="comb. nov.">
<emphasis id="B9441B74FFB3E6688ADFFC0A77806B2D" box="[1155,1171,1010,1035]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688AFFFC0A76166B2D" box="[1187,1285,1010,1035]" italics="true" pageId="8" pageNumber="45">lagrecai</emphasis>
<bibRefCitation id="EFA1BA97FFB3E6688B51FC0A76846B2A" author="Valle" box="[1293,1431,1010,1036]" pageId="8" pageNumber="53" refString="Valle, A. (1963) Intorno ad una nuova specie del genere Geholaspis, Berl. (Acari, Macrochelidae). Bollettino delle sedute dell'Accademia Gioenia di Scienze naturali in Catania, 7, 419 - 427." type="journal article" year="1963">Valle, 1963</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB3E6688ECBFBE072606B14" box="[151,371,1048,1074]" pageId="8" pageNumber="45">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB3E6688FDDFBE0701B6B14" authority="Balogh, 1958" authorityName="Balogh" authorityYear="1958" box="[385,776,1048,1074]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="longisetosus" status="comb. nov.">
<emphasis id="B9441B74FFB3E6688FDDFBE072826B17" box="[385,401,1048,1073]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688FFFFBE0712D6B17" box="[419,574,1048,1073]" italics="true" pageId="8" pageNumber="45">longisetosus</emphasis>
(
<bibRefCitation id="EFA1BA97FFB3E6688C0CFBE071ED6B14" author="Balogh" box="[592,766,1048,1074]" pageId="8" pageNumber="51" refString="Balogh, J. (1958) Macrocheliden aus Bulgarien (Acarina, Mesostigmata). Acta entomologica Musei Nationalis Prague, 32, 247 - 256." type="journal article" year="1958">Balogh, 1958</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB3E6688D4AFBE070E16B14" box="[790,1010,1048,1074]" pageId="8" pageNumber="45">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB3E6688A5CFBE076486B14" authority="Berlese, 1887" authorityName="Berlese" authorityYear="1887" box="[1024,1371,1048,1074]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="longulus" status="comb. nov.">
<emphasis id="B9441B74FFB3E6688A5CFBE077036B17" box="[1024,1040,1048,1073]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688A7EFBE0779E6B17" box="[1058,1165,1048,1073]" italics="true" pageId="8" pageNumber="45">longulus</emphasis>
(
<bibRefCitation id="EFA1BA97FFB3E6688AC3FBE076406B14" author="Berlese" box="[1183,1363,1048,1074]" pageId="8" pageNumber="51" refString="Berlese, A. (1887) Acari, Myriopoda et Scorpiones hucusque in Italiana reperta, 43, 9, 10." type="journal article" year="1887">Berlese, 1887</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB3E6688B35FBE072266B71" pageId="8" pageNumber="45">new combination</taxonomicNameLabel>
;
<taxonomicName id="4C30BCE5FFB3E6688F1EFBC570166B71" authority="Valle &amp; Mazzoleni, 1967" authorityName="Valle &amp; Mazzoleni" authorityYear="1967" box="[322,773,1085,1111]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="ranzii" status="comb. nov.">
<emphasis id="B9441B74FFB3E6688F1EFBC572416B70" box="[322,338,1085,1110]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688F38FBC572BE6B70" box="[356,429,1085,1110]" italics="true" pageId="8" pageNumber="45">ranzii</emphasis>
(
<bibRefCitation id="EFA1BA97FFB3E6688FE1FBC571EF6B71" author="Valle" box="[445,764,1085,1111]" pageId="8" pageNumber="53" refString="Valle, A., &amp; Mazzoleni, M. (1967) Nuove specie Mediterranee del genere Geholaspis Berl. e considerazioni intorno al sottogenere Longicheles Valle (Acari, Macrochelidae). Rendiconti Istituto Lombardo - accademia di Scienze e Lettere, 101 B, 385 - 403 + 5 plates." type="book chapter" year="1967">Valle &amp; Mazzoleni, 1967</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB3E6688D4FFBC570FE6B71" box="[787,1005,1085,1111]" pageId="8" pageNumber="45">new combination</taxonomicNameLabel>
.
<taxonomicName id="4C30BCE5FFB3E6688DA5FBC577D36B70" box="[1017,1216,1085,1111]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="mandibularis">
<emphasis id="B9441B74FFB3E6688DA5FBC5771A6B70" box="[1017,1033,1085,1110]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688A47FBC577D36B70" box="[1051,1216,1085,1110]" italics="true" pageId="8" pageNumber="45">mandibularis</emphasis>
</taxonomicName>
species group:
<taxonomicName id="4C30BCE5FFB3E6688BD8FBC572D66B5A" authority="Berlese, 1904" authorityName="Berlese" authorityYear="1904" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="hortorum">
<emphasis id="B9441B74FFB3E6688BD8FBC576876B70" box="[1412,1428,1085,1110]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688ECBFB9B72156B5A" box="[151,262,1123,1148]" italics="true" pageId="8" pageNumber="45">hortorum</emphasis>
(
<bibRefCitation id="EFA1BA97FFB3E6688F49FB9A72AF6B5A" author="Berlese" box="[277,444,1122,1148]" pageId="8" pageNumber="51" refString="Berlese, A. (1904) Acari nuovi. Manipulus II. Redia, 1, 258 - 280." type="journal article" year="1904">Berlese, 1904</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C30BCE5FFB3E6688F8DFB9B70586B5A" authority="Berlese, 1904" authorityName="Berlese" authorityYear="1904" box="[465,843,1122,1148]" class="Arachnida" family="Macrochelidae" genus="Longicheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="mandibularis" status="comb. nov.">
<emphasis id="B9441B74FFB3E6688F8DFB9B72F26B5A" box="[465,481,1123,1148]" italics="true" pageId="8" pageNumber="45">L</emphasis>
.
<emphasis id="B9441B74FFB3E6688FACFB9B719F6B5A" box="[496,652,1123,1148]" italics="true" pageId="8" pageNumber="45">mandibularis</emphasis>
(
<bibRefCitation id="EFA1BA97FFB3E6688CC0FB9A70516B5A" author="Berlese" box="[668,834,1122,1148]" pageId="8" pageNumber="51" refString="Berlese, A. (1904) Acari nuovi. Manipulus II. Redia, 1, 258 - 280." type="journal article" year="1904">Berlese, 1904</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A277A60FFFB3E6688D0BFB9A77356B5A" box="[855,1062,1122,1148]" pageId="8" pageNumber="45">new combination</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
</treatment>
</document>