226 lines
32 KiB
XML
226 lines
32 KiB
XML
<document id="AB2187A6A2736BE8BA8246E3C8B00945" ID-DOI="10.11646/zootaxa.3887.4.5" ID-GBIF-Dataset="e6fe4aa0-c179-467b-8a12-5155ae1551cf" ID-ISSN="1175-5326" ID-Zenodo-Dep="252267" ID-ZooBank="0ACD6410-2096-41C0-A499-5E59BC663116" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460980029017" checkinUser="plazi" docAuthor="Bilton, David T." docDate="2014" docId="03884A73FD0DD500FF5EFB63FBD3FE3F" docLanguage="en" docName="zt03887p480.pdf" docOrigin="Zootaxa 3887 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parhydraena faeni Bilton, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="477" masterDocId="FFB1320BFD09D506FFC9FFAEFFECFF92" masterDocTitle="Two new water beetles from the Hantamsberg, an inselberg in the Northern Cape of South Africa (Coleoptera, Hydraenidae)" masterLastPageNumber="480" masterPageNumber="471" pageNumber="475" updateTime="1698560054570" updateUser="plazi">
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<mods:titleInfo id="16D54E8BECA5B8B57DA79BA945E5C348">
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<mods:title id="9AA9188937AFA505E930DD03EAB1D9CE">Two new water beetles from the Hantamsberg, an inselberg in the Northern Cape of South Africa (Coleoptera, Hydraenidae)</mods:title>
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<mods:name id="4DA7F0A65C4CC99E0709A6D14FA54249" type="personal">
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<mods:roleTerm id="0636C61538B9978B86D57836F7F5C433">Author</mods:roleTerm>
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<mods:namePart id="25192756D65C576C8F46D42B066EF1FD">Bilton, David T.</mods:namePart>
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<mods:typeOfResource id="3188B4AAD638F249219FEB124BC60F14">text</mods:typeOfResource>
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<mods:title id="F46C885E22219154066841CBF5C765E9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6B913714E3922A30AD76C0A2C5781CFE">
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<mods:date id="84FE8719B147C88FF2BBFC81C29F8784">2014</mods:date>
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<mods:detail id="EE42C5C858E39A9CA9D934A41A4DCFE4" type="volume">
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<mods:number id="FCF8B7D693461C336DE066AD567B9961">3887</mods:number>
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</mods:detail>
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<mods:detail id="9BCF378E7C47BC2B7FD59DEBC0796320" type="issue">
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<mods:number id="54AFF4CA10F98187A63D5E40881485B0">3</mods:number>
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<mods:classification id="DE29D29B6849317A8E2B7CFDEF8A3C4C">journal article</mods:classification>
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<mods:identifier id="275942C57EE642E17EF97F3CC5DEA56B" type="DOI">10.11646/zootaxa.3887.4.5</mods:identifier>
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<mods:identifier id="82DEAA6DD42A29326EFAD4E8A1C29291" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03884A73FD0DD500FF5EFB63FBD3FE3F" ID-DOI="http://doi.org/10.5281/zenodo.5686940" ID-GBIF-Taxon="119594478" ID-Zenodo-Dep="5686940" LSID="urn:lsid:plazi:treatment:03884A73FD0DD500FF5EFB63FBD3FE3F" httpUri="http://treatment.plazi.org/id/03884A73FD0DD500FF5EFB63FBD3FE3F" lastPageId="6" lastPageNumber="477" pageId="4" pageNumber="475">
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<subSubSection id="C33BA8EEFD0DD502FF5EFB63FEE6FA9B" pageId="4" pageNumber="475" type="nomenclature">
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<paragraph id="8B9EFB65FD0DD502FF5EFB63FE33FB7A" blockId="4.[151,479,1229,1289]" box="[151,479,1229,1256]" pageId="4" pageNumber="475">
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<heading id="D0D64C09FD0DD502FF5EFB63FE33FB7A" bold="true" box="[151,479,1229,1256]" fontSize="11" level="1" pageId="4" pageNumber="475" reason="1">
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<emphasis id="B9552777FD0DD502FF5EFB63FE33FB7A" bold="true" box="[151,479,1229,1256]" pageId="4" pageNumber="475">
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<taxonomicName id="4C2180E6FD0DD502FF5EFB63FE96FB75" ID-CoL="4DLYQ" box="[151,378,1229,1255]" class="Insecta" family="Hydraenidae" genus="Parhydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="475" phylum="Arthropoda" rank="species" species="faeni" status="sp. nov.">
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<emphasis id="B9552777FD0DD502FF5EFB63FE96FB75" bold="true" box="[151,378,1229,1255]" italics="true" pageId="4" pageNumber="475">Parhydraena faeni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2669A0CFD0DD502FE48FB60FE33FB7A" box="[385,479,1230,1256]" pageId="4" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8B9EFB65FD0DD502FF5EFB5EFEE6FA9B" blockId="4.[151,479,1229,1289]" box="[151,266,1264,1289]" pageId="4" pageNumber="475">
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||
(
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||
<figureCitation id="131AE7E0FD0DD502FF56FB5EFEEDFA9B" box="[159,257,1264,1289]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1768,1790]" captionStartId-1="4.[151,250,1102,1124]" captionStartId-2="6.[151,250,1932,1954]" captionTargetBox-0="[175,1389,214,1672]" captionTargetBox-1="[151,1436,207,1080]" captionTargetBox-2="[162,1415,473,1899]" captionTargetId-0="figure@2.[151,1436,195,1743]" captionTargetId-1="figure@4.[151,1436,193,1080]" captionTargetId-2="figure@6.[162,1415,467,1901]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="FIGURE 1. Habitus and genitalia of Hantamsberg Hydraenidae. A) Mesoceration hantam sp. nov., holotype; B) Parhydraena faeni sp. nov., holotype; C) Mesoceration hantam sp. nov., aedeagus, ventral and lateral view; D) Parhydraena faeni sp. nov., aedeagus ventral and lateral views. Scale bar A – B = 1 mm; C – D = 0.25 mm." captionText-1="FIGURE 2. Southern margin of the Hantamsberg plateau, showing its isolation from the surrounding plains. The yellow flower in the foreground is likely Moraea reflexa Goldblatt, endemic to the Hantamsberg. Photo D. T. Bilton." captionText-2="FIGURE 3. Water beetle habitats on the Hantamsberg plateau, 1,500 m. A) Drying stream pool on gravel on northwest side of plateau; B) pool in stream 1 km north of mast; C) detail of residual pool in same stream system; D) temporary pool on clay / rock with Bulbinella nutans. Photos D. T. Bilton." httpUri-0="https://zenodo.org/record/252268/files/figure.png" httpUri-1="https://zenodo.org/record/252269/files/figure.png" httpUri-2="https://zenodo.org/record/252270/files/figure.png" pageId="4" pageNumber="475">Figs 1–3</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33BA8EEFD0DD502FF5EFA96FD24F9E3" pageId="4" pageNumber="475" type="materials_examined">
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||
<paragraph id="8B9EFB65FD0DD502FF5EFA96FC51FAE7" blockId="4.[151,1437,1336,2009]" pageId="4" pageNumber="475">
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||
<emphasis id="B9552777FD0DD502FF5EFA96FEC2FAC3" bold="true" box="[151,302,1336,1361]" pageId="4" pageNumber="475">
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||
<typeStatus id="549A45C7FD0DD502FF5EFA96FF3EFAC3" box="[151,210,1336,1361]" pageId="4" pageNumber="475">Type</typeStatus>
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locality
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||
</emphasis>
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||
.
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||
<emphasis id="B9552777FD0DD502FEF4FA96FE3BFAC3" bold="true" box="[317,471,1336,1361]" pageId="4" pageNumber="475">
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||
<collectingCountry id="F336BBF5FD0DD502FEF4FA96FE3BFAC3" box="[317,471,1336,1361]" name="South Africa" pageId="4" pageNumber="475">South Africa</collectingCountry>
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||
</emphasis>
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, Northern Cape, Calvinia, Hantamsberg, drying stream with pools
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<quantity id="4CD95680FD0DD502FB06FA96FAEDFAC3" box="[1231,1281,1336,1361]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="475" unit="km" value="1.0">1km</quantity>
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N of mast on summit plateau,
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<geoCoordinate id="EE159DA2FD0DD502FE98FAF3FDE6FAE7" box="[337,522,1373,1397]" direction="south" orientation="latitude" pageId="4" pageNumber="475" precision="1" value="-31.375895">31° 22’ 33.22’’S</geoCoordinate>
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||
<geoCoordinate id="EE159DA2FD0DD502FDD8FAF3FD2CFAE7" box="[529,704,1372,1397]" direction="east" orientation="longitude" pageId="4" pageNumber="475" precision="1" value="19.786528">19° 47’ 11.5’’E</geoCoordinate>
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||
,
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||
<quantity id="4CD95680FD0DD502FD05FAF2FCCBFAE6" box="[716,807,1372,1397]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="475" unit="m" value="1500.0">1,500 m</quantity>
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||
(
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||
<figureCitation id="131AE7E0FD0DD502FCFEFAF2FC99FAE7" box="[823,885,1372,1397]" captionStart="FIGURE 1" captionStartId="2.[151,250,1768,1790]" captionTargetBox="[175,1389,214,1672]" captionTargetId="figure@2.[151,1436,195,1743]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and genitalia of Hantamsberg Hydraenidae. A) Mesoceration hantam sp. nov., holotype; B) Parhydraena faeni sp. nov., holotype; C) Mesoceration hantam sp. nov., aedeagus, ventral and lateral view; D) Parhydraena faeni sp. nov., aedeagus ventral and lateral views. Scale bar A – B = 1 mm; C – D = 0.25 mm." httpUri="https://zenodo.org/record/252268/files/figure.png" pageId="4" pageNumber="475">Fig. 1</figureCitation>
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B–C).
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||
</paragraph>
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||
<paragraph id="8B9EFB65FD0DD502FF0EFA2EFEC7FA73" blockId="4.[151,1437,1336,2009]" pageId="4" pageNumber="475">
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<emphasis id="B9552777FD0DD502FF0EFA2EFE9DFA0B" bold="true" box="[199,369,1408,1433]" pageId="4" pageNumber="475">
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<typeStatus id="549A45C7FD0DD502FF0EFA2EFEEEFA0B" box="[199,258,1408,1433]" pageId="4" pageNumber="475">Type</typeStatus>
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||
material
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||
</emphasis>
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||
.
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||
<typeStatus id="549A45C7FD0DD502FE4AFA2EFE1CFA0B" box="[387,496,1408,1433]" pageId="4" pageNumber="475" type="holotype">
|
||
<emphasis id="B9552777FD0DD502FE4AFA2EFE1CFA0B" bold="true" box="[387,496,1408,1433]" pageId="4" pageNumber="475">Holotype</emphasis>
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||
</typeStatus>
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||
(male): “
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||
<date id="FF9FDDA5FD0DD502FDA8FA2EFD31FA0B" box="[609,733,1408,1433]" pageId="4" pageNumber="475" value="2010-09-18">18/ix/2010</date>
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||
<collectingCountry id="F336BBF5FD0DD502FD2EFA2EFC97FA0B" box="[743,891,1408,1433]" name="South Africa" pageId="4" pageNumber="475">South Africa</collectingCountry>
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||
NC// Calvinia Hantamsberg summit// pools in stream
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||
<quantity id="4CD95680FD0DD502FF21FA0BFECCFA2F" box="[232,288,1444,1469]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="475" unit="km" value="1.0">1 km</quantity>
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||
N of // mast
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||
<quantity id="4CD95680FD0DD502FE66FA0AFDE7FA2E" box="[431,523,1444,1469]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="475" unit="m" value="1500.0">1,500 m</quantity>
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||
D T Bilton leg.” (genitalia extracted and mounted on same card) and red
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||
<typeStatus id="549A45C7FD0DD502FAF3FA0AFA70FA2F" box="[1338,1436,1444,1469]" pageId="4" pageNumber="475" type="holotype">holotype</typeStatus>
|
||
label (
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||
<collectionCode id="ED3063A0FD0DD502FF15FA67FEF2FA73" box="[220,286,1481,1505]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="4" pageNumber="475">SAM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9EFB65FD0DD502FF0EFA42FD24F9E3" blockId="4.[151,1437,1336,2009]" pageId="4" pageNumber="475">
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||
<typeStatus id="549A45C7FD0DD502FF0EFA42FED3F997" box="[199,319,1516,1541]" pageId="4" pageNumber="475" type="paratype">
|
||
<emphasis id="B9552777FD0DD502FF0EFA42FED3F997" bold="true" box="[199,319,1516,1541]" pageId="4" pageNumber="475">Paratypes</emphasis>
|
||
</typeStatus>
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||
(15):
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||
<collectingCountry id="F336BBF5FD0DD502FE4AFA42FDF0F997" box="[387,540,1516,1541]" name="South Africa" pageId="4" pageNumber="475">
|
||
<emphasis id="B9552777FD0DD502FE4AFA42FDF0F997" bold="true" box="[387,540,1516,1541]" pageId="4" pageNumber="475">South Africa</emphasis>
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||
</collectingCountry>
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||
:
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||
<specimenCount id="9D2730ECFD0DD502FDE3FA43FDBAF996" box="[554,598,1517,1541]" pageId="4" pageNumber="475" type="generic">9 ♂</specimenCount>
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||
, 2 ♀ “
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||
<date id="FF9FDDA5FD0DD502FD54FA42FCF5F997" box="[669,793,1516,1541]" pageId="4" pageNumber="475" value="2010-09-18">18/ix/2010</date>
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||
<collectingCountry id="F336BBF5FD0DD502FCD6FA42FC43F997" box="[799,943,1516,1541]" name="South Africa" pageId="4" pageNumber="475">South Africa</collectingCountry>
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||
NC// Calvinia Hantamsberg summit// pools in stream
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||
<quantity id="4CD95680FD0DD502FECCF9BFFED2F9BB" box="[261,318,1552,1577]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="475" unit="km" value="1.0">1 km</quantity>
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||
N of // mast
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||
<quantity id="4CD95680FD0DD502FE18F9BEFDC1F9BA" box="[465,557,1552,1577]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="475" unit="m" value="1500.0">1,500 m</quantity>
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||
D T Bilton leg.” (
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||
<collectionCode id="ED3063A0FD0DD502FD30F9BFFCA4F9BB" box="[761,840,1553,1577]" pageId="4" pageNumber="475">CDTB</collectionCode>
|
||
,
|
||
<collectionCode id="ED3063A0FD0DD502FC9AF9BFFC77F9BA" box="[851,923,1553,1576]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="475">NMW</collectionCode>
|
||
,
|
||
<collectionCode id="ED3063A0FD0DD502FC6FF9BFFC1FF9BB" LSID="urn:lsid:biocol.org:col:34248" box="[934,1011,1553,1577]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="4" pageNumber="475">SANC</collectionCode>
|
||
,
|
||
<collectionCode id="ED3063A0FD0DD502FC36F9BFFBBEF9BB" box="[1023,1106,1553,1577]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="475">TMSA</collectionCode>
|
||
);
|
||
<specimenCount id="9D2730ECFD0DD502FBACF9BFFB7DF9BA" box="[1125,1169,1553,1577]" pageId="4" pageNumber="475" type="generic">1 ♂</specimenCount>
|
||
, 3 ♀ “
|
||
<date id="FF9FDDA5FD0DD502FB11F9BEFAB8F9BB" box="[1240,1364,1552,1577]" pageId="4" pageNumber="475" value="2010-09-18">18/ix/2010</date>
|
||
<collectingCountry id="F336BBF5FD0DD502FA92F9BEFF33F9DF" name="South Africa" pageId="4" pageNumber="475">South Africa</collectingCountry>
|
||
NC// Calvinia Hantamsberg summit// drying stream on gravel on NW side// of plateau—
|
||
<quantity id="4CD95680FD0DD502FB0DF99AFAF2F9DE" box="[1220,1310,1588,1613]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="475" unit="m" value="1500.0">1,500 m</quantity>
|
||
D T Bilton leg.” (
|
||
<collectionCode id="ED3063A0FD0DD502FF12F9F7FEC6F9E3" box="[219,298,1625,1649]" pageId="4" pageNumber="475">CDTB</collectionCode>
|
||
,
|
||
<collectionCode id="ED3063A0FD0DD502FEFCF9F7FE9BF9E3" box="[309,375,1625,1649]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="4" pageNumber="475">SAM</collectionCode>
|
||
). All with red
|
||
<typeStatus id="549A45C7FD0DD502FDD1F9F7FD94F9E3" box="[536,632,1625,1649]" pageId="4" pageNumber="475" type="paratype">paratype</typeStatus>
|
||
labels.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33BA8EEFD0DD500FF0EF9D2FE81FE8E" lastPageId="6" lastPageNumber="477" pageId="4" pageNumber="475" type="description">
|
||
<paragraph id="8B9EFB65FD0DD502FF0EF9D2FDBBF8FF" blockId="4.[151,1437,1336,2009]" pageId="4" pageNumber="475">
|
||
<emphasis id="B9552777FD0DD502FF0EF9D2FEBDF907" bold="true" box="[199,337,1660,1685]" pageId="4" pageNumber="475">Description</emphasis>
|
||
. Size:
|
||
<typeStatus id="549A45C7FD0DD502FE68F9D2FDE2F907" box="[417,526,1660,1685]" pageId="4" pageNumber="475" type="holotype">Holotype</typeStatus>
|
||
: BL
|
||
<quantity id="4CD95680FD0DD502FD8CF9D3FD42F906" box="[581,686,1661,1685]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="4" pageNumber="475" unit="mm" value="2.03">2.03 mm</quantity>
|
||
; EL
|
||
<quantity id="4CD95680FD0DD502FD2CF9D3FCD3F906" box="[741,831,1661,1685]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="475" unit="mm" value="1.3">1.3 mm</quantity>
|
||
; EW
|
||
<quantity id="4CD95680FD0DD502FC49F9D2FC04F906" box="[896,1000,1660,1685]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="475" unit="mm" value="0.75">0.75 mm</quantity>
|
||
.
|
||
<typeStatus id="549A45C7FD0DD502FC3DF9D3FB8AF907" box="[1012,1126,1661,1685]" pageId="4" pageNumber="475" type="paratype">Paratypes</typeStatus>
|
||
: ♂s BL
|
||
<quantity id="4CD95680FD0DD502FB0FF9D2FA81F906" box="[1222,1389,1660,1685]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.25" metricValueMin="2.05" pageId="4" pageNumber="475" unit="mm" value="2.15" valueMax="2.25" valueMin="2.05">2.05–2.25 mm</quantity>
|
||
; EL
|
||
<quantity id="4CD95680FD0DD502FF5EF90FFEC4F92A" box="[151,296,1697,1721]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="4" pageNumber="475" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
|
||
; EW
|
||
<quantity id="4CD95680FD0DD502FEBAF90FFDFEF92A" box="[371,530,1697,1721]" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.299999999999999" metricValueMin="8.0" pageId="4" pageNumber="475" unit="mm" value="0.815" valueMax="0.83" valueMin="0.8">0.8–0.83 mm</quantity>
|
||
. ♀s BL
|
||
<quantity id="4CD95680FD0DD502FD4AF90FFCCEF92A" box="[643,802,1697,1721]" metricMagnitude="-3" metricUnit="m" metricValue="2.265" metricValueMax="2.33" metricValueMin="2.2" pageId="4" pageNumber="475" unit="mm" value="2.265" valueMax="2.33" valueMin="2.2">2.2–2.33 mm</quantity>
|
||
; EL
|
||
<quantity id="4CD95680FD0DD502FCADF90EFBEFF92A" box="[868,1027,1696,1720]" metricMagnitude="-3" metricUnit="m" metricValue="1.4249999999999998" metricValueMax="1.45" metricValueMin="1.4" pageId="4" pageNumber="475" unit="mm" value="1.4249999999999998" valueMax="1.45" valueMin="1.4">1.4–1.45 mm</quantity>
|
||
; EW
|
||
<quantity id="4CD95680FD0DD502FB87F90FFB01F92A" box="[1102,1261,1697,1721]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" metricValueMax="9.8" metricValueMin="9.0" pageId="4" pageNumber="475" unit="mm" value="0.94" valueMax="0.98" valueMin="0.9">0.9–0.98 mm</quantity>
|
||
. Dorsum (
|
||
<figureCitation id="131AE7E0FD0DD502FAA6F90EFF4CF94F" captionStart="FIGURE 1" captionStartId="2.[151,250,1768,1790]" captionTargetBox="[175,1389,214,1672]" captionTargetId="figure@2.[151,1436,195,1743]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and genitalia of Hantamsberg Hydraenidae. A) Mesoceration hantam sp. nov., holotype; B) Parhydraena faeni sp. nov., holotype; C) Mesoceration hantam sp. nov., aedeagus, ventral and lateral view; D) Parhydraena faeni sp. nov., aedeagus ventral and lateral views. Scale bar A – B = 1 mm; C – D = 0.25 mm." httpUri="https://zenodo.org/record/252268/files/figure.png" pageId="4" pageNumber="475">Fig. 1</figureCitation>
|
||
B)—head dark piceous brown - black, pronotum dark straw yellow, with disc dark brown to black, elytral dark straw yellow with suture and lateral margin infuscated. Legs and maxillary palpi straw yellow; tarsi and apical palpal segment infuscated. Venter dark reddish brown to piceous, with silvery hydrofuge vestiture. Pronotal hypomeron dark straw yellow, as dorsum; inner 0.5 of bubble hypomeron dark pitchy brown. Elytral epipleurs and pseudepipleurs straw yellow as dorsum.
|
||
</paragraph>
|
||
<paragraph id="8B9EFB65FD0DD503FF0EF8D6FC74FE1A" blockId="4.[151,1437,1336,2009]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="476" pageId="4" pageNumber="475">Head: Labrum transverse, rounder apicolaterally, with broad, shallow apicomedian emargination, running approx. 0.15 of length, and broad, rounded notch either side of apicomedian emargination, approx. 0.5 as deep as central notch. Sides of apicomedian emargination very weakly raised. Surface of labrum shining, with distinct, weakly transverse - isodiametric microreticulation and with sparse, medium punctures, each bearing a long decumbent seta. Entire upper surface of head weakly shining, with strong isodiametric microreticulation and sparse coarse to medium punctures, some of which bear long decumbent setae. Punctures densest on clypeus and central parts of frons and vertex, more widely spaced elsewhere, and somewhat obsolete around inner margins of eyes. Fronto-clypeal suture strong, well impressed. Oblique sublateral impressions strong, deeper posteriorly, shallower anteriorly. Ocelli broad and flat, paler than rest of head. Eyes large, occupying almost 0.5 side of head; facets relatively flat. Maxillary palpi elongate, approx. 1.2x length of head.</paragraph>
|
||
<paragraph id="8B9EFB65FD0CD503FF0EFE3BFB55FD5E" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="476">Pronotum: Cordate, strongly transverse. Anterior margin straight, slightly indented in median third, where there is a trace of a narrow hyaline border. Anterior angles obtusely rounded. Sides rounded, broadest just in front of middle, gradually narrowing in an almost straight line to obtuse posterior angles. Sides distinctly crenulate, especially in posterior 0.5. Pronotum with fine, weakly impressed isodiametric microreticulation, somewhat shining. Entire surface punctate, with dense, shallow punctures. Each puncture with anterior margin extended posteriorly as narrow ridge, dividing puncture into a pair of two smaller punctures, most punctures with indistinct, short recumbent or decumbent seta arising from posterior extreme of dividing ridge. Punctures more widely spaced in centre of disc; closer, almost confluent, towards anterior and posterior margins, giving a rugulose appearance; larger and more elongate towards lateral margins. Discal impressions weak, shallow and open.</paragraph>
|
||
<paragraph id="8B9EFB65FD0CD503FF0EFD79FAA1FC12" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="476">Elytra: Elongate oval, sides weakly arcuate, broadest behind middle. Margins weakly explanate over posterior 0.8 of length, and minutely serrate. Elytra strongly and densely punctate, each puncture bearing a long recumbent white—yellowish seta (worn in places in older specimens). Each puncture with anterior margin extended posteriorly as narrow ridge which supports the seta. Elytral puncture rows not clearly discernable, punctation instead appearing subseriate to somewhat irregular. A single stria present alongside suture in posterior 0.33.</paragraph>
|
||
<paragraph id="8B9EFB65FD0CD503FF0EFC23FBC8F9C2" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="476">Venter: Mentum, submentum and genae shining. Mentum coarsely and rugosely punctate; punctures almost confluent towards lateral margins, and bearing long recumbent golden setae throughout, with visible traces of transverse microreticulation amongst punctures. Submentum with similar setose punctures; more widely spaced than on Mentum, with only faintest traces of microreticulation lines. Genae with rugose microreticulation towards sides; transverse corrugations internally. Gula dull, punctate, with rugose, isodiametric microreticulation; punctures bearing recurved golden setae. Prosternum with similar microreticulation and punctation in anterior 0.5; punctate only, and therefore shiny in posterior 0.5. Surface with long recurved setae, becoming denser anteriolaterally. Pronotal hypomera broad, shining. Wet hypomeron with open, weak, isodiametric reticulation; bubble hypomeron smooth, lacking vestiture in outer 0.5, rugose and pubescent in inner 0.5. Meso and metaventrites rugosely microreticulate, punctate, with long hydrofuge vestiture, densest laterally. Basal 0.5 of metaventrite with weak longitudinal carina, barely visible amonst the vestiture. Elytral pseudepipleurs broad, flat, shining, narrowed approx. 0.4 from base. Epipleurs narrow, with hydrofuge vestiture in anterior 0.4. Abdominal ventrites 1–4 microreticulate; covered in long hydrofuge vestiture; ventrite 5 with same vestiture and sculpture in anteriorly 0.33, sculpture and vestiture extending to 0.5 length of sclerite centrally. Posterior part of ventrite 5 glabrous and shining, with isodiametric microreticulation anteriorly, and transverse meshes posteriorly. Irregular transverse row of medium punctures bearing decumbent setae approx. 0.33 from apex of ventrite. Ventrite 6 entirely glabrous, shining, with weakly transeverse microreticulation over anterior 0.5; lacking microreticulation in posterior 0.5; two sections separated by a transverse ridge, and three irregular transverse rows of decumbent setae. Last ventrite asymmetrical, with rounded tooth-like projection on right, which fits between the lobes of the last tergite. Last tergite with apicomedian notch, flanked by tufts of short, stout setae.</paragraph>
|
||
<paragraph id="8B9EFB65FD0CD503FF0EF9F3FB15F9E6" blockId="5.[151,1437,151,2013]" box="[199,1273,1628,1653]" pageId="5" pageNumber="476">Legs: Moderately long and slender, basal 3 pro- and mesotarsomeres with pads of suction setae.</paragraph>
|
||
<paragraph id="8B9EFB65FD0CD503FF0EF9D1FF25F896" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="476">
|
||
Aedeagus: Strongly arcuate in lateral view; slightly sinuate in ventral view. Main piece with large, transverse, flattened ventral process. Distal piece complex, with articulated apical lobes. Gonopore tube distinct, relatively robust for the genus, and projecting beyond the apex of the distal piece. Parameres with long, robust setae (
|
||
<figureCitation id="131AE7E0FD0CD503FAA6F969FF4EF896" captionStart="FIGURE 1" captionStartId="2.[151,250,1768,1790]" captionTargetBox="[175,1389,214,1672]" captionTargetId="figure@2.[151,1436,195,1743]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and genitalia of Hantamsberg Hydraenidae. A) Mesoceration hantam sp. nov., holotype; B) Parhydraena faeni sp. nov., holotype; C) Mesoceration hantam sp. nov., aedeagus, ventral and lateral view; D) Parhydraena faeni sp. nov., aedeagus ventral and lateral views. Scale bar A – B = 1 mm; C – D = 0.25 mm." httpUri="https://zenodo.org/record/252268/files/figure.png" pageId="5" pageNumber="476">Fig. 1</figureCitation>
|
||
D).
|
||
</paragraph>
|
||
<paragraph id="8B9EFB65FD0CD503FF0EF8A1FD9AF8DE" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="476">Female: Last ventrite with simple, rounded apex; last tergite with slight apical emargination, but lacking obvious notch. Tarsi without suction setae.</paragraph>
|
||
<paragraph id="8B9EFB65FD0CD503FF0EF8F9FD30F807" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="476">
|
||
Variation:
|
||
<typeStatus id="549A45C7FD0CD503FE88F8F7FE42F8E3" box="[321,430,1881,1905]" pageId="5" pageNumber="476" type="paratype">Paratypes</typeStatus>
|
||
vary somewhat in colouration and in the depth of the discal impressions, which are sometimes more marked than those of the
|
||
<typeStatus id="549A45C7FD0CD503FDBBF8D2FD34F807" box="[626,728,1916,1941]" pageId="5" pageNumber="476" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9EFB65FD0CD500FF0EF80EFE81FE8E" blockId="5.[151,1437,151,2013]" lastBlockId="6.[151,1437,151,429]" lastPageId="6" lastPageNumber="477" pageId="5" pageNumber="476">
|
||
<emphasis id="B9552777FD0CD503FF0EF80EFE23F82B" bold="true" box="[199,463,1952,1977]" pageId="5" pageNumber="476">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C2180E6FD0CD503FE11F80EFD40F82A" box="[472,684,1952,1976]" class="Insecta" family="Hydraenidae" genus="Parhydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="476" phylum="Arthropoda" rank="species" species="faeni" status="sp. nov.">
|
||
<emphasis id="B9552777FD0CD503FE11F80EFD40F82A" box="[472,684,1952,1976]" italics="true" pageId="5" pageNumber="476">Parhydraena faeni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2669A0CFD0CD503FD7CF80EFCFCF82B" box="[693,784,1952,1977]" pageId="5" pageNumber="476" rank="species">
|
||
<emphasis id="B9552777FD0CD503FD7CF80EFCFCF82B" bold="true" box="[693,784,1952,1977]" pageId="5" pageNumber="476">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from other described species of the genus by the indistinct elytral series, which result in an irregularly punctate appearance. The pale colouration and male genitalia of the new species are also characteristic. The only other described species with similarly subseriate or irregular elytral punctures is
|
||
<taxonomicName id="4C2180E6FD0FD500FDD1FF12FC62FF47" authority="Perkins" authorityName="Perkins" box="[536,910,188,213]" class="Insecta" family="Hydraenidae" genus="Parhydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="477" phylum="Arthropoda" rank="species" species="sebastiani">
|
||
<emphasis id="B9552777FD0FD500FDD1FF12FCC6FF46" box="[536,810,188,212]" italics="true" pageId="6" pageNumber="477">Parhydraena sebastiani</emphasis>
|
||
Perkins
|
||
</taxonomicName>
|
||
, which has very different surface sculpture, colouration and aedeagus (
|
||
<bibRefCitation id="EFB08694FD0FD500FE0BFF71FD8FFF6A" author="Perkins" box="[450,611,223,248]" pageId="6" pageNumber="477" refString="Perkins, P. D. (2009) Revisions of the genera Parhydraena Orchymont, Protozantaena Perkins, Decarthrocerus Orchymont, and Parhydraenopsis nomen novum, aquatic and humicolous beetles from Africa and Madagascar, and comparative morphology of the tribe Parhydraenini (Coleoptera: Hydraenidae). Zootaxa, 2038, 1 - 119." type="journal article" year="2009">Perkins, 2009</bibRefCitation>
|
||
). This is apparently the largest known species of the genus, some females exceeding
|
||
<quantity id="4CD95680FD0FD500FEC6FEAAFE84FE8E" box="[271,360,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="477" unit="mm" value="2.3">2.3 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33BA8EEFD0FD500FF0EFE89FB16FE1A" pageId="6" pageNumber="477" type="distribution">
|
||
<paragraph id="8B9EFB65FD0FD500FF0EFE89FB16FE1A" blockId="6.[151,1437,151,429]" pageId="6" pageNumber="477">
|
||
<emphasis id="B9552777FD0FD500FF0EFE89FEB5FED2" bold="true" box="[199,345,295,320]" pageId="6" pageNumber="477">Distribution</emphasis>
|
||
. To date only known from two adjacent localities on the summit of the Hantamsberg, close to Calvinia, in
|
||
<taxonomicName id="4C2180E6FD0FD500FEE0FEE3FE6EFEF6" box="[297,386,333,356]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="479" phylum="Arthropoda" rank="species" species="hantam" status="sp. nov.">Hantam</taxonomicName>
|
||
Plateau Dolerite Renosterveld (sensu
|
||
<bibRefCitation id="EFB08694FD0FD500FC8AFEE2FB63FEF7" author="Mucina" box="[835,1167,332,357]" pageId="6" pageNumber="477" refString="Murcina, L. & Rutherford, M. C. (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19, 1 - 807." type="journal article" year="2006">Mucina & Rutherford, 2006</bibRefCitation>
|
||
). Here the species was abundant amongst gravel and marginal vegetation in pools in temporary stream systems (see
|
||
<figureCitation id="131AE7E0FD0FD500FB60FEC1FB00FE1A" box="[1193,1260,367,392]" captionStart="FIGURE 3" captionStartId="6.[151,250,1932,1954]" captionTargetBox="[162,1415,473,1899]" captionTargetId="figure@6.[162,1415,467,1901]" captionTargetPageId="6" captionText="FIGURE 3. Water beetle habitats on the Hantamsberg plateau, 1,500 m. A) Drying stream pool on gravel on northwest side of plateau; B) pool in stream 1 km north of mast; C) detail of residual pool in same stream system; D) temporary pool on clay / rock with Bulbinella nutans. Photos D. T. Bilton." httpUri="https://zenodo.org/record/252270/files/figure.png" pageId="6" pageNumber="477">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33BA8EEFD0FD500FF0EFE3AFBD3FE3F" box="[199,1087,404,429]" pageId="6" pageNumber="477" type="etymology">
|
||
<paragraph id="8B9EFB65FD0FD500FF0EFE3AFBD3FE3F" blockId="6.[151,1437,151,429]" box="[199,1087,404,429]" pageId="6" pageNumber="477">
|
||
<emphasis id="B9552777FD0FD500FF0EFE3AFEAAFE3F" bold="true" box="[199,326,404,429]" pageId="6" pageNumber="477">Etymology</emphasis>
|
||
. Named in reference to the straw yellow colouration of the dorsum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |