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<document id="616FC17C7116922235EE94C6883C1E54" ID-CLB-Dataset="25243" ID-DOI="10.11646/zootaxa.4686.1.7" ID-GBIF-Dataset="f1e0543c-796e-4d4c-beeb-97b178149afb" ID-ISSN="1175-5326" ID-Zenodo-Dep="3484397" ID-ZooBank="845277B2-ACCF-4CB2-8856-BE0164285D48" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1571037198940" checkinUser="plazi" docAuthor="Cheng, Zhi-Fei &amp; Peng, Zhong" docDate="2019" docId="03FDBC77FFF3FF93B2EF2CAA21FE15D3" docLanguage="en" docName="zootaxa.4686.1.7.pdf" docOrigin="Zootaxa 4686 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paederus (Gnathopaederus) zhangmuensis Cheng &amp; Peng 2019, new species" docType="treatment" docVersion="6" lastPageNumber="132" masterDocId="FFC4C40FFFF1FF96B278292522401463" masterDocTitle="A new species and additional records of brachypterous Paederus (Coleoptera, Staphylinidae) of mainland China and Taiwan" masterLastPageNumber="132" masterPageNumber="127" pageNumber="129" updateTime="1698758811933" updateUser="ExternalLinkService">
<mods:mods id="973B0ADCCF390295A2E98C4F265CA206" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0806DD735760091F76FD1E7DA65B25A6">
<mods:title id="74E520C5FA8BEE2F7D8D1DB4782EDDEE">A new species and additional records of brachypterous Paederus (Coleoptera, Staphylinidae) of mainland China and Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="8AC2B2C05B800E2A34D98265887EC074" type="personal">
<mods:role id="01B814C7660BCE48417EE79A0561822D">
<mods:roleTerm id="43F7EC4A42CA55E34D6FA194EA87CF37">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9D5A2B1A1F2DBB850B1C31B85D25BF81">Cheng, Zhi-Fei</mods:namePart>
</mods:name>
<mods:name id="3E0363AECC991F6FB2C94F35DF36B9F9" type="personal">
<mods:role id="8B9B45FDECCFFF5F554D782508CCAE61">
<mods:roleTerm id="560781B33D10E9D2CCA70303C588CBEB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="85BA52E39492990DE83FF4C10B72A143">Peng, Zhong</mods:namePart>
</mods:name>
<mods:typeOfResource id="C2F3CE71564C68A3AFA873D99FD3E0A0">text</mods:typeOfResource>
<mods:relatedItem id="66819BE269260A4B3C3F8D1EC5C5D909" type="host">
<mods:titleInfo id="BDF02A47450C592DB70B2336A19013EA">
<mods:title id="4E2D966B468B2D0E7DA14DEBB3156C92">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="490C688F45CE18EF4E0322D108DC655B">
<mods:date id="DB32083E614813139EBCDCE7B486746C">2019</mods:date>
<mods:detail id="5151451C07581E7EF5A09D2D98AE806E" type="pubDate">
<mods:number id="54CD9BAB8D96305825028A59DF85CBAB">2019-10-14</mods:number>
</mods:detail>
<mods:detail id="A919ED760FC4B5D3D448EA2C48BA1A50" type="volume">
<mods:number id="E4713C10E37751779E20EB050466F6EB">4686</mods:number>
</mods:detail>
<mods:detail id="97C996DDA8F1A0F55BC938B10016964C" type="issue">
<mods:number id="3B01D39DE4CE1EE5EA3823AC623B0268">1</mods:number>
</mods:detail>
<mods:extent id="FBD1DA1219727E0BD2FD16396F81E1E0" unit="page">
<mods:start id="3EE1435F8A602E13C8354F0F3CE0684B">127</mods:start>
<mods:end id="3C0ECDDD2EE913D0D0005D85606DB971">132</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9FDAAE8BDCD0E1795125A7734A385008">journal article</mods:classification>
<mods:identifier id="8572195F5EE0E56EEFFC553B0DB8C6A6" type="CLB-Dataset">25243</mods:identifier>
<mods:identifier id="6DD6720EFE8A987D4FB35CA82BA0B7F4" type="DOI">10.11646/zootaxa.4686.1.7</mods:identifier>
<mods:identifier id="120C7723030BAEB9598C07975B144965" type="GBIF-Dataset">f1e0543c-796e-4d4c-beeb-97b178149afb</mods:identifier>
<mods:identifier id="0F696164CBA7F047A0B13F9D689559EC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="765A61F67606A9D1D6AC1C18002A3C77" type="Zenodo-Dep">3484397</mods:identifier>
<mods:identifier id="353421AE74BF166D09B3AC699F8D5C31" type="ZooBank">845277B2-ACCF-4CB2-8856-BE0164285D48</mods:identifier>
</mods:mods>
<treatment id="03FDBC77FFF3FF93B2EF2CAA21FE15D3" ID-GBIF-Taxon="159709377" LSID="urn:lsid:plazi:treatment:03FDBC77FFF3FF93B2EF2CAA21FE15D3" httpUri="http://treatment.plazi.org/id/03FDBC77FFF3FF93B2EF2CAA21FE15D3" lastPageId="5" lastPageNumber="132" pageId="2" pageNumber="129">
<subSubSection id="C34E5EEAFFF3FF94B2EF2CAA265211CA" box="[151,1042,1423,1450]" pageId="2" pageNumber="129" type="nomenclature">
<paragraph id="8BEB0D61FFF3FF94B2EF2CAA265211CA" blockId="2.[151,1042,1423,1485]" box="[151,1042,1423,1450]" pageId="2" pageNumber="129">
<heading id="D0A3BA0DFFF3FF94B2EF2CAA265211CA" bold="true" box="[151,1042,1423,1450]" fontSize="11" level="1" pageId="2" pageNumber="129" reason="1">
<emphasis id="B920D173FFF3FF94B2EF2CAA265211CA" bold="true" box="[151,1042,1423,1450]" pageId="2" pageNumber="129">
<taxonomicName id="4C5476E2FFF3FF94B2EF2CAA213611CA" ID-CoL="8Y7P5" authority="Cheng and Peng" authorityName="Cheng &amp; Peng" authorityYear="2019" box="[151,886,1423,1450]" class="Insecta" family="Staphylinidae" genus="Paederus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="zhangmuensis" status="sp. nov." subGenus="Gnathopaederus">
<emphasis id="B920D173FFF3FF94B2EF2CAA234711C9" bold="true" box="[151,263,1423,1450]" italics="true" pageId="2" pageNumber="129">Paederus</emphasis>
(
<emphasis id="B920D173FFF3FF94B3612CAA23A111C9" bold="true" box="[281,481,1423,1450]" italics="true" pageId="2" pageNumber="129">Gnathopaederus</emphasis>
)
<emphasis id="B920D173FFF3FF94B38A2CAA20DF11C9" bold="true" box="[498,671,1423,1450]" italics="true" pageId="2" pageNumber="129">zhangmuensis</emphasis>
Cheng and Peng
</taxonomicName>
,
<taxonomicNameLabel id="A2136C08FFF3FF94B1FA2CAA265211CA" box="[898,1042,1423,1449]" pageId="2" pageNumber="129" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E5EEAFFF3FF94B2EF2C96234B11AE" box="[151,267,1459,1485]" pageId="2" pageNumber="129" type="description">
<paragraph id="8BEB0D61FFF3FF94B2EF2C96234B11AE" blockId="2.[151,1042,1423,1485]" box="[151,267,1459,1485]" pageId="2" pageNumber="129">
(
<figureCitation id="136F11E4FFF3FF94B2E72C9622A811AE" box="[159,232,1459,1485]" captionStart="FIGURE 2" captionStartId="3.[151,250,1398,1423]" captionTargetBox="[162,1388,175,1369]" captionTargetId="figure@3.[151,1401,166,1378]" captionTargetPageId="3" captionText="FIGURE 2. Paederus (Gnathopaederus) zhangmuensis. A—Habitus; B—female mouthparts; C—male mouthparts. D—male tergite V; E—male tergite VI; F—male sternite VII. Scale bars: A 3.0 mm, B–F 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484401" httpUri="https://zenodo.org/record/3484401/files/figure.png" pageId="2" pageNumber="129">Figs 2</figureCitation>
,
<figureCitation id="136F11E4FFF3FF94B28C2C96234211AE" box="[244,258,1459,1485]" captionStart="FIGURE 3" captionStartId="4.[151,250,1937,1962]" captionTargetBox="[200,1408,174,1909]" captionTargetId="figure@4.[179,1408,166,1917]" captionTargetPageId="4" captionText="FIGURE 3. Paederus (Gnathopaederus) zhangmuensis. A—male forebody; B—female tergite VIII; C—female sternite VIII. D—male tergite VIII; E—male sternite VIII; F—aedeagus in ventral view; G—aedeagus in lateral view; H—aedeagus in dorsal view. Scale bars: A 3.0 mm, B–H 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484403" httpUri="https://zenodo.org/record/3484403/files/figure.png" pageId="2" pageNumber="129">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E5EEAFFF3FF94B2EF2CDE273D12C6" pageId="2" pageNumber="129" type="materials_examined">
<paragraph id="8BEB0D61FFF3FF94B2EF2CDE273D12C6" blockId="2.[151,1437,1530,2025]" pageId="2" pageNumber="129">
<emphasis id="B920D173FFF3FF94B2EF2CDE23031276" bold="true" box="[151,323,1530,1557]" pageId="2" pageNumber="129">Type material.</emphasis>
<materialsCitation id="3B3C073CFFF3FF94B3302CDE232B125A" ID-GBIF-Occurrence="2425496252" collectingDate="2010-07-26" collectorName="Jian-Qing Zhu" country="China" county="Nielamu County" elevation="2600" location="Lixin" municipality="Zhangmu" pageId="2" pageNumber="129" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="54EFB3C3FFF3FF94B3302CDE23F51270" box="[328,437,1531,1557]" pageId="2" pageNumber="129" type="holotype">
<subScript id="17D00F24FFF3FF94B3302CDE23F51270" attach="left" box="[328,437,1531,1557]" fontSize="7" pageId="2" pageNumber="129">HOLOTYPE</subScript>
</typeStatus>
:
<specimenCount id="9D52C6E8FFF3FF94B3BA2CDE23991276" box="[450,473,1531,1557]" pageId="2" pageNumber="129" type="male">♂</specimenCount>
, labeled ‘
<collectingCountry id="F3434DF1FFF3FF94B0452CDE20C91276" box="[573,649,1531,1557]" name="China" pageId="2" pageNumber="129">China</collectingCountry>
:
<collectingRegion id="4990C383FFF3FF94B0E92CDE21641276" box="[657,804,1531,1557]" country="China" name="Xizang" pageId="2" pageNumber="129">Xizang Prov.</collectingRegion>
,
<collectingCounty id="628A75EDFFF3FF94B1552CDE21A61276" box="[813,998,1531,1557]" pageId="2" pageNumber="129">Nielamu County</collectingCounty>
,
<collectingMunicipality id="6B8F971BFFF3FF94B1962CDE261C1276" box="[1006,1116,1531,1557]" pageId="2" pageNumber="129">Zhangmu</collectingMunicipality>
,
<location id="8E8B5BBAFFF3FF94B61D2CDE26E41276" LSID="urn:lsid:plazi:treatment:03FDBC77FFF3FF93B2EF2CAA21FE15D3:8E8B5BBAFFF3FF94B61D2CDE26E41276" box="[1125,1188,1531,1557]" country="China" county="Nielamu County" municipality="Zhangmu" name="Lixin" pageId="2" pageNumber="129" stateProvince="Xizang">Lixin</location>
,
<quantity id="4CACA084FFF3FF94B6D62CDE27431276" box="[1198,1283,1531,1557]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="129" unit="m" value="2600.0">
<elevation id="0079EA52FFF3FF94B6D62CDE27431276" box="[1198,1283,1531,1557]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="129" unit="m" value="2600.0">2600 m</elevation>
</quantity>
,
<date id="FFEA2BA1FFF3FF94B7742CDE27D81276" box="[1292,1432,1531,1557]" pageId="2" pageNumber="129" value="2010-07-26">
<collectingDate id="EFAED249FFF3FF94B7742CDE27D81276" box="[1292,1432,1531,1557]" pageId="2" pageNumber="129" value="2010-07-26">26.VII.2010</collectingDate>
</date>
,
<collectorName id="26A168B7FFF3FF94B2EF2F3A2379125A" box="[151,313,1567,1593]" pageId="2" pageNumber="129">Jian-Qing Zhu</collectorName>
leg.
</materialsCitation>
’ /
<materialsCitation id="3B3C073CFFF3FF94B3FC2F3A2717125A" ID-GBIF-Occurrence="2425496253" box="[388,1367,1567,1594]" collectionCode="SNUC" collectorName="Peng" country="China" determinerName="Cheng" pageId="2" pageNumber="129" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="54EFB3C3FFF3FF94B3FC2F3A2058125A" box="[388,536,1567,1593]" pageId="2" pageNumber="129" type="holotype">HOLOTYPE</typeStatus>
(red),
<taxonomicName id="4C5476E2FFF3FF94B01A2F3A212F125A" authority="Cheng &amp; Peng, 2019" authorityName="Cheng &amp; Peng" authorityYear="2019" box="[610,879,1567,1593]" class="Insecta" family="Staphylinidae" genus="Paederus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="zhangmuensis" status="sp. nov.">
<emphasis id="B920D173FFF3FF94B01A2F3A212F125A" box="[610,879,1567,1593]" italics="true" pageId="2" pageNumber="129">Paederus zhangmuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2136C08FFF3FF94B10D2F3A218E125A" box="[885,974,1567,1593]" pageId="2" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
, det.
<determinerName id="7E37E1A5FFF3FF94B67F2F3A2610125A" box="[1031,1104,1567,1593]" pageId="2" pageNumber="129">Cheng</determinerName>
&amp;
<collectorName id="26A168B7FFF3FF94B60B2F3A26EE125A" box="[1139,1198,1567,1593]" pageId="2" pageNumber="129">Peng</collectorName>
, 2019 (
<collectionCode id="ED4595A4FFF3FF94B6872F3A2710125A" box="[1279,1360,1567,1593]" pageId="2" pageNumber="129">SNUC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54EFB3C3FFF3FF94B7192F3A2296123E" pageId="2" pageNumber="129" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D52C6E8FFF3FF94B2992F662362123E" box="[225,290,1603,1629]" pageId="2" pageNumber="129" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D52C6E8FFF3FF94B3552F662317123E" box="[301,343,1603,1629]" pageId="2" pageNumber="129" type="female">1 ♀</specimenCount>
, same label data as
<typeStatus id="54EFB3C3FFF3FF94B04E2F6620D8123E" box="[566,664,1603,1629]" pageId="2" pageNumber="129" type="holotype">holotype</typeStatus>
/
<materialsCitation id="3B3C073CFFF3FF94B0D52F6623E112E2" ID-GBIF-Occurrence="2425496251" collectionCode="SNUC" collectorName="Peng" country="China" determinerName="Cheng" pageId="2" pageNumber="129" specimenCount="1" stateProvince="Xizang" typeStatus="paratype">
<typeStatus id="54EFB3C3FFF3FF94B0D52F662178123E" box="[685,824,1603,1629]" pageId="2" pageNumber="129" type="paratype">PARATYPE</typeStatus>
(yellow),
<taxonomicName id="4C5476E2FFF3FF94B1D32F6626F9123E" authority="Cheng &amp; Peng, 2019" authorityName="Cheng &amp; Peng" authorityYear="2019" box="[939,1209,1603,1629]" class="Insecta" family="Staphylinidae" genus="Paederus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="zhangmuensis" status="sp. nov.">
<emphasis id="B920D173FFF3FF94B1D32F6626F9123E" box="[939,1209,1603,1629]" italics="true" pageId="2" pageNumber="129">Paederus zhangmuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2136C08FFF3FF94B6B82F662758123E" box="[1216,1304,1603,1629]" pageId="2" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
, det.
<determinerName id="7E37E1A5FFF3FF94B72B2F6627DC123E" box="[1363,1436,1603,1629]" pageId="2" pageNumber="129">Cheng</determinerName>
&amp;
<collectorName id="26A168B7FFF3FF94B2CE2F4222B112E2" box="[182,241,1639,1665]" pageId="2" pageNumber="129">Peng</collectorName>
, 2019 (
<collectionCode id="ED4595A4FFF3FF94B3302F4223D912E2" box="[328,409,1639,1665]" pageId="2" pageNumber="129">SNUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C073CFFF3FF94B3D62F42232412C6" ID-GBIF-Occurrence="2425496257" collectingDate="2010-07-24" collectorName="Wen-Xuan Bi" country="China" county="Nielamu County" elevation="3200" location="Quxian" municipality="Zhangmu" pageId="2" pageNumber="129" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Xizang" typeStatus="paratype">
<specimenCount id="9D52C6E8FFF3FF94B3D62F4223B212E2" box="[430,498,1639,1665]" pageId="2" pageNumber="129" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D52C6E8FFF3FF94B0782F42206E12E2" box="[512,558,1639,1665]" pageId="2" pageNumber="129" type="female">1 ♀</specimenCount>
: ‘
<collectingCountry id="F3434DF1FFF3FF94B03A2F4220CE12E2" box="[578,654,1639,1665]" name="China" pageId="2" pageNumber="129">China</collectingCountry>
:
<collectingRegion id="4990C383FFF3FF94B0E32F4220AF12E2" box="[667,751,1639,1665]" country="China" name="Xizang" pageId="2" pageNumber="129">Xizang</collectingRegion>
,
<collectingCounty id="628A75EDFFF3FF94B0842F4221FA12E2" box="[764,954,1639,1665]" pageId="2" pageNumber="129">Nielamu County</collectingCounty>
,
<collectingMunicipality id="6B8F971BFFF3FF94B1BF2F42267512E2" box="[967,1077,1639,1665]" pageId="2" pageNumber="129">Zhangmu</collectingMunicipality>
,
<location id="8E8B5BBAFFF3FF94B63A2F4226D712E2" LSID="urn:lsid:plazi:treatment:03FDBC77FFF3FF93B2EF2CAA21FE15D3:8E8B5BBAFFF3FF94B63A2F4226D712E2" box="[1090,1175,1639,1665]" country="China" county="Nielamu County" municipality="Zhangmu" name="Quxian" pageId="2" pageNumber="129" stateProvince="Xizang">Quxian</location>
,
<quantity id="4CACA084FFF3FF94B6DD2F4226BE12E2" box="[1189,1278,1639,1665]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="129" unit="m" value="3200.0">
<elevation id="0079EA52FFF3FF94B6DD2F4226BE12E2" box="[1189,1278,1639,1665]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="129" unit="m" value="3200.0">3200 m</elevation>
</quantity>
,
<date id="FFEA2BA1FFF3FF94B7742F4227D812E2" box="[1292,1432,1639,1665]" pageId="2" pageNumber="129" value="2010-07-24">
<collectingDate id="EFAED249FFF3FF94B7742F4227D812E2" box="[1292,1432,1639,1665]" pageId="2" pageNumber="129" value="2010-07-24">24.VII.2010</collectingDate>
</date>
,
<collectorName id="26A168B7FFF3FF94B2EF2FAE237112C6" box="[151,305,1675,1701]" pageId="2" pageNumber="129">Wen-Xuan Bi</collectorName>
leg.
</materialsCitation>
’ /
<materialsCitation id="3B3C073CFFF3FF94B3062FAE273912C6" ID-GBIF-Occurrence="2425496250" box="[382,1401,1675,1702]" collectionCode="SNUC" collectorName="Peng" country="China" determinerName="Cheng" pageId="2" pageNumber="129" specimenCount="1" stateProvince="Xizang" typeStatus="paratype">
<typeStatus id="54EFB3C3FFF3FF94B3062FAE204912C6" box="[382,521,1675,1701]" pageId="2" pageNumber="129" type="paratype">PARATYPE</typeStatus>
(yellow),
<taxonomicName id="4C5476E2FFF3FF94B0052FAE21CB12C6" authority="Cheng &amp; Peng, 2019" authorityName="Cheng &amp; Peng" authorityYear="2019" box="[637,907,1675,1701]" class="Insecta" family="Staphylinidae" genus="Paederus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="zhangmuensis" status="sp. nov.">
<emphasis id="B920D173FFF3FF94B0052FAE21CB12C6" box="[637,907,1675,1701]" italics="true" pageId="2" pageNumber="129">Paederus zhangmuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2136C08FFF3FF94B1EA2FAE21AB12C6" box="[914,1003,1675,1701]" pageId="2" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
, det.
<determinerName id="7E37E1A5FFF3FF94B65E2FAE262F12C6" box="[1062,1135,1675,1701]" pageId="2" pageNumber="129">Cheng</determinerName>
&amp;
<collectorName id="26A168B7FFF3FF94B6EB2FAE268E12C6" box="[1171,1230,1675,1701]" pageId="2" pageNumber="129">Peng</collectorName>
, 2019 (
<collectionCode id="ED4595A4FFF3FF94B7582FAE273112C6" box="[1312,1393,1675,1701]" pageId="2" pageNumber="129">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E5EEAFFF3FF93B2BF2F8B21131520" lastPageId="5" lastPageNumber="132" pageId="2" pageNumber="129" type="description">
<paragraph id="8BEB0D61FFF3FF94B2BF2F8B23231356" blockId="2.[151,1437,1530,2025]" pageId="2" pageNumber="129">
<emphasis id="B920D173FFF3FF94B2BF2F8B231912AA" bold="true" box="[199,345,1710,1737]" pageId="2" pageNumber="129">Description.</emphasis>
Measurements (in mm) and ratios: BL: 11.18–11.31; FL: 5.56–5.72; HL: 0.76–0.80; HW: 0.83– 0.91; AnL: 3.06–3.39; PL: 0.92–0.94; PW: 0.83–0.87; EL: 0.80–0.89; EW: 0.83–0.96; AW: 1.04–1.10; AL: 0.96– 1.06; HL/HW: 0.88–0.92; HW/PW: 1.00–1.05; HL/PL: 0.82–0.85; PL/PW: 1.08–1.12; EL/PL: 0.87–0.95; diameter of eye: 0.21–0.23.
</paragraph>
<paragraph id="8BEB0D61FFF3FF94B2BF2E1A20E913C2" blockId="2.[151,1437,1530,2025]" pageId="2" pageNumber="129">
Habitus as in
<figureCitation id="136F11E4FFF3FF94B3262E1A23F7133A" box="[350,439,1855,1881]" captionStart="FIGURE 2" captionStartId="3.[151,250,1398,1423]" captionTargetBox="[162,1388,175,1369]" captionTargetId="figure@3.[151,1401,166,1378]" captionTargetPageId="3" captionText="FIGURE 2. Paederus (Gnathopaederus) zhangmuensis. A—Habitus; B—female mouthparts; C—male mouthparts. D—male tergite V; E—male tergite VI; F—male sternite VII. Scale bars: A 3.0 mm, B–F 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484401" httpUri="https://zenodo.org/record/3484401/files/figure.png" pageId="2" pageNumber="129">Fig. 2A</figureCitation>
. Coloration: head and labrum black, labial palpi and antennae brown; pronotum brownish red; elytra metallic-blue; legs brown, with apical halves of the femora blackish; abdomen bicoloured with segments III-VI brownish red and segments VII-X black.
</paragraph>
<paragraph id="8BEB0D61FFF3FF94B2BF2E8E238A138A" blockId="2.[151,1437,1530,2025]" pageId="2" pageNumber="129">
Head wider than long, widest across eyes; punctation coarse and sparse; interstices glossy. Eyes distinctly convex. Antenna as in
<figureCitation id="136F11E4FFF3FF94B3152EEA2386138A" box="[365,454,1999,2025]" captionStart="FIGURE 2" captionStartId="3.[151,250,1398,1423]" captionTargetBox="[162,1388,175,1369]" captionTargetId="figure@3.[151,1401,166,1378]" captionTargetPageId="3" captionText="FIGURE 2. Paederus (Gnathopaederus) zhangmuensis. A—Habitus; B—female mouthparts; C—male mouthparts. D—male tergite V; E—male tergite VI; F—male sternite VII. Scale bars: A 3.0 mm, B–F 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484401" httpUri="https://zenodo.org/record/3484401/files/figure.png" pageId="2" pageNumber="129">Fig. 2A</figureCitation>
.
</paragraph>
<caption id="DF2B5DE9FFF2FF95B2EF2C53265B11D1" ID-DOI="http://doi.org/10.5281/zenodo.3484401" ID-Zenodo-Dep="3484401" httpUri="https://zenodo.org/record/3484401/files/figure.png" pageId="3" pageNumber="130" startId="3.[151,250,1398,1423]" targetBox="[162,1388,175,1369]" targetPageId="3">
<paragraph id="8BEB0D61FFF2FF95B2EF2C53265B11D1" blockId="3.[151,1437,1398,1459]" pageId="3" pageNumber="130">
<emphasis id="B920D173FFF2FF95B2EF2C53235411EC" bold="true" box="[151,276,1398,1423]" pageId="3" pageNumber="130">FIGURE 2.</emphasis>
<taxonomicName id="4C5476E2FFF2FF95B3632C53209411ED" authorityName="Cheng &amp; Peng" authorityYear="2019" box="[283,724,1398,1422]" class="Insecta" family="Staphylinidae" genus="Paederus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="zhangmuensis" subGenus="Gnathopaederus">
<emphasis id="B920D173FFF2FF95B3632C53233B11ED" box="[283,379,1398,1422]" italics="true" pageId="3" pageNumber="130">Paederus</emphasis>
(
<emphasis id="B920D173FFF2FF95B3F22C53207311ED" box="[394,563,1398,1422]" italics="true" pageId="3" pageNumber="130">Gnathopaederus</emphasis>
)
<emphasis id="B920D173FFF2FF95B03B2C53209411ED" box="[579,724,1398,1422]" italics="true" pageId="3" pageNumber="130">zhangmuensis</emphasis>
</taxonomicName>
<emphasis id="B920D173FFF2FF95B0AC2C53209811ED" box="[724,728,1398,1422]" italics="true" pageId="3" pageNumber="130">.</emphasis>
<emphasis id="B920D173FFF2FF95B0A52C5320AF11EC" bold="true" box="[733,751,1398,1423]" pageId="3" pageNumber="130">A</emphasis>
—Habitus;
<emphasis id="B920D173FFF2FF95B11D2C52213611EC" bold="true" box="[869,886,1399,1423]" pageId="3" pageNumber="130">B</emphasis>
—female mouthparts;
<emphasis id="B920D173FFF2FF95B6242C53262E11EC" bold="true" box="[1116,1134,1398,1423]" pageId="3" pageNumber="130">C</emphasis>
—male mouthparts.
<emphasis id="B920D173FFF2FF95B7472C52271111EC" bold="true" box="[1343,1361,1399,1423]" pageId="3" pageNumber="130">D</emphasis>
—male tergite V;
<emphasis id="B920D173FFF2FF95B2832CBE234C11D0" bold="true" box="[251,268,1435,1459]" pageId="3" pageNumber="130">E</emphasis>
—male tergite VI;
<emphasis id="B920D173FFF2FF95B3B32CBE239A11D0" bold="true" box="[459,474,1435,1459]" pageId="3" pageNumber="130">F</emphasis>
—male sternite VII. Scale bars:
<emphasis id="B920D173FFF2FF95B1592CBF217311D0" bold="true" box="[801,819,1434,1459]" pageId="3" pageNumber="130">A</emphasis>
3.0 mm,
<emphasis id="B920D173FFF2FF95B1EB2CBE218011D0" bold="true" box="[915,960,1435,1459]" pageId="3" pageNumber="130">B–F</emphasis>
0.5 mm.
</paragraph>
</caption>
<paragraph id="8BEB0D61FFF2FF95B2BF2CC42374127C" blockId="3.[151,1437,1505,2035]" pageId="3" pageNumber="130">Pronotum nearly oviform, strongly convex in cross-section; punctures sparser and somewhat shallower than those of head.</paragraph>
<paragraph id="8BEB0D61FFF2FF95B2BF2F0C20F91204" blockId="3.[151,1437,1505,2035]" pageId="3" pageNumber="130">Elytra nearly trapeziform; punctation coarse, defined and dense. Hind wings reduced. Metatarsomere I as long as combined length of metatarsomeres II and III.</paragraph>
<paragraph id="8BEB0D61FFF2FF95B2BF2F5420BB12CC" blockId="3.[151,1437,1505,2035]" pageId="3" pageNumber="130">Abdomen distinctly broader than elytra; punctation moderately coarse; interstices with shallow microsculpture; posterior margin of tergite VII without palisade fringe.</paragraph>
<paragraph id="8BEB0D61FFF2FF95B2BF2F9C21C81390" blockId="3.[151,1437,1505,2035]" pageId="3" pageNumber="130">
Male. Anterior margin of labrum (
<figureCitation id="136F11E4FFF2FF95B0372F9C20E712B0" box="[591,679,1721,1747]" captionStart="FIGURE 2" captionStartId="3.[151,250,1398,1423]" captionTargetBox="[162,1388,175,1369]" captionTargetId="figure@3.[151,1401,166,1378]" captionTargetPageId="3" captionText="FIGURE 2. Paederus (Gnathopaederus) zhangmuensis. A—Habitus; B—female mouthparts; C—male mouthparts. D—male tergite V; E—male tergite VI; F—male sternite VII. Scale bars: A 3.0 mm, B–F 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484401" httpUri="https://zenodo.org/record/3484401/files/figure.png" pageId="3" pageNumber="130">Fig. 2C</figureCitation>
) in middle with shallow excision. left mandible (
<figureCitation id="136F11E4FFF2FF95B6A52F9C277512B0" box="[1245,1333,1721,1747]" captionStart="FIGURE 2" captionStartId="3.[151,250,1398,1423]" captionTargetBox="[162,1388,175,1369]" captionTargetId="figure@3.[151,1401,166,1378]" captionTargetPageId="3" captionText="FIGURE 2. Paederus (Gnathopaederus) zhangmuensis. A—Habitus; B—female mouthparts; C—male mouthparts. D—male tergite V; E—male tergite VI; F—male sternite VII. Scale bars: A 3.0 mm, B–F 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484401" httpUri="https://zenodo.org/record/3484401/files/figure.png" pageId="3" pageNumber="130">Fig. 2C</figureCitation>
) without dorsal tooth, inner margin with one small tooth; right mandible (
<figureCitation id="136F11E4FFF2FF95B1272FF821F41294" box="[863,948,1757,1783]" captionStart="FIGURE 2" captionStartId="3.[151,250,1398,1423]" captionTargetBox="[162,1388,175,1369]" captionTargetId="figure@3.[151,1401,166,1378]" captionTargetPageId="3" captionText="FIGURE 2. Paederus (Gnathopaederus) zhangmuensis. A—Habitus; B—female mouthparts; C—male mouthparts. D—male tergite V; E—male tergite VI; F—male sternite VII. Scale bars: A 3.0 mm, B–F 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484401" httpUri="https://zenodo.org/record/3484401/files/figure.png" pageId="3" pageNumber="130">Fig. 2C</figureCitation>
) with one indistinct dorsal tooth, inner margin with single-pointed tooth. Posterior margin of tergite VIII (
<figureCitation id="136F11E4FFF2FF95B1232E2421F21378" box="[859,946,1793,1819]" captionStart="FIGURE 3" captionStartId="4.[151,250,1937,1962]" captionTargetBox="[200,1408,174,1909]" captionTargetId="figure@4.[179,1408,166,1917]" captionTargetPageId="4" captionText="FIGURE 3. Paederus (Gnathopaederus) zhangmuensis. A—male forebody; B—female tergite VIII; C—female sternite VIII. D—male tergite VIII; E—male sternite VIII; F—aedeagus in ventral view; G—aedeagus in lateral view; H—aedeagus in dorsal view. Scale bars: A 3.0 mm, B–H 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484403" httpUri="https://zenodo.org/record/3484403/files/figure.png" pageId="3" pageNumber="130">Fig. 3D</figureCitation>
) strongly convex; sternite V (
<figureCitation id="136F11E4FFF2FF95B77B2E24271B1378" box="[1283,1371,1793,1819]" captionStart="FIGURE 2" captionStartId="3.[151,250,1398,1423]" captionTargetBox="[162,1388,175,1369]" captionTargetId="figure@3.[151,1401,166,1378]" captionTargetPageId="3" captionText="FIGURE 2. Paederus (Gnathopaederus) zhangmuensis. A—Habitus; B—female mouthparts; C—male mouthparts. D—male tergite V; E—male tergite VI; F—male sternite VII. Scale bars: A 3.0 mm, B–F 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484401" httpUri="https://zenodo.org/record/3484401/files/figure.png" pageId="3" pageNumber="130">Fig. 2D</figureCitation>
) with shallow median impression posteriorly, this impression with weakly modified and dark setae; sternite VI (
<figureCitation id="136F11E4FFF2FF95B7472E0027D4135C" box="[1343,1428,1829,1855]" captionStart="FIGURE 2" captionStartId="3.[151,250,1398,1423]" captionTargetBox="[162,1388,175,1369]" captionTargetId="figure@3.[151,1401,166,1378]" captionTargetPageId="3" captionText="FIGURE 2. Paederus (Gnathopaederus) zhangmuensis. A—Habitus; B—female mouthparts; C—male mouthparts. D—male tergite V; E—male tergite VI; F—male sternite VII. Scale bars: A 3.0 mm, B–F 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484401" httpUri="https://zenodo.org/record/3484401/files/figure.png" pageId="3" pageNumber="130">Fig. 2E</figureCitation>
) with distinct median impression posteriorly, this impression with weakly modified and dark setae; sternite VII (
<figureCitation id="136F11E4FFF2FF95B7172E6C22F613E4" captionStart="FIGURE 2" captionStartId="3.[151,250,1398,1423]" captionTargetBox="[162,1388,175,1369]" captionTargetId="figure@3.[151,1401,166,1378]" captionTargetPageId="3" captionText="FIGURE 2. Paederus (Gnathopaederus) zhangmuensis. A—Habitus; B—female mouthparts; C—male mouthparts. D—male tergite V; E—male tergite VI; F—male sternite VII. Scale bars: A 3.0 mm, B–F 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484401" httpUri="https://zenodo.org/record/3484401/files/figure.png" pageId="3" pageNumber="130">Fig. 2F</figureCitation>
) strongly transverse, with distinct median impression posteriorly, this impression with numerous modified, short black setae, posterior excision rather shallow; sternite VIII (
<figureCitation id="136F11E4FFF2FF95B1542EB4213F13C8" box="[812,895,1937,1963]" captionStart="FIGURE 3" captionStartId="4.[151,250,1937,1962]" captionTargetBox="[200,1408,174,1909]" captionTargetId="figure@4.[179,1408,166,1917]" captionTargetPageId="4" captionText="FIGURE 3. Paederus (Gnathopaederus) zhangmuensis. A—male forebody; B—female tergite VIII; C—female sternite VIII. D—male tergite VIII; E—male sternite VIII; F—aedeagus in ventral view; G—aedeagus in lateral view; H—aedeagus in dorsal view. Scale bars: A 3.0 mm, B–H 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484403" httpUri="https://zenodo.org/record/3484403/files/figure.png" pageId="3" pageNumber="130">Fig. 3E</figureCitation>
) transverse and with deep posterior incision, this incision 0.5 times as long as sternite VIII; aedeagus as in
<figureCitation id="136F11E4FFF2FF95B16D2E90211813AC" box="[789,856,1973,1999]" captionStart="FIGURE 3" captionStartId="4.[151,250,1937,1962]" captionTargetBox="[200,1408,174,1909]" captionTargetId="figure@4.[179,1408,166,1917]" captionTargetPageId="4" captionText="FIGURE 3. Paederus (Gnathopaederus) zhangmuensis. A—male forebody; B—female tergite VIII; C—female sternite VIII. D—male tergite VIII; E—male sternite VIII; F—aedeagus in ventral view; G—aedeagus in lateral view; H—aedeagus in dorsal view. Scale bars: A 3.0 mm, B–H 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484403" httpUri="https://zenodo.org/record/3484403/files/figure.png" pageId="3" pageNumber="130">Figs 3</figureCitation>
F–H; dorsal plate apically acute; parameres apically very weakly curved; internal sac with single long sclerotized spine.
</paragraph>
<caption id="DF2B5DE9FFF5FF92B2EF2EB420071392" ID-DOI="http://doi.org/10.5281/zenodo.3484403" ID-Zenodo-Dep="3484403" httpUri="https://zenodo.org/record/3484403/files/figure.png" pageId="4" pageNumber="131" startId="4.[151,250,1937,1962]" targetBox="[200,1408,174,1909]" targetPageId="4">
<paragraph id="8BEB0D61FFF5FF92B2EF2EB420071392" blockId="4.[151,1436,1937,2034]" pageId="4" pageNumber="131">
<emphasis id="B920D173FFF5FF92B2EF2EB4235413C9" bold="true" box="[151,276,1937,1962]" pageId="4" pageNumber="131">FIGURE 3.</emphasis>
<taxonomicName id="4C5476E2FFF5FF92B3632EB7209113C9" authorityName="Cheng &amp; Peng" authorityYear="2019" box="[283,721,1938,1962]" class="Insecta" family="Staphylinidae" genus="Paederus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="131" phylum="Arthropoda" rank="species" species="zhangmuensis" subGenus="Gnathopaederus">
<emphasis id="B920D173FFF5FF92B3632EB7233B13C9" box="[283,379,1938,1962]" italics="true" pageId="4" pageNumber="131">Paederus</emphasis>
(
<emphasis id="B920D173FFF5FF92B3F22EB7207313C9" box="[394,563,1938,1962]" italics="true" pageId="4" pageNumber="131">Gnathopaederus</emphasis>
)
<emphasis id="B920D173FFF5FF92B03B2EB7209113C9" box="[579,721,1938,1962]" italics="true" pageId="4" pageNumber="131">zhangmuensis</emphasis>
</taxonomicName>
.
<emphasis id="B920D173FFF5FF92B0A52EB420AF13C9" bold="true" box="[733,751,1937,1962]" pageId="4" pageNumber="131">A</emphasis>
—male forebody;
<emphasis id="B920D173FFF5FF92B1D22EB721FB13C9" bold="true" box="[938,955,1938,1962]" pageId="4" pageNumber="131">B</emphasis>
—female tergite VIII;
<emphasis id="B920D173FFF5FF92B6D92EB426F313C9" bold="true" box="[1185,1203,1937,1962]" pageId="4" pageNumber="131">C</emphasis>
—female sternite VIII.
<emphasis id="B920D173FFF5FF92B2EF2E9322E913AD" bold="true" box="[151,169,1974,1998]" pageId="4" pageNumber="131">D</emphasis>
—male tergite VIII;
<emphasis id="B920D173FFF5FF92B30E2E9323C713AD" bold="true" box="[374,391,1974,1998]" pageId="4" pageNumber="131">E</emphasis>
—male sternite VIII;
<emphasis id="B920D173FFF5FF92B0272E93202E13AD" bold="true" box="[607,622,1974,1998]" pageId="4" pageNumber="131">F</emphasis>
—aedeagus in ventral view;
<emphasis id="B920D173FFF5FF92B1F42E9021E013AD" bold="true" box="[908,928,1973,1998]" pageId="4" pageNumber="131">G</emphasis>
—aedeagus in lateral view;
<emphasis id="B920D173FFF5FF92B6CE2E93268A13AD" bold="true" box="[1206,1226,1974,1998]" pageId="4" pageNumber="131">H</emphasis>
—aedeagus in dorsal view. Scale bars:
<emphasis id="B920D173FFF5FF92B3302EFC231A1391" bold="true" box="[328,346,2009,2034]" pageId="4" pageNumber="131">A</emphasis>
3.0 mm,
<emphasis id="B920D173FFF5FF92B3C32EFF23AC1391" bold="true" box="[443,492,2010,2034]" pageId="4" pageNumber="131">B–H</emphasis>
0.5 mm.
</paragraph>
</caption>
<paragraph id="8BEB0D61FFF4FF93B2BF29BC238A14B4" blockId="5.[151,1437,153,432]" pageId="5" pageNumber="132">
Female. Labrum as in
<figureCitation id="136F11E4FFF4FF93B3BB29BC205A14D0" box="[451,538,153,179]" captionStart="FIGURE 2" captionStartId="3.[151,250,1398,1423]" captionTargetBox="[162,1388,175,1369]" captionTargetId="figure@3.[151,1401,166,1378]" captionTargetPageId="3" captionText="FIGURE 2. Paederus (Gnathopaederus) zhangmuensis. A—Habitus; B—female mouthparts; C—male mouthparts. D—male tergite V; E—male tergite VI; F—male sternite VII. Scale bars: A 3.0 mm, B–F 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484401" httpUri="https://zenodo.org/record/3484401/files/figure.png" pageId="5" pageNumber="132">Fig. 2B</figureCitation>
. Posterior margin of tergite VIII (
<figureCitation id="136F11E4FFF4FF93B1EC29BC21AA14D0" box="[916,1002,153,179]" captionStart="FIGURE 3" captionStartId="4.[151,250,1937,1962]" captionTargetBox="[200,1408,174,1909]" captionTargetId="figure@4.[179,1408,166,1917]" captionTargetPageId="4" captionText="FIGURE 3. Paederus (Gnathopaederus) zhangmuensis. A—male forebody; B—female tergite VIII; C—female sternite VIII. D—male tergite VIII; E—male sternite VIII; F—aedeagus in ventral view; G—aedeagus in lateral view; H—aedeagus in dorsal view. Scale bars: A 3.0 mm, B–H 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484403" httpUri="https://zenodo.org/record/3484403/files/figure.png" pageId="5" pageNumber="132">Fig. 3B</figureCitation>
) convex; sternite VIII (
<figureCitation id="136F11E4FFF4FF93B68A29BC270814D0" box="[1266,1352,153,179]" captionStart="FIGURE 3" captionStartId="4.[151,250,1937,1962]" captionTargetBox="[200,1408,174,1909]" captionTargetId="figure@4.[179,1408,166,1917]" captionTargetPageId="4" captionText="FIGURE 3. Paederus (Gnathopaederus) zhangmuensis. A—male forebody; B—female tergite VIII; C—female sternite VIII. D—male tergite VIII; E—male sternite VIII; F—aedeagus in ventral view; G—aedeagus in lateral view; H—aedeagus in dorsal view. Scale bars: A 3.0 mm, B–H 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3484403" httpUri="https://zenodo.org/record/3484403/files/figure.png" pageId="5" pageNumber="132">Fig. 3C</figureCitation>
) poste- riorly with median process.
</paragraph>
<paragraph id="8BEB0D61FFF4FF93B2BF29C521131520" blockId="5.[151,1437,153,432]" pageId="5" pageNumber="132">
<emphasis id="B920D173FFF4FF93B2BF29C523F01498" bold="true" box="[199,432,224,251]" pageId="5" pageNumber="132">Comparative notes.</emphasis>
This new species is distinguished from other species of this subgenus by an indistinct dorsal tooth on the right mandible, the chaetotaxy of the male sternites V–VII and the structure of the aedeagus (shape of parameres; single long sclerotized spine in the internal sac).
</paragraph>
</subSubSection>
<subSubSection id="C34E5EEAFFF4FF93B2BF2869274215E8" pageId="5" pageNumber="132" type="distribution">
<paragraph id="8BEB0D61FFF4FF93B2BF2869274215E8" blockId="5.[151,1437,153,432]" pageId="5" pageNumber="132">
<emphasis id="B920D173FFF4FF93B2BF2869200F1504" bold="true" box="[199,591,332,359]" pageId="5" pageNumber="132">Distribution and natural history.</emphasis>
The
<typeStatus id="54EFB3C3FFF4FF93B0F0286820F81504" box="[648,696,333,359]" pageId="5" pageNumber="132">type</typeStatus>
locality is in Zhangmu to the south of Nielamu, southern
<collectingRegion id="4990C383FFF4FF93B73C286827D81504" box="[1348,1432,333,359]" country="China" name="Xizang" pageId="5" pageNumber="132">Xizang</collectingRegion>
. Some specimens were sifted from leaf litter from the floor of a mixed forest at an altitude of
<quantity id="4CACA084FFF4FF93B6DE285426BE15E8" box="[1190,1278,369,395]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="132" unit="m" value="2600.0">2600 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E5EEAFFF4FF93B2BF28B021FE15D3" box="[199,958,405,432]" pageId="5" pageNumber="132" type="etymology">
<paragraph id="8BEB0D61FFF4FF93B2BF28B021FE15D3" blockId="5.[151,1437,153,432]" box="[199,958,405,432]" pageId="5" pageNumber="132">
<emphasis id="B920D173FFF4FF93B2BF28B0230E15CC" bold="true" box="[199,334,405,431]" pageId="5" pageNumber="132">Etymology.</emphasis>
The species is named after its
<typeStatus id="54EFB3C3FFF4FF93B0DE28B0209615CC" box="[678,726,405,431]" pageId="5" pageNumber="132">type</typeStatus>
locality (Zhangmu).
</paragraph>
</subSubSection>
</treatment>
</document>
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