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<document id="5BB12AE6FB81433EC5A0A06B3B73E916" ID-CLB-Dataset="2470" ID-DOI="10.21248/contrib.entomol.67.2.213-246" ID-GBIF-Dataset="43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742339" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267510724" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03FD87A1F114FFE57640FCE357AE7B79" docLanguage="en" docName="ContribEntomol.67.2.213-246.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) hirtiventris Assing 2017, spec. nov." docType="treatment" docUuid="73489C67-FB52-4B48-A2E1-BA1F1E55F45D" docUuidSource="ZooBank" docVersion="10" lastPageNumber="238" masterDocId="FFC4FFD9F10CFFFC7525FFD054747F38" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="246" masterPageNumber="213" pageNumber="237" updateTime="1699471357828" updateUser="ExternalLinkService">
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<mods:title id="045D0294328DFB66FF33F8E69C457FF2">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:date id="9FDDAB59DC154BA3BB0BF464A4F6779D">2017</mods:date>
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<treatment id="03FD87A1F114FFE57640FCE357AE7B79" ID-DOI="http://doi.org/10.5281/zenodo.5888812" ID-GBIF-Taxon="192500464" ID-Zenodo-Dep="5888812" LSID="urn:lsid:zoobank.org:act:73489C67-FB52-4B48-A2E1-BA1F1E55F45D" httpUri="http://treatment.plazi.org/id/03FD87A1F114FFE57640FCE357AE7B79" lastPageId="25" lastPageNumber="238" pageId="24" pageNumber="237">
<subSubSection id="C34E653CF114FFE47640FCE3511E7C57" pageId="24" pageNumber="237" type="nomenclature">
<paragraph id="8BEB36B7F114FFE47640FCE350BD7C76" blockId="24.[869,1386,819,911]" box="[869,1225,819,846]" pageId="24" pageNumber="237">
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<taxonomicName id="4C544D34F114FFE47640FCE3502E7C76" ID-CoL="5DDCY" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[869,1114,819,846]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="237" phylum="Arthropoda" rank="species" species="hirtiventris" status="sp. nov." subGenus="Zyras">
<emphasis id="B920EAA5F114FFE47640FCE357EF7C76" box="[869,923,819,846]" italics="true" pageId="24" pageNumber="237">Zyras</emphasis>
(
<emphasis id="B920EAA5F114FFE4768CFCE357AB7C76" box="[937,991,819,846]" italics="true" pageId="24" pageNumber="237">Zyras</emphasis>
)
<emphasis id="B920EAA5F114FFE476C8FCE3502E7C76" box="[1005,1114,819,846]" italics="true" pageId="24" pageNumber="237">hirtiventris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357DEF114FFE47145FCE350BD7C76" box="[1120,1225,819,846]" pageId="24" pageNumber="237" rank="species">spec. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<subSubSection id="C34E653CF114FFE47640FCAB50247CB7" box="[869,1104,891,911]" pageId="24" pageNumber="237" type="description">
<paragraph id="8BEB36B7F114FFE47640FCAB50247CB7" blockId="24.[869,1386,819,911]" box="[869,1104,891,911]" pageId="24" pageNumber="237">
(
<figureCitation id="136F2A32F114FFE4764CFCAB57CB7CB7" box="[873,959,891,911]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="24" pageNumber="237">Figs 1112</figureCitation>
,
<figureCitation id="136F2A32F114FFE476E2FCAB57A87CB7" box="[967,988,891,911]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="24" pageNumber="237">32</figureCitation>
,
<figureCitation id="136F2A32F114FFE476C7FCAB50617CB7" box="[994,1045,891,911]" captionStart="Figs 4052" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="24" pageNumber="237">4950</figureCitation>
,
<figureCitation id="136F2A32F114FFE47139FCAB50387CB7" box="[1052,1100,891,911]" captionStart="Figs 87102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87102: Zyras parvilobatus (8789), Z. hlavaci (9095; 9092: holotype; 9395: paratype from Selangor), Z. hirtiventris, holotype (9698), Z. lunatus (99101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (8788, 9091, 9394, 9697, 99100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="24" pageNumber="237">9698</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF114FFE4761AFC7951F97A59" pageId="24" pageNumber="237" type="materials_examined">
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<emphasis id="B920EAA5F114FFE4761AFC79503F7CF9" bold="true" box="[831,1099,937,961]" pageId="24" pageNumber="237">Type material examined</emphasis>
:
<materialsCitation id="3B3C3CEAF114FFE4717CFC7951257B19" ID-GBIF-Occurrence="3450187305" collectingDate="2004-05-16" collectingDateMax="2004-05-25" collectingDateMin="2004-05-16" collectorName="Lao" country="Laos" determinerName="V. Assing" elevation="1200" location="Boun Tai" municipality="Boun Tai" pageId="24" pageNumber="237" specimenCount="1" stateProvince="Phongsali Prov." typeStatus="holotype">
<typeStatus id="54EF8815F114FFE4717CFC7950C87CF9" box="[1113,1212,937,961]" pageId="24" pageNumber="237" type="holotype">Holotype</typeStatus>
<specimenCount id="9D52FD3EF114FFE471E1FC7A50AD7CF9" box="[1220,1241,938,961]" pageId="24" pageNumber="237" type="generic"></specimenCount>
: “N
<collectingCountry id="F3437627F114FFE47028FC7951247CF9" box="[1293,1360,937,961]" name="Laos" pageId="24" pageNumber="237">LAOS</collectingCountry>
,
<quantity id="4CAC9B52F114FFE47079FC7957E47CD9" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="24" pageNumber="237" unit="m" value="1200.0" valueMax="1300.0" valueMin="1100.0">
<elevation id="0079D184F114FFE47079FC7957E47CD9" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="24" pageNumber="237" unit="m" value="1200.0" valueMax="1300.0" valueMin="1100.0">1100 1300 m</elevation>
</quantity>
,
<collectingRegion id="4990F855F114FFE47684FC19503B7CD9" box="[929,1103,969,993]" country="Laos" name="Phongsali" pageId="24" pageNumber="237">Phongsali Prov.</collectingRegion>
,
<collectingMunicipality id="6B8FACCDF114FFE4717AFC1A50B27CD9" box="[1119,1222,969,993]" pageId="24" pageNumber="237">Boun Tai</collectingMunicipality>
(
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<quantity id="4CAC9B52F114FFE471FEFC19516B7CD9" box="[1243,1311,969,993]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="237" unit="km" value="10.0">10 km</quantity>
SE
</locationDeviation>
),
<date id="FFEA1077F114FFE47046FC1957C17B3A" pageId="24" pageNumber="237" value="2004-05-16" valueMax="2004-05-25" valueMin="2004-05-16">
<collectingDate id="EFAEE99FF114FFE47046FC1957C17B3A" pageId="24" pageNumber="237" value="2004-05-16" valueMax="2004-05-25" valueMin="2004-05-16">2004, 1625 May</collectingDate>
</date>
, Lao collector leg. / Holotypus ♂
<taxonomicName id="4C544D34F114FFE47046FC3A57C77B19" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="237" phylum="Arthropoda" rank="species" species="hirtiventris" status="sp. nov.">
<emphasis id="B920EAA5F114FFE47046FC3A57C77B19" italics="true" pageId="24" pageNumber="237">Zyras hirtiventris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357DEF114FFE476E7FBDA578E7B19" box="[962,1018,1034,1057]" pageId="24" pageNumber="237" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="7E37DA73F114FFE47163FBDA50D37B19" box="[1094,1191,1033,1057]" pageId="24" pageNumber="237">
V.
<bibRefCitation id="EFC54B46F114FFE47145FBD950D37B19" author="ASSING, V." box="[1120,1191,1033,1057]" pageId="24" pageNumber="237" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
</determinerName>
2017” (cMar)
</materialsCitation>
.
<materialsCitation id="3B3C3CEAF114FFE47041FBDA503A7BB9" ID-GBIF-Occurrence="3450187334" collectingDate="2013-08-07" collectingDateMax="2016-05-12" collectingDateMin="2013-08-07" collectionCode="MZUF" collectorName="V. Assing" country="Vietnam" elevation="1810" latitude="22.360834" location="Vietnam" longLatPrecision="21" longitude="103.78861" municipality="Kon Chu Rang Nature Reserve" pageId="24" pageNumber="237" specimenCode="MZUF 18322" specimenCount="1" stateProvince="Gia Lai Province" typeStatus="paratype">
<typeStatus id="54EF8815F114FFE47041FBDA570E7B7A" pageId="24" pageNumber="237" type="paratype">Paratypes</typeStatus>
: 1♀: “N-Vietnam
<locationDeviation id="9A592F95F114FFE4717BFBF951677B79" box="[1118,1299,1065,1089]" location="Sa Pa" pageId="24" pageNumber="237">
<quantity id="4CAC9B52F114FFE4717BFBF950E07B79" box="[1118,1172,1065,1089]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="24" pageNumber="237" unit="km" value="6.0">6 km</quantity>
NW Sa Pa
</locationDeviation>
,
<geoCoordinate id="EE605070F114FFE47005FBF951EC7B79" box="[1312,1432,1065,1089]" degrees="22" direction="north" minutes="21" orientation="latitude" pageId="24" pageNumber="237" precision="15" seconds="39" value="22.360834">22°21'39&quot;N</geoCoordinate>
,
<geoCoordinate id="EE605070F114FFE47665FB9957CB7B59" box="[832,959,1097,1121]" degrees="103" direction="east" minutes="47" orientation="longitude" pageId="24" pageNumber="237" precision="15" seconds="19" value="103.78861">103°47'19&quot;E</geoCoordinate>
,
<quantity id="4CAC9B52F114FFE476F8FB9950597B59" box="[989,1069,1097,1121]" metricMagnitude="3" metricUnit="m" metricValue="1.81" pageId="24" pageNumber="237" unit="m" value="1810.0">
<elevation id="0079D184F114FFE476F8FB9950597B59" box="[989,1069,1097,1121]" metricMagnitude="3" metricUnit="m" metricValue="1.81" pageId="24" pageNumber="237" unit="m" value="1810.0">1810 m</elevation>
</quantity>
, pasture margin,
<date id="FFEA1077F114FFE47038FB9951ED7B59" box="[1309,1433,1097,1121]" pageId="24" pageNumber="237" value="2013-08-07">
<collectingDate id="EFAEE99FF114FFE47038FB9951ED7B59" box="[1309,1433,1097,1121]" pageId="24" pageNumber="237" value="2013-08-07">7.VIII.2013</collectingDate>
</date>
,
<collectorName id="26A15361F114FFE47665FBBA57D57BB9" box="[832,929,1129,1153]" pageId="24" pageNumber="237">V. Assing</collectorName>
[8+2]” (cAss);
</materialsCitation>
<materialsCitation id="3B3C3CEAF114FFE4717BFBB951FC7A59" ID-GBIF-Occurrence="3450187331" collectingDate="2016-05-08" collectingDateMax="2016-05-12" collectingDateMin="2016-05-08" collectionCode="MZUF" country="Vietnam" elevation="900" latitude="22.360834" location="Vietnam" longLatPrecision="21" longitude="103.78861" municipality="Kon Chu Rang Nature Reserve" pageId="24" pageNumber="237" specimenCode="MZUF 18322" specimenCount="1" stateProvince="Gia Lai Province" typeStatus="paratype">
1 ♂: “C
<collectingCountry id="F3437627F114FFE471E7FBB951497BB9" box="[1218,1341,1129,1153]" name="Vietnam" pageId="24" pageNumber="237">VIETNAM</collectingCountry>
:
<collectingRegion id="4990F855F114FFE47075FBB957D47B99" country="Vietnam" name="Gia Lai" pageId="24" pageNumber="237">Gia Lai Province</collectingRegion>
,
<collectingMunicipality id="6B8FACCDF114FFE47697FB5A51697B99" box="[946,1309,1161,1186]" country="Vietnam" latitude="22.360834" longLatPrecision="21" longitude="103.78861" name="Kon Chu Rang Nature Reserve" pageId="24" pageNumber="237" stateProvince="Gia Lai">Kon Chu Rang Nature Reserve</collectingMunicipality>
, surroundings HQ, about
<quantity id="4CAC9B52F114FFE476D4FB7950417BF9" box="[1009,1077,1193,1217]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="237" unit="m" value="900.0">
<elevation id="0079D184F114FFE476D4FB7950417BF9" box="[1009,1077,1193,1217]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="237" unit="m" value="900.0">900 m</elevation>
</quantity>
,
<geoCoordinate id="EE605070F114FFE47166FB7950B27BF9" box="[1091,1222,1193,1217]" degrees="14" direction="north" minutes="28,450" orientation="latitude" pageId="24" pageNumber="237" precision="1" value="14.474167">14°28,450'N</geoCoordinate>
,
<geoCoordinate id="EE605070F114FFE471F6FB79512F7BF9" box="[1235,1371,1193,1217]" degrees="108" direction="east" minutes="32,401" orientation="longitude" pageId="24" pageNumber="237" precision="1" value="108.540016">108°32,401'E</geoCoordinate>
/ leg.
<collectorName id="26A15361F114FFE47665FB1A57BC7BD9" box="[832,968,1225,1249]" pageId="24" pageNumber="237">L. Bartolozzi</collectorName>
,
<collectorName id="26A15361F114FFE476F4FB19502B7BD9" box="[977,1119,1225,1249]" pageId="24" pageNumber="237">A. Bandinelli</collectorName>
,
<collectorName id="26A15361F114FFE4714DFB1950B37BD9" box="[1128,1223,1225,1249]" pageId="24" pageNumber="237">S. Bambi</collectorName>
,
<collectorName id="26A15361F114FFE471F4FB1A513A7BD9" box="[1233,1358,1225,1249]" pageId="24" pageNumber="237">V. Sbordoni</collectorName>
at light
<date id="FFEA1077F114FFE47665FB3957B67A39" box="[832,962,1257,1281]" pageId="24" pageNumber="237" value="2016-05-08" valueMax="2016-05-12" valueMin="2016-05-08">
<collectingDate id="EFAEE99FF114FFE47665FB3957B67A39" box="[832,962,1257,1281]" pageId="24" pageNumber="237" value="2016-05-08" valueMax="2016-05-12" valueMin="2016-05-08">812.V.2016</collectingDate>
</date>
(n° Mag. 3078)” [registered as
<specimenCode id="DBF29ECCF114FFE47022FB3A51ED7A39" box="[1287,1433,1257,1281]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="24" pageNumber="237">MZUF 18322</specimenCode>
; collected during expeditions in
<location id="8E8B606CF114FFE471A2FAD950977A19" LSID="urn:lsid:plazi:treatment:03FD87A1F114FFE57640FCE357AE7B79:8E8B606CF114FFE471A2FAD950977A19" box="[1159,1251,1289,1313]" country="Vietnam" latitude="22.360834" longLatPrecision="21" longitude="103.78861" municipality="Kon Chu Rang Nature Reserve" name="Vietnam" pageId="24" pageNumber="237" stateProvince="Gia Lai Province">Vietnam</location>
in the framework of a
<location id="8E8B606CF114FFE47651FAF950EA7A59" LSID="urn:lsid:plazi:treatment:03FD87A1F114FFE57640FCE357AE7B79:8E8B606CF114FFE47651FAF950EA7A59" country="Vietnam" latitude="22.360834" longLatPrecision="21" longitude="103.78861" municipality="Kon Chu Rang Nature Reserve" name="Memorandum of Unterstanding between the Vietnam National Museum of Nature" pageId="24" pageNumber="237" stateProvince="Gia Lai Province">Memorandum of Unterstanding between the Vietnam National Museum of Nature</location>
and
<collectionCode id="ED45AE72F114FFE471F7FA9A516B7A59" box="[1234,1311,1354,1377]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="24" pageNumber="237">MZUF</collectionCode>
] (
<collectionCode id="ED45AE72F114FFE47017FA9951F67A59" box="[1330,1410,1353,1377]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="24" pageNumber="237">MZUF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF114FFE47665FA5951487939" pageId="24" pageNumber="237" type="etymology">
<paragraph id="8BEB36B7F114FFE47665FA5951487939" blockId="24.[832,1437,1417,1537]" pageId="24" pageNumber="237">
<emphasis id="B920EAA5F114FFE47665FA5957C37A99" bold="true" box="[832,951,1417,1441]" pageId="24" pageNumber="237">Etymology</emphasis>
: The specific epithet (with densely pubescent abdomen) is an adjective alluding to the dense setae on the abdominal sternites, as well as to the strong resemblance with
<taxonomicName id="4C544D34F114FFE476E5FA3A514C7939" authority="(CHAMPION, 1927)" baseAuthorityName="CHAMPION" baseAuthorityYear="1927" box="[960,1336,1513,1537]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="237" phylum="Arthropoda" rank="species" species="hirsutiventris">
<emphasis id="B920EAA5F114FFE476E5FA3A50127939" box="[960,1126,1513,1537]" italics="true" pageId="24" pageNumber="237">Z. hirsutiventris</emphasis>
(
<bibRefCitation id="EFC54B46F114FFE47150FA3951447939" author="CHAMPION, G. C." box="[1141,1328,1513,1537]" pageId="24" pageNumber="237" pagination="245 - 250" refId="ref26311" refString="CHAMPION, G. C. 1927: Some Indian Coleoptera (23). - The Entomologist's Monthly Magazine 63: 245 - 250." type="journal article" year="1927">CHAMPION, 1927</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF114FFE57665F9F950277C79" lastPageId="25" lastPageNumber="238" pageId="24" pageNumber="237" type="description">
<paragraph id="8BEB36B7F114FFE47665F9F9505378B9" blockId="24.[832,1437,1577,2081]" pageId="24" pageNumber="237">
<emphasis id="B920EAA5F114FFE47665F9F957B17979" bold="true" box="[832,965,1577,1601]" pageId="24" pageNumber="237">Description</emphasis>
: Large species; body length
<quantity id="4CAC9B52F114FFE47030F9F951EC7979" box="[1301,1432,1577,1601]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="8.9" metricValueMin="8.5" pageId="24" pageNumber="237" unit="mm" value="8.7" valueMax="8.9" valueMin="8.5">8.58.9 mm</quantity>
; length of forebody
<quantity id="4CAC9B52F114FFE47128F99950E67959" box="[1037,1170,1609,1633]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.7" metricValueMin="3.6" pageId="24" pageNumber="237" unit="mm" value="3.65" valueMax="3.7" valueMin="3.6">3.63.7 mm</quantity>
. Coloration (
<figureCitation id="136F2A32F114FFE4703AF99951E37959" box="[1311,1431,1609,1633]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="24" pageNumber="237">Figs 1112</figureCitation>
,
<figureCitation id="136F2A32F114FFE47665F9B9572879B9" box="[832,860,1641,1665]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="24" pageNumber="237">32</figureCitation>
,
<figureCitation id="136F2A32F114FFE47654F9B957CC79B9" box="[881,952,1641,1665]" captionStart="Figs 4052" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="24" pageNumber="237">4950</figureCitation>
): body black with the posterior margin of the abdominal segment VII and all, or the posterior portion, of segment VIII dark reddish-brown; legs yellowish-brown with the profemora dark-brown and the meso- and metafemora bicoloured, basally yellowish and apically blackish-brown; antennae darkbrown to blackish with the apical antennomere reddish to reddish-brown; maxillary palpi brown with the apical palpomere yellowish.
</paragraph>
<paragraph id="8BEB36B7F114FFE57665F859548A7E99" blockId="24.[832,1437,1577,2081]" lastBlockId="25.[151,756,233,1729]" lastPageId="25" lastPageNumber="238" pageId="24" pageNumber="237">
Head (
<figureCitation id="136F2A32F114FFE476AFF85957A07899" box="[906,980,1929,1953]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="24" pageNumber="237">Fig. 32</figureCitation>
) distinctly transverse; punctation moderately coarse and rather dense, median dorsal portion extensively impunctate; pubescence long and sub-erect to erect, brown to black. Eyes moderately large, slightly longer than postocular region in dorsal view. Antenna (
<figureCitation id="136F2A32F115FFE575BBFF3955617E39" box="[158,277,233,257]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="25" pageNumber="238">Figs 1112</figureCitation>
)
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long; antennomeres IV weakly oblong, VVI weakly oblong to weakly transverse, VIIIX weakly to distinctly transverse, X approximately 1.5 times as broad as long, and XI short and of distinctly conical shape, much shorter than the combined length of IX and X.
</paragraph>
<paragraph id="8BEB36B7F115FFE575B2FE7A56CB7DF9" blockId="25.[151,756,233,1729]" pageId="25" pageNumber="238">
Pronotum (
<figureCitation id="136F2A32F115FFE57436FE7955297EF9" box="[275,349,425,449]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="25" pageNumber="238">Fig. 32</figureCitation>
) 1.111.13 times as broad as long and 1.331.36 times as broad as head, broadest slightly anterior to middle, weakly convex in cross-section; posterior angles obtusely marked; lateral margins straight or very weakly sinuate in posterior half in dorsal view; punctation rather fine and dense, regularly distributed; midline without distinct impunctate band; pubescence long and sub-erect to erect, brown to black; lateral and anterior margins with longer and distinctly erect black setae.
</paragraph>
<paragraph id="8BEB36B7F115FFE575B2FD1954857CB9" blockId="25.[151,756,233,1729]" pageId="25" pageNumber="238">
Elytra (
<figureCitation id="136F2A32F115FFE575C9FD1955437DD9" box="[236,311,713,737]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="25" pageNumber="238">Fig. 32</figureCitation>
) 0.850.89 times as long as pronotum; punctation dense and moderately coarse, regularly distributed, and defined; pubescence dense, long, and brown to black. Hind wings fully developed. Metatarsomere I approximately as long as the combined length of IIIV.
</paragraph>
<paragraph id="8BEB36B7F115FFE575B2FC59567A7939" blockId="25.[151,756,233,1729]" pageId="25" pageNumber="238">
Abdomen (
<figureCitation id="136F2A32F115FFE5743DFC5955107C99" box="[280,356,905,929]" captionStart="Figs 4052" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="25" pageNumber="238">Fig. 49</figureCitation>
) approximately as broad as elytra, with moderately deep anterior impressions on tergites IIIV; tergites IIIV each with a transverse row of fine to moderately coarse non-setiferous punctures in anterior impressions, with several lateral setiferous punctures on either side, with 610 setiferous punctures at posterior margins, and with moderately sparse to moderately dense non-setiferous micropunctation on discs; tergites VIVII with extensive, dense, and coarse non-setiferous punctures anteriorly, with slightly sparser non-setiferous punctures on remainder of discs, and with some setiferous punctures laterally and at posterior margins (
<figureCitation id="136F2A32F115FFE5749CFAD956737A19" box="[441,519,1289,1313]" captionStart="Figs 4052" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="25" pageNumber="238">Fig. 50</figureCitation>
); posterior margin of tergite VII with palisade fringe; tergite VIII with dense and extensive non-setiferous punctation in posterior half and with dense and distinct setiferous punctation in posterior half, posterior margin convex, in the middle with very indistinct concavity; all sternites with long, dense, and sub-erect to erect pubescence in posterior portions (well visible from above).
</paragraph>
<paragraph id="8BEB36B7F115FFE575B2F9DA563279B9" blockId="25.[151,756,233,1729]" pageId="25" pageNumber="238">
♂: sternite VIII with convex posterior margin; median lobe of aedeagus approximately
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long and shaped as in
<figureCitation id="136F2A32F115FFE575EBF99955377959" box="[206,323,1609,1633]" captionStart="Figs 87102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87102: Zyras parvilobatus (8789), Z. hlavaci (9095; 9092: holotype; 9395: paratype from Selangor), Z. hirtiventris, holotype (9698), Z. lunatus (99101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (8788, 9091, 9394, 9697, 99100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="25" pageNumber="238">Figs 9697</figureCitation>
; paramere (
<figureCitation id="136F2A32F115FFE5749AF999567D7959" box="[447,521,1609,1633]" captionStart="Figs 87102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87102: Zyras parvilobatus (8789), Z. hlavaci (9095; 9092: holotype; 9395: paratype from Selangor), Z. hirtiventris, holotype (9698), Z. lunatus (99101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (8788, 9091, 9394, 9697, 99100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="25" pageNumber="238">Fig. 98</figureCitation>
)
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long, apical lobe short and with four long dark setae.
</paragraph>
<paragraph id="8BEB36B7F115FFE575B2F95A556579F9" blockId="25.[151,756,233,1729]" pageId="25" pageNumber="238">♀: posterior margin of sternite VIII shallowly concave in the middle.</paragraph>
<paragraph id="8BEB36B7F115FFE575B2F93951727E99" blockId="25.[151,757,1769,2081]" lastBlockId="25.[794,1399,233,833]" pageId="25" pageNumber="238">
<emphasis id="B920EAA5F115FFE575B2F93955117839" bold="true" box="[151,357,1769,1793]" pageId="25" pageNumber="238">Comparative notes</emphasis>
: Based on the external and the male sexual characters,
<taxonomicName id="4C544D34F115FFE5747BF8DA559E7819" authorityName="Assing" authorityYear="2017" box="[350,490,1801,1825]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirtiventris">
<emphasis id="B920EAA5F115FFE5747BF8DA559E7819" box="[350,490,1801,1825]" italics="true" pageId="25" pageNumber="238">Z. hirtiventris</emphasis>
</taxonomicName>
undoubtedly belongs to the
<taxonomicName id="4C544D34F115FFE575E5F8FA55637879" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[192,279,1833,1857]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B920EAA5F115FFE575E5F8FA55637879" box="[192,279,1833,1857]" italics="true" pageId="25" pageNumber="238">Z. hirtus</emphasis>
</taxonomicName>
group (see
<bibRefCitation id="EFC54B46F115FFE574B0F8F956557879" author="ASSING, V." box="[405,545,1833,1857]" pageId="25" pageNumber="238" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING 2017</bibRefCitation>
). In habitus, coloration, and other external and sexual characters, this species strongly resembles
<taxonomicName id="4C544D34F115FFE5744CF8BA567E78B9" baseAuthorityName="CHAMPION" baseAuthorityYear="1927" box="[361,522,1897,1921]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirsutiventris">
<emphasis id="B920EAA5F115FFE5744CF8BA567E78B9" box="[361,522,1897,1921]" italics="true" pageId="25" pageNumber="238">Z. hirsutiventris</emphasis>
</taxonomicName>
from the Himalayan region. It is distinguished from this species by larger body size (
<taxonomicName id="4C544D34F115FFE575F5F87A550578F9" baseAuthorityName="CHAMPION" baseAuthorityYear="1927" box="[208,369,1961,1985]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirsutiventris">
<emphasis id="B920EAA5F115FFE575F5F87A550578F9" box="[208,369,1961,1985]" italics="true" pageId="25" pageNumber="238">Z. hirsutiventris</emphasis>
</taxonomicName>
: body length
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; length of foreboedy
<quantity id="4CAC9B52F115FFE5742DF81955FC78D9" box="[264,392,1993,2017]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.3" metricValueMin="2.9" pageId="25" pageNumber="238" unit="mm" value="3.0999999999999996" valueMax="3.3" valueMin="2.9">2.93.3 mm</quantity>
), paler coloration of antennomere XI (black in
<taxonomicName id="4C544D34F115FFE57405F83A55B57739" baseAuthorityName="CHAMPION" baseAuthorityYear="1927" box="[288,449,2025,2049]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirsutiventris">
<emphasis id="B920EAA5F115FFE57405F83A55B57739" box="[288,449,2025,2049]" italics="true" pageId="25" pageNumber="238">Z. hirsutiventris</emphasis>
</taxonomicName>
), less fine and more defined punctation of the elytra, slightly darker coloration of the legs, denser and more extensive non-setiferous punctation of the posterior abdominal tergites, and a larger median lobe of the aedeagus with a ventral process of slightly different shape (
<taxonomicName id="4C544D34F115FFE576F7FE9A50007E59" baseAuthorityName="CHAMPION" baseAuthorityYear="1927" box="[978,1140,329,353]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirsutiventris">
<emphasis id="B920EAA5F115FFE576F7FE9A50007E59" box="[978,1140,329,353]" italics="true" pageId="25" pageNumber="238">Z. hirsutiventris</emphasis>
</taxonomicName>
: median lobe approximately
<quantity id="4CAC9B52F115FFE57643FEB957C27EB9" box="[870,950,361,385]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="238" unit="mm" value="0.8">0.8 mm</quantity>
long; ventral process apically longer, more acute, and less distinctly angled in lateral view).
</paragraph>
<paragraph id="8BEB36B7F115FFE5763FFE7950277C79" blockId="25.[794,1399,233,833]" pageId="25" pageNumber="238">
These differences may be clear-cut, though not pronounced, in the material examined thus far, but at present only
<specimenCount id="9D52FD3EF115FFE57691FE39501C7D39" box="[948,1128,489,513]" pageId="25" pageNumber="238" type="generic">three specimens</specimenCount>
of
<taxonomicName id="4C544D34F115FFE571BFFE3A51347D39" baseAuthorityName="CHAMPION" baseAuthorityYear="1927" box="[1178,1344,489,513]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirsutiventris">
<emphasis id="B920EAA5F115FFE571BFFE3A51347D39" box="[1178,1344,489,513]" italics="true" pageId="25" pageNumber="238">Z. hirsutiventris</emphasis>
</taxonomicName>
and three of
<taxonomicName id="4C544D34F115FFE57658FDDA50797D19" authorityName="Assing" authorityYear="2017" box="[893,1037,521,545]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirtiventris">
<emphasis id="B920EAA5F115FFE57658FDDA50797D19" box="[893,1037,521,545]" italics="true" pageId="25" pageNumber="238">Z. hirtiventris</emphasis>
</taxonomicName>
are known. The possibility that material with intermediate characters conditions will be found in the future and that the currently known material of
<taxonomicName id="4C544D34F115FFE57644FDBA50737DB9" baseAuthorityName="CHAMPION" baseAuthorityYear="1927" box="[865,1031,617,641]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirsutiventris">
<emphasis id="B920EAA5F115FFE57644FDBA50737DB9" box="[865,1031,617,641]" italics="true" pageId="25" pageNumber="238">Z. hirsutiventris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D34F115FFE57118FDBA50B97DB9" authorityName="Assing" authorityYear="2017" box="[1085,1229,617,641]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirtiventris">
<emphasis id="B920EAA5F115FFE57118FDBA50B97DB9" box="[1085,1229,617,641]" italics="true" pageId="25" pageNumber="238">Z. hirtiventris</emphasis>
</taxonomicName>
only represents extreme forms of the same species cannot be ruled out with certainty. At present, the distributions of both taxa are separated by a distance of more than
<quantity id="4CAC9B52F115FFE571F2FD1951427DD9" box="[1239,1334,713,737]" metricMagnitude="6" metricUnit="m" metricValue="1.1" pageId="25" pageNumber="238" unit="km" value="1100.0">1100 km</quantity>
. This, in combination with the morphological differences observed, suggests that
<taxonomicName id="4C544D34F115FFE5713AFCDA50B17C19" baseAuthorityName="CHAMPION" baseAuthorityYear="1927" box="[1055,1221,777,801]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirsutiventris">
<emphasis id="B920EAA5F115FFE5713AFCDA50B17C19" box="[1055,1221,777,801]" italics="true" pageId="25" pageNumber="238">Z. hirsutiventris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D34F115FFE57025FCDA57487C79" authorityName="Assing" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hirtiventris">
<emphasis id="B920EAA5F115FFE57025FCDA57487C79" italics="true" pageId="25" pageNumber="238">Z. hirtiventris</emphasis>
</taxonomicName>
represent distinct species.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF115FFE5763FFCB957AE7B79" pageId="25" pageNumber="238" type="distribution">
<paragraph id="8BEB36B7F115FFE5763FFCB957AE7B79" blockId="25.[794,1399,873,1089]" pageId="25" pageNumber="238">
<emphasis id="B920EAA5F115FFE5763FFCB950F67CB9" bold="true" box="[794,1154,873,897]" pageId="25" pageNumber="238">Distribution and natural history</emphasis>
: The currently known distribution is confined to two localities in North
<collectingCountry id="F3437627F115FFE57060FC5A51037C99" box="[1349,1399,906,929]" name="Laos" pageId="25" pageNumber="238">Laos</collectingCountry>
and North
<collectingCountry id="F3437627F115FFE576B3FC7957817CF9" box="[918,1013,937,961]" name="Vietnam" pageId="25" pageNumber="238">Vietnam</collectingCountry>
. The altitudes range from approximately
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. The female
<typeStatus id="54EF8815F115FFE57180FC1A51767CDA" box="[1189,1282,970,994]" pageId="25" pageNumber="238" type="paratype">paratype</typeStatus>
was sifted from litter and roots beneath bushes and small trees at the margin of a pasture. It was recorded as
<taxonomicName id="4C544D34F115FFE57190FBDA51357B19" authority="(Zyras)" box="[1205,1345,1033,1057]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="subGenus" subGenus="Zyras">
<emphasis id="B920EAA5F115FFE57190FBDA509B7B19" box="[1205,1263,1034,1057]" italics="true" pageId="25" pageNumber="238">Zyras</emphasis>
(
<emphasis id="B920EAA5F115FFE571DBFBDA514C7B19" box="[1278,1336,1034,1057]" italics="true" pageId="25" pageNumber="238">Zyras</emphasis>
)
</taxonomicName>
sp. 1 by
<bibRefCitation id="EFC54B46F115FFE5761CFBF957A37B79" author="ASSING, V." box="[825,983,1065,1089]" pageId="25" pageNumber="238" pagination="1257 - 1282" refId="ref25690" refString="ASSING, V. 2015: On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 47 (2): 1257 - 1282 - http: // www. zobodat. at / pdf / LBB _ 0047 _ 2 _ 1257 - 1282. pdf." type="journal article" year="2015">ASSING (2015)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>