treatments-xml/data/03/FD/87/03FD87925830FFC1FF56FD5E84764269.xml

<document id="3495781BD2ADB7FB0DBB543D14888BDF" ID-CLB-Dataset="106789" ID-DOI="10.11646/zootaxa.5169.1.1" ID-GBIF-Dataset="b76236d0-0336-4d17-804a-80f0344b097e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6911151" ID-ZooBank="3CB49021-90C5-46F0-AAAF-619EE8068F42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658900973851" checkinUser="plazi" docAuthor="Grall, Elena &amp; Jäger, Peter" docDate="2022" docId="03FD87925830FFC1FF56FD5E84764269" docLanguage="en" docName="zootaxa.5169.1.1.pdf" docOrigin="Zootaxa 5169 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Borniella parva Grall &amp; Jäger 2022, spec. nov." docType="treatment" docVersion="6" lastPageNumber="5" masterDocId="FFC4FFEA5832FFC5FFC1FFA480644008" masterDocTitle="Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="3" updateTime="1699362128362" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="39D48F1B1836C63F714D5D4CC1BCBE2B">Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae)</mods:title>
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<mods:namePart id="D43587B51693A2723144FE880F9F15A6">Grall, Elena</mods:namePart>
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<mods:affiliation id="DFC142247AD1B17B922FDADCF43B07D6">Arachnology, Senckenberg Research Institute, Mertonstrasse 17 - 21, 60325 Frankfurt. &amp; Institute of Ecology, Diversity and Evolution, Campus Riedberg, Goethe University, 60438 Frankfurt am Main, Germany; elena. grall @ senckenberg. de, https: // orcid. org / 0000 - 0002 - 8938 - 4105</mods:affiliation>
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<mods:namePart id="855CC74162FEE908249D68CDE2894A83">Jäger, Peter</mods:namePart>
<mods:nameIdentifier id="1F8AB44C9331B2990A2EB6D18D45B8B9" type="ORCID">0000-0003-1563-0147</mods:nameIdentifier>
<mods:affiliation id="E1C7319766D7E0A0448B1A0308B9485D">Arachnology, Senckenberg Research Institute, Mertonstrasse 17 - 21, 60325 Frankfurt. &amp; peter. jaeger @ senckenberg. de, https: // orcid. org / 0000 - 0003 - 1563 - 0147</mods:affiliation>
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<mods:title id="33E968F56BE40ABBDE7D7D3C41D436C0">Zootaxa</mods:title>
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<mods:date id="BCEF1E09252376F02EF972273A3A5756">2022</mods:date>
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<mods:number id="75C2378B03DB80F725DE1F96FFEB0AFB">2022-07-27</mods:number>
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<treatment id="03FD87925830FFC1FF56FD5E84764269" ID-DOI="http://doi.org/10.5281/zenodo.6911157" ID-GBIF-Taxon="197859761" ID-Zenodo-Dep="6911157" LSID="urn:lsid:plazi:treatment:03FD87925830FFC1FF56FD5E84764269" httpUri="http://treatment.plazi.org/id/03FD87925830FFC1FF56FD5E84764269" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
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<heading id="D0A381E85830FFC7FF56FD5E81B0431D" bold="true" box="[151,468,762,789]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B920EA965830FFC7FF56FD5E81B0431D" bold="true" box="[151,468,762,789]" pageId="2" pageNumber="3">
<taxonomicName id="4C544D075830FFC7FF56FD5E8131431D" authority="Grall &amp; Jäger, 2022" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[151,341,762,789]" class="Arachnida" family="Sparassidae" genus="Borniella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
<emphasis id="B920EA965830FFC7FF56FD5E8131431D" bold="true" box="[151,341,762,789]" italics="true" pageId="2" pageNumber="3">Borniella parva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357ED5830FFC7FE9DFD5F81B0431D" box="[348,468,763,789]" pageId="2" pageNumber="3" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34E650F5830FFC7FF56FCBB81E04331" box="[151,388,798,825]" pageId="2" pageNumber="3" type="description">
<paragraph id="8BEB36845830FFC7FF56FCBB81E04331" blockId="2.[151,468,762,825]" box="[151,388,798,825]" pageId="2" pageNumber="3">
<figureCitation id="136F2A015830FFC7FF56FCBB816C4331" box="[151,264,799,825]" captionStart-0="FIGURES 1–3" captionStart-1="FIGURES 4–7" captionStart-2="FIGURES 8–16" captionStartId-0="3.[151,264,1546,1570]" captionStartId-1="4.[151,264,1801,1825]" captionStartId-2="5.[151,264,1380,1404]" captionTargetBox-0="[151,1436,315,1519]" captionTargetBox-1="[151,1433,633,1778]" captionTargetBox-2="[151,1436,181,1357]" captionTargetId-0="figure-74@3.[151,1436,306,1519]" captionTargetId-1="figure-194@4.[151,1436,633,1778]" captionTargetId-2="figure-19@5.[151,1436,181,1357]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1–3. Borniella parva gen. &amp; spec. nov. holotype male from Sarawak, left male palp (1 prolateral, 2 ventral, 3 retrolateral). C—conductor, dTA— distal tegular apophysis, E—embolus, mTA—median tegular apophysis, pTA—proximal tegular apophysis, RTA—retrolateral tibial apophysis, S—spermophor, ST—subtegulum." captionText-1="FIGURES 4–7. Borniella parva gen. &amp; spec. nov., female paratype from Sarawak. 4, 6 Epigyne, ventral; 5 Vulva, dorsal (internal duct system and rim of copulatory opening in red, epigynal pocket and parts of the inner rims in green); 7 Schematic course of internal duct system, ventral. CO—copulatory opening, CP—copulatory plug, EF—epigynal field, EP—epigynal pocket, FD—fertilization duct, GA—glandular appendages, LS—lobal septum, MSu—membranous sac (unexpanded), SP— spermatheca." captionText-2="FIGURES 8–16. Borniella parva gen. &amp; spec. nov., holotype male (8, 10, 13, 15) and female paratype (9, 11–12, 14, 16) from Sarawak. 8–9 Eyes, dorsal. 10–12 Claws (10 right leg II, prolateral claw, prolateral; 11 right palp, prolateral; 12 right leg II, retrolateral claw, retrolateral). 13–14 Left chelicera, ventral. 15–16 Trilobate membrane, distal metatarsus, dorsal (15 right leg III, 16 right leg I). LP—lateral projections, MH—median hook." figureDoi-0="http://doi.org/10.5281/zenodo.6911153" figureDoi-1="http://doi.org/10.5281/zenodo.6911159" figureDoi-2="http://doi.org/10.5281/zenodo.6911161" httpUri-0="https://zenodo.org/record/6911153/files/figure.png" httpUri-1="https://zenodo.org/record/6911159/files/figure.png" httpUri-2="https://zenodo.org/record/6911161/files/figure.png" pageId="2" pageNumber="3">Figs 1–16</figureCitation>
, 
<figureCitation id="136F2A015830FFC7FED5FCBA81394331" box="[276,349,798,825]" captionStart="FIGURES 82–97" captionStartId="19.[151,264,1899,1923]" captionTargetBox="[152,1435,182,1873]" captionTargetId="figure-19@19.[151,1436,181,1875]" captionTargetPageId="19" captionText="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="2" pageNumber="3">82–85</figureCitation>
, 
<figureCitation id="136F2A015830FFC7FEA9FCBB81E04331" box="[360,388,799,825]" captionStart="FIGURE 98" captionStartId="20.[151,250,1436,1460]" captionTargetBox="[151,1436,956,1413]" captionTargetId="figure-372@20.[151,1436,956,1413]" captionTargetPageId="20" captionText="FIGURE 98. Distribution of the new genera and species in Malaysia, Brunei and Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.6911205" httpUri="https://zenodo.org/record/6911205/files/figure.png" pageId="2" pageNumber="3">98</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5830FFC7FF56FCC3816D44B1" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8BEB36845830FFC7FF56FCC384C043AB" blockId="2.[151,1437,870,2012]" pageId="2" pageNumber="3">
<emphasis id="B920EA965830FFC7FF56FCC381264388" bold="true" box="[151,322,870,897]" pageId="2" pageNumber="3">Type material.</emphasis>
<materialsCitation id="3B3C3CD95830FFC7FE86FCC284C043AB" ID-GBIF-Occurrence="3866517309" collectingDate="1978-05-24" collectionCode="NHM" collectorName="F. Wanless" country="Malaysia" county="Camp" elevation="330" latitude="4.0508003" location="Province" longLatPrecision="1" longitude="114.867004" municipality="Mulu Expedition" pageId="2" pageNumber="3" specimenCode="PJ 1227" specimenCount="1" typeStatus="holotype">
<emphasis id="B920EA965830FFC7FE86FCC283524389" bold="true" box="[327,822,870,897]" pageId="2" pageNumber="3">
<collectingCountry id="F34376145830FFC7FE86FCC281B94388" box="[327,477,870,896]" name="Malaysia" pageId="2" pageNumber="3">MALAYSIA</collectingCountry>
: 
<emphasis id="B920EA965830FFC7FE2BFCC282DF4389" bold="true" box="[490,699,870,897]" italics="true" pageId="2" pageNumber="3">
Sarawak 
<location id="8E8B605F5830FFC7FD95FCC382DF4389" LSID="urn:lsid:plazi:treatment:03FD87925830FFC1FF56FD5E84764269:8E8B605F5830FFC7FD95FCC382DF4389" box="[596,699,871,897]" country="Malaysia" county="Camp" latitude="4.0508003" longLatPrecision="1" longitude="114.867004" municipality="Mulu Expedition" name="Province" pageId="2" pageNumber="3">Province</location>
</emphasis>
: 
<typeStatus id="54EF88265830FFC7FD08FCC383524389" box="[713,822,871,897]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
</emphasis>
male (
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): 
<collectingMunicipality id="6B8FACFE5830FFC7FC2AFCC384CE4389" box="[1003,1194,871,897]" pageId="2" pageNumber="3">Mulu Expedition</collectingMunicipality>
, 
<collectingCounty id="628A4E085830FFC7FB73FCC384914389" box="[1202,1269,871,897]" pageId="2" pageNumber="3">Camp</collectingCounty>
1, 
<geoCoordinate id="EE6050435830FFC7FAD2FCC385F34389" box="[1299,1431,871,897]" degrees="4" direction="north" minutes="3" orientation="latitude" pageId="2" pageNumber="3" precision="1" seconds="2.88" value="4.0508003">4°3’2.88”N</geoCoordinate>
, 
<geoCoordinate id="EE6050435830FFC7FF56FC2D812643AC" box="[151,322,905,932]" degrees="114" direction="east" minutes="52" orientation="longitude" pageId="2" pageNumber="3" precision="1" seconds="1.20" value="114.867004">114°52’1.20”E</geoCoordinate>
, ca. 
<quantity id="4CAC9B615830FFC7FEB5FC2D81D943AC" box="[372,445,905,932]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="3" unit="m" value="330.0">
<elevation id="0079D1B75830FFC7FEB5FC2D81D943AC" box="[372,445,905,932]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="3" unit="m" value="330.0">330 m</elevation>
</quantity>
, MDF, leaf litter near waterfall, 
<date id="FFEA10445830FFC7FCE6FC2D83DB43AB" box="[807,959,905,932]" pageId="2" pageNumber="3" value="1978-05-24">
<collectingDate id="EFAEE9AC5830FFC7FCE6FC2D83DB43AB" box="[807,959,905,932]" pageId="2" pageNumber="3" value="1978-05-24">24 May 1978</collectingDate>
</date>
, 
<collectorName id="26A153525830FFC7FC0BFC2E842643AB" box="[970,1090,905,932]" pageId="2" pageNumber="3">F. Wanless</collectorName>
(
<collectionCode id="ED45AE415830FFC7FB90FC2D84FD43AB" box="[1105,1177,905,931]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="2" pageNumber="3" type="Herbarium">NHM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEB36845830FFC7FF06FC08848943E1" blockId="2.[151,1437,870,2012]" pageId="2" pageNumber="3">
<emphasis id="B920EA965830FFC7FF06FC0881B743CE" bold="true" box="[199,467,940,966]" pageId="2" pageNumber="3">
<typeStatus id="54EF88265830FFC7FF06FC08815B43CE" box="[199,319,940,966]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D52FD0D5830FFC7FE91FC0881A343CE" box="[336,455,940,966]" pageId="2" pageNumber="3" type="female">4 females</specimenCount>
):
</emphasis>
<materialsCitation id="3B3C3CD95830FFC7FE1AFC0885E643CE" ID-GBIF-Occurrence="3866517320" box="[475,1410,940,967]" collectingDate="1978-05-24" collectionCode="NHM" collectorName="F. Wanless" country="Malaysia" county="Camp" elevation="330" latitude="4.0508003" location="Province" longLatPrecision="1" longitude="114.867004" municipality="Mulu Expedition" pageId="2" pageNumber="3" specimenCode="PJ 1228-1229" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="9D52FD0D5830FFC7FE1AFC08822D43CE" box="[475,585,940,966]" pageId="2" pageNumber="3" type="female">2 females</specimenCount>
(
<specimenCode id="DBF29EFF5830FFC7FD98FC08829843CE" box="[601,764,940,966]" collectionCode="PJ" pageId="2" pageNumber="3">PJ 1228–1229</specimenCode>
) with same data as for holotype (
<collectionCode id="ED45AE415830FFC7FBB8FC0884DB43CE" box="[1145,1215,940,966]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="2" pageNumber="3" type="Herbarium">NHM</collectionCode>
; 
<specimenCount id="9D52FD0D5830FFC7FB0AFC08855743CE" box="[1227,1331,940,966]" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
: SMF)
</materialsCitation>
. 
<materialsCitation id="3B3C3CD95830FFC7FA4FFC08848D43E1" ID-GBIF-Occurrence="3866517336" collectingDate="1978-06-20" collectionCode="NHM" collectorName="F. Wanless" country="Malaysia" county="Camp" elevation="330" latitude="4.0508003" location="Province" longLatPrecision="1" longitude="114.867004" municipality="Mulu Expedition" pageId="2" pageNumber="3" specimenCode="PJ 1204-1205" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D52FD0D5830FFC7FA4FFC08808A43E1" pageId="2" pageNumber="3" type="female">2 females</specimenCount>
(
<specimenCode id="DBF29EFF5830FFC7FF3CFC6B81C443E1" box="[253,416,974,1001]" collectionCode="PJ" pageId="2" pageNumber="3">PJ 1204–1205</specimenCode>
) with same data as for holotype, except: shrub layer, 
<date id="FFEA10445830FFC7FC34FC6B84EE43E1" box="[1013,1162,975,1001]" pageId="2" pageNumber="3" value="1978-06-20">
<collectingDate id="EFAEE9AC5830FFC7FC34FC6B84EE43E1" box="[1013,1162,975,1001]" pageId="2" pageNumber="3" value="1978-06-20">20 June 1978</collectingDate>
</date>
(
<collectionCode id="ED45AE415830FFC7FB58FC6B848543E1" box="[1177,1249,975,1001]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="2" pageNumber="3" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEB36845830FFC7FF06FC55816D44B1" blockId="2.[151,1437,870,2012]" pageId="2" pageNumber="3">
<emphasis id="B920EA965830FFC7FF06FC5583104403" bold="true" box="[199,884,1009,1035]" pageId="2" pageNumber="3">
<materialsCitation id="3B3C3CD95830FFC7FF06FC55830F4403" ID-GBIF-Occurrence="3866517325" box="[199,875,1009,1035]" country="Malaysia" location="MALAYSIA" pageId="2" pageNumber="3" specimenCount="14" specimenCount-female="14" stateProvince="Province">
Additional material examined (
<specimenCount id="9D52FD0D5830FFC7FDFFFC5582A54403" box="[574,705,1009,1035]" pageId="2" pageNumber="3" type="female">14 females</specimenCount>
). 
<collectingCountry id="F34376145830FFC7FD14FC55830F4403" box="[725,875,1009,1035]" name="Malaysia" pageId="2" pageNumber="3">MALAYSIA</collectingCountry>
</materialsCitation>
:
</emphasis>
<materialsCitation id="3B3C3CD95830FFC7FCBDFC5581334426" ID-GBIF-Occurrence="3866517314" collectingDate="1978-05-24" collectionCode="NHM" collectorName="F. Wanless" country="Malaysia" county="Camp" elevation="330" latitude="4.0508003" location="Province" longLatPrecision="1" longitude="114.867004" municipality="Mulu Expedition" pageId="2" pageNumber="3" specimenCount="9" specimenCount-female="9" stateProvince="Province">
<emphasis id="B920EA965830FFC7FCBDFC55843C4403" bold="true" box="[892,1112,1009,1036]" pageId="2" pageNumber="3">
<emphasis id="B920EA965830FFC7FCBDFC55842B4403" bold="true" box="[892,1103,1009,1036]" italics="true" pageId="2" pageNumber="3">
<collectingCountry id="F34376145830FFC7FCBDFC5583854404" box="[892,993,1009,1036]" name="Malaysia" pageId="2" pageNumber="3">Sarawak</collectingCountry>
<collectingRegion id="4990F8665830FFC7FC29FC55842B4403" box="[1000,1103,1009,1035]" country="Malaysia" name="Province" pageId="2" pageNumber="3">Province</collectingRegion>
</emphasis>
:
</emphasis>
<specimenCount id="9D52FD0D5830FFC7FBA1FC5584A84403" box="[1120,1228,1009,1035]" pageId="2" pageNumber="3" type="female">9 females</specimenCount>
with same data as for holotype, but
</materialsCitation>
: 
<materialsCitation id="3B3C3CD95830FFC7FEA0FBB082534426" ID-GBIF-Occurrence="3866517326" box="[353,567,1044,1070]" collectionCode="PJ" country="Malaysia" pageId="2" pageNumber="3" specimenCode="PJ 1221" specimenCount="1" specimenCount-female="1" stateProvince="Province">
<specimenCount id="9D52FD0D5830FFC7FEA0FBB081A64426" box="[353,450,1044,1070]" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
(
<specimenCode id="DBF29EFF5830FFC7FE10FBB0824B4426" box="[465,559,1044,1070]" collectionCode="PJ" pageId="2" pageNumber="3">PJ 1221</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="3B3C3CD95830FFC7FD83FBB081E54459" ID-GBIF-Occurrence="3866517317" collectingDate="1978-06-05" collectionCode="NHM" country="Malaysia" elevation="100" latitude="4.059792" location="Tapin river" longLatPrecision="1" longitude="114.86328" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Province">
<location id="8E8B605F5830FFC7FD83FBB082A44426" LSID="urn:lsid:plazi:treatment:03FD87925830FFC1FF56FD5E84764269:8E8B605F5830FFC7FD83FBB082A44426" box="[578,704,1044,1070]" country="Malaysia" latitude="4.059792" longLatPrecision="1" longitude="114.86328" name="Tapin river" pageId="2" pageNumber="3" stateProvince="Province">Tapin river</location>
, 
<geoCoordinate id="EE6050435830FFC7FD0BFBB083394427" box="[714,861,1044,1071]" degrees="4" direction="north" minutes="3" orientation="latitude" pageId="2" pageNumber="3" precision="1" seconds="35.25" value="4.059792">4°3’35.25”N</geoCoordinate>
, 
<geoCoordinate id="EE6050435830FFC7FCA9FBB084454427" box="[872,1057,1044,1071]" degrees="114" direction="east" minutes="51" orientation="longitude" pageId="2" pageNumber="3" precision="1" seconds="47.83" value="114.86328">114°51’47.83”E</geoCoordinate>
, ca. 
<quantity id="4CAC9B615830FFC7FB92FBB084F84427" box="[1107,1180,1044,1071]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="100.0">
<elevation id="0079D1B75830FFC7FB92FBB084F84427" box="[1107,1180,1044,1071]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="100.0">100 m</elevation>
</quantity>
, waterside, vegetation, 
<date id="FFEA10445830FFC7FF56FB9281454459" box="[151,289,1078,1105]" pageId="2" pageNumber="3" value="1978-06-05">
<collectingDate id="EFAEE9AC5830FFC7FF56FB9281454459" box="[151,289,1078,1105]" pageId="2" pageNumber="3" value="1978-06-05">5 June 1978</collectingDate>
</date>
(
<collectionCode id="ED45AE415830FFC7FEF3FB93811D4459" box="[306,377,1079,1105]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="2" pageNumber="3" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B3C3CD95830FFC7FE4FFB9382DF4459" ID-GBIF-Occurrence="3866517339" box="[398,699,1079,1105]" collectionCode="PJ" country="Malaysia" pageId="2" pageNumber="3" specimenCode="PJ 1218-1220" specimenCount="3" specimenCount-female="3" stateProvince="Province">
<specimenCount id="9D52FD0D5830FFC7FE4FFB9381984459" box="[398,508,1079,1105]" pageId="2" pageNumber="3" type="female">3 females</specimenCount>
(
<specimenCode id="DBF29EFF5830FFC7FDCDFB9382D74459" box="[524,691,1079,1105]" collectionCode="PJ" pageId="2" pageNumber="3">PJ 1218–1220</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="3B3C3CD95830FFC7FD06FB9284E64459" ID-GBIF-Occurrence="3866517323" box="[711,1154,1078,1105]" collectionCode="NHM" country="Malaysia" elevation="150" location="Plot" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Province">
<quantity id="4CAC9B615830FFC7FD06FB9283764459" box="[711,786,1078,1105]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="3" unit="m" value="150.0">
<elevation id="0079D1B75830FFC7FD06FB9283764459" box="[711,786,1078,1105]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="3" unit="m" value="150.0">150 m</elevation>
</quantity>
, 
<location id="8E8B605F5830FFC7FCDEFB9383284459" LSID="urn:lsid:plazi:treatment:03FD87925830FFC1FF56FD5E84764269:8E8B605F5830FFC7FCDEFB9383284459" box="[799,844,1079,1105]" country="Malaysia" name="Plot" pageId="2" pageNumber="3" stateProvince="Province">Plot</location>
2, forest leaf litter (
<collectionCode id="ED45AE415830FFC7FBF3FB93841D4459" box="[1074,1145,1079,1105]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="2" pageNumber="3" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B3C3CD95830FFC7FB4EFB92817B447B" ID-GBIF-Occurrence="3866517303" collectionCode="SD, PJ" country="Malaysia" pageId="2" pageNumber="3" specimenCode="SD 1449, PJ 1222-1226" specimenCount="5" specimenCount-female="5" stateProvince="Province">
<specimenCount id="9D52FD0D5830FFC7FB4EFB9284984459" box="[1167,1276,1078,1105]" pageId="2" pageNumber="3" type="female">5 females</specimenCount>
(
<specimenCode id="DBF29EFF5830FFC7FACCFB9385114459" box="[1293,1397,1079,1105]" collectionCode="SD" pageId="2" pageNumber="3">SD 1449</specimenCode>
, 
<specimenCode id="DBF29EFF5830FFC7FA43FB938173447B" collectionCode="PJ" pageId="2" pageNumber="3">PJ 1222–1226</specimenCode>
)
</materialsCitation>
<materialsCitation id="3B3C3CD95830FFC7FEE9FBFD82BF447B" ID-GBIF-Occurrence="3866517322" box="[296,731,1113,1140]" collectingDate="1978-03-02" collectionCode="NHM" collectorName="H. W. Vallame" country="Malaysia" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Province">
<date id="FFEA10445830FFC7FEE9FBFD81A9447B" box="[296,461,1113,1139]" pageId="2" pageNumber="3" value="1978-03-02">
<collectingDate id="EFAEE9AC5830FFC7FEE9FBFD81A9447B" box="[296,461,1113,1139]" pageId="2" pageNumber="3" value="1978-03-02">2 March 1978</collectingDate>
</date>
, 
<collectorName id="26A153525830FFC7FE1AFBFE821E447B" box="[475,634,1113,1140]" pageId="2" pageNumber="3">H.W. Vallame</collectorName>
(
<collectionCode id="ED45AE415830FFC7FD4AFBFD82B7447B" box="[651,723,1113,1139]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="2" pageNumber="3" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B3C3CD95830FFC7FD29FBFD816144B1" ID-GBIF-Occurrence="3866517306" collectingDate="2013-02-01" collectingDateMax="2013-02-25" collectingDateMin="2013-02-01" collectionCode="SMF" collectorName="O. Machac" country="Brunei" county="Ulu Temburong National Park" elevation="147" latitude="4.5470304" location="Centre" longLatPrecision="1" longitude="115.155655" municipality="Kuala Belalong Field" pageId="2" pageNumber="3" specimenCount="2" specimenCount-female="2" stateProvince="Temburong">
<emphasis id="B920EA965830FFC7FD29FBFD8391447B" bold="true" box="[744,1013,1113,1140]" pageId="2" pageNumber="3">
<collectingCountry id="F34376145830FFC7FD29FBFD8331447B" box="[744,853,1113,1139]" name="Brunei" pageId="2" pageNumber="3">BRUNEI</collectingCountry>
: 
<emphasis id="B920EA965830FFC7FCA9FBFD8389447C" bold="true" box="[872,1005,1113,1140]" italics="true" pageId="2" pageNumber="3">
<collectingRegion id="4990F8665830FFC7FCA9FBFD8389447C" box="[872,1005,1113,1140]" country="Brunei" name="Temburong" pageId="2" pageNumber="3">Temburong</collectingRegion>
</emphasis>
:
</emphasis>
<specimenCount id="9D52FD0D5830FFC7FC3EFBFD8415447B" box="[1023,1137,1113,1139]" pageId="2" pageNumber="3" type="female">2 females</specimenCount>
, 
<collectingCounty id="628A4E085830FFC7FBBEFBFD80A9449E" pageId="2" pageNumber="3">Ulu Temburong National Park</collectingCounty>
, 
<collectingMunicipality id="6B8FACFE5830FFC7FF16FBD881A1449E" box="[215,453,1148,1174]" pageId="2" pageNumber="3">Kuala Belalong Field</collectingMunicipality>
study 
<location id="8E8B605F5830FFC7FDCFFBD88238449E" LSID="urn:lsid:plazi:treatment:03FD87925830FFC1FF56FD5E84764269:8E8B605F5830FFC7FDCFFBD88238449E" box="[526,604,1148,1174]" country="Brunei" county="Ulu Temburong National Park" latitude="4.5470304" longLatPrecision="1" longitude="115.155655" municipality="Kuala Belalong Field" name="Centre" pageId="2" pageNumber="3" stateProvince="Temburong">Centre</location>
, ca. 
<geoCoordinate id="EE6050435830FFC7FD4AFBD8834F449E" box="[651,811,1148,1174]" degrees="4" direction="north" minutes="32" orientation="latitude" pageId="2" pageNumber="3" precision="1" seconds="49.31" value="4.5470304">4°32’49.31”N</geoCoordinate>
, 
<geoCoordinate id="EE6050435830FFC7FCF7FBD88383449E" box="[822,999,1148,1175]" degrees="115" direction="east" minutes="9" orientation="longitude" pageId="2" pageNumber="3" precision="1" seconds="20.36" value="115.155655">115° 9’20.36”E</geoCoordinate>
, 
<quantity id="4CAC9B615830FFC7FC30FBD8845D449F" box="[1009,1081,1148,1175]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="2" pageNumber="3" unit="m" value="147.0">
<elevation id="0079D1B75830FFC7FC30FBD8845D449F" box="[1009,1081,1148,1175]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="2" pageNumber="3" unit="m" value="147.0">147 m</elevation>
</quantity>
, 
<date id="FFEA10445830FFC7FB85FBD8854C449E" box="[1092,1320,1148,1175]" pageId="2" pageNumber="3" value="2013-02-01" valueMax="2013-02-25" valueMin="2013-02-01">
<collectingDate id="EFAEE9AC5830FFC7FB85FBD8854C449E" box="[1092,1320,1148,1175]" pageId="2" pageNumber="3" value="2013-02-01" valueMax="2013-02-25" valueMin="2013-02-01">1–25 February 2013</collectingDate>
</date>
, 
<collectorName id="26A153525830FFC7FAF3FBD880D444B1" pageId="2" pageNumber="3">O. Machac</collectorName>
(
<collectionCode id="ED45AE415830FFC7FF7EFB3B809944B1" box="[191,253,1183,1209]" country="Germany" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="3">SMF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5830FFC7FF06FB65824744F6" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BEB36845830FFC7FF06FB65824744F6" blockId="2.[151,1437,870,2012]" pageId="2" pageNumber="3">
<emphasis id="B920EA965830FFC7FF06FB65812A44D3" bold="true" box="[199,334,1217,1243]" pageId="2" pageNumber="3">Etymology.</emphasis>
The species name is derived from the Latin word “parvus” meaning “small” and refers to the small body size of this species; adjective.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5830FFC7FF06FAA280B545F9" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BEB36845830FFC7FF06FAA280B545F9" blockId="2.[151,1437,870,2012]" pageId="2" pageNumber="3">
<emphasis id="B920EA965830FFC7FF06FAA281264528" bold="true" box="[199,322,1286,1312]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Small 
<taxonomicName id="4C544D075830FFC7FE4EFAA382774529" authorityName="Bertkau" authorityYear="1872" box="[399,531,1287,1313]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
with TL &lt;4.5 (
<figureCitation id="136F2A015830FFC7FD7EFAA383254529" box="[703,833,1286,1313]" captionStart="FIGURES 82–97" captionStartId="19.[151,264,1899,1923]" captionTargetBox="[152,1435,182,1873]" captionTargetId="figure-19@19.[151,1436,181,1875]" captionTargetPageId="19" captionText="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="2" pageNumber="3">Figs 82–85</figureCitation>
). 
<taxonomicName id="4C544D075830FFC7FC92FAA384604528" authority="Grall &amp; Jäger, 2022" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[851,1028,1287,1313]" class="Arachnida" family="Sparassidae" genus="Borniella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
<emphasis id="B920EA965830FFC7FC92FAA384604528" box="[851,1028,1287,1313]" italics="true" pageId="2" pageNumber="3">Borniella parva</emphasis>
</taxonomicName>
<emphasis id="B920EA965830FFC7FBCBFAA384184529" bold="true" box="[1034,1148,1287,1313]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A21357ED5830FFC7FBCBFAA384184529" box="[1034,1148,1287,1313]" pageId="2" pageNumber="3" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
may be recognised by the following combination of characters: Males (
<figureCitation id="136F2A015830FFC7FD4DFA8D8295454B" box="[652,753,1321,1347]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1546,1570]" captionTargetBox="[151,1436,315,1519]" captionTargetId="figure-74@3.[151,1436,306,1519]" captionTargetPageId="3" captionText="FIGURES 1–3. Borniella parva gen. &amp; spec. nov. holotype male from Sarawak, left male palp (1 prolateral, 2 ventral, 3 retrolateral). C—conductor, dTA— distal tegular apophysis, E—embolus, mTA—median tegular apophysis, pTA—proximal tegular apophysis, RTA—retrolateral tibial apophysis, S—spermophor, ST—subtegulum." figureDoi="http://doi.org/10.5281/zenodo.6911153" httpUri="https://zenodo.org/record/6911153/files/figure.png" pageId="2" pageNumber="3">Figs 1–3</figureCitation>
): 1. Single RTA long, reaching distally to retrolateral cymbial bulge, 2. Embolus arising from tegulum at 9-o’clock-position, running roughly a semicircle and 3. Distal TA largest with small tooth ventrally, median TA mid-range in size, triangular in prolateral view, proximal TA smallest, acute triangular, situated close to spermophor winding. Females (
<figureCitation id="136F2A015830FFC7FCF1FA3583FC45A3" box="[816,920,1425,1451]" captionStart="FIGURES 4–7" captionStartId="4.[151,264,1801,1825]" captionTargetBox="[151,1433,633,1778]" captionTargetId="figure-194@4.[151,1436,633,1778]" captionTargetPageId="4" captionText="FIGURES 4–7. Borniella parva gen. &amp; spec. nov., female paratype from Sarawak. 4, 6 Epigyne, ventral; 5 Vulva, dorsal (internal duct system and rim of copulatory opening in red, epigynal pocket and parts of the inner rims in green); 7 Schematic course of internal duct system, ventral. CO—copulatory opening, CP—copulatory plug, EF—epigynal field, EP—epigynal pocket, FD—fertilization duct, GA—glandular appendages, LS—lobal septum, MSu—membranous sac (unexpanded), SP— spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6911159" httpUri="https://zenodo.org/record/6911159/files/figure.png" pageId="2" pageNumber="3">Figs 4–7</figureCitation>
): 1. Posterior part of vulva with spermathecae distinctly wider than anterior part with glandular appendages and 2. Copulatory openings with roughly right-angled rims.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5830FFC1FF06FA5D85F44235" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="8BEB36845830FFC7FF06FA5D85194721" blockId="2.[151,1437,870,2012]" pageId="2" pageNumber="3">
<emphasis id="B920EA965830FFC7FF06FA5D8240461B" bold="true" box="[199,548,1529,1555]" pageId="2" pageNumber="3">
Description. Male (
<typeStatus id="54EF88265830FFC7FE6BFA5D827C461B" box="[426,536,1529,1555]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
):
</emphasis>
TL 3.3; PL 1.7, PW 1.7, AW 0.95; OL 1.6, OW 1.1. Eye measurements (
<figureCitation id="136F2A015830FFC7FA88FA5D85E9461B" box="[1353,1421,1529,1555]" captionStart="FIGURES 8–16" captionStartId="5.[151,264,1380,1404]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-19@5.[151,1436,181,1357]" captionTargetPageId="5" captionText="FIGURES 8–16. Borniella parva gen. &amp; spec. nov., holotype male (8, 10, 13, 15) and female paratype (9, 11–12, 14, 16) from Sarawak. 8–9 Eyes, dorsal. 10–12 Claws (10 right leg II, prolateral claw, prolateral; 11 right palp, prolateral; 12 right leg II, retrolateral claw, retrolateral). 13–14 Left chelicera, ventral. 15–16 Trilobate membrane, distal metatarsus, dorsal (15 right leg III, 16 right leg I). LP—lateral projections, MH—median hook." figureDoi="http://doi.org/10.5281/zenodo.6911161" httpUri="https://zenodo.org/record/6911161/files/figure.png" pageId="2" pageNumber="3">Fig. 8</figureCitation>
): AME 0.11; ALE 0.16; PME 0.10; PLE 0.22; AME–AME 0.06; AME–ALE 0.01; PME–PME 0.15; PME–PLE 0.18; AME–PME 0.15; ALE–PLE 0.16; clypeus AME 0.14; clypeus ALE 0.13. PLE on small humps. Measurements of palp and legs: palp: 2.55 (0.75, 0.40, 0.50, 0.90); I: 6.15 (1.65, 0.80, 1.60, 1.50, 0.60); II: 7.85 (2.20, 0.90, 2.10, 1.95, 0.70); III: 6.60 (1.90, 0.80, 1.70, 1.60, 0.60); IV: 6.70 (1.90, 0.70, 1.70, 1.75, 0.65). Leg formula: 2431. Spination: palp: 131, 101, 2110; legs: Fe I 322, Fe II &amp; III 222, Fe IV 231; Pa I–IV 000; Ti I &amp; 
<date id="FFEA10445830FFC7FB85F90384FD46C9" box="[1092,1177,1703,1729]" pageId="2" pageNumber="3" value="1016-02">II 1016</date>
, Ti III &amp; 
<date id="FFEA10445830FFC7FAC0F903850546C9" box="[1281,1377,1703,1729]" pageId="2" pageNumber="3" value="2226-04">IV 2226</date>
; Mt I &amp; II 0006, Mt 
<date id="FFEA10445830FFC7FEFBF96E81FC46EB" box="[314,408,1737,1764]" pageId="2" pageNumber="3" value="2036-03">III 2036</date>
, Mt IV 3036. Chelicerae with 3 promarginal, 5 retromarginal teeth, 13–14 denticles in patch close to promarginal teeth and 1 escort seta (
<figureCitation id="136F2A015830FFC7FD56F9488288470E" box="[663,748,1772,1798]" captionStart="FIGURES 8–16" captionStartId="5.[151,264,1380,1404]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-19@5.[151,1436,181,1357]" captionTargetPageId="5" captionText="FIGURES 8–16. Borniella parva gen. &amp; spec. nov., holotype male (8, 10, 13, 15) and female paratype (9, 11–12, 14, 16) from Sarawak. 8–9 Eyes, dorsal. 10–12 Claws (10 right leg II, prolateral claw, prolateral; 11 right palp, prolateral; 12 right leg II, retrolateral claw, retrolateral). 13–14 Left chelicera, ventral. 15–16 Trilobate membrane, distal metatarsus, dorsal (15 right leg III, 16 right leg I). LP—lateral projections, MH—median hook." figureDoi="http://doi.org/10.5281/zenodo.6911161" httpUri="https://zenodo.org/record/6911161/files/figure.png" pageId="2" pageNumber="3">Fig. 13</figureCitation>
). Serrula with 24 denticles. Prolateral claw of leg II with 17 teeth (
<figureCitation id="136F2A015830FFC7FF1CF8AB81544721" box="[221,304,1807,1833]" captionStart="FIGURES 8–16" captionStartId="5.[151,264,1380,1404]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-19@5.[151,1436,181,1357]" captionTargetPageId="5" captionText="FIGURES 8–16. Borniella parva gen. &amp; spec. nov., holotype male (8, 10, 13, 15) and female paratype (9, 11–12, 14, 16) from Sarawak. 8–9 Eyes, dorsal. 10–12 Claws (10 right leg II, prolateral claw, prolateral; 11 right palp, prolateral; 12 right leg II, retrolateral claw, retrolateral). 13–14 Left chelicera, ventral. 15–16 Trilobate membrane, distal metatarsus, dorsal (15 right leg III, 16 right leg I). LP—lateral projections, MH—median hook." figureDoi="http://doi.org/10.5281/zenodo.6911161" httpUri="https://zenodo.org/record/6911161/files/figure.png" pageId="2" pageNumber="3">Fig. 10</figureCitation>
). Trilobate membrane with narrow median hook, slightly shorter than lateral projections (
<figureCitation id="136F2A015830FFC7FADDF8AB85144721" box="[1308,1392,1806,1833]" captionStart="FIGURES 8–16" captionStartId="5.[151,264,1380,1404]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-19@5.[151,1436,181,1357]" captionTargetPageId="5" captionText="FIGURES 8–16. Borniella parva gen. &amp; spec. nov., holotype male (8, 10, 13, 15) and female paratype (9, 11–12, 14, 16) from Sarawak. 8–9 Eyes, dorsal. 10–12 Claws (10 right leg II, prolateral claw, prolateral; 11 right palp, prolateral; 12 right leg II, retrolateral claw, retrolateral). 13–14 Left chelicera, ventral. 15–16 Trilobate membrane, distal metatarsus, dorsal (15 right leg III, 16 right leg I). LP—lateral projections, MH—median hook." figureDoi="http://doi.org/10.5281/zenodo.6911161" httpUri="https://zenodo.org/record/6911161/files/figure.png" pageId="2" pageNumber="3">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BEB36845830FFC7FF06F895852B47D3" blockId="2.[151,1437,870,2012]" pageId="2" pageNumber="3">
Palp as in diagnosis (
<figureCitation id="136F2A015830FFC7FE79F895827B4743" box="[440,543,1841,1867]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1546,1570]" captionTargetBox="[151,1436,315,1519]" captionTargetId="figure-74@3.[151,1436,306,1519]" captionTargetPageId="3" captionText="FIGURES 1–3. Borniella parva gen. &amp; spec. nov. holotype male from Sarawak, left male palp (1 prolateral, 2 ventral, 3 retrolateral). C—conductor, dTA— distal tegular apophysis, E—embolus, mTA—median tegular apophysis, pTA—proximal tegular apophysis, RTA—retrolateral tibial apophysis, S—spermophor, ST—subtegulum." figureDoi="http://doi.org/10.5281/zenodo.6911153" httpUri="https://zenodo.org/record/6911153/files/figure.png" pageId="2" pageNumber="3">Figs 1–3</figureCitation>
). RTA long and tapering, almost straight in ventral view, moderately curved in retrolateral view. Conductor short and slender, arising at 12.30-o’clock-position from tegulum. Retrolateral part of tegulum kidney-shaped. Spermophor running retrolaterally submarginally along tegular margin, with distinct prolaterad U-turn proximally and another retrolaterad U-turn following directly after that. Embolus slender and semicircular. Distal tegular apophysis sail-shaped, connected directly with its prolateral rim to the embolus base.
</paragraph>
<paragraph id="8BEB36845831FFC6FF06FF33853E4115" blockId="3.[151,1437,151,285]" pageId="3" pageNumber="4">
Colouration in ethanol (
<figureCitation id="136F2A015831FFC6FE10FF33823140B9" box="[465,597,151,177]" captionStart="FIGURES 82–97" captionStartId="19.[151,264,1899,1923]" captionTargetBox="[152,1435,182,1873]" captionTargetId="figure-19@19.[151,1436,181,1875]" captionTargetPageId="19" captionText="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="3" pageNumber="4">Figs 82–83</figureCitation>
): Prosoma yellowish-white, posterior and lateral margins with brown band, medio-anteriorly light brown semi-circular pattern, medially with brown longitudinal stripe along fovea. Sternum yellowish-white. Chelicerae yellow-brown. Coxa I–III medio-dorsally with brown stripe. Palps and legs yellowish-white. Opisthosoma yellowish-white, laterally and posteriorly with brown spots, ventral part white, posteriorly brown.
</paragraph>
<caption id="DF2B660C5831FFC6FF56F9AE83B04662" ID-DOI="http://doi.org/10.5281/zenodo.6911153" ID-Zenodo-Dep="6911153" httpUri="https://zenodo.org/record/6911153/files/figure.png" pageId="3" pageNumber="4" startId="3.[151,264,1546,1570]" targetBox="[151,1436,315,1519]" targetPageId="3">
<paragraph id="8BEB36845831FFC6FF56F9AE83B04662" blockId="3.[151,1436,1546,1642]" pageId="3" pageNumber="4">
<emphasis id="B920EA965831FFC6FF56F9AE815E462A" bold="true" box="[151,314,1546,1570]" pageId="3" pageNumber="4">FIGURES 1–3.</emphasis>
<taxonomicName id="4C544D075831FFC6FE81F9AE8185462A" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[320,481,1546,1570]" class="Arachnida" family="Sparassidae" genus="Borniella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B920EA965831FFC6FE81F9AE8185462A" box="[320,481,1546,1570]" italics="true" pageId="3" pageNumber="4">Borniella parva</emphasis>
</taxonomicName>
<emphasis id="B920EA965831FFC6FE26F9AE82FF462A" bold="true" box="[487,667,1546,1570]" pageId="3" pageNumber="4">gen. &amp; spec. nov.</emphasis>
holotype male from Sarawak, left male palp (1 prolateral, 2 ventral, 3 retrolateral). C—conductor, dTA— distal tegular apophysis, E—embolus, mTA—median tegular apophysis, pTA—proximal tegular apophysis, RTA—retrolateral tibial apophysis, S—spermophor, ST—subtegulum.
</paragraph>
</caption>
<paragraph id="8BEB36845831FFC6FF06F93C83A247FE" blockId="3.[151,1437,1688,2039]" pageId="3" pageNumber="4">
<emphasis id="B920EA965831FFC6FF06F93C81C846BA" bold="true" box="[199,428,1688,1714]" pageId="3" pageNumber="4">
Female (
<typeStatus id="54EF88265831FFC6FEECF93C81FB46BA" box="[301,415,1688,1714]" pageId="3" pageNumber="4" type="paratype">paratype</typeStatus>
):
</emphasis>
TL 3.7; PL 1.8, PW 1.7, AW 1.0; OL 1.9, OW 1.2. Eye measurements (
<figureCitation id="136F2A015831FFC6FB06F93C856846BA" box="[1223,1292,1688,1714]" captionStart="FIGURES 8–16" captionStartId="5.[151,264,1380,1404]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-19@5.[151,1436,181,1357]" captionTargetPageId="5" captionText="FIGURES 8–16. Borniella parva gen. &amp; spec. nov., holotype male (8, 10, 13, 15) and female paratype (9, 11–12, 14, 16) from Sarawak. 8–9 Eyes, dorsal. 10–12 Claws (10 right leg II, prolateral claw, prolateral; 11 right palp, prolateral; 12 right leg II, retrolateral claw, retrolateral). 13–14 Left chelicera, ventral. 15–16 Trilobate membrane, distal metatarsus, dorsal (15 right leg III, 16 right leg I). LP—lateral projections, MH—median hook." figureDoi="http://doi.org/10.5281/zenodo.6911161" httpUri="https://zenodo.org/record/6911161/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
): AME 0.10; ALE 0.16; PME 0.09; PLE 0.19; AME–AME 0.10; AME–ALE 0.03; PME–PME 0.15; PME–PLE 0.21; AME–PME 0.15; ALE–PLE 0.19; clypeus AME 0.15; clypeus ALE 0.13. PLE on small humps. Leg formula: 2431; measurements of palp and legs: palp: 2.90 (0.80, 0.50, 0.70, 0.90); I: 5.85 (1.60, 0.70, 1.55, 1.45, 0.55); II: 7.25 (2.10, 0.80, 2.00, 1.75, 0.60); III: 6.20 (1.80, 0.70, 1.65, 1.50, 0.55); IV: 6.45 (1.80, 0.65, 1.70, 1.70, 0.60). Leg formula: 2431. Spination: palp: 131, 101, 2221, 1004; legs: Fe I 322, Fe II &amp; III 222, Fe IV 221; Pa I–IV 000; Ti I &amp; 
<date id="FFEA10445831FFC6FAC2F8E98533476E" box="[1283,1367,1868,1895]" pageId="3" pageNumber="4" value="1016-02">II 1016</date>
, Ti 
<date id="FFEA10445831FFC6FA41F8E980B64782" pageId="3" pageNumber="4" value="2006-03">III 2006</date>
, Ti 
<date id="FFEA10445831FFC6FF3DF8D581384782" box="[252,348,1904,1931]" pageId="3" pageNumber="4" value="2226-04">IV 2226</date>
; Mt I &amp; II 0006, Mt 
<date id="FFEA10445831FFC6FD85F8D582C64782" box="[580,674,1904,1931]" pageId="3" pageNumber="4" value="1016-03">III 1016</date>
, MT IV 3036. Chelicerae with 3 promarginal, 6 retromarginal teeth, 14–15 denticles in patch close to promarginal teeth and 1 escort seta (
<figureCitation id="136F2A015831FFC6FC54F830838347A6" box="[917,999,1940,1966]" captionStart="FIGURES 8–16" captionStartId="5.[151,264,1380,1404]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-19@5.[151,1436,181,1357]" captionTargetPageId="5" captionText="FIGURES 8–16. Borniella parva gen. &amp; spec. nov., holotype male (8, 10, 13, 15) and female paratype (9, 11–12, 14, 16) from Sarawak. 8–9 Eyes, dorsal. 10–12 Claws (10 right leg II, prolateral claw, prolateral; 11 right palp, prolateral; 12 right leg II, retrolateral claw, retrolateral). 13–14 Left chelicera, ventral. 15–16 Trilobate membrane, distal metatarsus, dorsal (15 right leg III, 16 right leg I). LP—lateral projections, MH—median hook." figureDoi="http://doi.org/10.5281/zenodo.6911161" httpUri="https://zenodo.org/record/6911161/files/figure.png" pageId="3" pageNumber="4">Fig. 14</figureCitation>
). Serrula with 26 denticles. Palpal claw with 5 long and 2–3 short teeth (
<figureCitation id="136F2A015831FFC6FDCBF81C823947DA" box="[522,605,1976,2002]" captionStart="FIGURES 8–16" captionStartId="5.[151,264,1380,1404]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-19@5.[151,1436,181,1357]" captionTargetPageId="5" captionText="FIGURES 8–16. Borniella parva gen. &amp; spec. nov., holotype male (8, 10, 13, 15) and female paratype (9, 11–12, 14, 16) from Sarawak. 8–9 Eyes, dorsal. 10–12 Claws (10 right leg II, prolateral claw, prolateral; 11 right palp, prolateral; 12 right leg II, retrolateral claw, retrolateral). 13–14 Left chelicera, ventral. 15–16 Trilobate membrane, distal metatarsus, dorsal (15 right leg III, 16 right leg I). LP—lateral projections, MH—median hook." figureDoi="http://doi.org/10.5281/zenodo.6911161" httpUri="https://zenodo.org/record/6911161/files/figure.png" pageId="3" pageNumber="4">Fig. 11</figureCitation>
). Retrolateral claw of leg III with 13 teeth (
<figureCitation id="136F2A015831FFC6FB8EF81C84C747DA" box="[1103,1187,1976,2002]" captionStart="FIGURES 8–16" captionStartId="5.[151,264,1380,1404]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-19@5.[151,1436,181,1357]" captionTargetPageId="5" captionText="FIGURES 8–16. Borniella parva gen. &amp; spec. nov., holotype male (8, 10, 13, 15) and female paratype (9, 11–12, 14, 16) from Sarawak. 8–9 Eyes, dorsal. 10–12 Claws (10 right leg II, prolateral claw, prolateral; 11 right palp, prolateral; 12 right leg II, retrolateral claw, retrolateral). 13–14 Left chelicera, ventral. 15–16 Trilobate membrane, distal metatarsus, dorsal (15 right leg III, 16 right leg I). LP—lateral projections, MH—median hook." figureDoi="http://doi.org/10.5281/zenodo.6911161" httpUri="https://zenodo.org/record/6911161/files/figure.png" pageId="3" pageNumber="4">Fig. 12</figureCitation>
). Trilobate membrane with narrow median hook, almost as long as lateral projections (
<figureCitation id="136F2A015831FFC6FCA7F87883DD47FE" box="[870,953,2012,2038]" captionStart="FIGURES 8–16" captionStartId="5.[151,264,1380,1404]" captionTargetBox="[151,1436,181,1357]" captionTargetId="figure-19@5.[151,1436,181,1357]" captionTargetPageId="5" captionText="FIGURES 8–16. Borniella parva gen. &amp; spec. nov., holotype male (8, 10, 13, 15) and female paratype (9, 11–12, 14, 16) from Sarawak. 8–9 Eyes, dorsal. 10–12 Claws (10 right leg II, prolateral claw, prolateral; 11 right palp, prolateral; 12 right leg II, retrolateral claw, retrolateral). 13–14 Left chelicera, ventral. 15–16 Trilobate membrane, distal metatarsus, dorsal (15 right leg III, 16 right leg I). LP—lateral projections, MH—median hook." figureDoi="http://doi.org/10.5281/zenodo.6911161" httpUri="https://zenodo.org/record/6911161/files/figure.png" pageId="3" pageNumber="4">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8BEB36845836FFC1FF06FF3381294115" blockId="4.[151,1437,151,609]" pageId="4" pageNumber="5">
Copulatory organ as in diagnosis (
<figureCitation id="136F2A015836FFC1FD8CFF3382D140B9" box="[589,693,151,177]" captionStart="FIGURES 4–7" captionStartId="4.[151,264,1801,1825]" captionTargetBox="[151,1433,633,1778]" captionTargetId="figure-194@4.[151,1436,633,1778]" captionTargetPageId="4" captionText="FIGURES 4–7. Borniella parva gen. &amp; spec. nov., female paratype from Sarawak. 4, 6 Epigyne, ventral; 5 Vulva, dorsal (internal duct system and rim of copulatory opening in red, epigynal pocket and parts of the inner rims in green); 7 Schematic course of internal duct system, ventral. CO—copulatory opening, CP—copulatory plug, EF—epigynal field, EP—epigynal pocket, FD—fertilization duct, GA—glandular appendages, LS—lobal septum, MSu—membranous sac (unexpanded), SP— spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6911159" httpUri="https://zenodo.org/record/6911159/files/figure.png" pageId="4" pageNumber="5">Figs 4–7</figureCitation>
). Epigynal field oval, wider than long, without slit sensilla or anterior bands. Internal duct system wider than long. Posterior part of internal duct system wider than anterior part. Fertilization ducts short and slender, arising postero-laterally. Membranous structure covers posterior part of internal duct system.
</paragraph>
<paragraph id="8BEB36845836FFC1FF06FE8381CA41D9" blockId="4.[151,1437,151,609]" pageId="4" pageNumber="5">
Colouration in ethanol (
<figureCitation id="136F2A015836FFC1FE0CFE8382344149" box="[461,592,294,321]" captionStart="FIGURES 82–97" captionStartId="19.[151,264,1899,1923]" captionTargetBox="[152,1435,182,1873]" captionTargetId="figure-19@19.[151,1436,181,1875]" captionTargetPageId="19" captionText="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="4" pageNumber="5">Figs 84–85</figureCitation>
): Prosoma yellowish-white, medio-anteriorly with semi-circular light brown pattern, medially with brown longitudinal stripe, posterior and lateral margin brown. Sternum white. Opisthosoma greyish-white with brown spots, posteriorly brown, ventral part greyish-white. Chelicerae yellowish-brown. Coxa I–III medio-dorsally with brown stripe. Palps yellowish-white. Legs yellowish-white; Fe proximally light brown; Pa ventrally light brown.
</paragraph>
<paragraph id="8BEB36845836FFC1FF06FE7E85F44235" blockId="4.[151,1437,151,609]" pageId="4" pageNumber="5">
<emphasis id="B920EA965836FFC1FF06FE7E819641FC" bold="true" box="[199,498,474,500]" pageId="4" pageNumber="5">Variation female (n=14).</emphasis>
TL 3.25–4.1; PL 1.65–1.9, PW 1.6–1.8, AW 1.0–1.1; OL 1.6–2.3, OW 0.9–1.7. Measurements leg I: total length 5.62–6.20, Fe 1.50–1.70, Pa 0.70–0.80, Ti 1.45–1.60, Mt 1.35–1.50, Ta 0.55–0.60. Spination: palps: Ti 2121; Mt 1014; legs: Fe II &amp; III 322; Ti 
<date id="FFEA10445836FFC1FCFFFD8783F84235" box="[830,924,547,573]" pageId="4" pageNumber="5" value="1016-03">III 1016</date>
, 2026; Mt 
<date id="FFEA10445836FFC1FBD7FD8784104235" box="[1046,1140,547,573]" pageId="4" pageNumber="5" value="1006-03">III 1006</date>
, 2006, 2016, 2026, 3026.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5836FFC1FF06FDE284764269" box="[199,1042,582,609]" pageId="4" pageNumber="5" type="distribution">
<paragraph id="8BEB36845836FFC1FF06FDE284764269" blockId="4.[151,1437,151,609]" box="[199,1042,582,609]" pageId="4" pageNumber="5">
<emphasis id="B920EA965836FFC1FF06FDE281054268" bold="true" box="[199,353,582,608]" pageId="4" pageNumber="5">Distribution.</emphasis>
Borneo (
<collectingCountry id="F34376145836FFC1FE08FDE382524269" box="[457,566,583,609]" name="Malaysia" pageId="4" pageNumber="5">Malaysia</collectingCountry>
: 
<collectingCountry id="F34376145836FFC1FD81FDE382C04269" box="[576,676,583,609]" name="Malaysia" pageId="4" pageNumber="5">Sarawak</collectingCountry>
; 
<collectingCountry id="F34376145836FFC1FD6EFDE382994269" box="[687,765,583,609]" name="Brunei" pageId="4" pageNumber="5">Brunei</collectingCountry>
) (
<figureCitation id="136F2A015836FFC1FCD5FDE3830C4269" box="[788,872,583,609]" captionStart="FIGURE 98" captionStartId="20.[151,250,1436,1460]" captionTargetBox="[151,1436,956,1413]" captionTargetId="figure-372@20.[151,1436,956,1413]" captionTargetPageId="20" captionText="FIGURE 98. Distribution of the new genera and species in Malaysia, Brunei and Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.6911205" httpUri="https://zenodo.org/record/6911205/files/figure.png" pageId="4" pageNumber="5">Fig. 98</figureCitation>
: green circles).
</paragraph>
</subSubSection>
</treatment>
</document>