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<document id="6B03F4015F11C286C3911A5FFAA2398A" ID-DOI="10.5281/zenodo.5341533" ID-ISSN="2345-7600" ID-Zenodo-Dep="5341533" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630190745178" checkinUser="diego" docAuthor="Cai, Yixiong &amp; Husana, Daniel Edison M." docDate="2009" docId="03FD8790FC3AFFA1FC2637A7B6C10C22" docLanguage="en" docName="57rbz0051-063.pdf" docOrigin="Raffles Bulletin of Zoology 57 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Edoneus atheatus Holthuis 1978" docType="treatment" docVersion="4" lastPageNumber="53" masterDocId="FFC4FFE8FC3BFFA3FF8B335BB2240E30" masterDocTitle="Cave Shrimps Of The Genus Edoneus Holthuis, 1978, From Luzon, The Philippines, With Descriptions Of Three New Species (Crustacea: Decapoda: Atyidae)" masterLastPageNumber="63" masterPageNumber="51" pageNumber="52" updateTime="1699181455100" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7B2B88137C1B1D203C60AC9661F61316">Cave Shrimps Of The Genus Edoneus Holthuis, 1978, From Luzon, The Philippines, With Descriptions Of Three New Species (Crustacea: Decapoda: Atyidae)</mods:title>
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<mods:namePart id="9D1004970DD6C898BA84D90E650F65C9">Husana, Daniel Edison M.</mods:namePart>
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<mods:title id="CEF03B3BD3D8B2C62AE37230FB4DDBD1">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="BB0F0F13D0D1B492FC74A4E86C505F42">2009</mods:date>
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<treatment id="03FD8790FC3AFFA1FC2637A7B6C10C22" ID-DOI="http://doi.org/10.5281/zenodo.5444122" ID-Zenodo-Dep="5444122" LSID="urn:lsid:plazi:treatment:03FD8790FC3AFFA1FC2637A7B6C10C22" httpUri="http://treatment.plazi.org/id/03FD8790FC3AFFA1FC2637A7B6C10C22" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
<subSubSection id="C34E650DFC3AFFA2FC2637A7B7060B24" box="[941,1314,1276,1300]" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="8BEB3686FC3AFFA2FC2637A7B7060B24" blockId="1.[941,1314,1276,1331]" box="[941,1314,1276,1300]" pageId="1" pageNumber="52">
<heading id="D0A381EAFC3AFFA2FC2637A7B7060B24" bold="true" box="[941,1314,1276,1300]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="52" reason="2">
<taxonomicName id="4C544D05FC3AFFA2FC2637A7B7060B24" ID-CoL="8VKJ4" authority="Holthuis, 1978" authorityName="Holthuis" authorityYear="1978" box="[941,1314,1276,1300]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="atheatus">
<emphasis id="B920EA94FC3AFFA2FC2637A7B7060B24" bold="true" box="[941,1314,1276,1300]" pageId="1" pageNumber="52">
<emphasis id="B920EA94FC3AFFA2FC2637A7B6570B24" bold="true" box="[941,1139,1276,1300]" italics="true" pageId="1" pageNumber="52">Edoneus atheatus</emphasis>
<bibRefCitation id="EFC54B77FC3AFFA2FBF737A7B7060B24" author="Holthuis, L" box="[1148,1314,1276,1300]" pageId="1" pageNumber="52" pagination="209 - 224" refId="ref7640" refString="Holthuis, L. B., 1978. Zoological results of the British Speleological Expedition to Papua New Guinea 1975. 7. Cavernicolous shrimps (Crustacea, Decapoda, Natantia) from New Ireland and the Philippines. Zoologische Mededelingen, 53: 209 - 224." type="journal article" year="1978">Holthuis, 1978</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E650DFC3AFFA2FBB43647B6B50B03" box="[1087,1169,1308,1331]" pageId="1" pageNumber="52" type="description">
<paragraph id="8BEB3686FC3AFFA2FBB43647B6B50B03" blockId="1.[941,1314,1276,1331]" box="[1087,1169,1308,1331]" pageId="1" pageNumber="52">
(
<figureCitation id="136F2A03FC3AFFA2FBCC3647B6A30B03" box="[1095,1159,1308,1331]" captionStart="Fig" captionStartId="2.[132,167,2057,2078]" captionTargetBox="[127,1460,625,2024]" captionTargetId="figure-8@2.[110,1488,602,2039]" captionTargetPageId="2" captionText="Fig. 1. Edoneus atheatus. A, telson; B, second maxilliped; C, fist pereiopod; D, second pereiopod; E, uropodal diaeresis. Scale bars: A, E = 0.1 mm; BD = 0.3 mm. (male, cl 2.9 mm, RMNH)." figureDoi="http://doi.org/10.5281/zenodo.5341535" httpUri="https://zenodo.org/record/5341535/files/figure.png" pageId="1" pageNumber="52">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E650DFC3AFFA2FCA33600B6270BBC" pageId="1" pageNumber="52" type="reference_group">
<paragraph id="8BEB3686FC3AFFA2FCA33600B6270BBC" blockId="1.[808,1447,1371,1420]" pageId="1" pageNumber="52">
<treatmentCitationGroup id="AB4411A8FC3AFFA2FCA33600B6270BBC" pageId="1" pageNumber="52">
<treatmentCitation id="0AF51097FC3AFFA2FCA33600B6E70B40" author="Holthuis, L" box="[808,1219,1371,1392]" page="220" pageId="1" pageNumber="52" year="1978">
<taxonomicName id="4C544D05FC3AFFA2FCA33600B6E70B40" ID-CoL="8VKJ4" authority="Holthuis, 1978: 220" authorityName="Holthuis" authorityPageNumber="220" authorityYear="1978" box="[808,1219,1371,1392]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="atheatus">
<emphasis id="B920EA94FC3AFFA2FCA33600B1C50B40" box="[808,993,1371,1392]" italics="true" pageId="1" pageNumber="52">Edoneus atheatus</emphasis>
<bibRefCitation id="EFC54B77FC3AFFA2FC603600B6E70B40" author="Holthuis, L" box="[1003,1219,1371,1392]" pageId="1" pageNumber="52" pagination="209 - 224" refId="ref7640" refString="Holthuis, L. B., 1978. Zoological results of the British Speleological Expedition to Papua New Guinea 1975. 7. Cavernicolous shrimps (Crustacea, Decapoda, Natantia) from New Ireland and the Philippines. Zoologische Mededelingen, 53: 209 - 224." type="journal article" year="1978">Holthuis, 1978: 220</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<figureCitation id="136F2A03FC3AFFA2FB5A3600B7380B40" box="[1233,1308,1371,1392]" captionStart="Fig" captionStartId="8.[132,167,2031,2052]" captionTargetBox="[151,1446,176,1989]" captionTargetId="figure-8@8.[141,1474,170,1997]" captionTargetPageId="8" captionText="Fig. 5. Edoneus sketi, new species: A, first pereiopod; B, second pereiopod; C, third pereiopod; D, dactylus of third pereiopod; E, fifth pereiopod; F, dactylus of fifth pereiopod; G, male first pleopod; H, male second pleopod; I, uropodal diaeresis. Scale bars: AC, E = 0.5 mm; D, F, I = 0.1 mm; G, H = 0.2 mm; (male, cl 4.1 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341543" httpUri="https://zenodo.org/record/5341543/files/figure.png" pageId="1" pageNumber="52">Figs. 5</figureCitation>
,
<figureCitation id="136F2A03FC3AFFA2FAA03600B71E0B40" box="[1323,1338,1371,1392]" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, FK = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="1" pageNumber="52">6</figureCitation>
; Balete &amp; Holthuis,1992: 99.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34E650DFC3AFFA2FCA336EAB646090C" pageId="1" pageNumber="52" type="materials_examined">
<paragraph id="8BEB3686FC3AFFA2FCA336EAB63F0866" blockId="1.[808,1448,1457,1622]" pageId="1" pageNumber="52">
<emphasis id="B920EA94FC3AFFA2FCA336EAB1D30BF6" bold="true" box="[808,1015,1457,1478]" italics="true" pageId="1" pageNumber="52">Material examined. </emphasis>
<materialsCitation id="3B3C3CDBFC3AFFA2FC7736EAB63F0866" accessionNumber="D 31898" collectionCode="RMNH" collectorName="Luzon &amp; Aglipay Municipality &amp; Barrio Palasian &amp; Sitio Disiluad &amp; Balete &amp; Holthuis &amp; F. Brandsma" country="Philippines" location="Maddela district" pageId="1" pageNumber="52" specimenCount="1" stateProvince="Isabela" typeStatus="holotype">
<typeStatus id="54EF8824FC3AFFA2FC7736EAB67D0BF6" box="[1020,1113,1457,1478]" pageId="1" pageNumber="52" type="holotype">Holotype</typeStatus>
: Male, cl
<quantity id="4CAC9B63FC3AFFA2FB3A36EAB6D80BF6" box="[1201,1276,1457,1478]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="52" unit="mm" value="4.6">4.6 mm</quantity>
,
<collectionCode id="ED45AE43FC3AFFA2FA8F36EAB7680BF6" box="[1284,1356,1457,1478]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="52" type="Museum">RMNH</collectionCode>
D 31898, the
<collectingCountry id="F3437616FC3AFFA2FCC43695B19B0BD3" box="[847,959,1486,1507]" name="Philippines" pageId="1" pageNumber="52">Philippines</collectingCountry>
, Luzon,
<collectingRegion id="4990F864FC3AFFA2FB933695B6980BD3" box="[1048,1212,1486,1507]" country="Philippines" name="Isabela" pageId="1" pageNumber="52">Isabela Province</collectingRegion>
<location id="8E8B605DFC3AFFA2FB4E3695B74D0BD3" LSID="urn:lsid:plazi:treatment:03FD8790FC3AFFA1FC2637A7B6C10C22:8E8B605DFC3AFFA2FB4E3695B74D0BD3" box="[1221,1385,1486,1507]" country="Philippines" name="Maddela district" pageId="1" pageNumber="52" stateProvince="Isabela">Maddela district</location>
, Cave near
<collectingRegion id="4990F864FC3AFFA2FCD036B0B1950830" box="[859,945,1515,1536]" country="Philippines" name="Santiago" pageId="1" pageNumber="52">Santiago</collectingRegion>
[recte: the
<collectingCountry id="F3437616FC3AFFA2FBA236B0B6BE0830" box="[1065,1178,1515,1536]" name="Philippines" pageId="1" pageNumber="52">Philippines</collectingCountry>
,
<collectorName id="26A15350FC3AFFA2FB2C36B0B6CC0830" box="[1191,1256,1515,1536]" pageId="1" pageNumber="52">Luzon</collectorName>
,
<collectingRegion id="4990F864FC3AFFA2FB7E36B0B7800830" box="[1269,1444,1515,1536]" country="Philippines" name="Quirino" pageId="1" pageNumber="52">Quirino Province</collectingRegion>
,
<collectorName id="26A15350FC3AFFA2FCA3355CB620082C" box="[808,1028,1543,1564]" pageId="1" pageNumber="52">Aglipay Municipality</collectorName>
,
<collectorName id="26A15350FC3AFFA2FB99355CB695082C" box="[1042,1201,1543,1564]" pageId="1" pageNumber="52">Barrio Palasian</collectorName>
, cave (no. 6) near
<collectorName id="26A15350FC3AFFA2FAF2355CB15A0809" pageId="1" pageNumber="52">Sitio Disiluad</collectorName>
, (see
<collectorName id="26A15350FC3AFFA2FC37357FB1DF0809" box="[956,1019,1572,1593]" pageId="1" pageNumber="52">Balete</collectorName>
&amp;
<collectorName id="26A15350FC3AFFA2FBAB357FB6530809" box="[1056,1143,1572,1593]" pageId="1" pageNumber="52">Holthuis</collectorName>
, 1992;
<bibRefCitation id="EFC54B77FC3AFFA2FB4D357FB7B10809" author="Fransen, C" box="[1222,1429,1572,1593]" pageId="1" pageNumber="52" pagination="1 - 334" refId="ref7507" refString="Fransen, C. H. J. M., L. B. Holthuis &amp; J. P. H. M. Adema, 1997. Type-catalogue of the decapod Crustacea in the collections of the Nationaal Natuurhistorisch Museum, with appendices of pre- 1900 collectors and material. Zoologische Verhandelingen, 311: 1 - 334." type="journal article" year="1997">Fransen et al., 1997</bibRefCitation>
)]; coll.
<collectorName id="26A15350FC3AFFA2FCD1351AB1FD0866" box="[858,985,1601,1622]" pageId="1" pageNumber="52">F. Brandsma</collectorName>
, 1974.
</materialsCitation>
</paragraph>
<paragraph id="8BEB3686FC3AFFA2FCA33520B1FC089C" blockId="1.[808,1447,1659,1708]" pageId="1" pageNumber="52">
<materialsCitation id="3B3C3CDBFC3AFFA2FCA33520B1F0089C" accessionNumber="D 31899" collectionCode="RMNH" collectorName="Luzon &amp; Aglipay Municipality &amp; Barrio Palasian &amp; Sitio Disiluad &amp; Balete &amp; Holthuis &amp; F. Brandsma" country="Philippines" location="Maddela district" pageId="1" pageNumber="52" specimenCount="1" stateProvince="Isabela" typeStatus="paratype">
<typeStatus id="54EF8824FC3AFFA2FCA33520B1AE08A0" box="[808,906,1659,1680]" pageId="1" pageNumber="52" type="paratype">Paratypes</typeStatus>
: Two females, cl
<quantity id="4CAC9B63FC3AFFA2FBCA3520B69F08A0" box="[1089,1211,1659,1680]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="4.9" metricValueMin="4.7" pageId="1" pageNumber="52" unit="mm" value="4.8" valueMax="4.9" valueMin="4.7">4.74.9 mm</quantity>
,
<collectionCode id="ED45AE43FC3AFFA2FB433520B73408A0" box="[1224,1296,1659,1680]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="52" type="Museum">RMNH</collectionCode>
D 31899, data same as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BEB3686FC3AFFA2FCA3358AB646090C" blockId="1.[808,1448,1745,1852]" pageId="1" pageNumber="52">
Other materials:
<materialsCitation id="3B3C3CDBFC3AFFA2FC47358AB67A090C" accessionNumber="D 41183" collectingDate="1985-12-30" collectionCode="RMNH" collectorName="D. S. Balete" country="Philippines" location="Sitio Disiluad. Cave" municipality="north Luzon" pageId="1" pageNumber="52" specimenCount="2" specimenCount-female="2" stateProvince="Quirino">
One male, cl
<quantity id="4CAC9B63FC3AFFA2FBDB358AB6BA08D6" box="[1104,1182,1745,1766]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="52" unit="mm" value="3.3">3.3 mm</quantity>
,
<specimenCount id="9D52FD0FFC3AFFA2FB22358AB72D08D6" box="[1193,1289,1745,1766]" pageId="1" pageNumber="52" type="female">2 females</specimenCount>
, cl
<quantity id="4CAC9B63FC3AFFA2FAA6358AB78008D6" box="[1325,1444,1745,1766]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="1" pageNumber="52" unit="mm" value="3.55" valueMax="3.6" valueMin="3.5">3.53.6 mm</quantity>
,
<collectionCode id="ED45AE43FC3AFFA2FCA335B5B1540933" box="[808,880,1774,1795]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="52" type="Museum">RMNH</collectionCode>
D 41183, the
<collectingCountry id="F3437616FC3AFFA2FC7635B5B6480933" box="[1021,1132,1774,1795]" name="Philippines" pageId="1" pageNumber="52">Philippines</collectingCountry>
,
<collectingMunicipality id="6B8FACFCFC3AFFA2FBFD35B5B6D40933" box="[1142,1264,1774,1795]" pageId="1" pageNumber="52">north Luzon</collectingMunicipality>
,
<collectingRegion id="4990F864FC3AFFA2FB7035B5B7800933" box="[1275,1444,1774,1795]" country="Philippines" name="Quirino" pageId="1" pageNumber="52">Quirino Province</collectingRegion>
, Aglipay Municipality, Barrio Palasian,
<location id="8E8B605DFC3AFFA2FB2D3450B74E0910" LSID="urn:lsid:plazi:treatment:03FD8790FC3AFFA1FC2637A7B6C10C22:8E8B605DFC3AFFA2FB2D3450B74E0910" box="[1190,1386,1803,1824]" country="Philippines" municipality="north Luzon" name="Sitio Disiluad. Cave" pageId="1" pageNumber="52" stateProvince="Quirino">Sitio Disiluad. Cave</location>
no. 6, coll.
<collectorName id="26A15350FC3AFFA2FCD1347CB1F0090C" box="[858,980,1831,1852]" pageId="1" pageNumber="52">D. S. Balete</collectorName>
,
<date id="FFEA1046FC3AFFA2FC54347CB67A090C" box="[991,1118,1831,1852]" pageId="1" pageNumber="52" value="1985-12-30">
<collectingDate id="EFAEE9AEFC3AFFA2FC54347CB67A090C" box="[991,1118,1831,1852]" pageId="1" pageNumber="52" value="1985-12-30">30 Dec.1985</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E650DFC3AFFA1FCA3343EB62E0EC2" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52" type="diagnosis">
<paragraph id="8BEB3686FC3AFFA1FCA3343EB35A0FC2" blockId="1.[808,1448,1893,2077]" lastBlockId="2.[132,771,187,498]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
<emphasis id="B920EA94FC3AFFA2FCA3343EB191094D" bold="true" box="[808,949,1893,1917]" italics="true" pageId="1" pageNumber="52">Diagnosis </emphasis>
Rostrum short, triangular in lateral view, unarmed, reaching nearly to end of eyes. Eyes small, unpigmented; pterygostomian margin broadly rounded; telson with a very tiny posteromedian projection; distal margin with intermediate pairs of setae much shorter than lateral pair of spines; carpus of first pereiopod moderately excavated anteriorly, twice as long as high; chela 1.7 times as long as broad; fingers as long as palm; carpus of second pereipod much slender than previous one, not excavated anteriorly, 6.2 times as long as high; chela 2.3 times as long as broad; fingers distinctly longer than palm; branchial formula incomplete, with 6 pairs of gills; no arthrobranch on pereiopods and maxillipeds; podobranch of second maxilliped lamina-shaped; uropod with diaeresis bearing about 1524 movable spinules, lateral angle much shorter than movable spinules.
</paragraph>
<paragraph id="8BEB3686FC39FFA1FCB533E0B62E0EC2" blockId="2.[830,1470,187,242]" pageId="2" pageNumber="53">
<emphasis id="B920EA94FC39FFA1FCB533E0B1890EE3" bold="true" box="[830,941,187,211]" italics="true" pageId="2" pageNumber="53">Habitat. </emphasis>
Collected from large flowing river inside a cave in northern Luzon.
</paragraph>
</subSubSection>
<subSubSection id="C34E650DFC39FFA1FCB53240B7810FE3" pageId="2" pageNumber="53" type="discussion">
<paragraph id="8BEB3686FC39FFA1FCB53240B7810FE3" blockId="2.[830,1470,283,467]" pageId="2" pageNumber="53">
<emphasis id="B920EA94FC39FFA1FCB53240B19B0F03" bold="true" box="[830,959,283,307]" italics="true" pageId="2" pageNumber="53">Remarks. </emphasis>
Found only from the
<typeStatus id="54EF8824FC39FFA1FB363240B6CF0F02" box="[1213,1259,283,306]" pageId="2" pageNumber="53">type</typeStatus>
locality (
<bibRefCitation id="EFC54B77FC39FFA1FAD33240B1520F62" author="Holthuis, L" pageId="2" pageNumber="53" pagination="209 - 224" refId="ref7640" refString="Holthuis, L. B., 1978. Zoological results of the British Speleological Expedition to Papua New Guinea 1975. 7. Cavernicolous shrimps (Crustacea, Decapoda, Natantia) from New Ireland and the Philippines. Zoologische Mededelingen, 53: 209 - 224." type="journal article" year="1978">Holthuis, 1978</bibRefCitation>
; Balete &amp; Holthuis, 1992).
<bibRefCitation id="EFC54B77FC39FFA1FB2F3260B76B0F62" author="Holthuis, L" box="[1188,1359,315,338]" pageId="2" pageNumber="53" pagination="209 - 224" refId="ref7640" refString="Holthuis, L. B., 1978. Zoological results of the British Speleological Expedition to Papua New Guinea 1975. 7. Cavernicolous shrimps (Crustacea, Decapoda, Natantia) from New Ireland and the Philippines. Zoologische Mededelingen, 53: 209 - 224." type="journal article" year="1978">Holthuis (1978)</bibRefCitation>
described the new genus and new species
<taxonomicName id="4C544D05FC39FFA1FB093207B71D0F43" authorityName="Holthuis" authorityYear="1978" box="[1154,1337,348,371]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="atheatus">
<emphasis id="B920EA94FC39FFA1FB093207B71D0F43" box="[1154,1337,348,371]" italics="true" pageId="2" pageNumber="53">Edoneus atheatus</emphasis>
</taxonomicName>
, on the basis of
<specimenCount id="9D52FD0FFC39FFA1FCD03220B62D0FA2" box="[859,1033,379,402]" pageId="2" pageNumber="53" type="generic">three specimens</specimenCount>
. Balete &amp; Holthuis (1992) reported some more specimens and provided more detailed information on the
<typeStatus id="54EF8824FC39FFA1FCE332E0B1B20FE2" box="[872,918,443,466]" pageId="2" pageNumber="53">type</typeStatus>
locality, as well as detailed habitat information.
</paragraph>
</subSubSection>
<subSubSection id="C34E650DFC39FFA1FCB532A0B6C10C22" box="[830,1253,507,531]" pageId="2" pageNumber="53" type="distribution">
<paragraph id="8BEB3686FC39FFA1FCB532A0B6C10C22" blockId="2.[830,1253,507,531]" box="[830,1253,507,531]" pageId="2" pageNumber="53">
<emphasis id="B920EA94FC39FFA1FCB532A0B1C40C23" bold="true" box="[830,992,507,531]" italics="true" pageId="2" pageNumber="53">Distribution. </emphasis>
Luzon, the
<collectingCountry id="F3437616FC39FFA1FBED32A0B6C60C22" box="[1126,1250,507,530]" name="Philippines" pageId="2" pageNumber="53">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>