142 lines
17 KiB
XML
142 lines
17 KiB
XML
<document id="538B39FAD19A2F41B7F0069C3C310A3E" ID-DOI="10.11646/zootaxa.5367.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10145106" ID-ZooBank="5A1E5321-D2BA-4B92-BA23-A7C1CDBA5723" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700167363318" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2023" docId="03FD878BFFFEFFDFFCBEFD8758953A8A" docLanguage="en" docName="zootaxa.5367.1.1.pdf" docOrigin="Zootaxa 5367 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena (Hydraenopsis) panelephanta Perkins 2023, new species" docType="treatment" docVersion="2" lastPageNumber="47" masterDocId="FFC4FFF3FFD0FFF1FC29FFA5596F3C37" masterDocTitle="New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)" masterLastPageNumber="86" masterPageNumber="1" pageNumber="47" updateTime="1700168056660" updateUser="ExternalLinkService">
|
||
<mods:mods id="6E9B1AD6F5C0E216C0012807D88C285E" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="9CA4EC7E5C813E75BD48A0B19EBCCA4E">
|
||
<mods:title id="14DE92C774E03329A418E9EAA8F60B54">New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="41DD5ACC5E23DC429798D62860B06C0B" type="personal">
|
||
<mods:role id="42060A066D0849E841FDE43935ED5198">
|
||
<mods:roleTerm id="11DE297891F42F3A573405A3F0547D32">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="31E0286D3CB34F882DD353642DB24ABD">Perkins, Philip D.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="4FFA0BBF9A9D5B373482999CA5DAD484">text</mods:typeOfResource>
|
||
<mods:relatedItem id="BAD45FA91423F105EC2D77C718221162" type="host">
|
||
<mods:titleInfo id="2B7A0B90BF30A395F9B110925AE0A1DD">
|
||
<mods:title id="C020785213D3366FD3D8E980E19E4800">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="D90532F6911BC0D68CD4A7CB3301F501">
|
||
<mods:date id="E6B5D6DC282A082A388CCD59337CEEF2">2023</mods:date>
|
||
<mods:detail id="1F3293C833D92EBA7B69759FDE64D01A" type="pubDate">
|
||
<mods:number id="7DBA15547B75C726DF7F57C402F308AD">2023-11-07</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="673CEAB93F7477D3A838006CDF4C1AF2" type="volume">
|
||
<mods:number id="CBB1CEF497249D04FEDC4AF2FF960131">5367</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A2DC5AB376BDC3897CA19F079BB60BD6" type="issue">
|
||
<mods:number id="3140BA918D486938491C5CEFCA4B0DAD">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="983FFD8ECA522B8C3E432FD343ECDB33" unit="page">
|
||
<mods:start id="D2A470D1C35A7018C549B2E71DC7F94E">1</mods:start>
|
||
<mods:end id="652D471261659DC36A15F361D72CD6E9">86</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="B8BD7B42374130E1751DDE6C51951451">
|
||
<mods:url id="4A4B7755A1454FAB35D497A9CF482A4D">https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="9AC7893B57A407B00A0A09DBE0953A3F">journal article</mods:classification>
|
||
<mods:identifier id="0F7F996A6CAC082C98DFC0D73C4EEFE1" type="DOI">10.11646/zootaxa.5367.1.1</mods:identifier>
|
||
<mods:identifier id="C1B6E795C87380732A2A40D2DC0A245B" type="ISSN">1175-5334</mods:identifier>
|
||
<mods:identifier id="9282CB6213D3C4DA7D779CA8593661C3" type="Zenodo-Dep">10145106</mods:identifier>
|
||
<mods:identifier id="B62D983BACC07FC4060501B774257D50" type="ZooBank">5A1E5321-D2BA-4B92-BA23-A7C1CDBA5723</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03FD878BFFFEFFDFFCBEFD8758953A8A" LSID="urn:lsid:plazi:treatment:03FD878BFFFEFFDFFCBEFD8758953A8A" httpUri="http://treatment.plazi.org/id/03FD878BFFFEFFDFFCBEFD8758953A8A" lastPageNumber="47" pageId="46" pageNumber="47">
|
||
<subSubSection id="C34E6516FFFEFFDFFCBEFD875A743E0B" box="[151,795,546,573]" pageId="46" pageNumber="47" type="nomenclature">
|
||
<paragraph id="8BEB369DFFFEFFDFFCBEFD875A743E0B" blockId="46.[151,795,546,609]" box="[151,795,546,573]" pageId="46" pageNumber="47">
|
||
<heading id="D0A381F1FFFEFFDFFCBEFD875A743E0B" bold="true" box="[151,795,546,573]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="4">
|
||
<emphasis id="B920EA8FFFFEFFDFFCBEFD875A743E0B" bold="true" box="[151,795,546,573]" pageId="46" pageNumber="47">
|
||
<taxonomicName id="4C544D1EFFFEFFDFFCBEFD875B123E0A" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[151,637,546,573]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="panelephanta" status="sp. nov." subGenus="Hydraenopsis">
|
||
<emphasis id="B920EA8FFFFEFFDFFCBEFD875B123E0A" bold="true" box="[151,637,546,573]" italics="true" pageId="46" pageNumber="47">Hydraena (Hydraenopsis) panelephanta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A21357F4FFFEFFDFFEA2FD865A743E0B" box="[651,795,546,573]" pageId="46" pageNumber="47" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E6516FFFEFFDFFCBEFDE25A653E56" box="[151,778,583,609]" pageId="46" pageNumber="47" type="description">
|
||
<paragraph id="8BEB369DFFFEFFDFFCBEFDE25A653E56" blockId="46.[151,795,546,609]" box="[151,778,583,609]" pageId="46" pageNumber="47">
|
||
<figureCitation id="136F2A18FFFEFFDFFCBEFDE2598C3E56" box="[151,227,583,609]" captionStart="FIGURE 2" captionStartId="6.[151,250,1867,1891]" captionTargetBox="[228,1360,181,1844]" captionTargetId="figure-35@6.[228,1360,181,1844]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.10145110" httpUri="https://zenodo.org/record/10145110/files/figure.png" pageId="46" pageNumber="47">Figs. 2</figureCitation>
|
||
(habitus comparison), 27 (habitus and aedeagus)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E6516FFFEFFDFFCBEFD2A581F3EFA" pageId="46" pageNumber="47" type="materials_examined">
|
||
<paragraph id="8BEB369DFFFEFFDFFCBEFD2A581F3EFA" blockId="46.[151,1437,654,1725]" pageId="46" pageNumber="47">
|
||
<emphasis id="B920EA8FFFFEFFDFFCBEFD2A58243E9F" bold="true" box="[151,331,654,681]" pageId="46" pageNumber="47">Type Material.</emphasis>
|
||
<materialsCitation id="3B3C3CC0FFFEFFDFFD7FFD2A581F3EFA" collectingDate="1975-03-01" collectionCode="MCZ" collectorName="J. F. Lawrence" country="Panama" location="Canal Zone" pageId="46" pageNumber="47" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B920EA8FFFFEFFDFFD7FFD2A58AC3E9E" bold="true" box="[342,451,655,681]" pageId="46" pageNumber="47">
|
||
<typeStatus id="54EF883FFFFEFFDFFD7FFD2A58AC3E9E" box="[342,451,655,681]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(male): “
|
||
<collectingCountry id="F343760DFFFEFFDFFE1BFD2A5BFE3E9E" box="[562,657,655,681]" name="Panama" pageId="46" pageNumber="47">Panama</collectingCountry>
|
||
:
|
||
<location id="8E8B6046FFFEFFDFFEB6FD2A5A463E9E" LSID="urn:lsid:plazi:treatment:03FD878BFFFEFFDFFCBEFD8758953A8A:8E8B6046FFFEFFDFFEB6FD2A5A463E9E" box="[671,809,655,681]" country="Panama" name="Canal Zone" pageId="46" pageNumber="47">Canal Zone</location>
|
||
:
|
||
<locationDeviation id="9A592FBFFFFEFFDFFF1EFD2A5D683E9E" box="[823,1031,655,681]" location="Gatun" pageId="46" pageNumber="47">
|
||
<quantity id="4CAC9B78FFFEFFDFFF1EFD2A5AED3E9E" box="[823,898,655,681]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="46" pageNumber="47" unit="mi" value="10.0">10 mi.</quantity>
|
||
SW Gatun
|
||
</locationDeviation>
|
||
,
|
||
<date id="FFEA105DFFFEFFDFF83FFD2B5DEA3E9E" box="[1046,1157,654,681]" pageId="46" pageNumber="47" value="1975-03-01">
|
||
<collectingDate id="EFAEE9B5FFFEFFDFF83FFD2B5DEA3E9E" box="[1046,1157,654,681]" pageId="46" pageNumber="47" value="1975-03-01">1.III.1975</collectingDate>
|
||
</date>
|
||
// litter by stream,
|
||
<collectorName id="26A1534BFFFEFFDFF945FD2A58673EFA" pageId="46" pageNumber="47">J. F. Lawrence</collectorName>
|
||
” (
|
||
<collectionCode id="ED45AE58FFFEFFDFFD08FD16580B3EFA" box="[289,356,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="46" pageNumber="47" type="Museum">MCZ</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E6516FFFEFFDFFCEEFD735B463F92" pageId="46" pageNumber="47" type="diagnosis">
|
||
<paragraph id="8BEB369DFFFEFFDFFCEEFD735B463F92" blockId="46.[151,1437,654,1725]" pageId="46" pageNumber="47">
|
||
<emphasis id="B920EA8FFFFEFFDFFCEEFD7358BB3EC7" bold="true" box="[199,468,726,752]" pageId="46" pageNumber="47">Differential Diagnosis.</emphasis>
|
||
This new species, which is a member of the
|
||
<taxonomicName id="4C544D1EFFFEFFDFFFF5FD725C653EC6" authority="Subgroup" authorityName="Subgroup" box="[988,1290,727,753]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="marginicollis">
|
||
<emphasis id="B920EA8FFFFEFFDFFFF5FD725DFB3EC6" box="[988,1172,727,753]" italics="true" pageId="46" pageNumber="47">H. marginicollis</emphasis>
|
||
Subgroup
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFC54B6CFFFEFFDFF961FD7259923F22" author="Perkins, P. D." pageId="46" pageNumber="47" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins 1980: 69</bibRefCitation>
|
||
), can be differentiated from others by the combination of the small body size (ca.
|
||
<quantity id="4CAC9B78FFFEFFDFF8A5FD5E5D9B3F22" box="[1164,1268,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="46" pageNumber="47" unit="mm" value="1.3">1.30 mm</quantity>
|
||
), the pronotum with distinctive dark brown macula surrounded by wide testaceous margins, the pronotum very finely very sparsely punctulate (interstices on disc ca. 3–6xpd or slightly larger), and the aedeagus (
|
||
<figureCitation id="136F2A18FFFEFFDFF82DFCE65D373F6A" box="[1028,1112,835,861]" captionStart="FIGURE 27" captionStartId="47.[151,250,1792,1816]" captionTargetBox="[291,1295,181,1768]" captionTargetId="figure-18@47.[291,1295,181,1768]" captionTargetPageId="47" captionText="FIGURE 27. Hydraena (Hydraenopsis) panelephanta, new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145158" httpUri="https://zenodo.org/record/10145158/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
|
||
). Based on the aedeagus, this species appears to be a distant relative of
|
||
<taxonomicName id="4C544D1EFFFEFFDFFE41FCC25A703FB6" box="[616,799,871,897]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="marginicollis">
|
||
<emphasis id="B920EA8FFFFEFFDFFE41FCC25A703FB6" box="[616,799,871,897]" italics="true" pageId="46" pageNumber="47">H. marginicollis</emphasis>
|
||
</taxonomicName>
|
||
, and others with similar basic aedeagal morphology (see figures 53 and
|
||
<quantity id="4CAC9B78FFFEFFDFFD17FC2F58183F92" box="[318,375,906,933]" metricMagnitude="0" metricUnit="m" metricValue="1.3969999999999998" pageId="46" pageNumber="47" unit="in" value="55.0">55 in</quantity>
|
||
<bibRefCitation id="EFC54B6CFFFEFFDFFD57FC2E5B733F92" author="Perkins, P. D." box="[382,540,907,933]" pageId="46" pageNumber="47" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins, 1980</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E6516FFFEFFDFFCEEFC0B59B23A66" pageId="46" pageNumber="47" type="description">
|
||
<paragraph id="8BEB369DFFFEFFDFFCEEFC0B5C2438F2" blockId="46.[151,1437,654,1725]" pageId="46" pageNumber="47">
|
||
<emphasis id="B920EA8FFFFEFFDFFCEEFC0B583E3FFF" bold="true" box="[199,337,942,968]" pageId="46" pageNumber="47">Description</emphasis>
|
||
. Size:
|
||
<typeStatus id="54EF883FFFFEFFDFFD8DFC0A5B693FFE" box="[420,518,943,969]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
(length/width, mm): body (length to elytral apices) 1.30/0.56; head width 0.34; pronotum 0.32/0.44, PA 0.37, PB 0.40; elytra 0.82/0.56. Habitus as illustrated (
|
||
<figureCitation id="136F2A18FFFEFFDFF80AFC765D163FDA" box="[1059,1145,979,1005]" captionStart="FIGURE 27" captionStartId="47.[151,250,1792,1816]" captionTargetBox="[291,1295,181,1768]" captionTargetId="figure-18@47.[291,1295,181,1768]" captionTargetPageId="47" captionText="FIGURE 27. Hydraena (Hydraenopsis) panelephanta, new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145158" httpUri="https://zenodo.org/record/10145158/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
|
||
). Dorsum with head dark brown to black, pronotum with distinctive dark brown macula surrounded by wide testaceous margins, elytra brown, palpi testaceous, tip of last palpomere not darker. Dorsum of head and pronotum very finely very sparsely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices strongly shining; punctures of frons disc and pronotal disc ca. 1xef, interstices on pronotal disc ca. 3–6xpd or slightly larger; punctures of clypeus very fine, lateral areas microreticulate. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal fovea shallow, other usual foveae absent.
|
||
</paragraph>
|
||
<paragraph id="8BEB369DFFFEFFDFFCEEFB6A5DAE3962" blockId="46.[151,1437,654,1725]" pageId="46" pageNumber="47">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
|
||
<paragraph id="8BEB369DFFFEFFDFFCEEFAFA59B23A66" blockId="46.[151,1437,654,1725]" pageId="46" pageNumber="47">Venter:Mentum moderately densely very finely punctulate,each puncture with moderately long seta; postmentum very finely densely micropunctulate. Genae raised, dull, without posterior ridge, with shallow midlongitudinal impression. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/3/3. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques moderately narrow, straight, parallel, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 2.5x P2. All tibiae slender, straight. Abdominal apex symmetrical; last tergite with apicomedian notch.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E6516FFFEFFDFFCEEF9FF5A323A42" box="[199,861,1626,1653]" pageId="46" pageNumber="47" type="distribution">
|
||
<paragraph id="8BEB369DFFFEFFDFFCEEF9FF5A323A42" blockId="46.[151,1437,654,1725]" box="[199,861,1626,1653]" pageId="46" pageNumber="47">
|
||
<emphasis id="B920EA8FFFFEFFDFFCEEF9FF580E3A43" bold="true" box="[199,353,1626,1652]" pageId="46" pageNumber="47">Distribution.</emphasis>
|
||
Currently known only from the
|
||
<typeStatus id="54EF883FFFFEFFDFFEE5F9FE5B933A42" box="[716,764,1627,1653]" pageId="46" pageNumber="47">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E6516FFFEFFDFFCEEF9DA58953A8A" pageId="46" pageNumber="47" type="etymology">
|
||
<paragraph id="8BEB369DFFFEFFDFFCEEF9DA58953A8A" blockId="46.[151,1437,654,1725]" pageId="46" pageNumber="47">
|
||
<emphasis id="B920EA8FFFFEFFDFFCEEF9DA58213AAE" bold="true" box="[199,334,1663,1689]" pageId="46" pageNumber="47">Etymology.</emphasis>
|
||
Named in reference to the known geographical distribution and the shape of the aedeagus, with a long gonopore-bearing process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |