treatments-xml/data/03/FD/87/03FD878AA269FFD95B45FCCB8A1B2F1C.xml

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<mods:title id="B8B027E23C8726E2AF2F7CE9580898FA">Two new troglobitic cockroach species of the genus Spelaeoblatta (Blattaria: Nocticolidae) from North Thailand</mods:title>
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<mods:namePart id="CC41F299505289BACAC1FB8313375CA2">Vidlička, Ľubomír</mods:namePart>
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<mods:namePart id="90FA722C4D4D4A30C4D5735B78669104">Vršanský, Peter</mods:namePart>
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<mods:title id="8BEA6CC482DB3163D71F5F40BFC0DDA0">Journal of Natural History</mods:title>
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<paragraph id="8BEB369CA269FFDA5B45FCCB8DAD2A70" blockId="5.[614,967,827,884]" box="[614,967,827,852]" pageId="5" pageNumber="111">
<heading id="D0A381F0A269FFDA5B45FCCB8DAD2A70" box="[614,967,827,852]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="111" reason="3">
<taxonomicName id="4C544D1FA269FFDA5B45FCCB8DEA2A77" ID-CoL="6Z93B" authority="Vidlička &amp; Vršanský &amp; Shcherbakov, 2003" authorityName="Vidlička &amp; Vršanský &amp; Shcherbakov" authorityYear="2003" box="[614,896,827,851]" class="Insecta" family="Nocticolidae" genus="Spelaeoblatta" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="111" phylum="Arthropoda" rank="species" species="thailandica" status="sp. nov.">
<emphasis id="B920EA8EA269FFDA5B45FCCB8DEA2A77" bold="true" box="[614,896,827,851]" italics="true" pageId="5" pageNumber="111">Spelaeoblatta thailandica</emphasis>
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<paragraph id="8BEB369CA269FFDA5BEBFCAC8D0C2A50" blockId="5.[614,967,827,884]" box="[712,870,860,884]" pageId="5" pageNumber="111">
(
<figureCitation id="136F2A19A269FFDA5BECFCAC8D492A50" box="[719,803,860,884]" captionStart="FIG" captionStartId="6.[313,329,978,999]" captionTargetBox="[314,1268,179,950]" captionTargetId="figure-10@6.[313,1268,177,951]" captionTargetPageId="6" captionText="FIG. 5. Spelaeoblatta thailandica, male. (A) Habitus; (B) abdomen (lateral view); (C) subgenital plate (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.4653258" httpUri="https://zenodo.org/record/4653258/files/figure.png" pageId="5" pageNumber="111">figure 5</figureCitation>
A–C)
</paragraph>
<paragraph id="8BEB369CA269FFDA5842FC7F8A9E2A82" blockId="5.[313,1268,910,1029]" box="[353,1268,910,934]" pageId="5" pageNumber="111">
<emphasis id="B920EA8EA269FFDA5842FC7F8F952A82" box="[353,511,910,934]" italics="true" pageId="5" pageNumber="111">Type material</emphasis>
. 
<materialsCitation id="3B3C3CC1A269FFDA5B33FC7E8A9E2A82" box="[528,1268,910,934]" country="Thailand" location="Red Cliff Cave" pageId="5" pageNumber="111" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EF883EA269FFDA5B33FC7E8CE02A82" box="[528,650,910,934]" pageId="5" pageNumber="111" type="holotype">HOLOTYPE</typeStatus>
: 
<emphasis id="B920EA8EA269FFDA5BB5FC7F8CC22A82" bold="true" box="[662,680,911,934]" pageId="5" pageNumber="111">³</emphasis>
, north-west 
<collectingCountry id="F343760CA269FFDA5A1EFC7E8DCD2A82" box="[829,935,910,934]" name="Thailand" pageId="5" pageNumber="111">Thailand</collectingCountry>
: 
<location id="8E8B6047A269FFDA5A97FC7E8A0B2A82" LSID="urn:lsid:plazi:treatment:03FD878AA269FFD95B45FCCB8A1B2F1C:8E8B6047A269FFDA5A97FC7E8A0B2A82" box="[948,1121,910,934]" country="Thailand" name="Red Cliff Cave" pageId="5" pageNumber="111">Red Cliff Cave</location>
(
<collectingRegion id="4990F87EA269FFDA5D54FC7E8A9E2A82" box="[1143,1268,910,934]" country="Thailand" name="Mae Hong Son" pageId="5" pageNumber="111">Mae Hong</collectingRegion>
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</paragraph>
<paragraph id="8BEB369CA269FFDA581AFC5E8A9E2AE2" blockId="5.[313,1268,910,1029]" box="[313,1268,942,966]" pageId="5" pageNumber="111">
Son Province, Nam Lang region, 19°36∞N, 98°04∞E), 
<date id="FFEA105CA269FFDA5A96FC5E8A012AE2" box="[949,1131,942,966]" pageId="5" pageNumber="111" value="1997-03-24">24 March 1997</date>
, N. Myuge
</paragraph>
<paragraph id="8BEB369CA269FFDA581AFC3E8A752D21" blockId="5.[313,1268,910,1029]" pageId="5" pageNumber="111">
(H. Miore) leg., coll. SNM-NHMB. 
<emphasis id="B920EA8EA269FFDA5842FC1D8FB32D21" box="[353,473,1005,1029]" italics="true" pageId="5" pageNumber="111">Etymology</emphasis>
. The specific name refers to the country of origin.
</paragraph>
<paragraph id="8BEB369CA269FFDA581AFBC38FD22D6E" blockId="5.[313,1269,1075,1736]" box="[313,440,1075,1098]" pageId="5" pageNumber="111">
<emphasis id="B920EA8EA269FFDA581AFBC38FD22D6E" box="[313,440,1075,1098]" italics="true" pageId="5" pageNumber="111">Description</emphasis>
</paragraph>
<paragraph id="8BEB369CA269FFDA5842FBA28FB92DAE" blockId="5.[313,1269,1075,1736]" pageId="5" pageNumber="111">
<emphasis id="B920EA8EA269FFDA5842FBA28FE52D4D" box="[353,399,1106,1129]" italics="true" pageId="5" pageNumber="111">Size</emphasis>
(mm). Body length: 
<emphasis id="B920EA8EA269FFDA5BA1FBA38CFE2D4E" bold="true" box="[642,660,1107,1130]" pageId="5" pageNumber="111">³</emphasis>
8.75; pronotum length×width, 
<emphasis id="B920EA8EA269FFDA5D2BFBA38A702D4E" bold="true" box="[1032,1050,1107,1130]" pageId="5" pageNumber="111">³</emphasis>
2.4×3.4; forewing length, 
<emphasis id="B920EA8EA269FFDA58ACFB838FCB2DAE" bold="true" box="[399,417,1139,1162]" pageId="5" pageNumber="111">³</emphasis>
5.0.
</paragraph>
<paragraph id="8BEB369CA269FFDA5842FB628CAF2FAC" blockId="5.[313,1269,1075,1736]" pageId="5" pageNumber="111">
<emphasis id="B920EA8EA269FFDA5842FB628FF12D8E" box="[353,411,1170,1194]" italics="true" pageId="5" pageNumber="111">Male</emphasis>
(
<figureCitation id="136F2A19A269FFDA588DFB628C692D8E" box="[430,515,1170,1194]" captionStart="FIG" captionStartId="6.[313,329,978,999]" captionTargetBox="[314,1268,179,950]" captionTargetId="figure-10@6.[313,1268,177,951]" captionTargetPageId="6" captionText="FIG. 5. Spelaeoblatta thailandica, male. (A) Habitus; (B) abdomen (lateral view); (C) subgenital plate (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.4653258" httpUri="https://zenodo.org/record/4653258/files/figure.png" pageId="5" pageNumber="111">figure 5</figureCitation>
A–C). Head longitudinal, oval, partly visible from above; compound eyes strongly reduced but present; ocelli absent; apex of fifth maxillary palpomere rounded. Antennae long, slender, about 60 segments, scapus and third segment long, other segments markedly shorter. Pronotum unicoloured, parabolic, the middle of hind margin straight, lateral corners obliquely truncated. Tegmina reaching to the end of fifth tergite, obliquely truncate, surface covered by rather sparse trichia, especially on apical part of venation. Hind wings reduced, only lobate, reaching to a half of the metanotum length. Ten abdominal terga visible. The posterior margin of T2 medially concave. T3 has a medial, inverted Y-like glandular fossa, with hind margin medially elongated and elevated. T4 with a small medial indentation, and T4–T7 with collar-like elevations on the hind margins. Hind margin of supraanal plate is oval with medial shallow depression. Cerci are composed of 10 segments, surface dorsally smooth, ventrally with long setae, apically with long sensilla. Subgenital plate symmetrical with two similar setose styles, interstylar margin plane contains small medial elevation (
<figureCitation id="136F2A19A269FFDA5A77F9A08DD72F4C" box="[852,957,1616,1640]" captionStart="FIG" captionStartId="6.[313,329,978,999]" captionTargetBox="[314,1268,179,950]" captionTargetId="figure-10@6.[313,1268,177,951]" captionTargetPageId="6" captionText="FIG. 5. Spelaeoblatta thailandica, male. (A) Habitus; (B) abdomen (lateral view); (C) subgenital plate (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.4653258" httpUri="https://zenodo.org/record/4653258/files/figure.png" pageId="5" pageNumber="111">figure 5C</figureCitation>
). The genital hook on the left side, the apex strongly curved.
</paragraph>
<paragraph id="8BEB369CA269FFDA5842F9608A3A2F8C" blockId="5.[313,1269,1075,1736]" box="[353,1104,1680,1704]" pageId="5" pageNumber="111">Colour: the specimen is pale brownish yellow, nearly translucent.</paragraph>
<paragraph id="8BEB369CA269FFD95842F9408A3E2DCD" blockId="5.[313,1269,1075,1736]" lastBlockId="6.[313,1269,1122,1305]" lastPageId="6" lastPageNumber="112" pageId="5" pageNumber="111">
<emphasis id="B920EA8EA269FFDA5842F9408F9B2FEC" box="[353,497,1712,1736]" italics="true" pageId="5" pageNumber="111">Male nymph</emphasis>
. Size (mm): body length: 6.9; pronotum length×width, 1.9×2.5; front femur length, 1.9. Head with compound eyes having approximately 100 separated ommatidia that become smaller towards the antennae. Wing pads on meso- and metanotum markedly long. Anteroventral margin of front femur with three large proximal spines followed by a row of piliform spinules (
<typeStatus id="54EF883EA26AFFD95ADEFB318A5A2DFD" box="[1021,1072,1217,1241]" pageId="6" pageNumber="112">type</typeStatus>
B); tarsal claws 
<emphasis id="B920EA8EA26AFFD95D69FB218A3E2DCD" bold="true" box="[1098,1108,1233,1257]" pageId="6" pageNumber="112">1</emphasis>
</paragraph>
<caption id="DF2B6614A26AFFD9581AFC228DA32D26" ID-DOI="http://doi.org/10.5281/zenodo.4653258" ID-Zenodo-Dep="4653258" httpUri="https://zenodo.org/record/4653258/files/figure.png" pageId="6" pageNumber="112" startId="6.[313,329,978,999]" targetBox="[314,1268,179,950]" targetPageId="6">
<paragraph id="8BEB369CA26AFFD9581AFC228DA32D26" blockId="6.[313,1269,977,1026]" pageId="6" pageNumber="112">
FIG. 5. 
<taxonomicName id="4C544D1FA26AFFD958B6FC228CF42AC3" authorityName="Vidlička &amp; Vršanský &amp; Shcherbakov" authorityYear="2003" box="[405,670,978,999]" class="Insecta" family="Nocticolidae" genus="Spelaeoblatta" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="112" phylum="Arthropoda" rank="species" species="thailandica">
<emphasis id="B920EA8EA26AFFD958B6FC228CF42AC3" box="[405,670,978,999]" italics="true" pageId="6" pageNumber="112">Spelaeoblatta thailandica</emphasis>
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, male. (A) Habitus; (B) abdomen (lateral view); (C) subgenital plate (dorsal view).
</paragraph>
</caption>
<paragraph id="8BEB369CA26AFFD9581AFB118D522C3D" blockId="6.[313,1269,1122,1305]" pageId="6" pageNumber="112">simple, symmetrical, pulvilli and arolia absent. T3 medially elongated as in the adult male. Subgenital plate with two small styles.</paragraph>
<paragraph id="8BEB369CA26AFFD9581AFAB18FF62C7D" blockId="6.[313,1269,1345,1592]" box="[313,412,1345,1369]" pageId="6" pageNumber="112">
<emphasis id="B920EA8EA26AFFD9581AFAB18FF62C7D" box="[313,412,1345,1369]" italics="true" pageId="6" pageNumber="112">Remarks</emphasis>
</paragraph>
<paragraph id="8BEB369CA26AFFD95842FA918A1B2F1C" blockId="6.[313,1269,1345,1592]" pageId="6" pageNumber="112">
The species 
<taxonomicName id="4C544D1FA26AFFD958CFFA918CEF2C5D" authorityName="Vidlička &amp; Vršanský &amp; Shcherbakov" authorityYear="2003" box="[492,645,1377,1401]" class="Insecta" family="Nocticolidae" genus="Spelaeoblatta" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="112" phylum="Arthropoda" rank="species" species="thailandica">
<emphasis id="B920EA8EA26AFFD958CFFA918CEF2C5D" box="[492,645,1377,1401]" italics="true" pageId="6" pageNumber="112">S. thailandica</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C544D1FA26AFFD95BE2FA918D462C5D" authorityName="Bolivar" authorityYear="1897" box="[705,812,1377,1401]" class="Insecta" family="Nocticolidae" genus="Spelaeoblatta" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="112" phylum="Arthropoda" rank="species" species="gestroi">
<emphasis id="B920EA8EA26AFFD95BE2FA918D462C5D" box="[705,812,1377,1401]" italics="true" pageId="6" pageNumber="112">S. gestroi</emphasis>
</taxonomicName>
were only described from a single male and a single female, respectively. So far, their opposite sexes are not known. Until the male of 
<taxonomicName id="4C544D1FA26AFFD958EBFA518C5E2C9C" authorityName="Bolivar" authorityYear="1897" box="[456,564,1441,1464]" class="Insecta" family="Nocticolidae" genus="Spelaeoblatta" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="112" phylum="Arthropoda" rank="species" species="gestroi">
<emphasis id="B920EA8EA26AFFD958EBFA518C5E2C9C" box="[456,564,1441,1464]" italics="true" pageId="6" pageNumber="112">S. gestroi</emphasis>
</taxonomicName>
is found, we cannot exclude a possibility that 
<taxonomicName id="4C544D1FA26AFFD95D7AFA518A9F2C9D" authorityName="Vidlička &amp; Vršanský &amp; Shcherbakov" authorityYear="2003" box="[1113,1269,1441,1465]" class="Insecta" family="Nocticolidae" genus="Spelaeoblatta" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="112" phylum="Arthropoda" rank="species" species="thailandica">
<emphasis id="B920EA8EA26AFFD95D7AFA518A9F2C9D" box="[1113,1269,1441,1465]" italics="true" pageId="6" pageNumber="112">S. thailandica</emphasis>
</taxonomicName>
is a male of 
<taxonomicName id="4C544D1FA26AFFD958F3FA318C572CFC" authorityName="Bolivar" authorityYear="1897" box="[464,573,1473,1496]" class="Insecta" family="Nocticolidae" genus="Spelaeoblatta" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="112" phylum="Arthropoda" rank="species" species="gestroi">
<emphasis id="B920EA8EA26AFFD958F3FA318C572CFC" box="[464,573,1473,1496]" italics="true" pageId="6" pageNumber="112">S. gestroi</emphasis>
</taxonomicName>
. However, 
<taxonomicName id="4C544D1FA26AFFD95BE5FA318D592CFC" authorityName="Bolivar" authorityYear="1897" box="[710,819,1473,1496]" class="Insecta" family="Nocticolidae" genus="Spelaeoblatta" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="112" phylum="Arthropoda" rank="species" species="gestroi">
<emphasis id="B920EA8EA26AFFD95BE5FA318D592CFC" box="[710,819,1473,1496]" italics="true" pageId="6" pageNumber="112">S. gestroi</emphasis>
</taxonomicName>
was captured about 
<quantity id="4CAC9B79A26AFFD95D0DFA318A142CFD" box="[1070,1150,1473,1497]" metricMagnitude="5" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="112" unit="km" value="170.0">170 km</quantity>
SW from the locality inhabited by 
<taxonomicName id="4C544D1FA26AFFD95B71FA118C812CDC" authorityName="Vidlička &amp; Vršanský &amp; Shcherbakov" authorityYear="2003" box="[594,747,1504,1528]" class="Insecta" family="Nocticolidae" genus="Spelaeoblatta" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="112" phylum="Arthropoda" rank="species" species="thailandica">
<emphasis id="B920EA8EA26AFFD95B71FA118C812CDC" box="[594,747,1504,1528]" italics="true" pageId="6" pageNumber="112">S. thailandica</emphasis>
</taxonomicName>
. Known cave-dwelling cockroaches have only very limited areas of distribution (
<bibRefCitation id="EFC54B6DA26AFFD95BEAF9F08D372F3D" author="VANDEL, A." box="[713,861,1536,1561]" pageId="6" pageNumber="112" refId="ref2706" refString="VANDEL, A., 1965, Biospeleology. The Biology of Cavernicolous Animals (Oxford: Pergamon Press), 524 pp." type="book" year="1965">Vandel, 1965</bibRefCitation>
; 
<bibRefCitation id="EFC54B6DA26AFFD95A4EF9F08ACD2F3C" author="IZQUIERDO, I. &amp; OROMI, P." box="[877,1191,1536,1560]" pageId="6" pageNumber="112" pagination="295 - 300" refId="ref2612" refString="IZQUIERDO, I. and OROMI, P., 1994, Dictyoptera-Blattaria, in Ch. Juberthie and V. Decu (eds) Encyclopaedia Biospeologica, Vol. 1 (Moulis, Bucarest: Societe de Biospeologie), pp. 295 - 300." type="journal article" year="1994">Izquierdo and Oromi, 1994</bibRefCitation>
), thus we believe that both described sexes are very probably different species.
</paragraph>
</subSubSection>
</treatment>
</document>