237 lines
31 KiB
XML
237 lines
31 KiB
XML
<document id="D347ECB44B82576C2C3AD8AD26206C34" ID-DOI="10.5281/zenodo.6391857" ID-ISSN="1942-1354" ID-Zenodo-Dep="6391857" ID-ZooBank="E80F6A2A-2E1A-479F-BBEF-A5E3BB84E7C9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648503991815" checkinUser="felipe" docAuthor="Muona, Jyrki" docDate="2022" docId="03F5F45FFF996308FF1CFD18FD2D593E" docLanguage="en" docName="InsectaMundi.2022.0912.1-11.pdf" docOrigin="Insecta Mundi 2022 (912)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Neusiokia appalachiensis Muona 2022, new species" docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="FFCC8C27FF9D630FFFDCFFCDFFAD5B51" masterDocTitle="New fossil false click-beetles from the Americas (Coleoptera: Eucnemidae)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="3" updateTime="1699308786968" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="12697FF68F0042047558A4D8026DEE5E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B327E85C45DA0205D54FD067D39172BB">
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<mods:title id="0E7218B5FBD06AC48C5069E3F75A9D24">New fossil false click-beetles from the Americas (Coleoptera: Eucnemidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A8C8620FD9565B558340D6B721EA88F3" type="personal">
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<mods:role id="5728C8B9F7D456F4E6AA66DDD9C9894C">
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<mods:roleTerm id="D8A80287012FC31268802A0912ACB565">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="93886E28F65B0F6A85C239CB6DD021BB">Muona, Jyrki</mods:namePart>
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<mods:affiliation id="4321B4A0E12E6FCFEF6EE166D323FE8E">Finnish Museum of Natural History P. O. Box 17 FIN- 00014 University of Helsinki Helsinki, Finland</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="0A210F57E59B5A9A005AA9538AA8D32A">text</mods:typeOfResource>
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<mods:relatedItem id="8ADD5E439F0FB1DDFBDF51A7B9CFB4C6" type="host">
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<mods:titleInfo id="5DB95306F69E58D2AB6F33DA2CAAA705">
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<mods:title id="7BE9F732A731FA23D38259AF9F414B61">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="FB11BCEA74F0EEF5E270D4264F5AA408">
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<mods:date id="C4741A9BB58689F12E2A82C77B8CF5FF">2022</mods:date>
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<mods:detail id="C8D56B92270F849B0FDF88A89870C586" type="pubDate">
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<mods:number id="9028708B3BC4F0BC2F02220571307F15">2022-02-24</mods:number>
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</mods:detail>
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<mods:detail id="EAAD54A0F0A7DAB0574C6A471CEA9E6F" type="volume">
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<mods:number id="F7308C26FDE2BCD64798E7CAAE8A0CA4">2022</mods:number>
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</mods:detail>
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<mods:detail id="A86D3C51AF86FF8201C05FCEC91EFCFD" type="issue">
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<mods:number id="71DC1BC4FDD5041745DB8A55CECAC829">912</mods:number>
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</mods:detail>
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<mods:extent id="CA6DE55C8C60A5450D83BAE434D32426" unit="page">
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<mods:start id="C1EB87D60F8CA9790DF57867A36F9EC2">1</mods:start>
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<mods:end id="FC0C3628384E600A82494BA1EC596FB0">11</mods:end>
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<mods:classification id="027A5DA5AE3784ABC106C0FE8F438E7F">journal article</mods:classification>
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<mods:identifier id="0546504DD016902B6AC6B7A977AF7804" type="DOI">10.5281/zenodo.6391857</mods:identifier>
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<mods:identifier id="CF9522DDEBD1D09551CB9D24DBDA35F0" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="EC81D7755E37FE29949E21EDA8A00AB3" type="Zenodo-Dep">6391857</mods:identifier>
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<mods:identifier id="B6706EFA93EC85D2C82448CEB29C9AE2" type="ZooBank">E80F6A2A-2E1A-479F-BBEF-A5E3BB84E7C9</mods:identifier>
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</mods:mods>
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<treatment id="03F5F45FFF996308FF1CFD18FD2D593E" LSID="urn:lsid:plazi:treatment:03F5F45FFF996308FF1CFD18FD2D593E" httpUri="http://treatment.plazi.org/id/03F5F45FFF996308FF1CFD18FD2D593E" lastPageId="7" lastPageNumber="6" pageId="4" pageNumber="3">
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<subSubSection id="C34616C2FF99630BFF1CFD18FCAF59A3" box="[192,770,725,754]" pageId="4" pageNumber="3" type="nomenclature">
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<paragraph id="8BE34549FF99630BFF1CFD18FCAF59A3" blockId="4.[192,770,725,788]" box="[192,770,725,754]" pageId="4" pageNumber="3">
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<heading id="D0ABF225FF99630BFF1CFD18FCAF59A3" box="[192,770,725,754]" fontSize="12" level="2" pageId="4" pageNumber="3" reason="2">
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<emphasis id="B928995BFF99630BFF1CFD18FCAF59A3" bold="true" box="[192,770,725,754]" pageId="4" pageNumber="3">
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<taxonomicName id="4C5C3ECAFF99630BFF1CFD18FDCC59A3" authority="Muona" authorityName="Muona" authorityYear="2022" box="[192,609,725,754]" class="Insecta" family="Eucnemidae" genus="Neusiokia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="appalachiensis" status="sp. nov.">
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<emphasis id="B928995BFF99630BFF1CFD18FE5059A3" bold="true" box="[192,509,725,754]" italics="true" pageId="4" pageNumber="3">Neusiokia appalachiensis</emphasis>
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Muona
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21B2420FF99630BFDB2FD1BFCAF59A3" box="[622,770,725,754]" pageId="4" pageNumber="3" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34616C2FF99630BFF1CFD34FEE45845" box="[192,329,761,788]" pageId="4" pageNumber="3" type="description">
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<paragraph id="8BE34549FF99630BFF1CFD34FEE45845" blockId="4.[192,770,725,788]" box="[192,329,761,788]" pageId="4" pageNumber="3">
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<figureCitation id="136759CCFF99630BFF1CFD34FEE85845" box="[192,325,761,788]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="3.[192,268,1900,1926]" captionStartId-1="4.[192,268,1900,1926]" captionStartId-2="5.[193,269,1276,1302]" captionStartId-3="6.[192,268,1020,1046]" captionStartId-4="6.[192,268,1900,1926]" captionTargetBox-0="[192,1440,1136,1870]" captionTargetBox-1="[192,1440,1034,1849]" captionTargetBox-2="[192,1440,213,1247]" captionTargetBox-3="[192,1440,213,990]" captionTargetBox-4="[192,1440,1099,1870]" captionTargetId-0="figure-178@3.[192,1440,1136,1872]" captionTargetId-1="figure-297@4.[192,1440,1034,1864]" captionTargetId-2="figure-287@5.[192,1440,213,1247]" captionTargetId-3="figure-45@6.[192,1440,213,993]" captionTargetId-4="figure-47@6.[192,1440,1096,1873]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionText-0="Figure 1. Neusiokia appalachiensis n. sp., holotype, ventrolateral view." captionText-1="Figure 2. Neusiokia appalachiensis n. sp., holotype, dorsal view." captionText-2="Figure 3. Neusiokia appalachiensis n. sp., holotype, ventral view of tips of protibiae and protarsi. Left arrow points at the divided protibial spur, right arrow points at the second protibial spur." captionText-3="Figure 4. Neusiokia appalachiensis n. sp., holotype, ventral view of head, pro- and mesothorax." captionText-4="Figure 5. Neusiokia appalachiensis n. sp., holotype, ventrolateral view of pronotum and head." figureDoi-0="http://doi.org/10.5281/zenodo.6391859" figureDoi-1="http://doi.org/10.5281/zenodo.6391861" figureDoi-2="http://doi.org/10.5281/zenodo.6391865" figureDoi-3="http://doi.org/10.5281/zenodo.6391867" figureDoi-4="http://doi.org/10.5281/zenodo.6391871" httpUri-0="https://zenodo.org/record/6391859/files/figure.png" httpUri-1="https://zenodo.org/record/6391861/files/figure.png" httpUri-2="https://zenodo.org/record/6391865/files/figure.png" httpUri-3="https://zenodo.org/record/6391867/files/figure.png" httpUri-4="https://zenodo.org/record/6391871/files/figure.png" pageId="4" pageNumber="3">Figures 1–5</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34616C2FF99630BFF1CFCEEFB21586D" box="[192,1164,802,829]" pageId="4" pageNumber="3" type="etymology">
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<paragraph id="8BE34549FF99630BFF1CFCEEFB21586D" blockId="4.[192,1164,802,829]" box="[192,1164,802,829]" pageId="4" pageNumber="3">
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<emphasis id="B928995BFF99630BFF1CFCEEFEE9586C" bold="true" box="[192,324,803,829]" pageId="4" pageNumber="3">Etymology.</emphasis>
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Named after the eastern Cretaceous part of the North American continent.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34616C2FF99630BFF1CFC81FEAB58F8" pageId="4" pageNumber="3" type="materials_examined">
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<paragraph id="8BE34549FF99630BFF1CFC81FEAB58F8" blockId="4.[192,1442,844,937]" pageId="4" pageNumber="3">
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<materialsCitation id="3B344F14FF99630BFF1CFC81FEAB58F8" county="Embedded" location="Cretaceous Black Creek Formation" municipality="Found" pageId="4" pageNumber="3" specimenCount="1" stateProvince="North Carolina" typeStatus="holotype">
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<emphasis id="B928995BFF99630BFF1CFC81FE995837" bold="true" box="[192,308,844,870]" pageId="4" pageNumber="3">
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<typeStatus id="54E7FBEBFF99630BFF1CFC81FE825837" box="[192,303,844,870]" pageId="4" pageNumber="3" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingCounty id="62823DC5FF99630BFEE0FC81FE185837" box="[316,437,844,870]" pageId="4" pageNumber="3">Embedded</collectingCounty>
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in a triangular
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<collectingRegion id="49988BABFF99630BFDB5FC81FCBA5837" box="[617,791,844,870]" country="United States of America" name="North Carolina" pageId="4" pageNumber="3">North Carolina</collectingRegion>
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resin piece, 17 × 10 ×
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<quantity id="4CA4E8ACFF99630BFBC2FC81FBC95836" box="[1054,1124,844,871]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="3" unit="mm" value="9.0">9 mm</quantity>
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, sex unknown.
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<collectingMunicipality id="6B87DF33FF99630BFACBFC81FACD5837" box="[1303,1376,844,870]" pageId="4" pageNumber="3">Found</collectingMunicipality>
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from locality
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<locationDeviation id="9A515C6BFF99630BFEC1FCA3FE7658D9" box="[285,475,878,904]" location="Goldsboro" pageId="4" pageNumber="3">
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<quantity id="4CA4E8ACFF99630BFEC1FCA3FEF458D9" box="[285,345,878,904]" metricMagnitude="-1" metricUnit="m" metricValue="8.636" pageId="4" pageNumber="3" unit="in" value="34.0">34 in</quantity>
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Goldsboro
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</locationDeviation>
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, a
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<location id="8E831392FF99630BFE22FCA3FC2358D9" LSID="urn:lsid:plazi:treatment:03F5F45FFF996308FF1CFD18FD2D593E:8E831392FF99630BFE22FCA3FC2358D9" box="[510,910,878,904]" county="Embedded" municipality="Found" name="Cretaceous Black Creek Formation" pageId="4" pageNumber="3" stateProvince="North Carolina">
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<location id="8E831392FF99630BFE22FCA3FCA058D9" LSID="urn:lsid:plazi:treatment:03F5F45FFF996308FF1CFD18FD2D593E:8E831392FF99630BFE22FCA3FCA058D9" box="[510,781,878,904]" county="Embedded" municipality="Found" name="Cretaceous Black Creek" pageId="4" pageNumber="3" stateProvince="North Carolina">Cretaceous Black Creek</location>
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Formation
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</location>
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exposure along the
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<location id="8E831392FF99630BFBA7FCA3FAA858D9" LSID="urn:lsid:plazi:treatment:03F5F45FFF996308FF1CFD18FD2D593E:8E831392FF99630BFBA7FCA3FAA858D9" box="[1147,1285,878,904]" county="Embedded" municipality="Found" name="Neuse River" pageId="4" pageNumber="3" stateProvince="North Carolina">Neuse River</location>
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(
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<bibRefCitation id="EFCD38B8FF99630BFACAFCA3FF5758F8" author="Carter JG & Gallagher PE & Valone RE & Rossbach TJ" pageId="4" pageNumber="3" refId="ref4102" refString="Carter JG, Gallagher PE, Valone RE, Rossbach TJ. 1988. Fossil collecting in North Carolina. Bulletin 89. Department of Natural Resources and Community Development; Raleigh, NC. 86 p." type="book" year="1988">Carter et al. 1988</bibRefCitation>
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).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34616C2FF99630BFF1CFC74FA905882" box="[192,1341,953,979]" pageId="4" pageNumber="3" type="diagnosis">
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<paragraph id="8BE34549FF99630BFF1CFC74FA905882" blockId="4.[192,1341,953,979]" box="[192,1341,953,979]" pageId="4" pageNumber="3">
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<emphasis id="B928995BFF99630BFF1CFC74FE915882" bold="true" box="[192,316,953,979]" pageId="4" pageNumber="3">Diagnosis.</emphasis>
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Characterized by the enlarged f7–f
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<quantity id="4CA4E8ACFF99630BFD19FC74FD425882" box="[709,751,953,979]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="4" pageNumber="3" unit="in" value="9.0">9 in</quantity>
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combination with two protibial spurs and small size.
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</paragraph>
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</subSubSection>
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<caption id="DF2315C1FF99630BFF1CF8A1FC2E5CD7" ID-DOI="http://doi.org/10.5281/zenodo.6391861" ID-Zenodo-Dep="6391861" box="[192,899,1900,1927]" httpUri="https://zenodo.org/record/6391861/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,268,1900,1926]" targetBox="[192,1440,1034,1849]" targetPageId="4">
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<paragraph id="8BE34549FF99630BFF1CF8A1FC2E5CD7" blockId="4.[192,899,1900,1927]" box="[192,899,1900,1927]" pageId="4" pageNumber="3">
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<emphasis id="B928995BFF99630BFF1CF8A1FE855CD7" bold="true" box="[192,296,1900,1926]" pageId="4" pageNumber="3">Figure 2.</emphasis>
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<taxonomicName id="4C5C3ECAFF99630BFEF2F8A1FD925CD7" authority="Muona, 2022" authorityName="Muona" authorityYear="2022" box="[302,575,1900,1926]" class="Insecta" family="Eucnemidae" genus="Neusiokia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="appalachiensis" status="sp. nov.">
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<emphasis id="B928995BFF99630BFEF2F8A1FD925CD7" box="[302,575,1900,1926]" italics="true" pageId="4" pageNumber="3">Neusiokia appalachiensis</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B928995BFF99630BFD99F8A1FD2E5CD7" bold="true" box="[581,643,1900,1926]" pageId="4" pageNumber="3">
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<taxonomicNameLabel id="A21B2420FF99630BFD99F8A1FD2E5CD7" box="[581,643,1900,1926]" pageId="4" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, holotype, dorsal view.
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</paragraph>
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</caption>
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<caption id="DF2315C1FF98630AFF1DFB31FBAA5E68" ID-DOI="http://doi.org/10.5281/zenodo.6391865" ID-Zenodo-Dep="6391865" httpUri="https://zenodo.org/record/6391865/files/figure.png" pageId="5" pageNumber="4" startId="5.[193,269,1276,1302]" targetBox="[192,1440,213,1247]" targetPageId="5">
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||
<paragraph id="8BE34549FF98630AFF1DFB31FBAA5E68" blockId="5.[191,1440,1276,1337]" pageId="5" pageNumber="4">
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||
<emphasis id="B928995BFF98630AFF1DFB31FE855E46" bold="true" box="[193,296,1276,1303]" pageId="5" pageNumber="4">Figure 3.</emphasis>
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<taxonomicName id="4C5C3ECAFF98630AFEF0FB30FD965E46" authority="Muona, 2022" authorityName="Muona" authorityYear="2022" box="[300,571,1277,1303]" class="Insecta" family="Eucnemidae" genus="Neusiokia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="appalachiensis" status="sp. nov.">
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<emphasis id="B928995BFF98630AFEF0FB30FD965E46" box="[300,571,1277,1303]" italics="true" pageId="5" pageNumber="4">Neusiokia appalachiensis</emphasis>
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</taxonomicName>
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||
<emphasis id="B928995BFF98630AFD9DFB30FDD05E46" bold="true" box="[577,637,1277,1303]" pageId="5" pageNumber="4">
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||
<taxonomicNameLabel id="A21B2420FF98630AFD9DFB30FDD05E46" box="[577,637,1277,1303]" pageId="5" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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, holotype, ventral view of tips of protibiae and protarsi. Left arrow points at the divided protibial spur, right arrow points at the second protibial spur.
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</paragraph>
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</caption>
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||
<subSubSection id="C34616C2FF986308FF1CFABEFE745A40" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4" type="description">
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||
<paragraph id="8BE34549FF98630AFF1CFABEFA985EDF" blockId="5.[191,1441,1395,1926]" box="[192,1333,1395,1422]" pageId="5" pageNumber="4">
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||
<emphasis id="B928995BFF98630AFF1CFABEFEFF5EDC" bold="true" box="[192,338,1395,1421]" pageId="5" pageNumber="4">Description.</emphasis>
|
||
Length
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||
<quantity id="4CA4E8ACFF98630AFE72FABEFDAB5EDF" box="[430,518,1395,1422]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="4" unit="mm" value="4.5">4.5 mm</quantity>
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||
.
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<taxonomicName id="4C5C3ECAFF98630AFDCCFABEFD235EDC" authorityName="Leach" authorityYear="1815" box="[528,654,1395,1421]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="2" phylum="Arthropoda" rank="superFamily" superFamily="Elateroidea">Elateroidea</taxonomicName>
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synapomorphy: pro-mesocoxal clicking mechanism present.
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||
</paragraph>
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<paragraph id="8BE34549FF98630AFEDCFA58FB115DED" blockId="5.[191,1441,1395,1926]" pageId="5" pageNumber="4">
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||
<taxonomicName id="4C5C3ECAFF98630AFEDCFA58FE265EFE" authorityName="Eschscholtz" authorityYear="1829" box="[256,395,1429,1455]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="family">Eucnemidae</taxonomicName>
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synapomorphies: pedicel attached subapically to scape, labrum hidden and sutural striae with excretory punctures apically. Head rounded, very densely punctate, eyes medium sized, frontoclypeal region wide, about 1.5 times as wide apically as between antennal sockets, apical margin straight with caudally projecting lateral apices, mandibles elongated with simple apex (
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<figureCitation id="136759CCFF98630AFC9BFA34FC245D45" box="[839,905,1529,1556]" captionStart="Figure 5" captionStartId="6.[192,268,1900,1926]" captionTargetBox="[192,1440,1099,1870]" captionTargetId="figure-47@6.[192,1440,1096,1873]" captionTargetPageId="6" captionText="Figure 5. Neusiokia appalachiensis n. sp., holotype, ventrolateral view of pronotum and head." figureDoi="http://doi.org/10.5281/zenodo.6391871" httpUri="https://zenodo.org/record/6391871/files/figure.png" pageId="5" pageNumber="4">Fig. 5</figureCitation>
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). Pronotum about as wide as long, sides evenly and strongly narrowing craniad, with rounded hind angles, lateral and frontal margins simple, scutellum wider than long, bluntly pointed caudad, parallel-sided on basal half, elytra with rounded sides, striae distinct, sutural ones strong apically with excretory punctures, interstices nearly flat, moderately densely punctate (
|
||
<figureCitation id="136759CCFF98630AFB21F993FA935D29" box="[1277,1342,1630,1656]" captionStart="Figure 2" captionStartId="4.[192,268,1900,1926]" captionTargetBox="[192,1440,1034,1849]" captionTargetId="figure-297@4.[192,1440,1034,1864]" captionTargetPageId="4" captionText="Figure 2. Neusiokia appalachiensis n. sp., holotype, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6391861" httpUri="https://zenodo.org/record/6391861/files/figure.png" pageId="5" pageNumber="4">Fig. 2</figureCitation>
|
||
). Antennal fairly short, f9 reaching pronotal hind angles, scape elongate, nearly as long as pedicel and f1–f2 combined, f2–f6 twice as long as wide, apically expanded, f7–f9 forming loose, fattened club (
|
||
<figureCitation id="136759CCFF98630AFB8EF96CFB3E5DED" box="[1106,1171,1697,1724]" captionStart="Figure 4" captionStartId="6.[192,268,1020,1046]" captionTargetBox="[192,1440,213,990]" captionTargetId="figure-45@6.[192,1440,213,993]" captionTargetPageId="6" captionText="Figure 4. Neusiokia appalachiensis n. sp., holotype, ventral view of head, pro- and mesothorax." figureDoi="http://doi.org/10.5281/zenodo.6391867" httpUri="https://zenodo.org/record/6391867/files/figure.png" pageId="5" pageNumber="4">Fig. 4</figureCitation>
|
||
,
|
||
<figureCitation id="136759CCFF98630AFB42F96CFB005DED" box="[1182,1197,1697,1724]" captionStart="Figure 5" captionStartId="6.[192,268,1900,1926]" captionTargetBox="[192,1440,1099,1870]" captionTargetId="figure-47@6.[192,1440,1096,1873]" captionTargetPageId="6" captionText="Figure 5. Neusiokia appalachiensis n. sp., holotype, ventrolateral view of pronotum and head." figureDoi="http://doi.org/10.5281/zenodo.6391871" httpUri="https://zenodo.org/record/6391871/files/figure.png" pageId="5" pageNumber="4">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE34549FF986308FEDCF909FE745A40" blockId="5.[191,1441,1395,1926]" lastBlockId="7.[192,1440,214,274]" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4">
|
||
Hypomera simple, smooth, widely and shallowly grooved, delimited by notosternal suture medially and pronotal ridge laterally, prosternal peg wide, short, prosternum narrow, metasternum densely punctate, without mesocoxal lines. Metacoxal plates densely punctate, elongate, widest close to midline, here about twice as wide as on sides (
|
||
<figureCitation id="136759CCFF98630AFEF4F8E5FECA5C13" box="[296,359,1832,1858]" captionStart="Figure 1" captionStartId="3.[192,268,1900,1926]" captionTargetBox="[192,1440,1136,1870]" captionTargetId="figure-178@3.[192,1440,1136,1872]" captionTargetPageId="3" captionText="Figure 1. Neusiokia appalachiensis n. sp., holotype, ventrolateral view." figureDoi="http://doi.org/10.5281/zenodo.6391859" httpUri="https://zenodo.org/record/6391859/files/figure.png" pageId="5" pageNumber="4">Fig. 1</figureCitation>
|
||
). Legs long, femora stout, fairly short, meso- and metatibiae long, strongly built, with well-developed spine-combs, meso- and metatarsi shorter than tibiae, protibia short, powerful, apically expanded, apex with two spurs, dorsal apical spur split in two (
|
||
<figureCitation id="136759CCFF98630AFD0EF8A6FCB95CD4" box="[722,788,1899,1925]" captionStart="Figure 3" captionStartId="5.[193,269,1276,1302]" captionTargetBox="[192,1440,213,1247]" captionTargetId="figure-287@5.[192,1440,213,1247]" captionTargetPageId="5" captionText="Figure 3. Neusiokia appalachiensis n. sp., holotype, ventral view of tips of protibiae and protarsi. Left arrow points at the divided protibial spur, right arrow points at the second protibial spur." figureDoi="http://doi.org/10.5281/zenodo.6391865" httpUri="https://zenodo.org/record/6391865/files/figure.png" pageId="5" pageNumber="4">Fig. 3</figureCitation>
|
||
), first protarsomere without sex-comb, fifth protarsomere exceptionally long (
|
||
<figureCitation id="136759CCFF9A6308FE7CFF1BFE4B5BA1" box="[416,486,214,240]" captionStart="Figure 4" captionStartId="6.[192,268,1020,1046]" captionTargetBox="[192,1440,213,990]" captionTargetId="figure-45@6.[192,1440,213,993]" captionTargetPageId="6" captionText="Figure 4. Neusiokia appalachiensis n. sp., holotype, ventral view of head, pro- and mesothorax." figureDoi="http://doi.org/10.5281/zenodo.6391867" httpUri="https://zenodo.org/record/6391867/files/figure.png" pageId="7" pageNumber="6">Fig. 4</figureCitation>
|
||
), all tarsomeres 4 simple, claws simple (
|
||
<figureCitation id="136759CCFF9A6308FC6BFF1BFC575BA1" box="[951,1018,214,240]" captionStart="Figure 1" captionStartId="3.[192,268,1900,1926]" captionTargetBox="[192,1440,1136,1870]" captionTargetId="figure-178@3.[192,1440,1136,1872]" captionTargetPageId="3" captionText="Figure 1. Neusiokia appalachiensis n. sp., holotype, ventrolateral view." figureDoi="http://doi.org/10.5281/zenodo.6391859" httpUri="https://zenodo.org/record/6391859/files/figure.png" pageId="7" pageNumber="6">Fig. 1</figureCitation>
|
||
,
|
||
<figureCitation id="136759CCFF9A6308FBD6FF1BFBB45BA1" box="[1034,1049,214,240]" captionStart="Figure 3" captionStartId="5.[193,269,1276,1302]" captionTargetBox="[192,1440,213,1247]" captionTargetId="figure-287@5.[192,1440,213,1247]" captionTargetPageId="5" captionText="Figure 3. Neusiokia appalachiensis n. sp., holotype, ventral view of tips of protibiae and protarsi. Left arrow points at the divided protibial spur, right arrow points at the second protibial spur." figureDoi="http://doi.org/10.5281/zenodo.6391865" httpUri="https://zenodo.org/record/6391865/files/figure.png" pageId="7" pageNumber="6">3</figureCitation>
|
||
,
|
||
<figureCitation id="136759CCFF9A6308FBF4FF1BFB955BA1" box="[1064,1080,214,240]" captionStart="Figure 4" captionStartId="6.[192,268,1020,1046]" captionTargetBox="[192,1440,213,990]" captionTargetId="figure-45@6.[192,1440,213,993]" captionTargetPageId="6" captionText="Figure 4. Neusiokia appalachiensis n. sp., holotype, ventral view of head, pro- and mesothorax." figureDoi="http://doi.org/10.5281/zenodo.6391867" httpUri="https://zenodo.org/record/6391867/files/figure.png" pageId="7" pageNumber="6">4</figureCitation>
|
||
). Abdominal ventrites connate, densely punctate (
|
||
<figureCitation id="136759CCFF9A6308FE56FF3AFE645A40" box="[394,457,247,273]" captionStart="Figure 1" captionStartId="3.[192,268,1900,1926]" captionTargetBox="[192,1440,1136,1870]" captionTargetId="figure-178@3.[192,1440,1136,1872]" captionTargetPageId="3" captionText="Figure 1. Neusiokia appalachiensis n. sp., holotype, ventrolateral view." figureDoi="http://doi.org/10.5281/zenodo.6391859" httpUri="https://zenodo.org/record/6391859/files/figure.png" pageId="7" pageNumber="6">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2315C1FF9B6309FF1CFC31FB725F47" ID-DOI="http://doi.org/10.5281/zenodo.6391867" ID-Zenodo-Dep="6391867" box="[192,1247,1020,1047]" httpUri="https://zenodo.org/record/6391867/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,268,1020,1046]" targetBox="[192,1440,213,990]" targetPageId="6">
|
||
<paragraph id="8BE34549FF9B6309FF1CFC31FB725F47" blockId="6.[192,1247,1020,1047]" box="[192,1247,1020,1047]" pageId="6" pageNumber="5">
|
||
<emphasis id="B928995BFF9B6309FF1CFC31FE855F47" bold="true" box="[192,296,1020,1046]" pageId="6" pageNumber="5">Figure 4.</emphasis>
|
||
<taxonomicName id="4C5C3ECAFF9B6309FEF2FC31FD925F47" authority="Muona, 2022" authorityName="Muona" authorityYear="2022" box="[302,575,1020,1046]" class="Insecta" family="Eucnemidae" genus="Neusiokia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="appalachiensis" status="sp. nov.">
|
||
<emphasis id="B928995BFF9B6309FEF2FC31FD925F47" box="[302,575,1020,1046]" italics="true" pageId="6" pageNumber="5">Neusiokia appalachiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928995BFF9B6309FD99FC30FD2E5F46" bold="true" box="[581,643,1021,1047]" pageId="6" pageNumber="5">
|
||
<taxonomicNameLabel id="A21B2420FF9B6309FD99FC30FD2E5F46" box="[581,643,1021,1047]" pageId="6" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, ventral view of head, pro- and mesothorax.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2315C1FF9B6309FF1CF8A1FB635CD7" ID-DOI="http://doi.org/10.5281/zenodo.6391871" ID-Zenodo-Dep="6391871" box="[192,1230,1900,1927]" httpUri="https://zenodo.org/record/6391871/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,268,1900,1926]" targetBox="[192,1440,1099,1870]" targetPageId="6">
|
||
<paragraph id="8BE34549FF9B6309FF1CF8A1FB635CD7" blockId="6.[192,1230,1900,1927]" box="[192,1230,1900,1927]" pageId="6" pageNumber="5">
|
||
<emphasis id="B928995BFF9B6309FF1CF8A1FE855CD7" bold="true" box="[192,296,1900,1926]" pageId="6" pageNumber="5">Figure 5.</emphasis>
|
||
<taxonomicName id="4C5C3ECAFF9B6309FEF2F8A1FD925CD7" authority="Muona, 2022" authorityName="Muona" authorityYear="2022" box="[302,575,1900,1926]" class="Insecta" family="Eucnemidae" genus="Neusiokia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="appalachiensis" status="sp. nov.">
|
||
<emphasis id="B928995BFF9B6309FEF2F8A1FD925CD7" box="[302,575,1900,1926]" italics="true" pageId="6" pageNumber="5">Neusiokia appalachiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928995BFF9B6309FD99F8A1FD2E5CD7" bold="true" box="[581,643,1900,1926]" pageId="6" pageNumber="5">
|
||
<taxonomicNameLabel id="A21B2420FF9B6309FD99F8A1FD2E5CD7" box="[581,643,1900,1926]" pageId="6" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, ventrolateral view of pronotum and head.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34616C2FF9A6308FF1CFEECFD2D593E" pageId="7" pageNumber="6" type="discussion">
|
||
<paragraph id="8BE34549FF9A6308FF1CFEECFDCD5AF1" blockId="7.[191,1440,289,416]" pageId="7" pageNumber="6">
|
||
<emphasis id="B928995BFF9A6308FF1CFEECFE825A6A" bold="true" box="[192,303,289,315]" pageId="7" pageNumber="6">Remarks.</emphasis>
|
||
<taxonomicName id="4C5C3ECAFF9A6308FEE4FEECFDE15A6A" authorityName="Muona" authorityYear="2022" box="[312,588,289,315]" class="Insecta" family="Eucnemidae" genus="Neusiokia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="appalachiensis">
|
||
<emphasis id="B928995BFF9A6308FEE4FEECFDE15A6A" box="[312,588,289,315]" italics="true" pageId="7" pageNumber="6">Neusiokia appalachiensis</emphasis>
|
||
</taxonomicName>
|
||
challenges the present classification, partly in the same manner as
|
||
<taxonomicName id="4C5C3ECAFF9A6308FA8AFEECFE965A0D" class="Insecta" family="Eucnemidae" genus="Erdaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="bisipnulosa">
|
||
<emphasis id="B928995BFF9A6308FA8AFEECFE965A0D" italics="true" pageId="7" pageNumber="6">Erdaia bisipnulosa</emphasis>
|
||
</taxonomicName>
|
||
did (
|
||
<bibRefCitation id="EFCD38B8FF9A6308FEA0FE8EFDBE5A0C" author="Muona J." box="[380,531,323,349]" pageId="7" pageNumber="6" pagination="90 - 113" refId="ref4568" refString="Muona J. 2021. False click-beetles from Baltic amber (Coleoptera, Eucnemidae). Entomologische Blatter fur Biologie und Systematik der Kafer 117: 90 - 113." type="journal article" year="2021">Muona 2021</bibRefCitation>
|
||
). On the basis of the presently accepted synapomorphies, they both appear to belong in the subfamily
|
||
<taxonomicName id="4C5C3ECAFF9A6308FE16FEA9FDC05A2F" authorityName="Muona" authorityYear="1993" box="[458,621,356,382]" pageId="7" pageNumber="6" rank="subFamily" subFamily="Palaeoxeninae">Palaeoxeninae</taxonomicName>
|
||
. As a new analysis is needed to clarify this question, I have chosen to place
|
||
<taxonomicName id="4C5C3ECAFF9A6308FF63FE4BFE815AF1" authorityName="Muona" authorityYear="2022" box="[191,300,390,416]" class="Insecta" family="Eucnemidae" genus="Neusiokia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928995BFF9A6308FF63FE4BFE815AF1" box="[191,300,390,416]" italics="true" pageId="7" pageNumber="6">Neusiokia</emphasis>
|
||
</taxonomicName>
|
||
provisionally in
|
||
<taxonomicName id="4C5C3ECAFF9A6308FE34FE4BFDF15AF1" authorityName="Muona" authorityYear="1993" box="[488,604,390,416]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Orodotini">Orodotini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE34549FF9A6308FF1CFE05FD0B5AB4" blockId="7.[192,678,456,485]" box="[192,678,456,485]" pageId="7" pageNumber="6">
|
||
<emphasis id="B928995BFF9A6308FF1CFE05FD0B5AB4" bold="true" box="[192,678,456,485]" pageId="7" pageNumber="6">
|
||
Family
|
||
<taxonomicName id="4C5C3ECAFF9A6308FEC3FE05FD0B5AB4" authority="Eschscholtz, 1829" authorityName="Eschscholtz" authorityYear="1829" box="[287,678,456,485]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="family">Eucnemidae Eschscholtz, 1829</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE34549FF9A6308FF1CFDC0FD7D597B" blockId="7.[192,720,525,554]" box="[192,720,525,554]" pageId="7" pageNumber="6">
|
||
<emphasis id="B928995BFF9A6308FF1CFDC0FD7D597B" bold="true" box="[192,720,525,554]" pageId="7" pageNumber="6">
|
||
Subfamily
|
||
<taxonomicName id="4C5C3ECAFF9A6308FE95FDC0FD7D597B" authority="Fleutiaux, 1922" authorityName="Fleutiaux" authorityYear="1922" box="[329,720,525,554]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="subFamily" subFamily="Macraulacinae">Macraulacinae Fleutiaux, 1922</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BE34549FF9A6308FF1CFD9FFD2D593E" blockId="7.[192,640,594,623]" box="[192,640,594,623]" pageId="7" pageNumber="6">
|
||
<emphasis id="B928995BFF9A6308FF1CFD9FFD2D593E" bold="true" box="[192,640,594,623]" pageId="7" pageNumber="6">
|
||
Tribe
|
||
<taxonomicName id="4C5C3ECAFF9A6308FED7FD9FFD2D593E" authority="Fleutiaux, 1922" authorityName="Fleutiaux" authorityYear="1922" box="[267,640,594,623]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Macraulacini">Macraulacini Fleutiaux, 1922</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |