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<document id="F9865D2E1CEA52A45439FD9A29C45E43" ID-DOI="10.5281/zenodo.195431" ID-GBIF-Dataset="2df8bd5a-e125-42aa-aa6f-ae3e78b83929" ID-ISSN="1175-5326" ID-Zenodo-Dep="195431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226955823" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2010" docId="03F39207FFB7FFCD7EFCFA6DF780D9D6" docLanguage="en" docName="zt02481p036.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tortopus bellus Lugo-Ortiz &amp; McCafferty" docType="treatment" docVersion="10" lastPageNumber="19" masterDocId="FFCAEA7FFFB9FFDE7E6BFF82F50BD901" masterDocTitle="A cladistic revision of Tor top us Needham &amp; Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="15" updateTime="1698238640945" updateUser="plazi">
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<mods:title id="00385553910A4832EEB04241F2FA84E1">A cladistic revision of Tor top us Needham &amp; Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
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<mods:date id="5FCCEAEDF85733BD99BD50D7F44A26D7">2010</mods:date>
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<treatment id="03F39207FFB7FFCD7EFCFA6DF780D9D6" ID-DOI="http://doi.org/10.5281/zenodo.5690567" ID-GBIF-Taxon="119386656" ID-Zenodo-Dep="5690567" LSID="urn:lsid:plazi:treatment:03F39207FFB7FFCD7EFCFA6DF780D9D6" httpUri="http://treatment.plazi.org/id/03F39207FFB7FFCD7EFCFA6DF780D9D6" lastPageId="19" lastPageNumber="19" pageId="14" pageNumber="15">
<subSubSection id="C340709AFFB7FFD07EFCFA6DF794DF08" box="[151,671,1519,1545]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BE52311FFB7FFD07EFCFA6DF794DF08" blockId="14.[151,671,1519,1545]" box="[151,671,1519,1545]" pageId="14" pageNumber="15">
<heading id="D0AD947DFFB7FFD07EFCFA6DF794DF08" bold="true" box="[151,671,1519,1545]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="4C5A5892FFB7FFD07EFCFA6DF794DF08" ID-CoL="57H2N" authority="Lugo-Ortiz &amp; McCafferty" authorityName="Lugo-Ortiz &amp; McCafferty" box="[151,671,1519,1545]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="B92EFF03FFB7FFD07EFCFA6DF794DF08" bold="true" box="[151,671,1519,1545]" pageId="14" pageNumber="15">
<emphasis id="B92EFF03FFB7FFD07EFCFA6DF45BDF08" bold="true" box="[151,336,1519,1545]" italics="true" pageId="14" pageNumber="15">Tortopus bellus</emphasis>
Lugo-Ortiz &amp; McCafferty
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C340709AFFB7FFD07EFCF9B5F7C8DF4F" box="[151,707,1591,1614]" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BE52311FFB7FFD07EFCF9B5F7C8DF4F" blockId="14.[151,707,1591,1614]" box="[151,707,1591,1614]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB4A043FFFB7FFD07EFCF9B5F7C8DF4F" box="[151,707,1591,1614]" pageId="14" pageNumber="15">
<taxonomicName id="4C5A5892FFB7FFD07EFCF9B5F7CBDF4F" ID-CoL="57H2N" authority="Lugo-Ortiz &amp; McCafferty 1996: 24" authorityName="Lugo-Ortiz &amp; McCafferty" authorityPageNumber="24" authorityYear="1996" box="[151,704,1591,1614]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="B92EFF03FFB7FFD07EFCF9B5F430DF4F" box="[151,315,1591,1614]" italics="true" pageId="14" pageNumber="15">Tortopus bellus</emphasis>
<treatmentCitation id="0AFB0500FFB7FFD07F29F9B5F7CBDF4F" author="Lugo-Ortiz" box="[322,704,1591,1614]" page="24" pageId="14" pageNumber="15" year="1996">
<bibRefCitation id="EFCB5EE0FFB7FFD07F29F9B5F79CDF4F" author="Lugo-Ortiz" box="[322,663,1591,1614]" pageId="14" pageNumber="34" refString="Lugo-Ortiz, C. R. &amp; McCafferty W. P. (1996) Central American Tortopus (Ephemeroptera: Polymitarcyidae): a unique new species and new country records. Entomological News, 107, 23 - 27." type="journal article" year="1996">Lugo-Ortiz &amp; McCafferty 1996</bibRefCitation>
: 24
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C340709AFFB7FFCD7EFCF9FFF780D9D6" lastPageId="19" lastPageNumber="20" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BE52311FFB7FFD07EFCF9FFF433DE06" blockId="14.[151,1437,1661,2023]" pageId="14" pageNumber="15">
Material (
<collectionCode id="ED4BBBD4FFB7FFD07F61F9FFF46FDF96" box="[266,356,1661,1687]" pageId="14" pageNumber="15">FAMU</collectionCode>
):
<specimenCount id="9D5CE898FFB7FFD07F13F9FFF4CCDF96" box="[376,455,1661,1687]" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
(genitalia on slide 1116) and
<specimenCount id="9D5CE898FFB7FFD07D4DF9FFF686DF96" box="[806,909,1661,1687]" pageId="14" pageNumber="15" type="female">1 female</specimenCount>
imagos from
<collectingCountry id="F34D6381FFB7FFD07A5AF9FFF1BDDF96" box="[1073,1206,1661,1687]" name="Costa Rica" pageId="14" pageNumber="15">Costa Rica</collectingCountry>
: Heredia, Est. Biol. La Selva, río Puerto Viejo,
<date id="FFE405D1FFB7FFD07FB3F920F76DDFBD" box="[472,614,1698,1724]" pageId="14" pageNumber="15" value="1986-06-19">19-VI-1986</date>
, Holzenthal, Heyn &amp; Armitage cols. Attempts to borrow the
<typeStatus id="54E19DB3FFB7FFD07B5EF920F090DFBD" box="[1333,1435,1698,1724]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
male from Purdue Entomological Collection were not successful.
<typeStatus id="54E19DB3FFB7FFD07DF3F945F6D9DFE0" box="[920,978,1735,1761]" pageId="14" pageNumber="15">Type</typeStatus>
locality:
<collectingCountry id="F34D6381FFB7FFD07A2BF945F1C8DFE0" box="[1088,1219,1735,1761]" name="Costa Rica" pageId="14" pageNumber="15">Costa Rica</collectingCountry>
, Heredia (no more locality data).
</paragraph>
<paragraph id="8BE52311FFB7FFCF7EADF890F6C5D83D" blockId="14.[151,1437,1661,2023]" lastBlockId="17.[151,1437,152,316]" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
Male imago. Length (mm): body, 10.5 (
<quantity id="4CA28EF4FFB7FFD07CD9F890F7F0DE2D" box="[690,763,1810,1836]" metricMagnitude="-1" metricUnit="m" metricValue="2.1082" pageId="14" pageNumber="15" unit="in" value="8.3">8.3 in</quantity>
original description); fore wing, 12.9 (
<quantity id="4CA28EF4FFB7FFD07AB5F890F03EDE2D" box="[1246,1333,1810,1836]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" pageId="14" pageNumber="15" unit="in" value="10.5">10.5 in</quantity>
original description); hind wing, 6.2. General coloration yellowish brown. Head yellowish brown shaded black on dorsum almost entirely, shaded heavier between ocelli. Antennae yellowish shaded gray on scape, black on pedicel, flagellum hyaline. Thorax orangeish brown shaded black mainly on pronotum. Pleura and sterna slightly paler shaded black on mesofurcasternal membrane and metasternum. Fore legs yellowish shaded with black except at articulations of segments and claws, pale; legs II–III whitish yellow shaded gray on femora and tibiae. Wings membrane hyaline translucent, veins brownish yellow, lighter toward apex. Abdomen yellowish completely shaded brownish gray except some pale markings, segment X slightly paler than the rest. Genitalia (
<figureCitation id="13613F94FFA8FFCF7F33FF5FF4FAD9F6" box="[344,497,221,247]" captionStart="FIGURES 10 – 13" captionStartId="10.[151,270,1119,1143]" captionTargetBox="[169,1402,195,1092]" captionTargetId="figure@10.[151,1436,175,1095]" captionTargetPageId="10" captionText="FIGURES 10 – 13. Tortopus bellus: 10, male genitalia, v. v.; 11, details of apex of penes; 12, detail of pedestal, v. v.; 13, sockets on female abdominal sternum VIII." httpUri="https://zenodo.org/record/195433/files/figure.png" pageId="17" pageNumber="18">Figs. 10–12</figureCitation>
): sternum IX shaded with black on median 1/3, stronger on median furrow; pedestals and short parastyli (
<figureCitation id="13613F94FFA8FFCF7F9BFE82F74FD81B" box="[496,580,256,282]" captionStart="FIGURES 10 – 13" captionStartId="10.[151,270,1119,1143]" captionTargetBox="[169,1402,195,1092]" captionTargetId="figure@10.[151,1436,175,1095]" captionTargetPageId="10" captionText="FIGURES 10 – 13. Tortopus bellus: 10, male genitalia, v. v.; 11, details of apex of penes; 12, detail of pedestal, v. v.; 13, sockets on female abdominal sternum VIII." httpUri="https://zenodo.org/record/195433/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
) yellowish translucent; forceps whitish yellow shaded almost completely with black; penes translucent yellowish white, apex rounded (
<figureCitation id="13613F94FFA8FFCF7D01FEA0F6CBD83D" box="[874,960,290,316]" captionStart="FIGURES 10 – 13" captionStartId="10.[151,270,1119,1143]" captionTargetBox="[169,1402,195,1092]" captionTargetId="figure@10.[151,1436,175,1095]" captionTargetPageId="10" captionText="FIGURES 10 – 13. Tortopus bellus: 10, male genitalia, v. v.; 11, details of apex of penes; 12, detail of pedestal, v. v.; 13, sockets on female abdominal sternum VIII." httpUri="https://zenodo.org/record/195433/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
).
</paragraph>
<caption id="DF257399FFB6FFD17EFCF8BCF682DE96" httpUri="https://zenodo.org/record/195437/files/figure.png" pageId="15" pageNumber="16" targetBox="[191,1384,197,1829]" targetPageId="15">
<paragraph id="8BE52311FFB6FFD17EFCF8BCF682DE96" blockId="15.[151,1436,1854,1943]" pageId="15" pageNumber="16">
<emphasis id="B92EFF03FFB6FFD17EFCF8BCF455DE57" bold="true" box="[151,350,1854,1878]" pageId="15" pageNumber="16">FIGURES 31–40.</emphasis>
<taxonomicName id="4C5A5892FFB6FFD17F0EF8BDF72EDE57" box="[357,549,1855,1878]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="arenales">
<emphasis id="B92EFF03FFB6FFD17F0EF8BDF72EDE57" box="[357,549,1855,1878]" italics="true" pageId="15" pageNumber="16">Tortopus arenales</emphasis>
</taxonomicName>
. 31, male head, d.v.; 32, male abdominal terga VI–VII; 33, male abdominal sterna VII–VIII; 34, male mesofurcasternal plates; 35, female abdominal sternum VIII; 36, male genitalia, v.v.; 37, same, l.v.; 38, penes after treatment with potash; 39–40, detail of apex of forceps.
</paragraph>
</caption>
<caption id="DF257399FFA9FFCE7EFCF838F6D3DEF0" httpUri="https://zenodo.org/record/195438/files/figure.png" pageId="16" pageNumber="17" targetBox="[221,1357,197,1936]" targetPageId="16">
<paragraph id="8BE52311FFA9FFCE7EFCF838F6D3DEF0" blockId="16.[151,1436,1978,2033]" pageId="16" pageNumber="17">
<emphasis id="B92EFF03FFA9FFCE7EFCF838F46FDED3" bold="true" box="[151,356,1978,2002]" pageId="16" pageNumber="17">FIGURES 41–43.</emphasis>
<taxonomicName id="4C5A5892FFA9FFCE7F07F838F763DED0" box="[364,616,1978,2001]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="unguiculatus">
<emphasis id="B92EFF03FFA9FFCE7F07F838F763DED0" box="[364,616,1978,2001]" italics="true" pageId="16" pageNumber="17">Tortopsis unguiculatus</emphasis>
</taxonomicName>
, type male genitalia, photographs and interpretations (line drawings). 41, ventral view; 42, lateral view; 43, laterodorsal view (details of apex of penes).
</paragraph>
</caption>
<caption id="DF257399FFA8FFCF7EFCF807F6D3DEDD" httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="17" pageNumber="18" targetBox="[285,1301,367,1892]" targetPageId="17">
<paragraph id="8BE52311FFA8FFCF7EFCF807F6D3DEDD" blockId="17.[151,1436,1925,2013]" pageId="17" pageNumber="18">
<emphasis id="B92EFF03FFA8FFCF7EFCF807F46CDE9C" bold="true" box="[151,359,1925,1949]" pageId="17" pageNumber="18">FIGURES 44–52.</emphasis>
<taxonomicName id="4C5A5892FFA8FFCF7F04F804F4DCDE9C" box="[367,471,1926,1949]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFA8FFCF7F04F804F4DCDE9C" box="[367,471,1926,1949]" italics="true" pageId="17" pageNumber="18">Tortopsis</emphasis>
</taxonomicName>
, female imago. 44,
<taxonomicName id="4C5A5892FFA8FFCF7CD7F804F67CDE9C" box="[700,887,1926,1949]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="B92EFF03FFA8FFCF7CD7F804F67CDE9C" box="[700,887,1926,1949]" italics="true" pageId="17" pageNumber="18">T. obscuripennis</emphasis>
</taxonomicName>
, fore wing.
<taxonomicName id="4C5A5892FFA8FFCF7A68F804F154DE9C" box="[1027,1119,1926,1949]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sarae">
<emphasis id="B92EFF03FFA8FFCF7A68F804F154DE9C" box="[1027,1119,1926,1949]" italics="true" pageId="17" pageNumber="18">T. sarae</emphasis>
</taxonomicName>
: 45, mesofurcasternum; 46, abdominal sternum VIII,v.v.; 47, same, l.v.
<taxonomicName id="4C5A5892FFA8FFCF7C00F824F617DEBC" box="[619,796,1958,1981]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="obscuripennis">
<emphasis id="B92EFF03FFA8FFCF7C00F824F617DEBC" box="[619,796,1958,1981]" italics="true" pageId="17" pageNumber="18">T. obscuripennis</emphasis>
</taxonomicName>
: 48, abdominal sternum VIII, l.v.
<taxonomicName id="4C5A5892FFA8FFCF7AE5F824F1F8DEBC" box="[1166,1267,1958,1981]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="primus">
<emphasis id="B92EFF03FFA8FFCF7AE5F824F1F8DEBC" box="[1166,1267,1958,1981]" italics="true" pageId="17" pageNumber="18">T. primus</emphasis>
</taxonomicName>
: 49, abdominal sternum VIII, l.v.
<taxonomicName id="4C5A5892FFA8FFCF7F3EF844F4BADEDC" box="[341,433,1990,2013]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="puella">
<emphasis id="B92EFF03FFA8FFCF7F3EF844F4BADEDC" box="[341,433,1990,2013]" italics="true" pageId="17" pageNumber="18">T. puella</emphasis>
</taxonomicName>
: 50, abdominal sternum VIII,v.v.; 51–52, same, l.v.
</paragraph>
</caption>
<caption id="DF257399FFABFFCC7EFCFABCF1BEDC77" httpUri="https://zenodo.org/record/195440/files/figure.png" pageId="18" pageNumber="19" targetBox="[202,1387,197,1318]" targetPageId="18">
<paragraph id="8BE52311FFABFFCC7EFCFABCF1BEDC77" blockId="18.[151,1436,1342,1398]" pageId="18" pageNumber="19">
<emphasis id="B92EFF03FFABFFCC7EFCFABCF469DC57" bold="true" box="[151,354,1342,1366]" pageId="18" pageNumber="19">FIGURES 53–59.</emphasis>
<taxonomicName id="4C5A5892FFABFFCC7F02FABDF75EDC57" box="[361,597,1343,1366]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="limoncocha">
<emphasis id="B92EFF03FFABFFCC7F02FABDF75EDC57" box="[361,597,1343,1366]" italics="true" pageId="18" pageNumber="19">Tortopsis limoncocha</emphasis>
</taxonomicName>
. 53, male head, v.v.; 54, male mesofurcasternal plate; 55, female abdominal sternum VIII, l.v.; 56, same, v.v.; 57, male genitalia, v.v.; 58, same, l.v.; 59, detail of apex of penes.
</paragraph>
</caption>
<paragraph id="8BE52311FFABFFCC7EADFA1DF14FDF75" blockId="18.[151,1437,1439,2025]" pageId="18" pageNumber="19">
Female imago. Length (mm): body (shrunken abdomen), 10.5; fore wing, 16.0; hind wing, 6.9; cerci, 4.5. As in male, exceptions follow. Antennae with scape and pedicel brownish black. Thorax shaded more markedly with brownish gray. Legs yellowish white shaded brownish, turning lighter towards apex. Wings as male except usual sexual dimorphism (thicker veins). Abdomen yellowish strongly shaded brownish gray. Sternum VIII shaded strongly with black, lighter anteriorly; sockets blackish, relatively short and with more or less transverse opening (
<figureCitation id="13613F94FFABFFCC7FBDF9D8F727DF75" box="[470,556,1626,1652]" captionStart="FIGURES 10 – 13" captionStartId="10.[151,270,1119,1143]" captionTargetBox="[169,1402,195,1092]" captionTargetId="figure@10.[151,1436,175,1095]" captionTargetPageId="10" captionText="FIGURES 10 – 13. Tortopus bellus: 10, male genitalia, v. v.; 11, details of apex of penes; 12, detail of pedestal, v. v.; 13, sockets on female abdominal sternum VIII." httpUri="https://zenodo.org/record/195433/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
), anterior depressed area paler. Cerci whitish.
</paragraph>
<paragraph id="8BE52311FFABFFCD7EADF9FDF780D9D6" blockId="18.[151,1437,1439,2025]" lastBlockId="19.[151,1435,152,215]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Discussion and diagnosis. A short description of the male is given to complete the original description by
<bibRefCitation id="EFCB5EE0FFABFFCC7EFCF926F72FDFBF" author="Lugo-Ortiz" box="[151,548,1700,1726]" pageId="18" pageNumber="34" refString="Lugo-Ortiz, C. R. &amp; McCafferty W. P. (1996) Central American Tortopus (Ephemeroptera: Polymitarcyidae): a unique new species and new country records. Entomological News, 107, 23 - 27." type="journal article" year="1996">Lugo-Ortiz &amp; McCafferty (1996)</bibRefCitation>
with some new characters for the group. The female adult is described for the first time.
<taxonomicName id="4C5A5892FFABFFCC7F2AF948F70ADFE2" box="[321,513,1738,1763]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="B92EFF03FFABFFCC7F2AF948F70ADFE2" box="[321,513,1738,1763]" italics="true" pageId="18" pageNumber="19">Tortopus bellus</emphasis>
</taxonomicName>
can be distinguished from other species in the genus by: 1) fore wing length
<quantity id="4CA28EF4FFABFFCC7EFCF96DF449DE08" box="[151,322,1775,1801]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" metricValueMax="1.29" metricValueMin="1.05" pageId="18" pageNumber="19" unit="mm" value="11.7" valueMax="12.9" valueMin="10.5">10.5–12.9 mm</quantity>
(female 16.0 mm); 2) parastyli very short and pointed (
<figureCitation id="13613F94FFABFFCC7DBEF96DF151DE08" box="[981,1114,1775,1801]" captionStart="FIGURES 10 – 13" captionStartId="10.[151,270,1119,1143]" captionTargetBox="[169,1402,195,1092]" captionTargetId="figure@10.[151,1436,175,1095]" captionTargetPageId="10" captionText="FIGURES 10 – 13. Tortopus bellus: 10, male genitalia, v. v.; 11, details of apex of penes; 12, detail of pedestal, v. v.; 13, sockets on female abdominal sternum VIII." httpUri="https://zenodo.org/record/195433/files/figure.png" pageId="18" pageNumber="19">Fig. 10, 12</figureCitation>
); 3) ventral knob relatively small (
<figureCitation id="13613F94FFABFFCC7E8CF896F434DE2F" box="[231,319,1812,1838]" captionStart="FIGURES 10 – 13" captionStartId="10.[151,270,1119,1143]" captionTargetBox="[169,1402,195,1092]" captionTargetId="figure@10.[151,1436,175,1095]" captionTargetPageId="10" captionText="FIGURES 10 – 13. Tortopus bellus: 10, male genitalia, v. v.; 11, details of apex of penes; 12, detail of pedestal, v. v.; 13, sockets on female abdominal sternum VIII." httpUri="https://zenodo.org/record/195433/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
); 4) penes relatively wide (
<figureCitation id="13613F94FFABFFCC7CEAF896F7D2DE2F" box="[641,729,1812,1838]" captionStart="FIGURES 10 – 13" captionStartId="10.[151,270,1119,1143]" captionTargetBox="[169,1402,195,1092]" captionTargetId="figure@10.[151,1436,175,1095]" captionTargetPageId="10" captionText="FIGURES 10 – 13. Tortopus bellus: 10, male genitalia, v. v.; 11, details of apex of penes; 12, detail of pedestal, v. v.; 13, sockets on female abdominal sternum VIII." httpUri="https://zenodo.org/record/195433/files/figure.png" pageId="18" pageNumber="19">Fig. 10</figureCitation>
); 5) female with sockets on s. VIII relatively wide and with transverse opening (
<figureCitation id="13613F94FFABFFCC7FE8F8B8F4D1DE55" box="[387,474,1850,1876]" captionStart="FIGURES 10 – 13" captionStartId="10.[151,270,1119,1143]" captionTargetBox="[169,1402,195,1092]" captionTargetId="figure@10.[151,1436,175,1095]" captionTargetPageId="10" captionText="FIGURES 10 – 13. Tortopus bellus: 10, male genitalia, v. v.; 11, details of apex of penes; 12, detail of pedestal, v. v.; 13, sockets on female abdominal sternum VIII." httpUri="https://zenodo.org/record/195433/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
); and 6) a relatively dark species, dorsum of head almost entirely shaded black. Male genitalia of
<taxonomicName id="4C5A5892FFABFFCC7F07F8DDF728DE79" box="[364,547,1887,1912]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="B92EFF03FFABFFCC7F07F8DDF728DE79" box="[364,547,1887,1912]" italics="true" pageId="18" pageNumber="19">Tortopus bellus</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C5A5892FFABFFCC7C94F8DDF6CADE79" box="[767,961,1887,1912]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="harrisi">
<emphasis id="B92EFF03FFABFFCC7C94F8DDF6CADE79" box="[767,961,1887,1912]" italics="true" pageId="18" pageNumber="19">Tortopus harrisi</emphasis>
</taxonomicName>
, but in
<taxonomicName id="4C5A5892FFABFFCC7A75F8DDF188DE79" box="[1054,1155,1887,1912]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="B92EFF03FFABFFCC7A75F8DDF188DE79" box="[1054,1155,1887,1912]" italics="true" pageId="18" pageNumber="19">T. bellus</emphasis>
</taxonomicName>
the knob at the base of forceps is reduced, pedestals have a widely rounded posteromedial corner (opposite to parastylus), forceps are completely shaded black and penes are much wider. Further differences between both species include: a darker coloration in
<taxonomicName id="4C5A5892FFABFFCC7FE4F84DF4FDDEE9" box="[399,502,1999,2024]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="B92EFF03FFABFFCC7FE4F84DF4FDDEE9" box="[399,502,1999,2024]" italics="true" pageId="18" pageNumber="19">T. bellus</emphasis>
</taxonomicName>
with abdominal terga and sterna shaded strongly with brownish gray (sterna unmarked in
<taxonomicName id="4C5A5892FFAAFFCD7F52FF1AF4A0D9B0" box="[313,427,152,177]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="harrisi">
<emphasis id="B92EFF03FFAAFFCD7F52FF1AF4A0D9B0" box="[313,427,152,177]" italics="true" pageId="19" pageNumber="20">T. harrisi</emphasis>
</taxonomicName>
), and wing venation without short marginal intercalaries in MA and R regions (
<taxonomicName id="4C5A5892FFAAFFCD7BEDFF1AF5ECD9D7" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="harrisi">
<emphasis id="B92EFF03FFAAFFCD7BEDFF1AF5ECD9D7" italics="true" pageId="19" pageNumber="20">T. harrisi</emphasis>
</taxonomicName>
<typeStatus id="54E19DB3FFAAFFCD7E86FF3FF45FD9D6" box="[237,340,189,215]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
presents 3 of these veins).
</paragraph>
</subSubSection>
</treatment>
</document>
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