129 lines
16 KiB
XML
129 lines
16 KiB
XML
<document id="B4EB137DB91B280D767C51E322C39745" ID-DOI="10.5281/zenodo.195431" ID-GBIF-Dataset="2df8bd5a-e125-42aa-aa6f-ae3e78b83929" ID-ISSN="1175-5326" ID-Zenodo-Dep="195431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226955823" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2010" docId="03F39207FFA3FFC47EFCFD6FF62FDE53" docLanguage="en" docName="zt02481p036.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tortopsis puella Pictet, New combination" docType="treatment" docVersion="10" lastPageNumber="27" masterDocId="FFCAEA7FFFB9FFDE7E6BFF82F50BD901" masterDocTitle="A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="27" updateTime="1698238640945" updateUser="plazi">
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<mods:title id="896A2C3BD39CB441DFE58A14B6341866">A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
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<mods:namePart id="9C7F29DC5B8654BBFB2160EB354624C9">Molineri, Carlos</mods:namePart>
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<mods:title id="E1370F2661967E8EB3EA2078C7475A0B">Zootaxa</mods:title>
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<mods:date id="4FD79F2DFBB9D78AD370C9AE2DC013A7">2010</mods:date>
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<mods:number id="AB239CDEBCA278B0BE1780B8DE604C70">2481</mods:number>
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<mods:identifier id="543B94717F8749E4F0A2795F52CBA199" type="DOI">10.5281/zenodo.195431</mods:identifier>
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<treatment id="03F39207FFA3FFC47EFCFD6FF62FDE53" ID-DOI="http://doi.org/10.5281/zenodo.5690589" ID-GBIF-Taxon="119386670" ID-Zenodo-Dep="5690589" LSID="urn:lsid:plazi:treatment:03F39207FFA3FFC47EFCFD6FF62FDE53" httpUri="http://treatment.plazi.org/id/03F39207FFA3FFC47EFCFD6FF62FDE53" lastPageNumber="27" pageId="26" pageNumber="27">
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<subSubSection id="C340709AFFA3FFC47EFCFD6FF747DA4D" pageId="26" pageNumber="27" type="nomenclature">
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<paragraph id="8BE52311FFA3FFC47EFCFD6FF790DA06" blockId="26.[151,667,749,775]" box="[151,667,749,775]" pageId="26" pageNumber="27">
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<heading id="D0AD947DFFA3FFC47EFCFD6FF790DA06" bold="true" box="[151,667,749,775]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
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<emphasis id="B92EFF03FFA3FFC47EFCFD6FF790DA06" bold="true" box="[151,667,749,775]" pageId="26" pageNumber="27">
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<taxonomicName id="4C5A5892FFA3FFC47EFCFD6FF4B2DA06" ID-CoL="57H2H" authority="Pictet" authorityName="Pictet" box="[151,441,749,775]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="puella" status="comb. nov.">
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<emphasis id="B92EFF03FFA3FFC47EFCFD6FF45CDA06" bold="true" box="[151,343,749,775]" italics="true" pageId="26" pageNumber="27">Tortopsis puella</emphasis>
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(Pictet)
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</taxonomicName>
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<taxonomicNameLabel id="A21D4278FFA3FFC47FABFD6FF790DA06" box="[448,667,749,775]" pageId="26" pageNumber="27">New combination</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE52311FFA3FFC47EFCFCB7F747DA4D" blockId="26.[151,770,821,940]" box="[151,588,821,844]" pageId="26" pageNumber="27">
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<taxonomicName id="4C5A5892FFA3FFC47EFCFCB7F4FADA4D" authority="(Pictet), 1843" authorityName="Pictet" baseAuthorityName="Pictet" baseAuthorityYear="1843" box="[151,497,821,844]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="puella">
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<emphasis id="B92EFF03FFA3FFC47EFCFCB7F451DA4D" box="[151,346,821,844]" italics="true" pageId="26" pageNumber="27">Campsurus puella</emphasis>
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(
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<bibRefCitation id="EFCB5EE0FFA3FFC47F03FCB7F4BADA4D" author="Pictet" box="[360,433,821,844]" pageId="26" pageNumber="35" refString="Pictet, F. J. (1843 - 1845) Histoire naturelle generale et particuliere des insectes nevropteres. Famille des Ephemerines. Bailliere edit., Paris (Kessmann et Cherbuliez edit., aussi a Geneve), 300 pp." type="book" year="1843">Pictet)</bibRefCitation>
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, 1843
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</taxonomicName>
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(comb.)
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</paragraph>
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</subSubSection>
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<subSubSection id="C340709AFFA3FFC47EFCFCD7F7F2DAAD" pageId="26" pageNumber="27" type="reference_group">
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<paragraph id="8BE52311FFA3FFC47EFCFCD7F72EDA8D" blockId="26.[151,770,821,940]" pageId="26" pageNumber="27">
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<taxonomicName id="4C5A5892FFA3FFC47EFCFCD7F4B7DA6D" authority="Traver" authorityName="Traver" box="[151,444,853,876]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="incertus">
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<emphasis id="B92EFF03FFA3FFC47EFCFCD7F466DA6D" box="[151,365,853,876]" italics="true" pageId="26" pageNumber="27">Campsurus incertus</emphasis>
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Traver
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</taxonomicName>
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, in
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<bibRefCitation id="EFCB5EE0FFA3FFC47F8AFCD7F7B5DA6D" author="Needham" box="[481,702,853,876]" pageId="26" pageNumber="35" refString="Needham, J. G., Traver, J. R. & Hsu, Y. C. (1935) The Biology of Mayflies. Comstock Pub. Inc. New York, 759 pp." type="book" year="1935">Needham et al. 1935</bibRefCitation>
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(syn.)
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<taxonomicName id="4C5A5892FFA3FFC47EFCFCF7F4D0DA8D" authority="Pictet, 1843" authorityName="Pictet" authorityYear="1843" box="[151,475,885,908]" class="Insecta" family="Palingeniidae" genus="Palingenia" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="puella">
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<emphasis id="B92EFF03FFA3FFC47EFCFCF7F45EDA8D" box="[151,341,885,908]" italics="true" pageId="26" pageNumber="27">Palingenia puella</emphasis>
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<bibRefCitation id="EFCB5EE0FFA3FFC47F37FCF7F4D0DA8D" author="Pictet" box="[348,475,885,908]" pageId="26" pageNumber="35" refString="Pictet, F. J. (1843 - 1845) Histoire naturelle generale et particuliere des insectes nevropteres. Famille des Ephemerines. Bailliere edit., Paris (Kessmann et Cherbuliez edit., aussi a Geneve), 300 pp." type="book" year="1843">Pictet, 1843</bibRefCitation>
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</taxonomicName>
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(orig.)
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</paragraph>
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<paragraph id="8BE52311FFA3FFC47EFCFC17F7F2DAAD" blockId="26.[151,770,821,940]" box="[151,761,917,940]" pageId="26" pageNumber="27">
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<taxonomicName id="4C5A5892FFA3FFC47EFCFC17F4B9DAAD" authority="Traver" authorityName="Traver" box="[151,434,917,940]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="incertus">
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<emphasis id="B92EFF03FFA3FFC47EFCFC17F45ADAAD" box="[151,337,917,940]" italics="true" pageId="26" pageNumber="27">Tortopus incertus</emphasis>
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(Traver)
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</taxonomicName>
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, in
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<bibRefCitation id="EFCB5EE0FFA3FFC47FB3FC17F7BEDAAD" author="Needham" box="[472,693,917,940]" pageId="26" pageNumber="35" refString="Needham, J. G., Traver, J. R. & Hsu, Y. C. (1935) The Biology of Mayflies. Comstock Pub. Inc. New York, 759 pp." type="book" year="1935">Needham et al. 1935</bibRefCitation>
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(syn.)
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</paragraph>
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</subSubSection>
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<subSubSection id="C340709AFFA3FFC47EFCFC58F62FDE53" pageId="26" pageNumber="27" type="materials_examined">
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<paragraph id="8BE52311FFA3FFC47EFCFC58F4B0DD3E" blockId="26.[151,1437,986,1874]" pageId="26" pageNumber="27">
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Material (
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<collectionCode id="ED4BBBD4FFA3FFC47F67FC58F442DAF5" box="[268,329,986,1012]" httpUri="http://grbio.org/cool/f5t5-0jbj" name="Instituto Miguel Lillo" pageId="26" pageNumber="27">IML</collectionCode>
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):
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<specimenCount id="9D5CE898FFA3FFC47F36FC58F4A5DAF5" box="[349,430,986,1012]" pageId="26" pageNumber="27" type="male">3 male</specimenCount>
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imagos from
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<collectingCountry id="F34D6381FFA3FFC47C3EFC58F79EDAF5" box="[597,661,986,1012]" name="United States of America" pageId="26" pageNumber="27">USA</collectingCountry>
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: Florida, Liberty Co., Apalachicola R. at Hwy. 20, Bristol,
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<date id="FFE405D1FFA3FFC47B31FC58F5DFDD1B" pageId="26" pageNumber="27" value="1972-09-05">5-IX- 1972</date>
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, P.H. Carlson;
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<specimenCount id="9D5CE898FFA3FFC47FEAFB82F4EDDD1B" box="[385,486,1024,1050]" pageId="26" pageNumber="27" type="female">7 female</specimenCount>
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imagos same data except date
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<date id="FFE405D1FFA3FFC47D27FB82F6DBDD1B" box="[844,976,1024,1050]" pageId="26" pageNumber="27" value="1973-10-10">10-X-1973</date>
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;
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<specimenCount id="9D5CE898FFA3FFC47DB7FB82F121DD1B" box="[988,1066,1024,1050]" pageId="26" pageNumber="27" type="male">4 male</specimenCount>
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and
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<specimenCount id="9D5CE898FFA3FFC47A0AFB82F1CCDD1B" box="[1121,1223,1024,1050]" pageId="26" pageNumber="27" type="female">6 female</specimenCount>
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imagos same data but no date nor collector.
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</paragraph>
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<paragraph id="8BE52311FFA3FFC47EADFBC8F069DCB6" blockId="26.[151,1437,986,1874]" pageId="26" pageNumber="27">
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Female adult. Length (mm): body, 9.0–13.0; fore wing, 12.0–16.0; hind wing, 5.0–6.2; cerci, 5.0–6.0. Head: whitish around median ocellus, blackish between lateral ocelli, occiput with gray anastomosed marks mainly on lateral thirds. Thorax: pronotum with anterior ring widely shaded black, and posterior ring shaded grayish black on central area, lateral to this shading there is an isolated narrow gray mark, and also shaded near posterolateral corner. Wings: membrane whitish hyaline, veins whitish, except those near costa shaded gray; some females show a weak and short (1 cell) vein between R2+3 and IR, formed by the coalescence of 2 cross veins. Abdomen: terga with wide median band darker than the rest, except on tergum I whitish medially, and thin medial blackish line on terga III–IX. Sternum VIII with large sockets, the sclerotized anterior margin of the socket is somewhat sinuous (
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<figureCitation id="13613F94FFA3FFC47C51FAFAF7C3DC93" box="[570,712,1400,1426]" captionStart="FIGURES 44 – 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 – 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 – 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="26" pageNumber="27">Figs. 50–52</figureCitation>
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) and reaches the anterior margin of the sternum; anterior and posterior margins of socket are more or less parallel; the depression delimited by the socket is elongated.
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</paragraph>
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<paragraph id="8BE52311FFA3FFC47EADFA40F4F4DF03" blockId="26.[151,1437,986,1874]" pageId="26" pageNumber="27">Egg. Suboval, no attachment structures. Length, 450–490 µm; width, 370–390 µm. The chorion is sculptured with small grooves.</paragraph>
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<paragraph id="8BE52311FFA3FFC47EADF98FF62FDE53" blockId="26.[151,1437,986,1874]" pageId="26" pageNumber="27">
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Discussion and diagnosis. This species is known from adults of both sexes and nymphs from SE and SW
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<collectingCountry id="F34D6381FFA3FFC47EFCF9B0F5DDDF4D" box="[151,214,1586,1612]" name="United States of America" pageId="26" pageNumber="27">USA</collectingCountry>
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. A short description of the female adult is presented, enumerating those characters useful in separating the female adults of this species from those of other species of the genus. Male adults and nymphs were sufficiently treated elsewhere (
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<bibRefCitation id="EFCB5EE0FFA3FFC47F94F9FFF7FCDF96" author="Needham" box="[511,759,1661,1687]" pageId="26" pageNumber="35" refString="Needham, J. G., Traver, J. R. & Hsu, Y. C. (1935) The Biology of Mayflies. Comstock Pub. Inc. New York, 759 pp." type="book" year="1935">Needham et al. 1935</bibRefCitation>
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,
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<bibRefCitation id="EFCB5EE0FFA3FFC47D6FF9FFF6CDDF96" author="Scott" box="[772,966,1661,1687]" pageId="26" pageNumber="35" refString="Scott, D. C., Berner, L., Hirsch, A. (1959) The nymph of the mayfly genus Tortopus (Ephemeroptera: Polymitarcidae). Annals of the Entomological Society of America, 52, 205 - 213." type="journal article" year="1959">Scott et al. 1959</bibRefCitation>
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,
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<bibRefCitation id="EFCB5EE0FFA3FFC47DB8F9FFF1ABDF96" author="McCafferty" box="[979,1184,1661,1687]" pageId="26" pageNumber="34" refString="McCafferty, W. P. (1975) The burrowing mayflies (Ephemeroptera: Ephemeroidea) of the United States. Transactions of the American Entomological Society, 101, 447 - 504." type="journal article" year="1975">McCafferty 1975</bibRefCitation>
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,
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<bibRefCitation id="EFCB5EE0FFA3FFC47AC7F9FFF050DF96" author="Molineri" box="[1196,1371,1661,1687]" pageId="26" pageNumber="35" refString="Molineri, C. (2008) The larvae of the burrowing mayfly genus Tortopus (Ephemeroptera: Polymitarcyidae). Aquatic Insects, 30, 7 - 19." type="journal article" year="2008">Molineri 2008</bibRefCitation>
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). The adult stage of
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<taxonomicName id="4C5A5892FFA3FFC47F2CF921F705DFBD" box="[327,526,1699,1724]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="puella">
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<emphasis id="B92EFF03FFA3FFC47F2CF921F705DFBD" box="[327,526,1699,1724]" italics="true" pageId="26" pageNumber="27">Tortopsis puella</emphasis>
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</taxonomicName>
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can be separated by: 1) fore wing length 14.0 mm (male), 12.0–16.0 mm (female); 2) wing veins dark, mainly on basal half and costal margin; 3) parastyli moderately curved; 4) penes cylindrical; 5) female parastylus receptors large, elongated, with sinuous medial margin (
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<figureCitation id="13613F94FFA3FFC47ABCF96FF066DE06" box="[1239,1389,1773,1799]" captionStart="FIGURES 44 – 52" captionStartId="17.[151,274,1925,1949]" captionTargetBox="[285,1301,367,1892]" captionTargetId="figure@17.[285,1302,367,1904]" captionTargetPageId="17" captionText="FIGURES 44 – 52. Tortopsis, female imago. 44, T. obscuripennis, fore wing. T. sarae: 45, mesofurcasternum; 46, abdominal sternum VIII, v. v.; 47, same, l. v. T. obscuripennis: 48, abdominal sternum VIII, l. v. T. primus: 49, abdominal sternum VIII, l. v. T. puella: 50, abdominal sternum VIII, v. v.; 51 – 52, same, l. v." httpUri="https://zenodo.org/record/195439/files/figure.png" pageId="26" pageNumber="27">Figs. 50–52</figureCitation>
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); 6) female head not shaded black behind median ocellus; 7) female pronotum color pattern similar to other species, but with a gray band near posterolateral corner.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |