treatments-xml/data/03/E9/E6/03E9E636E25A5138948B09EAB972EE14.xml
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.225.99356" ID-Pensoft-Pub="1314-2003-225-115" ID-Pensoft-UUID="AB75E2E9609C5A7583F34B48EA61B976" ModsDocID="1314-2003-225-115" checkinTime="1682624511249" checkinUser="pensoft" docAuthor="Khamar, Hamid, Oualidi, Jalal El, Touhami, Amina Ouazzani &amp; Civeyrel, Laure" docDate="2023" docId="03E9E636E25A5138948B09EAB972EE14" docLanguage="en" docName="PhytoKeys 225: 115-152" docOrigin="PhytoKeys 225" docPubDate="2023-04-27" docSource="http://dx.doi.org/10.3897/phytokeys.225.99356" docTitle="Verbascum faurei subsp. acanthifolium (Pau) Benedí &amp; J. M. Monts., Lagascalia 20: 169. 1997." docType="treatment" docVersion="1" id="AB75E2E9609C5A7583F34B48EA61B976" lastPageNumber="115" masterDocId="AB75E2E9609C5A7583F34B48EA61B976" masterDocTitle="Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa" masterLastPageNumber="152" masterPageNumber="115" pageNumber="115" updateTime="1682624511249" updateUser="pensoft">
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<mods:title>Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Khamar, Hamid</mods:namePart>
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<mods:affiliation>Department of Botany and Plant Ecology, Scientific Institute, University Mohammed V in Rabat, B. P. 703, Rabat 10106, Morocco</mods:affiliation>
<mods:nameIdentifier type="email">khamarhamid90@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Oualidi, Jalal El</mods:namePart>
<mods:affiliation>Department of Botany and Plant Ecology, Scientific Institute, University Mohammed V in Rabat, B. P. 703, Rabat 10106, Morocco</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Touhami, Amina Ouazzani</mods:namePart>
<mods:affiliation>Plant, Animal Productions and Agro-industry Laboratory, Department of Biology, Faculty of Sciences, Ibn Tofail University, B. P. 133 14000, Kenitra, Morocco</mods:affiliation>
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<mods:namePart>Civeyrel, Laure</mods:namePart>
<mods:affiliation>Laboratoire Evolution &amp; Diversite Biologique (EDB UMR 5174), Universite de Toulouse, CNRS, IRD, UPS. 118 route de Narbonne, Bat 4 R 1, 31062 Toulouse cedex 9, France</mods:affiliation>
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<mods:date>2023</mods:date>
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<mods:number>2023-04-27</mods:number>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.225.99356</mods:identifier>
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<taxonomicName LSID="03E9E636-E25A-5138-948B-09EAB972EE14" authority="(Pau) Benedi &amp; J. M. Monts., Lagascalia 20: 169. 1997." authorityName="(Pau) Benedí &amp; J. M. Monts., Lagascalia 20: 169. 1997." class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Verbascum faurei subsp. acanthifolium" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="subSpecies" species="faurei" subSpecies="acanthifolium">
Verbascum faurei subsp. acanthifolium (Pau)
<normalizedToken originalValue="Benedí">Benedi</normalizedToken>
&amp; J.M.Monts., Lagascalia 20: 169. 1997.
</taxonomicName>
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<subSubSection pageId="0" pageNumber="115" type="reference_group">
<paragraph pageId="0" pageNumber="115">
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="acanthifolia">Celsia acanthifolia</taxonomicName>
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="acanthifolia">Celsia acanthifolia</taxonomicName>
Pau in Font Quer. Iter Maroc. n° 566. 1927. Type: [Morocco]. pr.
<normalizedToken originalValue="Badú">Badu</normalizedToken>
(Atlante rhiphaeo); Hab. in saxosis, solo schistose, 6 July 1927,
<emphasis italics="true" pageId="0" pageNumber="115">P. Font Quer, 566</emphasis>
(lectotype, selected by
<bibRefCitation author="Benedi, C" journalOrPublisher="Lagascalia" pageId="0" pageNumber="115" pagination="167 - 169" refId="B17" refString="Benedi, C, Montserrat, JM, 1997. Verbascum faurei (Murb.) Hub.-Mor. (Scrophulariaceae) y especies afines en el norte de Africa. Lagascalia 20 (1): 167 - 169" title="Verbascum faurei (Murb.) Hub. - Mor. (Scrophulariaceae) y especies afines en el norte de Africa." volume="20" year="1997">
<normalizedToken originalValue="Benedí">Benedi</normalizedToken>
and Montserrat 1997
</bibRefCitation>
, pg. 169, first step
<normalizedToken originalValue="“type”">&quot;type&quot;</normalizedToken>
, as &quot;
<emphasis italics="true" pageId="0" pageNumber="115">Font Quer 566-1927</emphasis>
&quot;; second step, designated here: BC [BC43694]!; isolectotypes: LD [LD1222710]!, GDA [GDA39155]!, G [G00015117]!, MPU [MPU009627, MPU009628]!, BCN [BCN18029]!, BM [BM000930561]!). [image available at http://psimg.jstor.org/fsi/img/pdf/i0/10.5555/al.ap.specimen.bc43694_normal.pdf]
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="acanthifolia">Celsia acanthifolia</taxonomicName>
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="acanthifolia">Celsia acanthifolia</taxonomicName>
Pau, in Cavanillesia 1: 47. 1928, nom. nud.
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia faurei" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="faurei">Celsia faurei</taxonomicName>
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia faurei subsp. var. var. acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="variety" species="faurei" subSpecies="var." variety="acanthifolia">Celsia faurei var. acanthifolia</taxonomicName>
(Pau) Maire in Jahand. &amp; Maire. Cat. PI. Maroc.3: 669 (1934).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="notes">
<paragraph pageId="0" pageNumber="115">Notes.</paragraph>
<paragraph pageId="0" pageNumber="115">
<bibRefCitation author="Font Quer, P" journalOrPublisher="Feddes Repertorium" pageId="0" pageNumber="115" refId="B38" refString="Font Quer, P, 1927. Iter Maroccanum. Exsiccata schedula. Natural Science Museum, Barcelona, N° 566." title="Iter Maroccanum. Exsiccata schedula. Natural Science Museum, Barcelona, N ° 566." year="1927">Font Quer (1927)</bibRefCitation>
based his species on a collection of plants from the Rif region that he has gathered during his 1927 campaign in Morocco. Font Quer gave the following diagnosis &quot;Folia Acanthi mollis L., superiora oblonga, sinuato-dentata, floribus longe pedunculatis, pedicellis glanduliferis, racemosis; bractae lanceolatae brevissimae, calycis laciniae lanceolatae, margine integro, capsula globosa, obtusa calyce triplo longior&quot; with the collection locality &quot;hab. in saxosis, pr.
<normalizedToken originalValue="Badù">Badu</normalizedToken>
(Atlante rhiphaeo), 1500 m. alt., solo schistoso, 6 julii&quot;. This protologue has been validly published by
<bibRefCitation author="Gonzalez-Bueno, A" journalOrPublisher="Treballs de l'Institut Botanic de Barcelona" pageId="0" pageNumber="115" pagination="5 - 173" refId="B40" refString="Gonzalez-Bueno, A, 1988. Les campanyes botaniques de Pius Font i Quer al Nord d'Àfrica. Treballs de l'Institut Botanic de Barcelona 12: 5 - 173" title="Les campanyes botaniques de Pius Font i Quer al Nord d'Àfrica." volume="12" year="1988">
<normalizedToken originalValue="Gonzâlez-Bueno">Gonzalez-Bueno</normalizedToken>
(1988)
</bibRefCitation>
. According to
<bibRefCitation author="Nualart, N" journalOrPublisher="17), Oxford University Press, New York" pageId="0" pageNumber="115" publicationUrl="http://hdl.handle.net/2445/117964" refId="B84" refString="Nualart, N, 2017. Els herbaris, fonts per al coneixement de la flora. Aplicacions en conservacio i taxonomia. PhD Thesis, University of Barcelona, Spain. http://hdl.handle.net/2445/117964" title="Els herbaris, fonts per al coneixement de la flora. Aplicacions en conservacio i taxonomia. PhD Thesis, University of Barcelona, Spain." url="http://hdl.handle.net/2445/117964" year="2017">Nualart (2017)</bibRefCitation>
,
<bibRefCitation author="Benedi, C" journalOrPublisher="Lagascalia" pageId="0" pageNumber="115" pagination="167 - 169" refId="B17" refString="Benedi, C, Montserrat, JM, 1997. Verbascum faurei (Murb.) Hub.-Mor. (Scrophulariaceae) y especies afines en el norte de Africa. Lagascalia 20 (1): 167 - 169" title="Verbascum faurei (Murb.) Hub. - Mor. (Scrophulariaceae) y especies afines en el norte de Africa." volume="20" year="1997">
<normalizedToken originalValue="Benedí">Benedi</normalizedToken>
and Montserrat (1997)
</bibRefCitation>
wrongly cited the collection number of the lectotype as
<normalizedToken originalValue="“BC436934”">&quot;BC436934&quot;</normalizedToken>
instead of
<normalizedToken originalValue="“BC43694”">&quot;BC43694&quot;</normalizedToken>
. So, in addition to the specimen kept at Herb. BC (BC43694), seven other specimens are in agreement with the description in the protologue. They are housed at Herbs LD (LD1222710), GDA (GDA39155), G (G00015117), MPU (MPU009627, MPU009628), BCN (BCN18029), and BM (BM000930561). According to Art. 9.6 of the ICN (
<bibRefCitation DOI="https://doi.org/10.12705/Code.2018" author="Turland, NJ" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="115" refId="B108" refString="Turland, NJ, Wiersema, JH, Barrie, FR, Greuter, W, Hawksworth, DL, Herendeen, PS, Knapp, S, Kusber, WH, Li, DZ, Marhold, K, May, TW, McNeill, J, Monro, AM, Prado, J, Price, MJ, Smith, GF [Eds], 2018. International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten. https://doi.org/10.12705/Code.2018" title="International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten." url="https://doi.org/10.12705/Code.2018" year="2018">Turland et al. 2018</bibRefCitation>
), all of these specimens and those persevered at Herb. BC must be treated as syntypes. The designation by
<bibRefCitation author="Benedi, C" journalOrPublisher="Lagascalia" pageId="0" pageNumber="115" pagination="167 - 169" refId="B17" refString="Benedi, C, Montserrat, JM, 1997. Verbascum faurei (Murb.) Hub.-Mor. (Scrophulariaceae) y especies afines en el norte de Africa. Lagascalia 20 (1): 167 - 169" title="Verbascum faurei (Murb.) Hub. - Mor. (Scrophulariaceae) y especies afines en el norte de Africa." volume="20" year="1997">
<normalizedToken originalValue="Benedí">Benedi</normalizedToken>
and Montserrat (1997)
</bibRefCitation>
can be considered here as a first-step typification (see Art. 9.17 of the ICN,
<bibRefCitation DOI="https://doi.org/10.12705/Code.2018" author="Turland, NJ" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="115" refId="B108" refString="Turland, NJ, Wiersema, JH, Barrie, FR, Greuter, W, Hawksworth, DL, Herendeen, PS, Knapp, S, Kusber, WH, Li, DZ, Marhold, K, May, TW, McNeill, J, Monro, AM, Prado, J, Price, MJ, Smith, GF [Eds], 2018. International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten. https://doi.org/10.12705/Code.2018" title="International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten." url="https://doi.org/10.12705/Code.2018" year="2018">Turland et al. 2018</bibRefCitation>
). As the herbarium specimen BC43694 [mounted on three sheets]) kept at Herb. BC is the one showing the best quality of preservation of morphological features described in the protologue, we select it here as second-step lectotype of the name.
</paragraph>
<paragraph pageId="0" pageNumber="115">
The taxonomic status of
<taxonomicName class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia acanthifolia" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="acanthifolia">
<emphasis italics="true" pageId="0" pageNumber="115">Celsia acanthifolia</emphasis>
</taxonomicName>
has been assessed by some authors. Maire (in
<bibRefCitation author="Jahandiez, E" journalOrPublisher="Archives des Sciences" pageId="0" pageNumber="115" refId="B56" refString="Jahandiez, E, Maire, R, 1934. Catalogue des plantes du Maroc (Vol. 3). Imprimerie Minerva, Algiers, 663-669." title="Catalogue des plantes du Maroc (Vol. 3). Imprimerie Minerva, Algiers, 663 - 669." year="1934">Jahandiez and Maire 1934</bibRefCitation>
) regarded it as a variety under
<taxonomicName genus="Ceslia" lsidName="Ceslia faurei" pageId="0" pageNumber="115" rank="species" species="faurei">
<emphasis italics="true" pageId="0" pageNumber="115">Ceslia faurei</emphasis>
</taxonomicName>
Murb. [
<taxonomicName family="Scrophulariaceae" kingdom="Plantae" lsidName="C. faurei subsp. var. var. acanthifolia" order="Lamiales" pageId="0" pageNumber="115" rank="variety" species="faurei" subSpecies="var." variety="acanthifolia">
<emphasis italics="true" pageId="0" pageNumber="115">C. faurei var. acanthifolia</emphasis>
</taxonomicName>
(Pau) Maire]. Later, Benedi and Montserrat (1997) raised it to the rank of subspecies and they published it as a new combination,
<taxonomicName class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Verbascum faurei subsp. acanthifolium" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="subSpecies" species="faurei" subSpecies="acanthifolium">Verbascum faurei (Murb.) Hub.-Morath subsp. acanthifolium</taxonomicName>
(Pau)
<normalizedToken originalValue="Benedí">Benedi</normalizedToken>
&amp; J.M.Monts.
</paragraph>
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