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<document ID-DOI="http://doi.org/10.5281/zenodo.3792110" ID-GBIF-Dataset="09827415-df1c-4129-9bc1-39321f933d5d" ID-GBIF-Taxon="163794395" ID-ISSN="13132970" ID-Zenodo-Dep="3792110" ID-ZooBank="531CF885-FED5-4B1B-806B-02C9E5051929" LSID="urn:lsid:zoobank.org:act:859FD1E9-018A-4EDC-8E67-B485CDAC3B77" checkinTime="1588009417727" checkinUser="plazi" docAuthor="Giupponi, Alessandro, de Vasconcelos, Eduardo &amp; Lourenco, Wilson" docDate="2009" docId="03DC0060FF893C7BDC8CFF15599DFD16" docLanguage="en" docName="ZK_article_1998.pdf" docOrigin="ZooKeys 13 (13)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@18c8ecf5" docStyleName="zookeys.2008.journal_article" docTitle="Ananteris bernabei Giupponi &amp; de Vasconcelos &amp; Lourenco 2009, sp. n." docType="treatment" docUuid="859FD1E9-018A-4EDC-8E67-B485CDAC3B77" docUuidSource="ZooBank" docVersion="9" lastPageId="11" lastPageNumber="39" masterDocId="FFE57818FF803C70DC00FFB35804FFC7" masterDocTitle="The genus Ananteris Thorell, 1891 (Scorpiones, Buthidae) in southeast Brazil, with the description of three new species" masterLastPageNumber="41" masterPageNumber="29" pageId="9" pageNumber="38" updateTime="1636545320677" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo>
<mods:title>The genus Ananteris Thorell, 1891 (Scorpiones, Buthidae) in southeast Brazil, with the description of three new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Giupponi, Alessandro</mods:namePart>
<mods:affiliation>,, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>de Vasconcelos, Eduardo</mods:namePart>
<mods:affiliation>Museu Nacional - UFRJ, Rio de Janeiro ,, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lourenco, Wilson</mods:namePart>
<mods:affiliation>Museum national d Histoire naturelle, Paris, France</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-07-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>13</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>13</mods:number>
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<mods:extent unit="page">
<mods:start>29</mods:start>
<mods:end>41</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.13.125</mods:identifier>
<mods:identifier type="GBIF-Dataset">09827415-df1c-4129-9bc1-39321f933d5d</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576464</mods:identifier>
<mods:identifier type="ZooBank">531CF885-FED5-4B1B-806B-02C9E5051929</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792110" ID-GBIF-Taxon="163794395" ID-Zenodo-Dep="3792110" LSID="urn:lsid:zoobank.org:act:859FD1E9-018A-4EDC-8E67-B485CDAC3B77" httpUri="http://treatment.plazi.org/id/03DC0060FF893C7BDC8CFF15599DFD16" lastPageId="11" lastPageNumber="39" pageId="9" pageNumber="38">
<subSubSection box="[140,440,166,193]" pageId="9" pageNumber="38" type="nomenclature">
<paragraph blockId="9.[140,991,166,263]" box="[140,440,166,193]" pageId="9" pageNumber="38">
<heading box="[140,440,166,193]" fontSize="11" level="2" pageId="9" pageNumber="38" reason="3">
<emphasis bold="true" box="[140,440,166,193]" pageId="9" pageNumber="38">
<taxonomicName authority="Giupponi &amp; de Vasconcelos &amp; Lourenco, 2009" authorityName="Giupponi &amp; de Vasconcelos &amp; Lourenco" authorityYear="2009" box="[140,367,166,193]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="bernabei" status="sp. n.">
<emphasis bold="true" box="[140,367,166,193]" italics="true" pageId="9" pageNumber="38">
<emphasis box="[140,255,166,192]" inLineHeading="true" pageId="9" pageNumber="38" reason="1">Ananteris</emphasis>
bernabei
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[375,440,167,193]" pageId="9" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="38" type="description">
<paragraph blockId="9.[140,991,166,263]" box="[140,991,201,228]" pageId="9" pageNumber="38">
<uri box="[140,991,201,228]" pageId="9" pageNumber="38">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph blockId="9.[140,991,166,263]" box="[140,260,237,263]" pageId="9" pageNumber="38">
<figureCitation box="[140,260,237,263]" captionStart-0="Figures 19-27" captionStart-1="Figure 28" captionStartId-0="10.[140,222,1395,1417]" captionStartId-1="7.[139,211,1576,1598]" captionTargetBox-0="[156,1094,165,1360]" captionTargetBox-1="[138,1108,164,1558]" captionTargetId-0="figure@10.[140,1108,165,1375]" captionTargetId-1="figure@7.[139,1108,165,1556]" captionTargetPageId-0="10" captionTargetPageId-1="7" captionText-0="Figures 19-27. Ananteris barnabei sp. n., male holotype. 19 Carapace dorsal view. 20 Pectines. 21 Chelicera, dorsal view. 22 Metasoma, segments IV-V and telson, lateral view. 23 Chela, dorsal view. 24 Manus, ventral view. 25 Pedipalpal femur, dorsal view. 26-27 Pedipalpal patella, dorsal and external view. Scale bars: 1mm." captionText-1="Figure 28. Map of distribution of the species of the genus Ananteris in Brazil." figureDoi-0="http://doi.org/10.5281/zenodo.3814819" figureDoi-1="http://doi.org/10.5281/zenodo.3814817" httpUri-0="https://zenodo.org/record/3814819/files/figure.png" httpUri-1="https://zenodo.org/record/3814817/files/figure.png" pageId="9" pageNumber="38">Figs 19-28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="38" type="materials_examined">
<paragraph blockId="9.[140,1108,307,1671]" pageId="9" pageNumber="38">
<emphasis bold="true" box="[140,303,307,333]" pageId="9" pageNumber="38">
<emphasis box="[140,198,307,333]" inLineHeading="true" pageId="9" pageNumber="38" reason="1">Type</emphasis>
material
</emphasis>
.
<materialsCitation ID-GBIF-Occurrence="2610746301" collectingDate="2006-04-20" collectionCode="T, MNRJ, REBIO" country="Brazil" location="Reserva Augusto Ruschi" municipality="Santa Tereza" pageId="9" pageNumber="38" specimenCount="4" specimenCount-female="2" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Espirito Santo" typeStatus="holotype">
<emphasis bold="true" box="[316,513,307,333]" pageId="9" pageNumber="38">
<specimenCount box="[316,400,307,333]" pageId="9" pageNumber="38" type="female">Female</specimenCount>
<typeStatus box="[407,513,307,333]" pageId="9" pageNumber="38">holotype</typeStatus>
</emphasis>
.
<collectingCountry box="[526,594,307,333]" name="Brazil" pageId="9" pageNumber="38">Brazil</collectingCountry>
, State of
<collectingRegion box="[695,859,307,333]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="38">Espírito Santo</collectingRegion>
,
<collectingMunicipality box="[869,1011,307,334]" pageId="9" pageNumber="38">Santa Tereza</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03DC0060FF893C7BDC8CFF15599DFD16:8EAAE7ADFF893C79DFFEFE87593AFEB7" country="Brazil" municipality="Santa Tereza" name="Reserva Augusto Ruschi" pageId="9" pageNumber="38" stateProvince="Espirito Santo">Reserva Augusto Ruschi</location>
(
<collectionCode box="[334,354,343,369]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="38">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:03DC0060FF893C7BDC8CFF15599DFD16:8EAAE7ADFF893C79DD6EFEE559CEFEB7" box="[366,458,342,368]" country="Brazil" municipality="Santa Tereza" name="Bernabe" pageId="9" pageNumber="38" stateProvince="Espirito Santo">Bernabe</location>
leg.), 2006.
<typeStatus box="[602,712,343,369]" pageId="9" pageNumber="38">Paratypes</typeStatus>
.
<specimenCount box="[723,799,342,369]" pageId="9" pageNumber="38" type="male">1 male</specimenCount>
(
<specimenCount box="[814,909,342,369]" pageId="9" pageNumber="38" type="juvenile">juvenile</specimenCount>
), same data as for the
<typeStatus box="[183,286,378,404]" pageId="9" pageNumber="38">holotype</typeStatus>
.
<location LSID="urn:lsid:plazi:treatment:03DC0060FF893C7BDC8CFF15599DFD16:8EAAE7ADFF893C79DD2AFEC95966FE53" box="[298,354,378,404]" country="Brazil" municipality="Santa Tereza" name="Both" pageId="9" pageNumber="38" stateProvince="Espirito Santo">Both</location>
deposited in the arachnological collection of the
<location LSID="urn:lsid:plazi:treatment:03DC0060FF893C7BDC8CFF15599DFD16:8EAAE7ADFF893C79DF92FEC95C4BFE53" box="[914,1103,378,404]" country="Brazil" municipality="Santa Tereza" name="Museu Nacional" pageId="9" pageNumber="38" stateProvince="Espirito Santo">Museu Nacional</location>
,
<collectingRegion box="[140,308,413,439]" country="Brazil" name="Rio de Janeiro" pageId="9" pageNumber="38">Rio de Janeiro</collectingRegion>
,
<collectingCountry box="[321,386,413,439]" name="Brazil" pageId="9" pageNumber="38">Brazil</collectingCountry>
(
<collectionCode box="[402,482,413,439]" collectionName="Brazil, Rio de Janeiro, Sao Cristovao, Universidade do Rio Janeiro, Museu Nacional" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="9" pageNumber="38">MNRJ</collectionCode>
11305).
<specimenCount box="[614,715,413,439]" pageId="9" pageNumber="38" type="female">1 female</specimenCount>
,
<collectingRegion box="[728,895,413,439]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="38">Espírito Santo</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03DC0060FF893C7BDC8CFF15599DFD16:8EAAE7ADFF893C79DF8CFE2E5C50FE70" box="[908,1108,413,439]" country="Brazil" municipality="Santa Tereza" name="Reserva Biologica" pageId="9" pageNumber="38" stateProvince="Espirito Santo">Reserva Biológica</location>
(
<collectionCode box="[148,238,448,474]" pageId="9" pageNumber="38">REBIO</collectionCode>
)
<location LSID="urn:lsid:plazi:treatment:03DC0060FF893C7BDC8CFF15599DFD16:8EAAE7ADFF893C79DD12FE735989FE1D" box="[274,397,448,474]" country="Brazil" municipality="Santa Tereza" name="Sooretama" pageId="9" pageNumber="38" stateProvince="Espirito Santo">Sooretama</location>
,
<location LSID="urn:lsid:plazi:treatment:03DC0060FF893C7BDC8CFF15599DFD16:8EAAE7ADFF893C79DD98FE735A64FE1D" box="[408,608,448,474]" country="Brazil" municipality="Santa Tereza" name="Porteira Quirinao" pageId="9" pageNumber="38" stateProvince="Espirito Santo">Porteira Quirinão</location>
(
<location LSID="urn:lsid:plazi:treatment:03DC0060FF893C7BDC8CFF15599DFD16:8EAAE7ADFF893C79DE6FFE735B0DFE1D" box="[623,777,448,474]" country="Brazil" municipality="Santa Tereza" name="Exp. Arachne" pageId="9" pageNumber="38" stateProvince="Espirito Santo">Exp. Arachné</location>
col.),
<date box="[848,988,448,474]" pageId="9" pageNumber="38" value="2006-04-20">
<collectingDate box="[848,988,448,474]" pageId="9" pageNumber="38" value="2006-04-20">20/IV/2006</collectingDate>
</date>
, deposited in the
<location LSID="urn:lsid:plazi:treatment:03DC0060FF893C7BDC8CFF15599DFD16:8EAAE7ADFF893C79DCD5FE50593DFE3A" box="[213,313,483,509]" country="Brazil" municipality="Santa Tereza" name="Museum" pageId="9" pageNumber="38" stateProvince="Espirito Santo">Muséum</location>
national dHistoire naturelle,
<location LSID="urn:lsid:plazi:treatment:03DC0060FF893C7BDC8CFF15599DFD16:8EAAE7ADFF893C79DE8EFE505ACCFE3A" box="[654,712,483,509]" country="Brazil" municipality="Santa Tereza" name="Paris" pageId="9" pageNumber="38" stateProvince="Espirito Santo">Paris</location>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="38" type="diagnosis">
<paragraph blockId="9.[140,1108,307,1671]" pageId="9" pageNumber="38">
<emphasis bold="true" box="[185,304,518,544]" inLineHeading="true" pageId="9" pageNumber="38" reason="1">Diagnosis</emphasis>
. Species of moderate to large size when compared with the average size of the other species of the genus (
<quantity box="[510,616,554,580]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="9" pageNumber="38" unit="mm" value="27.6">27.6 mm</quantity>
in total length; see
<tableCitation box="[832,912,554,580]" captionStart="Table I" captionStartId="8.[140,200,166,188]" captionTargetBox="[147,1074,211,1131]" captionTargetPageId="8" captionText="Table I. Morphometric values (in mm) of the new Ananteris species described in this paper" httpUri="http://table.plazi.org/id/DF0AE1FEFF883C78DC8CFF155BD8FF7C" pageId="9" pageNumber="38" tableUuid="DF0AE1FEFF883C78DC8CFF155BD8FF7C">Table I</tableCitation>
). General coloration yellow to reddish-yellow with variegated dark pigmentation. Pedipalps slender; fingers with 6-7 rows of granules; male and female pectines with 13-14 teeth. The new species can be distinguished from other known species of the genus, also distributed in the Atlantic Forest region of
<collectingCountry box="[463,531,694,720]" name="Brazil" pageId="9" pageNumber="38">Brazil</collectingCountry>
, by: (i) a paler, reddish-yellow coloration and a distinct pattern of pigmentation on the chelicerae; these are not pigmented in the central zone, (ii) pedipalp fingers with have 6-7 rows of granules, (iii) male and female pectines with 13-14 teeth, (iv) anterior margin of the carapace with a sharp and convex projection; this character is better marked in adults. Th e new species is possibly an endemic element to the Atlantic Forest region.
</paragraph>
</subSubSection>
<subSubSection box="[185,1026,906,932]" pageId="9" pageNumber="38" type="etymology">
<paragraph blockId="9.[140,1108,307,1671]" box="[185,1026,906,932]" pageId="9" pageNumber="38">
<emphasis bold="true" box="[185,320,906,932]" pageId="9" pageNumber="38">
<emphasis box="[185,315,906,932]" inLineHeading="true" pageId="9" pageNumber="38" reason="1">Etymology</emphasis>
.
</emphasis>
Patronym is in honor of biologist Tiago Nascimento Bernabé.
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="40" pageId="9" pageNumber="38" type="description">
<paragraph blockId="9.[140,1108,307,1671]" box="[185,941,941,967]" pageId="9" pageNumber="38">
<emphasis bold="true" box="[185,335,941,967]" pageId="9" pageNumber="38">
<emphasis box="[185,330,941,967]" inLineHeading="true" pageId="9" pageNumber="38" reason="1">Description</emphasis>
.
</emphasis>
Based on female
<typeStatus box="[533,632,941,967]" pageId="9" pageNumber="38">holotype</typeStatus>
(measurements in
<tableCitation box="[846,927,941,967]" captionStart="Table I" captionStartId="8.[140,200,166,188]" captionTargetBox="[147,1074,211,1131]" captionTargetPageId="8" captionText="Table I. Morphometric values (in mm) of the new Ananteris species described in this paper" httpUri="http://table.plazi.org/id/DF0AE1FEFF883C78DC8CFF155BD8FF7C" pageId="9" pageNumber="38" tableUuid="DF0AE1FEFF883C78DC8CFF155BD8FF7C">Table I</tableCitation>
).
</paragraph>
<paragraph blockId="9.[140,1108,307,1671]" pageId="9" pageNumber="38">
<emphasis bold="true" box="[185,324,976,1002]" pageId="9" pageNumber="38">
<emphasis box="[185,319,976,1002]" inLineHeading="true" pageId="9" pageNumber="38" reason="1">Coloration</emphasis>
.
</emphasis>
Generally yellow to reddish-yellow with dark brown to blackish pigmented zones on the body and its appendages. Prosoma: carapace reddish-yellow with blackish-brown spots on the anterior, lateral and posterior edges; eyes surrounded by black pigment. Mesosoma: reddish-yellow with confluent blackish zones on the posterior and lateral edges of tergites. Metasoma: segments I to III yellowish; IV-V reddish-yellow; all segments intensely marked with blackish-brown spots. Vesicle reddishyellow without any spots; the base of the aculeus reddish-yellow and the tip reddish. Venter yellowish; sternites with very diffused brownish spots. Chelicerae yellowish with variegated blackish-brown spots over almost the entire surface, with the exception of the central zone; fingers with blackish spots; teeth reddish. Pedipalps: yellowish; femur and patella with densely marked blackish-brown spots; chela hand almost entirely blackish-brown; fingers yellowish with dark on the extremities. Legs yellowish, with several blackish-brown spots.
</paragraph>
<paragraph blockId="9.[140,1108,307,1671]" pageId="9" pageNumber="38">Morphology. Carapace with coarse intense granulation; anterior margin not emarginated, almost straight, but with a sharp convex projection. Anterior median superciliary and posterior median carinae weak or absent. All furrows moderate to weak. Median ocular tubercle distinctly anterior to the centre of the carapace; median eyes separated by approximately 0.7 of one ocular diameter. Th ree pairs of lateral eyes. Sternum subpentagonal. Mesosoma: tergites with coarse intense granulation. Median carina moderate in all tergites. Tergite VII pentacarinate. Venter: genital operculum di-</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3814819" ID-Zenodo-Dep="3814819" httpUri="https://zenodo.org/record/3814819/files/figure.png" pageId="10" pageNumber="39" startId="10.[140,222,1395,1417]" targetBox="[156,1094,165,1360]" targetPageId="10">
<paragraph blockId="10.[140,1109,1395,1513]" pageId="10" pageNumber="39">
<emphasis bold="true" box="[140,299,1395,1417]" pageId="10" pageNumber="39">Figures 19-27.</emphasis>
<taxonomicName authority="Giupponi &amp; de Vasconcelos &amp; Lourenco, 2009" authorityName="Giupponi &amp; de Vasconcelos &amp; Lourenco" authorityYear="2009" box="[308,481,1395,1416]" class="Arachnida" family="Buthidae" genus="Ananteris" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="barnabei" status="sp. n.">
<emphasis box="[308,481,1395,1416]" italics="true" pageId="10" pageNumber="39">Ananteris barnabei</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[490,544,1396,1417]" pageId="10" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
, male
<typeStatus box="[611,695,1395,1417]" pageId="10" pageNumber="39">holotype</typeStatus>
.
<emphasis bold="true" box="[707,733,1395,1417]" pageId="10" pageNumber="39">1 9</emphasis>
Carapace dorsal view.
<emphasis bold="true" box="[956,982,1395,1417]" pageId="10" pageNumber="39">20</emphasis>
Pectines.
<emphasis bold="true" box="[1081,1107,1395,1417]" pageId="10" pageNumber="39">2 1</emphasis>
Chelicera, dorsal view.
<emphasis bold="true" box="[355,381,1427,1449]" pageId="10" pageNumber="39">22</emphasis>
Metasoma, segments IV-V and telson, lateral view.
<emphasis bold="true" box="[866,892,1427,1449]" pageId="10" pageNumber="39">23</emphasis>
Chela, dorsal view.
<emphasis bold="true" box="[1081,1107,1427,1449]" pageId="10" pageNumber="39">24</emphasis>
Manus, ventral view.
<emphasis bold="true" box="[335,361,1459,1481]" pageId="10" pageNumber="39">25</emphasis>
Pedipalpal femur, dorsal view.
<emphasis bold="true" box="[645,705,1459,1481]" pageId="10" pageNumber="39">26-27</emphasis>
Pedipalpal patella, dorsal and external view. Scale bars:
<quantity box="[242,294,1491,1512]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="39" unit="mm" value="1.0">1mm</quantity>
.
</paragraph>
</caption>
<paragraph blockId="10.[140,1108,1574,1671]" lastBlockId="11.[140,1108,166,721]" lastPageId="11" lastPageNumber="40" pageId="10" pageNumber="39">
vided longitudinally, each plate more or less sub-triangular in shape. Pectines: pectinal tooth count 14-
<quantity box="[322,383,1610,1636]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="10" pageNumber="39" unit="in" value="14.0">14 in</quantity>
<typeStatus box="[392,494,1610,1636]" pageId="10" pageNumber="39">holotype</typeStatus>
; 13-13 and 14-
<quantity box="[673,734,1610,1636]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="10" pageNumber="39" unit="in" value="14.0">14 in</quantity>
<typeStatus box="[742,851,1610,1636]" pageId="10" pageNumber="39">paratypes</typeStatus>
; basal middle lamellae of the pectines not dilated; fulcra absent. Sternites V to VII slightly granular; stigmata moderately elongate; setation moderate; sternite VII more intensely granulated and with four weakly marked carinae. Metasoma: segments I and II with 10 carinae, moderately crenulate. Segments III and IV with 8 carinae, moderately crenulate. Intercarinal spaces slightly granular, almost smooth. Segment V slightly rounded with 5 carinae. Telson moderately elongated, without granulations, smooth; with one ventral carina weakly marked; aculeus short and weakly curved; subaculear tooth strong and spinoid. Cheliceral dentition characteristic of the family
<taxonomicName authority="(Vachon 1963)" baseAuthorityName="Vachon" baseAuthorityYear="1963" box="[683,968,378,405]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="40" phylum="Arthropoda" rank="family">
Buthidae (
<bibRefCitation author="Vachon M" box="[801,959,378,405]" pageId="11" pageNumber="40" pagination="161 - 166" refId="ref5096" refString="Vachon M (1963) De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris 2 e ser., 35 (2): 161 - 166." type="journal article" year="1963">Vachon 1963</bibRefCitation>
)
</taxonomicName>
; fixed finger with two moderate basal teeth; movable finger with two weak basal teeth; ventral aspect of both finger and manus with dense, long setae. Pedipalps: femur pentacarinate; patella and chela with weak to vestigial carinae; internal face of patella with 4-5 minute spinoid granules; all faces slightly granular, almost smooth. Fixed and movable fingers with 6-7 almost linear rows of granules; two small external and one internal accessory granule present at the base of each row; three granules in the extremity of the fingers; Trichobothriotaxy; orthobothriotaxy A-ss-beta (
<bibRefCitation author="Vachon M" box="[672,828,624,650]" pageId="11" pageNumber="40" pagination="857 - 958" refId="ref5139" refString="Vachon M (1974) Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris, 3 e ser., n ° 140, Zool. 104: 857 - 958." type="journal article" year="1974">Vachon 1974</bibRefCitation>
,
<bibRefCitation author="Vachon M" box="[839,902,624,651]" pageId="11" pageNumber="40" pagination="1597 - 1599" refId="ref5204" refString="Vachon M (1975) Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des Scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus des seances de l'Academie des Sciences, Paris, ser. D, 281: 1597 - 1599." type="journal article" year="1975">1975</bibRefCitation>
). Legs: tarsus with very numerous fine median setae ventrally. Tibial spurs moderately to strongly developed on legs III and IV.
</paragraph>
</subSubSection>
</treatment>
</document>