311 lines
36 KiB
XML
311 lines
36 KiB
XML
<document id="533C8FC02335BCAA5FE9923AE996FFF9" ID-DOI="10.11646/zootaxa.3955.4.3" ID-GBIF-Dataset="ac08f47a-d395-406a-8461-b1a65d65fc38" ID-ISSN="1175-5326" ID-Zenodo-Dep="244417" ID-ZooBank="B4E21D99-203B-4818-904E-BBA145E5CD33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461052425946" checkinUser="plazi" docAuthor="Silva, Fernando A. B. & Vaz-De-Mello, Fernando" docDate="2015" docId="03D8893BB174CC332390FB6E48C6FA95" docLanguage="en" docName="zt03955p520.pdf" docOrigin="Zootaxa 3955 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aphengium cupreum Shipp 1897" docType="treatment" docVersion="8" lastPageNumber="510" masterDocId="FFE1F143B170CC362307FFA14A72FFA5" masterDocTitle="A revision of the genus Aphengium Harold, 1868 (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini)" masterLastPageNumber="520" masterPageNumber="505" pageNumber="509" updateTime="1698579553482" updateUser="plazi">
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<mods:mods id="38AC6D342546EE7DCBFED118C9998F2F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="32FDD5C117AEE7189CB23373B9B9E75B">
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<mods:title id="C9A64BF3736E13D0E0DA962D5B5CC1C3">A revision of the genus Aphengium Harold, 1868 (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini)</mods:title>
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<mods:name id="A0F747198F7B427FB0975751CA8ECE02" type="personal">
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<mods:namePart id="671BC644D764298592359D55B27684AB">Silva, Fernando A. B.</mods:namePart>
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<mods:name id="DAAA19147539D4C739DE9A6E53A01328" type="personal">
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<mods:roleTerm id="E1452B13F40250828CCF7425CFDB6153">Author</mods:roleTerm>
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<mods:namePart id="EA4B9605C927935D8ED84EE805CE5FD0">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:typeOfResource id="DC8F81FCD9F62002FE5E5E6DC770D2EE">text</mods:typeOfResource>
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<mods:title id="1EFF616BA88A424479ABAC5816B75DCC">Zootaxa</mods:title>
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<mods:part id="9F91A806E00E0324A416F0DFFB63DB4E">
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<mods:date id="64C3C9BB8D202D22E693C1AA04A185BD">2015</mods:date>
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<mods:detail id="D3B42BA47AADAC4B40CBCEA0DA12B7AA" type="volume">
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<mods:number id="9B44246AF5636930A36BA459B4C6DC90">3955</mods:number>
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<mods:number id="924715B97C0C9A414C2910C50F0718A9">4</mods:number>
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<mods:start id="5995909E2EC3A4CEDD704EE116AB3E2B">505</mods:start>
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<mods:identifier id="C2A2029616C814CED13371786DD874DD" type="DOI">10.11646/zootaxa.3955.4.3</mods:identifier>
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<mods:identifier id="2456AC9400221F19855F1C4953258739" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03D8893BB174CC332390FB6E48C6FA95" ID-DOI="http://doi.org/10.5281/zenodo.6122148" ID-GBIF-Taxon="119612288" ID-Zenodo-Dep="6122148" LSID="urn:lsid:plazi:treatment:03D8893BB174CC332390FB6E48C6FA95" httpUri="http://treatment.plazi.org/id/03D8893BB174CC332390FB6E48C6FA95" lastPageId="5" lastPageNumber="510" pageId="4" pageNumber="509">
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<subSubSection id="C36B6BA6B174CC322390FB6E4BA5FAA9" pageId="4" pageNumber="509" type="nomenclature">
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<paragraph id="8BCE382DB174CC322390FB6E4840FB4F" blockId="4.[151,562,1231,1293]" box="[151,562,1231,1258]" pageId="4" pageNumber="509">
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<heading id="D0868F41B174CC322390FB6E4840FB4F" bold="true" box="[151,562,1231,1258]" fontSize="11" level="1" pageId="4" pageNumber="509" reason="1">
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<taxonomicName id="4C7143AEB174CC322390FB6E4840FB4F" ID-CoL="FD4C" authority="Shipp 1897" authorityName="Shipp" authorityYear="1897" box="[151,562,1231,1258]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="cupreum">
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<emphasis id="B905E43FB174CC322390FB6E4840FB4F" bold="true" box="[151,562,1231,1258]" pageId="4" pageNumber="509">
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<emphasis id="B905E43FB174CC322390FB6E4BEFFB4C" bold="true" box="[151,413,1231,1257]" italics="true" pageId="4" pageNumber="509">Aphengium cupreum</emphasis>
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<bibRefCitation id="EFE045DCB174CC3222A3FB714840FB4F" author="Shipp" box="[420,562,1232,1258]" pageId="4" pageNumber="509" refString="Shipp, J. W. (1897) On three undescribed coprophagous lamellicorns in the Tring Museum. Novitates Zoologicae, 4, 185 - 187." type="journal article" year="1897">Shipp 1897</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BCE382DB174CC322390FB554BA5FAA9" blockId="4.[151,562,1231,1293]" box="[151,471,1268,1293]" pageId="4" pageNumber="509">
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(
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<figureCitation id="134A24A8B174CC322398FB554B51FAA9" box="[159,291,1268,1293]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 – 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="4" pageNumber="509">Figs 13–18</figureCitation>
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,
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<figureCitation id="134A24A8B174CC322228FB554B3CFAA9" box="[303,334,1268,1292]" captionStart="FIGURES 31 – 34" captionStartId="12.[151,264,1136,1158]" captionTargetBox="[165,1436,193,1114]" captionTargetId="figure@12.[150,1436,193,1115]" captionTargetPageId="12" captionText="FIGURES 31 – 34. Aedeagus of Aphengium; 31. A. sordidum (arrow shows dorsal margin of parameres curved inward at its apical third); 32. A. cupreum; 33. A. ibateguara sp. nov.; 34. A. curtum sp. nov." httpUri="https://zenodo.org/record/244423/files/figure.png" pageId="4" pageNumber="509">32</figureCitation>
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,
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<figureCitation id="134A24A8B174CC32225DFB554B0BFAA8" box="[346,377,1268,1293]" captionStart="FIGURES 35 – 36" captionStartId="13.[151,264,1955,1977]" captionTargetBox="[217,1390,211,1904]" captionTargetId="figure@13.[196,1390,195,1933]" captionTargetPageId="13" captionText="FIGURES 35 – 36. Superior right peripheral sclerite (SRP) of Aphengium; 35. A. sordidum; 36. A. cupreum; Figures 37 – 40. Fronto-lateral peripheral sclerite (FLP) of Aphengium; 37. A. sordidum; 38. A. cupreum; 39. A. ibateguara sp. nov.; 40. A. curtum sp. nov." httpUri="https://zenodo.org/record/244424/files/figure.png" pageId="4" pageNumber="509">36</figureCitation>
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, 38, 41)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C36B6BA6B174CC322390FA9D4973FAF7" box="[151,769,1340,1362]" pageId="4" pageNumber="509" type="reference_group">
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<paragraph id="8BCE382DB174CC322390FA9D4973FAF7" blockId="4.[151,769,1340,1362]" box="[151,769,1340,1362]" pageId="4" pageNumber="509">
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<treatmentCitationGroup id="AB611F03B174CC322390FA9D4973FAF7" box="[151,769,1340,1362]" pageId="4" pageNumber="509">
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<taxonomicName id="4C7143AEB174CC322390FA9D486CFAF7" ID-CoL="FD4C" authority="Shipp 1897: 187" authorityName="Shipp" authorityPageNumber="187" authorityYear="1897" box="[151,542,1340,1362]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="cupreum">
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<emphasis id="B905E43FB174CC322390FA9D4B1EFAF7" box="[151,364,1340,1362]" italics="true" pageId="4" pageNumber="509">Aphengium cupreum</emphasis>
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<treatmentCitation id="0AD01E3CB174CC322274FA9D486CFAF7" author="Shipp" box="[371,542,1340,1362]" page="187" pageId="4" pageNumber="509" year="1897">
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<bibRefCitation id="EFE045DCB174CC322274FA9D4B9FFAF7" author="Shipp" box="[371,493,1340,1362]" pageId="4" pageNumber="509" refString="Shipp, J. W. (1897) On three undescribed coprophagous lamellicorns in the Tring Museum. Novitates Zoologicae, 4, 185 - 187." type="journal article" year="1897">Shipp 1897</bibRefCitation>
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: 187
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</treatmentCitation>
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</taxonomicName>
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(original description)
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B6BA6B174CC322390FA25497FF9F5" pageId="4" pageNumber="509" type="diagnosis">
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<paragraph id="8BCE382DB174CC322390FA25497FF9F5" blockId="4.[151,1437,1412,2013]" pageId="4" pageNumber="509">
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<emphasis id="B905E43FB174CC322390FA254B7FFA38" bold="true" box="[151,269,1412,1437]" pageId="4" pageNumber="509">Diagnosis</emphasis>
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. Male of
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<taxonomicName id="4C7143AEB174CC322283FA254878FA39" box="[388,522,1412,1436]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="cupreum">
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<emphasis id="B905E43FB174CC322283FA254878FA39" box="[388,522,1412,1436]" italics="true" pageId="4" pageNumber="509">A. cupreum</emphasis>
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</taxonomicName>
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is readily distinguishable from all other
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<taxonomicName id="4C7143AEB174CC3220F7FA254E00FA39" box="[1008,1138,1412,1436]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="genus">
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<emphasis id="B905E43FB174CC3220F7FA254E00FA39" box="[1008,1138,1412,1436]" italics="true" pageId="4" pageNumber="509">Aphengium</emphasis>
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</taxonomicName>
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species by the following character: dorsal surface of head with foveiform punctures separated by distance equivalent to at least two times puncture diameter along most of midline (
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<figureCitation id="134A24A8B174CC322192FA6D489DFA40" box="[661,751,1484,1509]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 – 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="4" pageNumber="509">Fig. 15</figureCitation>
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). Furthermore, the aedeagus and fronto-lateral peripheral sclerite (FLP) are unique among
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<taxonomicName id="4C7143AEB174CC322117FA5148E0F9AD" box="[528,658,1520,1544]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="genus">
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<emphasis id="B905E43FB174CC322117FA5148E0F9AD" box="[528,658,1520,1544]" italics="true" pageId="4" pageNumber="509">Aphengium</emphasis>
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</taxonomicName>
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species (
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<figureCitation id="134A24A8B174CC3221FBFA4E492EF9AD" box="[764,860,1519,1544]" captionStart="FIGURES 31 – 34" captionStartId="12.[151,264,1136,1158]" captionTargetBox="[165,1436,193,1114]" captionTargetId="figure@12.[150,1436,193,1115]" captionTargetPageId="12" captionText="FIGURES 31 – 34. Aedeagus of Aphengium; 31. A. sordidum (arrow shows dorsal margin of parameres curved inward at its apical third); 32. A. cupreum; 33. A. ibateguara sp. nov.; 34. A. curtum sp. nov." httpUri="https://zenodo.org/record/244423/files/figure.png" pageId="4" pageNumber="509">Figs. 32</figureCitation>
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, 38). Female can be immediately distinguished by the combination of two characters: punctures on dorsal surface of head usually contiguous (
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<figureCitation id="134A24A8B174CC3227B5F9B54F7AF988" box="[1202,1288,1556,1581]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 – 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="4" pageNumber="509">Fig. 16</figureCitation>
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); central and anterior portions of head with rugose surface (
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<figureCitation id="134A24A8B174CC3221ADF996488DF9F5" box="[682,767,1591,1616]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 – 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="4" pageNumber="509">Fig. 16</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B6BA6B174CC3323C0F9FD4B9BFEB9" lastPageId="5" lastPageNumber="510" pageId="4" pageNumber="509" type="description">
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<paragraph id="8BCE382DB174CC3323C0F9FD4B9BFEB9" blockId="4.[151,1437,1412,2013]" lastBlockId="5.[151,1437,151,1328]" lastPageId="5" lastPageNumber="510" pageId="4" pageNumber="509">
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<emphasis id="B905E43FB174CC3223C0F9FD4B26F9D0" bold="true" box="[199,340,1628,1653]" pageId="4" pageNumber="509">Description</emphasis>
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.
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<emphasis id="B905E43FB174CC322262F9FD4BCEF9D0" bold="true" box="[357,444,1628,1653]" pageId="4" pageNumber="509">Length</emphasis>
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. Small (
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<quantity id="4C8995C8B174CC322124F9FD48F4F9D1" box="[547,646,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="4" pageNumber="509" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
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) (
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<figureCitation id="134A24A8B174CC3221A6F9FD4940F9D1" box="[673,818,1628,1653]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 – 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="4" pageNumber="509">Figs. 13, 14</figureCitation>
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||
).
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<emphasis id="B905E43FB174CC32204EF9FD49F8F9D0" bold="true" box="[841,906,1628,1653]" pageId="4" pageNumber="509">Head</emphasis>
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||
. Foveiform punctures dense, not contiguous, separated by at least two times puncture diameter (
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<figureCitation id="134A24A8B174CC3221FFF9DE4922F93D" box="[760,848,1663,1688]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 – 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="4" pageNumber="509">Fig. 15</figureCitation>
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); punctures often elliptical. Punctures on anterior portion, near to clypeal teeth, weakly impressed.
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<emphasis id="B905E43FB174CC3221C8F9054938F918" bold="true" box="[719,842,1700,1725]" pageId="4" pageNumber="509">Pronotum</emphasis>
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. Covered by foveiform, setose punctures; larger on lateral and posterior portions, not contiguous, usually separated by less than one puncture diameter (
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<figureCitation id="134A24A8B174CC322630F9664FFDF945" box="[1335,1423,1735,1760]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 – 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="4" pageNumber="509">Fig. 17</figureCitation>
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). Disc of pronotum slightly convex. Anterior angle of pronotum approximately 70°. Lateral angle greater than 90°; margin not explanate. Margin between lateral and posterior angles straight.
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<emphasis id="B905E43FB174CC322756F8AE4ED4F88D" bold="true" box="[1105,1190,1807,1832]" pageId="4" pageNumber="509">Elytra.</emphasis>
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Elongate-oval shape (
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<figureCitation id="134A24A8B174CC322398F8954B49F8E9" box="[159,315,1844,1869]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 – 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="4" pageNumber="509">Figs. 13, 14</figureCitation>
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). Disc slightly convex, with dense and conspicuous setae. Lateral margins slightly curved outwards. Foveiform punctures separated on the elytral disc by distance equivalent to at least two times puncture diameter.
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<emphasis id="B905E43FB174CC32220FF8DD481EF830" bold="true" box="[264,620,1916,1941]" pageId="4" pageNumber="509">Secondary sexual characters.</emphasis>
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Besides the sexual dimorphism described for the genus, females differ from males in the following aspects: punctures on dorsal surface of head usually contiguous (
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<figureCitation id="134A24A8B174CC3227DDF8014F40F81C" box="[1242,1330,1952,1977]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 – 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="4" pageNumber="509">Fig. 16</figureCitation>
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). Central and anterior portions of head with rugose surface. Punctures on central portion of pronotum small and weakly impressed (
|
||
<figureCitation id="134A24A8B175CC332221FF364BF7FF15" box="[294,389,151,176]" captionStart="FIGURES 13 – 18" captionStartId="9.[151,264,1955,1977]" captionTargetBox="[183,1402,209,1920]" captionTargetId="figure@9.[164,1422,193,1934]" captionTargetPageId="9" captionText="FIGURES 13 – 18. Aphengium cupreum Shipp; 13. Male (dorsal view); 14. Female (dorsal view); 15. Male head; 16. Female head; 17. Male pronotum; 18. Female pronotum." httpUri="https://zenodo.org/record/244420/files/figure.png" pageId="5" pageNumber="510">Fig. 18</figureCitation>
|
||
); punctures more dispersed than those in males.
|
||
<emphasis id="B905E43FB175CC33270DFF364EF5FF15" bold="true" box="[1034,1159,151,176]" pageId="5" pageNumber="510">Aedeagus.</emphasis>
|
||
Length of parameres approximately one-half that of phallobase; ventral margin slightly curved inward; dorsal margin almost straight; apex acuminate (
|
||
<figureCitation id="134A24A8B175CC332264FF7E4BCCFF5D" box="[355,446,223,248]" captionStart="FIGURES 31 – 34" captionStartId="12.[151,264,1136,1158]" captionTargetBox="[165,1436,193,1114]" captionTargetId="figure@12.[150,1436,193,1115]" captionTargetPageId="12" captionText="FIGURES 31 – 34. Aedeagus of Aphengium; 31. A. sordidum (arrow shows dorsal margin of parameres curved inward at its apical third); 32. A. cupreum; 33. A. ibateguara sp. nov.; 34. A. curtum sp. nov." httpUri="https://zenodo.org/record/244423/files/figure.png" pageId="5" pageNumber="510">Fig. 32</figureCitation>
|
||
). Superior right peripheral sclerite (SRP) as in figure 36. Fronto-lateral peripheral sclerite (FLP) as in figure 38.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6BA6B175CC3323C0FE864EFCFB3B" pageId="5" pageNumber="510" type="materials_examined">
|
||
<paragraph id="8BCE382DB175CC3323C0FE864E66FEC1" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="510">
|
||
<emphasis id="B905E43FB175CC3323C0FE864BE3FEE5" bold="true" box="[199,401,295,320]" pageId="5" pageNumber="510">Material studied</emphasis>
|
||
.
|
||
<emphasis id="B905E43FB175CC3322A7FE86483AFEE5" bold="true" box="[416,584,295,320]" pageId="5" pageNumber="510">
|
||
<typeStatus id="54CA868FB175CC3322A7FE864BAEFEE5" box="[416,476,295,320]" pageId="5" pageNumber="510">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B905E43FB175CC33215FFE8648CEFEE5" bold="true" box="[600,700,295,320]" pageId="5" pageNumber="510">
|
||
<typeStatus id="54CA868FB175CC33215FFE8648CEFEE5" box="[600,700,295,320]" pageId="5" pageNumber="510" type="neotype">Neotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="ED60A0E8B175CC3321CCFE894950FEE5" box="[715,802,296,320]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="5" pageNumber="510">NHML</collectionCode>
|
||
) [
|
||
<emphasis id="B905E43FB175CC33203CFE864988FEE5" bold="true" box="[827,1018,295,320]" pageId="5" pageNumber="510">here designated</emphasis>
|
||
]: ♂.
|
||
<collectingCountry id="F36678BDB175CC332731FE864ED1FEE5" box="[1078,1187,295,320]" name="Brazil" pageId="5" pageNumber="510">
|
||
<emphasis id="B905E43FB175CC332731FE864ED1FEE5" bold="true" box="[1078,1187,295,320]" pageId="5" pageNumber="510">BRAZIL</emphasis>
|
||
</collectingCountry>
|
||
: ESPÍRITO
|
||
<collectionCode id="ED60A0E8B175CC332631FE894FEBFEE5" box="[1334,1433,296,320]" pageId="5" pageNumber="510">SANTO</collectionCode>
|
||
: Linhares, Reserva Natural Vale,
|
||
<date id="FFCF1EEDB175CC33210BFEED48A3FEC0" box="[524,721,332,357]" pageId="5" pageNumber="510" value="2012-08-04" valueMax="2012-08-06" valueMin="2012-08-04">04–06/VIII/2012</date>
|
||
. R.C. Lima col (ex-CEMT).
|
||
</paragraph>
|
||
<paragraph id="8BCE382DB175CC3323C0FECE4F1AFD99" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="510">
|
||
<emphasis id="B905E43FB175CC3323C0FECE4B8BFE2D" bold="true" box="[199,505,367,392]" pageId="5" pageNumber="510">Non-type material [38]</emphasis>
|
||
.
|
||
<collectingCountry id="F36678BDB175CC332112FECE48F0FE2D" box="[533,642,367,392]" name="Brazil" pageId="5" pageNumber="510">
|
||
<emphasis id="B905E43FB175CC332112FECE48F0FE2D" bold="true" box="[533,642,367,392]" pageId="5" pageNumber="510">BRAZIL</emphasis>
|
||
</collectingCountry>
|
||
: ESPÍRITO
|
||
<collectionCode id="ED60A0E8B175CC332036FED149E6FE2D" box="[817,916,368,392]" pageId="5" pageNumber="510">SANTO</collectionCode>
|
||
: Cachoeira de Itapemirim, [
|
||
<geoCoordinate id="EE455EEAB175CC33260FFED14FEAFE2D" box="[1288,1432,368,392]" direction="west" orientation="longitude" pageId="5" pageNumber="510" precision="15" value="-41.120277">41°07'13"W</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE455EEAB175CC332390FE354B6CFE08" box="[151,286,404,429]" direction="south" orientation="latitude" pageId="5" pageNumber="510" precision="15" value="-20.85">20°51'00"S</geoCoordinate>
|
||
],
|
||
<date id="FFCF1EEDB175CC332235FE354BFBFE09" box="[306,393,404,428]" pageId="5" pageNumber="510" value="1997-02">II.1997</date>
|
||
, J. Louzada (1♀
|
||
<collectionCode id="ED60A0E8B175CC332156FE3548D7FE09" box="[593,677,404,428]" pageId="5" pageNumber="510">CEMT</collectionCode>
|
||
); Castelo, [
|
||
<geoCoordinate id="EE455EEAB175CC33202FFE3549C5FE09" box="[808,951,404,428]" direction="west" orientation="longitude" pageId="5" pageNumber="510" precision="15" value="-41.203613">41°12'13"W</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE455EEAB175CC3320C3FE354E39FE08" box="[964,1099,404,429]" direction="south" orientation="latitude" pageId="5" pageNumber="510" precision="15" value="-20.603611">20°36'13"S</geoCoordinate>
|
||
],
|
||
<date id="FFCF1EEDB175CC332758FE354EB8FE08" box="[1119,1226,404,429]" pageId="5" pageNumber="510" value="1996-12">XII.1996</date>
|
||
, J. N. C. Louzada (
|
||
<specimenCount id="9D77F3A4B175CC3323A7FE194AB1FE75" box="[160,195,440,464]" pageId="5" pageNumber="510" type="generic">9♂</specimenCount>
|
||
11♀
|
||
<collectionCode id="ED60A0E8B175CC332213FE194B17FE75" box="[276,357,440,464]" pageId="5" pageNumber="510">CEMT</collectionCode>
|
||
;
|
||
<specimenCount id="9D77F3A4B175CC33227EFE194BD9FE6A" box="[377,427,440,464]" pageId="5" pageNumber="510" type="generic">1 ♂</specimenCount>
|
||
1 ♀
|
||
<collectionCode id="ED60A0E8B175CC3322F9FE19482EFE75" box="[510,604,440,464]" pageId="5" pageNumber="510">CERPE</collectionCode>
|
||
); Linhares, Faz. Lagoa do Macuco, prim. lowland Atlantic Forest, (
|
||
<geoCoordinate id="EE455EEAB175CC332398FE7D4B5FFE50" box="[159,301,476,501]" direction="west" orientation="longitude" pageId="5" pageNumber="510" precision="15" value="-39.97861">39°58'43"W</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE455EEAB175CC33223EFE7D4BCDFE50" box="[313,447,476,501]" direction="south" orientation="latitude" pageId="5" pageNumber="510" precision="15" value="-19.063889">19°03'50"S</geoCoordinate>
|
||
),
|
||
<date id="FFCF1EEDB175CC3322D4FE7D4836FE51" box="[467,580,476,500]" pageId="5" pageNumber="510" value="2000-01-27">27.I.2000</date>
|
||
, pitfall, F. Génier and S. Ide (
|
||
<specimenCount id="9D77F3A4B175CC33209DFE7D49CFFE51" box="[922,957,476,500]" pageId="5" pageNumber="510" type="generic">1♂</specimenCount>
|
||
2♀
|
||
<collectionCode id="ED60A0E8B175CC3320F7FE7D4E36FE51" box="[1008,1092,476,500]" pageId="5" pageNumber="510">CEMT</collectionCode>
|
||
); same, Reserva Natural Vale, [
|
||
<geoCoordinate id="EE455EEAB175CC332399FE5E4B5FFDBD" box="[158,301,511,536]" direction="west" orientation="longitude" pageId="5" pageNumber="510" precision="15" value="-40.06389">40°03'50"W</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE455EEAB175CC33223CFDA14BB0FDBD" box="[315,450,512,536]" direction="south" orientation="latitude" pageId="5" pageNumber="510" precision="15" value="-19.141111">19°08'28"S</geoCoordinate>
|
||
],
|
||
<date id="FFCF1EEDB175CC3322D0FDA1483CFDBD" box="[471,590,512,536]" pageId="5" pageNumber="510" value="2012-06-09">9.VI.2012</date>
|
||
, pitfall pig faeces, R. C. Lima (
|
||
<specimenCount id="9D77F3A4B175CC3320C2FE5E499BFDBD" box="[965,1001,511,536]" pageId="5" pageNumber="510" type="generic">5♂</specimenCount>
|
||
5♀
|
||
<collectionCode id="ED60A0E8B175CC332718FDA14E01FDBD" box="[1055,1139,512,536]" pageId="5" pageNumber="510">CEMT</collectionCode>
|
||
); same,
|
||
<date id="FFCF1EEDB175CC3327D3FE5E4FEAFDBD" box="[1236,1432,511,536]" pageId="5" pageNumber="510" value="2012-08-04" valueMax="2012-08-06" valueMin="2012-08-04">04–06.VIII.2012</date>
|
||
, (1♀
|
||
<collectionCode id="ED60A0E8B175CC3323CCFD854B6DFD99" box="[203,287,548,572]" pageId="5" pageNumber="510">CEMT</collectionCode>
|
||
);
|
||
<collectionCode id="ED60A0E8B175CC332234FD854BFDFD99" box="[307,399,548,572]" pageId="5" pageNumber="510">MINAS</collectionCode>
|
||
GERAIS: Açucena, [
|
||
<geoCoordinate id="EE455EEAB175CC332182FD854967FD99" box="[645,789,548,572]" direction="west" orientation="longitude" pageId="5" pageNumber="510" precision="15" value="-42.542778">42°32'34"W</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE455EEAB175CC332026FD8549DAFD99" box="[801,936,548,572]" direction="south" orientation="latitude" pageId="5" pageNumber="510" precision="15" value="-19.074167">19°04'27"S</geoCoordinate>
|
||
],
|
||
<date id="FFCF1EEDB175CC3320BCFD854E60FD98" box="[955,1042,548,573]" pageId="5" pageNumber="510" value="1952-02">II.1952</date>
|
||
, A. B. Machado (1♀
|
||
<collectionCode id="ED60A0E8B175CC33260FFD854F2FFD99" box="[1288,1373,548,572]" pageId="5" pageNumber="510">CEMT</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCE382DB175CC3323C0FDE64850FB90" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="510">
|
||
<emphasis id="B905E43FB175CC3323C0FDE64BEEFDC5" bold="true" box="[199,412,583,608]" pageId="5" pageNumber="510">Taxonomic notes.</emphasis>
|
||
The
|
||
<typeStatus id="54CA868FB175CC3322DEFDE94878FDC5" box="[473,522,584,608]" pageId="5" pageNumber="510">type</typeStatus>
|
||
of this species as described in the original description (
|
||
<bibRefCitation id="EFE045DCB175CC332793FDE64F50FDC5" author="Shipp" box="[1172,1314,583,608]" pageId="5" pageNumber="510" refString="Shipp, J. W. (1897) On three undescribed coprophagous lamellicorns in the Tring Museum. Novitates Zoologicae, 4, 185 - 187." type="journal article" year="1897">Shipp, 1897</bibRefCitation>
|
||
) is said to be in the Tring Museum. The Tring Museum insect collection is now deposited in
|
||
<collectionCode id="ED60A0E8B175CC33276CFDCC4ECDFD21" box="[1131,1215,621,644]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="5" pageNumber="510">NHML</collectionCode>
|
||
(M. Barclay, pers. com.), but no specimen of
|
||
<taxonomicName id="4C7143AEB175CC3322D6FD314824FD0D" box="[465,598,656,680]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="cupreum">
|
||
<emphasis id="B905E43FB175CC3322D6FD314824FD0D" box="[465,598,656,680]" italics="true" pageId="5" pageNumber="510">A. cupreum</emphasis>
|
||
</taxonomicName>
|
||
was found in that latter institution that could be considered a
|
||
<typeStatus id="54CA868FB175CC332629FD314F11FD0D" box="[1326,1379,656,680]" pageId="5" pageNumber="510">type</typeStatus>
|
||
. Nor has any specimen identified under this name that could have come from either Tring or Shipp been found in the Oxford University Museum of Natural History, or in any other consulted museums. The
|
||
<typeStatus id="54CA868FB175CC3327A7FD794EA3FD55" box="[1184,1233,728,752]" pageId="5" pageNumber="510">type</typeStatus>
|
||
specimen is then considered by us to be irretrievably lost (
|
||
<collectionCode id="ED60A0E8B175CC332185FD5D48B5FCB1" box="[642,711,764,788]" pageId="5" pageNumber="510">ICZN</collectionCode>
|
||
article 75.3.4). The original description mentions differences between this species and the older
|
||
<taxonomicName id="4C7143AEB175CC332121FC8148C3FC9D" box="[550,689,800,824]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="sordidum">
|
||
<emphasis id="B905E43FB175CC332121FC8148C3FC9D" box="[550,689,800,824]" italics="true" pageId="5" pageNumber="510">A. sordidum</emphasis>
|
||
</taxonomicName>
|
||
that are of secondary sexual nature (e.g clypeal teeth shorter and thorax broader than
|
||
<taxonomicName id="4C7143AEB175CC332286FCE5487FFCF9" box="[385,525,836,860]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="sordidum">
|
||
<emphasis id="B905E43FB175CC332286FCE5487FFCF9" box="[385,525,836,860]" italics="true" pageId="5" pageNumber="510">A. sordidum</emphasis>
|
||
</taxonomicName>
|
||
), making this species hard to characterize from the original description except for the
|
||
<typeStatus id="54CA868FB175CC3323ECFCC94B6FFC25" box="[235,285,872,896]" pageId="5" pageNumber="510">type</typeStatus>
|
||
locality given there. Hence, the exceptional need to clarify its identity (
|
||
<collectionCode id="ED60A0E8B175CC332758FCC94ED1FC25" box="[1119,1187,872,896]" pageId="5" pageNumber="510">ICZN</collectionCode>
|
||
article 75.3.1). Since no name-bearing
|
||
<typeStatus id="54CA868FB175CC332261FC2C4BE5FC00" box="[358,407,909,933]" pageId="5" pageNumber="510">type</typeStatus>
|
||
specimen is believed to be extant and a name-bearing
|
||
<typeStatus id="54CA868FB175CC33272BFC2C4E2CFC00" box="[1068,1118,909,933]" pageId="5" pageNumber="510">type</typeStatus>
|
||
is necessary to define this species objectively, we herein designate a
|
||
<typeStatus id="54CA868FB175CC33217CFC1148A8FC6D" box="[635,730,944,968]" pageId="5" pageNumber="510" type="neotype">neotype</typeStatus>
|
||
. So, in order to maintain the stability of nomenclature, a male from Reserva Natural Vale, Linhares, Espírito Santo,
|
||
<collectingCountry id="F36678BDB175CC33201EFC754910FC48" box="[793,866,980,1005]" name="Brazil" pageId="5" pageNumber="510">Brazil</collectingCountry>
|
||
,
|
||
<date id="FFCF1EEDB175CC332075FC754E45FC48" box="[882,1079,980,1005]" pageId="5" pageNumber="510" value="2012-08-04" valueMax="2012-08-06" valueMin="2012-08-04">04–06/VIII/2012</date>
|
||
, R.C. Lima leg (
|
||
<collectionCode id="ED60A0E8B175CC332603FC754F3BFC49" box="[1284,1353,980,1004]" pageId="5" pageNumber="510">ICNZ</collectionCode>
|
||
article 75.3.3), coming from a locality described as the original
|
||
<typeStatus id="54CA868FB175CC332057FC5949F3FBB5" box="[848,897,1016,1040]" pageId="5" pageNumber="510">type</typeStatus>
|
||
locality of this species (
|
||
<collectionCode id="ED60A0E8B175CC3327A1FC594E99FBB5" box="[1190,1259,1016,1040]" pageId="5" pageNumber="510">ICZN</collectionCode>
|
||
article 75.3.6), and
|
||
<emphasis id="B905E43FB175CC3323C0FBBD4850FB90" bold="true" box="[199,546,1052,1077]" pageId="5" pageNumber="510">corresponding to its original</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCE382DB175CC332390FBE14EFCFB3B" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="510">
|
||
<emphasis id="B905E43FB175CC332390FBE14B61FBF3" bold="true" box="[151,275,1088,1110]" pageId="5" pageNumber="510">description</emphasis>
|
||
(see
|
||
<bibRefCitation id="EFE045DCB175CC33224BFBE14BBDFBF3" author="Shipp" box="[332,463,1088,1110]" pageId="5" pageNumber="510" refString="Shipp, J. W. (1897) On three undescribed coprophagous lamellicorns in the Tring Museum. Novitates Zoologicae, 4, 185 - 187." type="journal article" year="1897">Shipp, 1897</bibRefCitation>
|
||
;
|
||
<collectionCode id="ED60A0E8B175CC3322DCFBE14865FBF3" box="[475,535,1088,1110]" pageId="5" pageNumber="510">ICNZ</collectionCode>
|
||
articles 75.3.2 and 75.3.5) was chosen to be the
|
||
<typeStatus id="54CA868FB175CC332719FBE04E01FBF3" box="[1054,1139,1089,1110]" pageId="5" pageNumber="510" type="neotype">neotype</typeStatus>
|
||
, being here deposited in the institution where the original
|
||
<typeStatus id="54CA868FB175CC3322C9FBC44B88FBDF" box="[462,506,1125,1146]" pageId="5" pageNumber="510">type</typeStatus>
|
||
specimen was alleged to be (
|
||
<collectionCode id="ED60A0E8B175CC332028FBC549F2FBDF" box="[815,896,1124,1146]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="5" pageNumber="510">NHML</collectionCode>
|
||
), a recognized scientific institution that maintains a research collection with proper facilities for preserving name-bearing
|
||
<typeStatus id="54CA868FB175CC33206AFB2849D6FB3B" box="[877,932,1161,1182]" pageId="5" pageNumber="510">types</typeStatus>
|
||
(
|
||
<collectionCode id="ED60A0E8B175CC3320B5FB294982FB3B" box="[946,1008,1160,1182]" pageId="5" pageNumber="510">ICZN</collectionCode>
|
||
article 75.3.7).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B6BA6B175CC3323C0FB0D48C6FA95" pageId="5" pageNumber="510" type="distribution">
|
||
<paragraph id="8BCE382DB175CC3323C0FB0D48C6FA95" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="510">
|
||
<emphasis id="B905E43FB175CC3323C0FB0D4B2EFB60" bold="true" box="[199,348,1196,1221]" pageId="5" pageNumber="510">Distribution</emphasis>
|
||
. This species is known from lowland Atlantic Forest from Espírito Santo and Minas Gerais states. It has not been recorded yet but could also be present in northern Rio de Janeiro state (Campos area) and extreme southern Bahia (around Porto Seguro). Numerous field trips have been made to the latter location but no
|
||
<taxonomicName id="4C7143AEB175CC332390FAB94B6BFA95" box="[151,281,1304,1328]" class="Insecta" family="Scarabaeidae" genus="Aphengium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905E43FB175CC332390FAB94B6BFA95" box="[151,281,1304,1328]" italics="true" pageId="5" pageNumber="510">Aphengium</emphasis>
|
||
</taxonomicName>
|
||
specimens have been caught so far.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |