308 lines
37 KiB
XML
308 lines
37 KiB
XML
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<mods:title id="7AD3BD36519C2B80E2D515A718330332">Revision of the rare centipede genus Sterropristes Attems, 1934, with description of a new species from Thailand (Chilopoda: Scolopendromorpha: Scolopendridae)</mods:title>
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<mods:namePart id="D1769F9CC7FE74074989E0F8E86A15CB">Muadsub, Sawitree</mods:namePart>
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<mods:namePart id="71F4753F128498F17E22B9B9F666A831">Sutcharit, Chirasak</mods:namePart>
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<mods:namePart id="30D46D53E7036D03E95BB83917BBECA1">Pimvichai, Piyatida</mods:namePart>
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<mods:namePart id="3566B4FF3AF1DF70F31751560BAC1A5B">Enghoff, Henrik</mods:namePart>
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<mods:typeOfResource id="ABC46EA021315BB4071C385E065980B3">text</mods:typeOfResource>
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<mods:title id="F91B0CAC83546E785D66AEF08FAF2860">Zootaxa</mods:title>
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<mods:date id="CB6874AE27513E5A6682E86402DB78EC">2012</mods:date>
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<mods:detail id="6E07A396F9EFE9F76399169DED686232" type="volume">
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<mods:number id="FBE26E5C31A9930BA473295680F81139">3484</mods:number>
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<treatment id="03D8764453613175EEAC5A34FD2FFAF9" ID-DOI="http://doi.org/10.5281/zenodo.5694287" ID-GBIF-Taxon="119526885" ID-Zenodo-Dep="5694287" LSID="urn:lsid:plazi:treatment:03D8764453613175EEAC5A34FD2FFAF9" httpUri="http://treatment.plazi.org/id/03D8764453613175EEAC5A34FD2FFAF9" lastPageId="2" lastPageNumber="37" pageId="1" pageNumber="36">
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<subSubSection id="C36B94D953613176EEAC5A34FDE9F9D9" box="[151,560,1554,1580]" pageId="1" pageNumber="36" type="nomenclature">
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<paragraph id="8BCEC75253613176EEAC5A34FDE9F9D9" blockId="1.[151,560,1554,1580]" box="[151,560,1554,1580]" pageId="1" pageNumber="36">
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<heading id="D086703E53613176EEAC5A34FDE9F9D9" bold="true" box="[151,560,1554,1580]" fontSize="11" level="1" pageId="1" pageNumber="36" reason="1">
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<emphasis id="B9051B4053613176EEAC5A34FDE9F9D9" bold="true" box="[151,560,1554,1580]" pageId="1" pageNumber="36">
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Genus
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<taxonomicName id="4C71BCD153613176EED55A34FDE9F9D9" ID-CoL="7NB8" ID-ENA="1418208" authority="Attems, 1934" authorityName="Attems" authorityYear="1934" box="[238,560,1554,1580]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="36" phylum="Arthropoda" rank="genus">
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<emphasis id="B9051B4053613176EED55A34FE5FF9D9" bold="true" box="[238,390,1554,1580]" italics="true" pageId="1" pageNumber="36">Sterropristes</emphasis>
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<bibRefCitation id="EFE0BAA353613176EFB75A34FDE9F9D9" author="Attems" box="[396,560,1554,1580]" pageId="1" pageNumber="36" refString="Attems, C. (1934) Neue Myriopoden des Museums Basel. Verhandlungen der Naturforschenden Gesellschaft in Basel, 45, 43 - 62." type="journal article" year="1934">Attems, 1934</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B94D953613176EEAC5A7CFE2AF959" pageId="1" pageNumber="36" type="reference_group">
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<paragraph id="8BCEC75253613176EEAC5A7CFE2AF959" blockId="1.[151,1435,1625,1709]" pageId="1" pageNumber="36">
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<treatmentCitationGroup id="AB61E07C53613176EEAC5A7CFE2AF959" pageId="1" pageNumber="36">
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<taxonomicName id="4C71BCD153613176EEAC5A7CFE17F99A" ID-CoL="7NB8" authority="Attems, 1934: 43" authorityName="Attems" authorityPageNumber="43" authorityYear="1934" box="[151,462,1626,1647]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="36" phylum="Arthropoda" rank="genus">
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<emphasis id="B9051B4053613176EEAC5A7CFEC1F99A" box="[151,280,1626,1647]" italics="true" pageId="1" pageNumber="36">Sterropristes</emphasis>
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<treatmentCitation id="0AD0E14353613176EF255A7CFE17F99A" author="Attems" box="[286,462,1626,1647]" page="43" pageId="1" pageNumber="36" year="1934">
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<bibRefCitation id="EFE0BAA353613176EF255A7CFE73F99A" author="Attems" box="[286,426,1626,1647]" pageId="1" pageNumber="36" refString="Attems, C. (1934) Neue Myriopoden des Museums Basel. Verhandlungen der Naturforschenden Gesellschaft in Basel, 45, 43 - 62." type="journal article" year="1934">Attems, 1934</bibRefCitation>
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: 43
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</treatmentCitation>
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</taxonomicName>
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–47.
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<treatmentCitation id="0AD0E14353613176EC3B5A7FFD15F99A" author="Verhoeff" box="[512,716,1625,1647]" page="201" pageId="1" pageNumber="36" year="1937">
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<bibRefCitation id="EFE0BAA353613176EC3B5A7FFD42F99A" author="Verhoeff" box="[512,667,1625,1647]" pageId="1" pageNumber="36" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270." type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
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||
: 201
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</treatmentCitation>
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, 202.
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<treatmentCitation id="0AD0E14353613176ED315A7FFC01F99A" author="Schileyko" box="[778,984,1625,1647]" page="34" pageId="1" pageNumber="36" year="1997">
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<bibRefCitation id="EFE0BAA353613176ED315A7FFC6AF99A" author="Schileyko" box="[778,947,1625,1647]" pageId="1" pageNumber="36" refString="Schileyko, A. A. & Pavlinov, I. J. (1997) A cladistic analysis of the order Scolopendromorpha (Chilopoda). Entomologica Scandinavica Supplementum, 51, 33 - 40." type="journal article" year="1997">Schileyko, 1997</bibRefCitation>
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: 34
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</treatmentCitation>
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.
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<taxonomicName id="4C71BCD153613176EEAC5A5EFDD1F978" ID-CoL="5KC7" authority="Verhoeff, 1937: 201" authorityName="Verhoeff" authorityPageNumber="201" authorityYear="1937" box="[151,520,1656,1678]" class="Chilopoda" family="Scolopendridae" genus="Malaccolabis" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="36" phylum="Arthropoda" rank="genus">
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<emphasis id="B9051B4053613176EEAC5A5EFEF8F97B" box="[151,289,1656,1678]" italics="true" pageId="1" pageNumber="36">Malaccolabis</emphasis>
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<treatmentCitation id="0AD0E14353613176EF165A5EFDD1F978" author="Verhoeff" box="[301,520,1656,1678]" page="201" pageId="1" pageNumber="36" year="1937">
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<bibRefCitation id="EFE0BAA353613176EF165A5EFE16F978" author="Verhoeff" box="[301,463,1656,1678]" pageId="1" pageNumber="36" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270." type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
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: 201
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</treatmentCitation>
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</taxonomicName>
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.
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<typeStatus id="54CA79F053613176EC235A5FFD93F97B" box="[536,586,1657,1678]" pageId="1" pageNumber="36">Type</typeStatus>
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species:
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<taxonomicName id="4C71BCD153613176EC8F5A5EFB8FF978" ID-CoL="3XK8P" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" box="[692,1110,1656,1678]" class="Chilopoda" family="Scolopendridae" genus="Malaccolabis" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="36" phylum="Arthropoda" rank="species" species="metallica">
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<emphasis id="B9051B4053613176EC8F5A5EFC70F97B" box="[692,937,1656,1678]" italics="true" pageId="1" pageNumber="36">Malaccolabis metallica</emphasis>
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<bibRefCitation id="EFE0BAA353613176ED8D5A5EFB8FF978" author="Verhoeff" box="[950,1110,1656,1678]" pageId="1" pageNumber="36" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270." type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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<treatmentCitation id="0AD0E14353613176EB3E5A5EFE51F959" author="Schileyko" page="34" pageId="1" pageNumber="36" year="1997">
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<bibRefCitation id="EFE0BAA353613176EB3E5A5EFEBBF959" author="Schileyko" pageId="1" pageNumber="36" refString="Schileyko, A. A. & Pavlinov, I. J. (1997) A cladistic analysis of the order Scolopendromorpha (Chilopoda). Entomologica Scandinavica Supplementum, 51, 33 - 40." type="journal article" year="1997">Schileyko and Pavlinov, 1997</bibRefCitation>
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: 34
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</treatmentCitation>
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||
.
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<emphasis id="B9051B4053613176EFA95AB0FE2AF959" bold="true" box="[402,499,1686,1709]" pageId="1" pageNumber="36">Syn. nov.</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36B94D953613176EEAC5AFCFC7BF907" box="[151,930,1753,1778]" pageId="1" pageNumber="36" type="materials_examined">
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<paragraph id="8BCEC75253613176EEAC5AFCFC7BF907" blockId="1.[151,1436,1753,2021]" box="[151,930,1753,1778]" pageId="1" pageNumber="36">
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<emphasis id="B9051B4053613176EEAC5AFCFEEBF907" bold="true" box="[151,306,1753,1778]" pageId="1" pageNumber="36">
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<typeStatus id="54CA79F053613176EEAC5AFCFF0BF907" box="[151,210,1754,1778]" pageId="1" pageNumber="36">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C71BCD153613176EF035AFCFD24F904" authority="Attems, 1934" authorityName="Attems" authorityYear="1934" box="[312,765,1753,1778]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="36" phylum="Arthropoda" rank="species" species="sarasinorum">
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<emphasis id="B9051B4053613176EF035AFCFD82F904" box="[312,603,1754,1777]" italics="true" pageId="1" pageNumber="36">Sterropristes sarasinorum</emphasis>
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<bibRefCitation id="EFE0BAA353613176EC585AFCFD24F904" author="Attems" box="[611,765,1753,1778]" pageId="1" pageNumber="36" refString="Attems, C. (1934) Neue Myriopoden des Museums Basel. Verhandlungen der Naturforschenden Gesellschaft in Basel, 45, 43 - 62." type="journal article" year="1934">Attems, 1934</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C36B94D953613176EEFC5ADBFB01F810" pageId="1" pageNumber="36" type="diagnosis">
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<paragraph id="8BCEC75253613176EEFC5ADBFB01F810" blockId="1.[151,1436,1753,2021]" pageId="1" pageNumber="36">
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<emphasis id="B9051B4053613176EEFC5ADBFE99F8E3" bold="true" box="[199,320,1789,1814]" pageId="1" pageNumber="36">Diagnosis.</emphasis>
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The unique distinguishing character of
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<taxonomicName id="4C71BCD153613176ECC05AD8FC51F8E0" box="[763,904,1790,1813]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="1" pageNumber="36" phylum="Arthropoda" rank="genus">
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<emphasis id="B9051B4053613176ECC05AD8FC51F8E0" box="[763,904,1790,1813]" italics="true" pageId="1" pageNumber="36">Sterropristes</emphasis>
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</taxonomicName>
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is the saw-like internal margin of the forcipular tarsungula. Other characters include basally wide, dorso-ventrally flattened antennae; tergite 1 abutting the cephalic plate; forcipular coxosternal tooth plate with 4+4 teeth; forcipular trochanteroprefemora without processes; tergites from at least T6 to T20 and sternites from at least S13 to S19 with complete paramedian sutures; spiracular apertures oval, spiracles present on trunk segment 7; coxopleura of ultimate legs truncated, without processes and spines; ultimate legs short, thickened; prefemora without spines; distal part of femora with a groove on their dorsal sides; ventral side of either tibiae or tarsi (or both) prominently convex; pretarsi longer than second tarsi.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B94D953623175EEFC5CBEFD2FFAF9" pageId="2" pageNumber="37" type="discussion">
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<paragraph id="8BCEC75253623175EEFC5CBEFC27FCE1" blockId="2.[153,1436,151,1292]" pageId="2" pageNumber="37">
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<emphasis id="B9051B4053623175EEFC5CBEFEE2FF45" bold="true" box="[199,315,152,176]" pageId="2" pageNumber="37">Remarks.</emphasis>
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The traditional classification of the
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<taxonomicName id="4C71BCD153623175ECCE5CB1FC7AFF5A" box="[757,931,151,175]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="family">Scolopendridae</taxonomicName>
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is based on morphological characters, for example, the shape of the spiracle and the presence or absence of three-flapped valves in it, the presence and shape of coxopleural processes, and details of spinulation of the ultimate legs (
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<bibRefCitation id="EFE0BAA353623175EDCD5CF9FB4EFF02" author="Attems" box="[1014,1175,223,247]" pageId="2" pageNumber="37" refString="Attems, C. (1930) Myriapoda. 2. Scolopendromorpha. Das Tierreich, Walter de Gruyter & Co, Berlin, Leipzig, Vol. 54, pp. 1 - 308." type="journal article" year="1930">Attems, 1930</bibRefCitation>
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). The classification of Sterropristini as a group at the tribe or subfamily level particularly emphasizes serrated margins of the forcipular tarsungula (
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<bibRefCitation id="EFE0BAA353623175EF255D01FE17FECA" author="Verhoeff" box="[286,462,295,319]" pageId="2" pageNumber="37" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270." type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
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||
).
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<bibRefCitation id="EFE0BAA353623175EFD85D01FD7EFECA" author="Schileyko" box="[483,679,295,319]" pageId="2" pageNumber="37" refString="Schileyko, A. A. (1992) Scolopenders of Viet-Nam and some aspects of the system of Scolopendromorpha (Chilopoda, Epimorpha), Part 1. Arthropoda Selecta, 1, 5 - 19." type="journal article" year="1992">Schileyko (1992)</bibRefCitation>
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proposed a new classification of the
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<taxonomicName id="4C71BCD153623175EA6D5D01FAE5FECA" box="[1110,1340,295,319]" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="order">Scolopendromorpha</taxonomicName>
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, placing particular weight on the number of body segments and presence of a spiracle on trunk segment 7 (
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<figureCitation id="134ADBD753623175EAC45D6AFA99FE91" box="[1279,1344,332,356]" captionStart="FIGURE 1" captionStartId="3.[151,250,1830,1853]" captionTargetBox="[157,1428,253,1752]" captionTargetId="figure@3.[151,1436,245,1757]" captionTargetPageId="3" captionText="FIGURE 1. Spiracles of segments 3, 5 and 7 (left to right, respectively). A – C. Holotype NMB 432 a of Sterropristes sarasinorum Attems, 1934. D – F. Specimen CUMZ 3205 of Sterropristes metallicus from Penang Botanical Gardens, Penang, Malaysia. G – I. Holotype CUMZ 3207 of Sterropristes violaceus Muadsub and Panha n. sp." httpUri="https://zenodo.org/record/282389/files/figure.png" pageId="2" pageNumber="37">Fig. 1</figureCitation>
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C, F, I). The name Sterropristinae was applied therein to a group united by a spiracle on segment 7, that group then being subdivided into Ethmostigmini (
|
||
<taxonomicName id="4C71BCD153623175EC175DB2FD12FE59" box="[556,715,404,428]" class="Chilopoda" family="Scolopendridae" genus="Ethmostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175EC175DB2FD12FE59" box="[556,715,404,428]" italics="true" pageId="2" pageNumber="37">Ethmostigmus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C71BCD153623175ECDC5DB2FC98FE59" box="[743,833,404,428]" class="Chilopoda" family="Scolopendridae" genus="Rhysida" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175ECDC5DB2FC98FE59" box="[743,833,404,428]" italics="true" pageId="2" pageNumber="37">Rhysida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C71BCD153623175EDA95DB2FC26FE59" box="[914,1023,404,428]" class="Chilopoda" family="Scolopendridae" genus="Alluropus" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175EDA95DB2FC26FE59" box="[914,1023,404,428]" italics="true" pageId="2" pageNumber="37">Alluropus</emphasis>
|
||
</taxonomicName>
|
||
) and Arrhabdotini (
|
||
<taxonomicName id="4C71BCD153623175EB325DB2FA4CFE59" box="[1289,1429,404,428]" class="Chilopoda" family="Scolopendridae" genus="Edentistoma" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175EB325DB2FA4CFE59" box="[1289,1429,404,428]" italics="true" pageId="2" pageNumber="37">Edentistoma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C71BCD153623175EEA25D9EFEF1FE3A" box="[153,296,440,463]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175EEA25D9EFEF1FE3A" box="[153,296,440,463]" italics="true" pageId="2" pageNumber="37">Sterropristes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C71BCD153623175EFB95D9EFDC3FE25" box="[386,538,440,464]" class="Chilopoda" family="Scolopendridae" genus="Malaccolabis" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175EFB95D9EFDC3FE25" box="[386,538,440,464]" italics="true" pageId="2" pageNumber="37">Malaccolabis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<bibRefCitation id="EFE0BAA353623175EC7F5D91FC0EFE3A" author="Schileyko" box="[580,983,439,463]" pageId="2" pageNumber="37" refString="Schileyko, A. A. & Pavlinov, I. J. (1997) A cladistic analysis of the order Scolopendromorpha (Chilopoda). Entomologica Scandinavica Supplementum, 51, 33 - 40." type="journal article" year="1997">Schileyko and Pavlinov (1997)</bibRefCitation>
|
||
presented a cladistic analysis of
|
||
<taxonomicName id="4C71BCD153623175EEA25DFAFEA5FE01" box="[153,380,476,500]" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="order">Scolopendromorpha</taxonomicName>
|
||
in which
|
||
<taxonomicName id="4C71BCD153623175EFD75DFBFDA3FE01" box="[492,634,477,500]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175EFD75DFBFDA3FE01" box="[492,634,477,500]" italics="true" pageId="2" pageNumber="37">Sterropristes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C71BCD153623175EC885DFAFC92FE01" box="[691,843,476,500]" class="Chilopoda" family="Scolopendridae" genus="Malaccolabis" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175EC885DFAFC92FE01" box="[691,843,476,500]" italics="true" pageId="2" pageNumber="37">Malaccolabis</emphasis>
|
||
</taxonomicName>
|
||
formed a clade that was sister-group to
|
||
<taxonomicName id="4C71BCD153623175EB2B5DFAFEBDFDE2" authority="Tomosvary, 1882" authorityName="Tomosvary" authorityYear="1882" class="Chilopoda" family="Scolopendridae" genus="Edentistoma" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175EB2B5DFAFA45FE01" box="[1296,1436,476,500]" italics="true" pageId="2" pageNumber="37">Edentistoma</emphasis>
|
||
Tömösváry, 1882
|
||
</taxonomicName>
|
||
(=
|
||
<emphasis id="B9051B4053623175EFB05E26FDC1FDED" box="[395,536,512,536]" italics="true" pageId="2" pageNumber="37">Arrhabdotus</emphasis>
|
||
<bibRefCitation id="EFE0BAA353623175EC1E5E26FD11FDE2" author="Attems" box="[549,712,511,536]" pageId="2" pageNumber="37" refString="Attems, C. (1930) Myriapoda. 2. Scolopendromorpha. Das Tierreich, Walter de Gruyter & Co, Berlin, Leipzig, Vol. 54, pp. 1 - 308." type="journal article" year="1930">Attems, 1930</bibRefCitation>
|
||
), another deviating monotypic scolopendromorph genus (see
|
||
<bibRefCitation id="EFE0BAA353623175EEA25E02FEECFDC9" author="Lewis" box="[153,309,548,572]" pageId="2" pageNumber="37" refString="Lewis, J. G. E. (1981 b) Observations on the morphology and habits of the bizarre Borneo centipede Arrhabdotus octosulcatus (Tomosvary) (Chilopoda, Scolopendromorpha). Entomologist's Monthly Magazine, 117, 245 - 248." type="journal article" year="1981" yearSuffix="b">Lewis, 1981b</bibRefCitation>
|
||
). Recent analyses of either morphological data (
|
||
<bibRefCitation id="EFE0BAA353623175ED755E02FB51FDC9" author="Edgecombe" box="[846,1160,548,572]" pageId="2" pageNumber="37" refString="Edgecombe, G. D. & Koch, M. (2008) Phylogeny of scolopendromorph centipedes (Chilopoda): morphological analysis featuring characters from the peristomatic area. Cladistics, 24, 872 - 901." type="journal article" year="2008">Edgecombe and Koch 2008</bibRefCitation>
|
||
, 2009; Koch
|
||
<emphasis id="B9051B4053623175EB265E03FA97FDC9" box="[1309,1358,548,572]" italics="true" pageId="2" pageNumber="37">et al</emphasis>
|
||
. 2009;
|
||
<bibRefCitation id="EFE0BAA353623175EEA25E61FEA7FDAA" author="Vahtera" box="[153,382,583,608]" pageId="2" pageNumber="37" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2012 b) Spiracle structure in scolopendromorph centipedes (Chilopoda: Scolopendromorpha) and its contribution to phylogenetics. Zoomorphology, 131, in press (DOI 10.1007 / s 00435 - 012 - 0157 - 0)." type="book chapter" year="2012" yearSuffix="b">
|
||
Vahtera
|
||
<emphasis id="B9051B4053623175EECD5E6FFEFEFD95" box="[246,295,584,608]" italics="true" pageId="2" pageNumber="37">et al</emphasis>
|
||
. 2012b
|
||
</bibRefCitation>
|
||
) or molecular data analyzed either on their own or in combination with morphology (
|
||
<bibRefCitation id="EFE0BAA353623175EB7F5E61FEC4FD71" author="Vahtera" pageId="2" pageNumber="37" refString="Vahtera, V., Edgecombe, G. D. & Giribet, G. (2012 a) Evolution of blindness in scolopendromorph centipedes (Chilopoda: Scolopendromorpha): insight from an expanded sampling of molecular data. Cladistics, 28, 4 - 20." type="journal article" year="2012" yearSuffix="a">
|
||
Vahtera
|
||
<emphasis id="B9051B4053623175EEA25E4BFF13FD71" box="[153,202,620,644]" italics="true" pageId="2" pageNumber="37">et al</emphasis>
|
||
. 2012a
|
||
</bibRefCitation>
|
||
) have not supported Schileyko and Pavlinov’s findings with respect to a taxonomic separation between Otostigmini and “Ethmostigmini”, and the value of a spiracle on segment 7 as a high-level taxonomic character in
|
||
<taxonomicName id="4C71BCD153623175EEA25E92FEA5FD39" box="[153,380,692,716]" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="order">Scolopendromorpha</taxonomicName>
|
||
has been called into question (
|
||
<bibRefCitation id="EFE0BAA353623175ECE35E92FCA2FD39" author="Di" box="[728,891,692,716]" pageId="2" pageNumber="37" refString="Di, Z. Y., Cao, Z. - J., Wu, Y. - L., Yin, S. - J., Edgecombe, G. D. & Li, W. - X. (2010) Discovery of the centipede family Plutoniumidae (Chilopoda) in Asia: a new species of Theatops from China, and the taxonomic value of spiracle distributions in Scolopendromorpha. Zootaxa, 2667, 51 - 63." type="journal article" year="2010">
|
||
<collectingCountry id="F36687C253623175ECE35E92FD2FFD39" box="[728,758,692,716]" name="Iceland" pageId="2" pageNumber="37">Di</collectingCountry>
|
||
<emphasis id="B9051B4053623175ECC65E93FCF6FD39" box="[765,815,692,716]" italics="true" pageId="2" pageNumber="37">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
). Although the exact position of Sterropristini in the phylogeny of
|
||
<taxonomicName id="4C71BCD153623175EF585EF1FDC8FD1A" box="[355,529,727,751]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="family">Scolopendridae</taxonomicName>
|
||
remains unsettled, the ovate outline of the spiracles and their strongly humped atrial wall and floor (
|
||
<figureCitation id="134ADBD753623175EFBD5EDAFE1FFCE1" box="[390,454,764,788]" captionStart="FIGURE 1" captionStartId="3.[151,250,1830,1853]" captionTargetBox="[157,1428,253,1752]" captionTargetId="figure@3.[151,1436,245,1757]" captionTargetPageId="3" captionText="FIGURE 1. Spiracles of segments 3, 5 and 7 (left to right, respectively). A – C. Holotype NMB 432 a of Sterropristes sarasinorum Attems, 1934. D – F. Specimen CUMZ 3205 of Sterropristes metallicus from Penang Botanical Gardens, Penang, Malaysia. G – I. Holotype CUMZ 3207 of Sterropristes violaceus Muadsub and Panha n. sp." httpUri="https://zenodo.org/record/282389/files/figure.png" pageId="2" pageNumber="37">Fig. 1</figureCitation>
|
||
) are consistent with membership in Otostigminae.
|
||
</paragraph>
|
||
<paragraph id="8BCEC75253623175EEFE5F39FD2FFAF9" blockId="2.[153,1436,151,1292]" pageId="2" pageNumber="37">
|
||
<bibRefCitation id="EFE0BAA353623175EEFE5F39FEA0FCC2" author="Verhoeff" box="[197,377,799,823]" pageId="2" pageNumber="37" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270." type="journal article" year="1937">Verhoeff (1937)</bibRefCitation>
|
||
erected the monotypic genus
|
||
<taxonomicName id="4C71BCD153623175ECF65F06FCBCFCCD" box="[717,869,800,824]" class="Chilopoda" family="Scolopendridae" genus="Malaccolabis" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175ECF65F06FCBCFCCD" box="[717,869,800,824]" italics="true" pageId="2" pageNumber="37">Malaccolabis</emphasis>
|
||
</taxonomicName>
|
||
for material from Penang Hill, Penang,
|
||
<collectingCountry id="F36687C253623175EB155F39FA41FCC2" box="[1326,1432,799,823]" name="Malaysia" pageId="2" pageNumber="37">Malaysia</collectingCountry>
|
||
. However, on the basis of the newly collected specimens (including topotypes of
|
||
<taxonomicName id="4C71BCD153623175EA7C5F63FB01FCA9" box="[1095,1240,836,860]" class="Chilopoda" family="Scolopendridae" genus="Malaccolabis" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="metallica">
|
||
<emphasis id="B9051B4053623175EA7C5F63FB01FCA9" box="[1095,1240,836,860]" italics="true" pageId="2" pageNumber="37">M. metallica</emphasis>
|
||
</taxonomicName>
|
||
) and re-study of pertinent
|
||
<typeStatus id="54CA79F053623175EF335F4FFEE1FC74" box="[264,312,873,897]" pageId="2" pageNumber="37">type</typeStatus>
|
||
material we treat
|
||
<taxonomicName id="4C71BCD153623175EC2E5F4EFCBCFC8A" authority="Verhoeff, 1937" authorityName="Verhoeff" authorityYear="1937" box="[533,869,871,896]" class="Chilopoda" family="Scolopendridae" genus="Malaccolabis" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175EC2E5F4EFD74FC75" box="[533,685,872,896]" italics="true" pageId="2" pageNumber="37">Malaccolabis</emphasis>
|
||
<bibRefCitation id="EFE0BAA353623175EC825F41FCBCFC8A" author="Verhoeff" box="[697,869,871,895]" pageId="2" pageNumber="37" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270." type="journal article" year="1937">Verhoeff, 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
as a junior subjective synonym of
|
||
<taxonomicName id="4C71BCD153623175EB365F4EFEEFFC51" authority="Attems, 1934" authorityName="Attems" authorityYear="1934" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175EB365F4EFA45FC8A" box="[1293,1436,872,895]" italics="true" pageId="2" pageNumber="37">Sterropristes</emphasis>
|
||
<bibRefCitation id="EFE0BAA353623175EEA25FAAFEEFFC51" author="Attems" box="[153,310,908,932]" pageId="2" pageNumber="37" refString="Attems, C. (1934) Neue Myriopoden des Museums Basel. Verhandlungen der Naturforschenden Gesellschaft in Basel, 45, 43 - 62." type="journal article" year="1934">Attems, 1934</bibRefCitation>
|
||
</taxonomicName>
|
||
. Although the
|
||
<typeStatus id="54CA79F053623175EFDF5FABFDCDFC50" box="[484,532,909,933]" pageId="2" pageNumber="37">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C71BCD153623175ECA05FAAFCEAFC51" box="[667,819,908,932]" class="Chilopoda" family="Scolopendridae" genus="Malaccolabis" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175ECA05FAAFCEAFC51" box="[667,819,908,932]" italics="true" pageId="2" pageNumber="37">Malaccolabis</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="4C71BCD153623175EA1E5FABFB6DFC51" box="[1061,1204,909,932]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175EA1E5FABFB6DFC51" box="[1061,1204,909,932]" italics="true" pageId="2" pageNumber="37">Sterropristes</emphasis>
|
||
</taxonomicName>
|
||
in some characters, notably the number of sparsely setose antennal articles, these do not warrant separation at the genus level (
|
||
<tableCitation id="C6F3F2E953623175EB0C5F89FA53FC32" box="[1335,1418,943,967]" captionStart="TABLE 1" captionStartId="15.[151,235,327,348]" captionTargetBox="[151,1436,478,1600]" captionTargetPageId="15" captionText="TABLE 1. Morphological comparison among Sterropristes species. All data are taken from the holotypes and the original" httpUri="http://table.plazi.org/id/DF0E97DA536F3178EEAC5D61FA45FEA9" pageId="2" pageNumber="37" tableUuid="DF0E97DA536F3178EEAC5D61FA45FEA9">Table 1</tableCitation>
|
||
). Some of the purportedly diagnostic differences of
|
||
<taxonomicName id="4C71BCD153623175ECEF5FF2FCB5FC19" box="[724,876,980,1004]" class="Chilopoda" family="Scolopendridae" genus="Malaccolabis" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9051B4053623175ECEF5FF2FCB5FC19" box="[724,876,980,1004]" italics="true" pageId="2" pageNumber="37">Malaccolabis</emphasis>
|
||
</taxonomicName>
|
||
exhibit variability now that additional specimens of its
|
||
<typeStatus id="54CA79F053623175EEDA5FDFFEC8FBE4" box="[225,273,1017,1041]" pageId="2" pageNumber="37">type</typeStatus>
|
||
species are known, with the range of variation overlapping with
|
||
<taxonomicName id="4C71BCD153623175EA2C5FDEFB1DFBE5" box="[1047,1220,1016,1040]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="sarasinorum">
|
||
<emphasis id="B9051B4053623175EA2C5FDEFB1DFBE5" box="[1047,1220,1016,1040]" italics="true" pageId="2" pageNumber="37">S. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
. For example, the difference in number of antennal articles (17 versus 12–15) is likely a result of the
|
||
<typeStatus id="54CA79F053623175EA04583AFB78FBC1" box="[1087,1185,1052,1076]" pageId="2" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C71BCD153623175EAFC583BFA8FFBC1" box="[1223,1366,1052,1076]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="metallicus">
|
||
<emphasis id="B9051B4053623175EAFC583BFA8FFBC1" box="[1223,1366,1052,1076]" italics="true" pageId="2" pageNumber="37">S. metallicus</emphasis>
|
||
</taxonomicName>
|
||
being teratological or due to damage and repair with respect to this character because the new topotypes of
|
||
<taxonomicName id="4C71BCD153623175EB375866FA45FBAD" box="[1292,1436,1088,1112]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="metallicus">
|
||
<emphasis id="B9051B4053623175EB375866FA45FBAD" box="[1292,1436,1088,1112]" italics="true" pageId="2" pageNumber="37">S. metallicus</emphasis>
|
||
</taxonomicName>
|
||
have 17 articles as in
|
||
<taxonomicName id="4C71BCD153623175EFA05843FD9EFB89" box="[411,583,1125,1148]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="sarasinorum">
|
||
<emphasis id="B9051B4053623175EFA05843FD9EFB89" box="[411,583,1125,1148]" italics="true" pageId="2" pageNumber="37">S. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
. A supposedly diagnostic difference in number of teeth on the forcipular tarsungula (9 versus 13) can likewise be dismissed because topotypes of
|
||
<taxonomicName id="4C71BCD153623175EDD358AEFBA3FB55" box="[1000,1146,1160,1184]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="metallicus">
|
||
<emphasis id="B9051B4053623175EDD358AEFBA3FB55" box="[1000,1146,1160,1184]" italics="true" pageId="2" pageNumber="37">S. metallicus</emphasis>
|
||
</taxonomicName>
|
||
have as few teeth as the
|
||
<typeStatus id="54CA79F053623175EEA2588AFF22FB31" box="[153,251,1196,1220]" pageId="2" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C71BCD153623175EF1F588BFE09FB31" box="[292,464,1197,1220]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="sarasinorum">
|
||
<emphasis id="B9051B4053623175EF1F588BFE09FB31" box="[292,464,1197,1220]" italics="true" pageId="2" pageNumber="37">S. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
. The presence or absence of a dorsal groove on the ultimate leg femur was cited by
|
||
<bibRefCitation id="EFE0BAA353623175EEA258F6FE96FB1D" author="Verhoeff" box="[153,335,1232,1256]" pageId="2" pageNumber="37" refString="Verhoeff, K. W. (1937) Chilopoden aus Malacca, nach den Objecten des Raffles Museum in Singapore. Bulletin of the Raffles Museum, 13, 198 - 270." type="journal article" year="1937">Verhoeff (1937)</bibRefCitation>
|
||
as diagnostic for two genera, but this furrow is in fact present in
|
||
<taxonomicName id="4C71BCD153623175EA6758F6FAD1FB1D" box="[1116,1288,1232,1256]" class="Chilopoda" family="Scolopendridae" genus="Sterropristes" kingdom="Animalia" order="Scolopendromorpha" pageId="2" pageNumber="37" phylum="Arthropoda" rank="species" species="sarasinorum">
|
||
<emphasis id="B9051B4053623175EA6758F6FAD1FB1D" box="[1116,1288,1232,1256]" italics="true" pageId="2" pageNumber="37">S. sarasinorum</emphasis>
|
||
</taxonomicName>
|
||
but was not described or illustrated by
|
||
<bibRefCitation id="EFE0BAA353623175EFF858D2FDBFFAF9" author="Attems" box="[451,614,1268,1292]" pageId="2" pageNumber="37" refString="Attems, C. (1934) Neue Myriopoden des Museums Basel. Verhandlungen der Naturforschenden Gesellschaft in Basel, 45, 43 - 62." type="journal article" year="1934">Attems (1934)</bibRefCitation>
|
||
(see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |