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<document id="740F523CE8A6CDBC7087FE250FEA7009" ID-DOI="10.37828/em.2020.38.7" ID-ISSN="2336-9744" ID-Zenodo-Dep="13232632" ID-ZooBank="urn:lsid:zoobank.org:pub:4B85DA98-957E-4788-BC8B-25526047DC8B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722914106896" checkinUser="felipe" docAuthor="Saldaitis, Aidas &amp; Volynkin, Anton V." docDate="2020" docId="03CBF90BFF8C4B68C7B098CD14265CBC" docLanguage="en" docName="EcolMontenegrina.38.51-62.pdf" docOrigin="Ecologica Montenegrina 38" docSource="http://dx.doi.org/10.37828/em.2020.38.7" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Chelis gratiosa subsp. kartashovi Saldaitis &amp; Volynkin, 2020, ssp. nov." docType="treatment" docVersion="2" lastPageNumber="56" masterDocId="FFF28173FF8E4B6DC7279A2D16565C62" masterDocTitle="On the taxonomy of the Chelis glaphyra (Eversmann, 1843) species-group, with description of a new subspecies of Chelis gratiosa (Grum-Grshimailo, 1890) from Kyrgyzstan, brachipterous females of Chelis kashmirica (Ferguson, 1985) and Chelis golbecki (Dubatolov, 1996), and the checklist of the species-group (Lepidoptera: Erebidae: Arctiinae: Arctiini)" masterLastPageNumber="62" masterPageNumber="51" pageNumber="53" updateTime="1722917066501" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2D4529F3255BDE775E1D79FA7B0FCF93">On the taxonomy of the Chelis glaphyra (Eversmann, 1843) species-group, with description of a new subspecies of Chelis gratiosa (Grum-Grshimailo, 1890) from Kyrgyzstan, brachipterous females of Chelis kashmirica (Ferguson, 1985) and Chelis golbecki (Dubatolov, 1996), and the checklist of the species-group (Lepidoptera: Erebidae: Arctiinae: Arctiini)</mods:title>
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<mods:name id="CFEE04FEAEEC2C7995BDFF8559F02537" type="personal">
<mods:role id="0371B4CBA8B368D7EC4D699AA870A4AF">
<mods:roleTerm id="72E3D0A7BC2946BDAFBEED42DD1A8F5D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5C19DD016E3A28F11770BEC23BCF1B31">Saldaitis, Aidas</mods:namePart>
<mods:affiliation id="196F786F28ADC17C4295944FD877C09B">Nature Research Centre, Akademijos str., 2, LT- 08412, Vilnius- 21, Lithuania. E-mail: saldrasa @ gmail. com &amp; Corresponding author</mods:affiliation>
</mods:name>
<mods:name id="9C59A5DACBD0F64ED2C0642436CF3D25" type="personal">
<mods:role id="E48BE829D5892E60D98EAF1A047AEBA6">
<mods:roleTerm id="67CD4733ED2F6003FC98000210ED5C91">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0ECDE0C16DD8D660272A4742D4EC49D2">Volynkin, Anton V.</mods:namePart>
<mods:affiliation id="E37C5BE55AE7A95DDD5C619A919F1B34">Altai State University, Lenina Avenue, 61, RF- 656049, Barnaul, Russia. E-mails: monstruncusarctia @ gmail. com; volynkin _ a @ mail. ru &amp; National Research Tomsk State University, Lenina Avenue, 36, RF- 634050, Tomsk, Russia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="CDD470000516505E48A167F74D803156">text</mods:typeOfResource>
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<mods:title id="8015B0BCE935B800FA324FBC5A5DF552">Ecologica Montenegrina</mods:title>
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<mods:part id="97BE22845AF046FF47D5E6F1BD78E41D">
<mods:date id="400B4C6D50BF1C3D464CB33A027DD302">2020</mods:date>
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<mods:number id="6404380899CE9DF0F8C28C9A96A14717">2020-12-01</mods:number>
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<mods:number id="B90BAEA3DA8AB27599569456A132844D">38</mods:number>
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<mods:url id="204E648D7044A8E25EC081C4EE0605C5">http://dx.doi.org/10.37828/em.2020.38.7</mods:url>
</mods:location>
<mods:classification id="908AB7648F2FD72044FA21A6173D1755">journal article</mods:classification>
<mods:identifier id="D1F7BF75FCA83D0B91783CE51C084313" type="DOI">10.37828/em.2020.38.7</mods:identifier>
<mods:identifier id="0157D932ECC7093C81E53102FC07367A" type="ISSN">2336-9744</mods:identifier>
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<treatment id="03CBF90BFF8C4B68C7B098CD14265CBC" LSID="urn:lsid:plazi:treatment:03CBF90BFF8C4B68C7B098CD14265CBC" httpUri="http://treatment.plazi.org/id/03CBF90BFF8C4B68C7B098CD14265CBC" lastPageId="5" lastPageNumber="56" pageId="2" pageNumber="53">
<subSubSection id="C3781B96FF8C4B6FC7B098CD14135E98" box="[151,581,736,762]" pageId="2" pageNumber="53" type="nomenclature">
<paragraph id="8BDD481DFF8C4B6FC7B098CD14135E98" blockId="2.[151,581,736,796]" box="[151,581,736,762]" pageId="2" pageNumber="53">
<heading id="D095FF71FF8C4B6FC7B098CD14135E98" box="[151,581,736,762]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="53" reason="2">
<emphasis id="B916940FFF8C4B6FC7B098CD14135E98" bold="true" box="[151,581,736,762]" pageId="2" pageNumber="53">
<taxonomicName id="4C62339EFF8C4B6FC7B098CD17855E98" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" box="[151,467,736,762]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="gratiosa" status="subsp. nov." subSpecies="kartashovi">
<emphasis id="B916940FFF8C4B6FC7B098CD17855E98" bold="true" box="[151,467,736,762]" italics="true" pageId="2" pageNumber="53">Chelis gratiosa kartashovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2252974FF8C4B6FC6FC98CD14135E98" box="[475,581,736,762]" pageId="2" pageNumber="53" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3781B96FF8C4B6FC7B0992C17D05F7E" box="[151,390,769,796]" pageId="2" pageNumber="53" type="description">
<paragraph id="8BDD481DFF8C4B6FC7B0992C17D05F7E" blockId="2.[151,581,736,796]" box="[151,390,769,796]" pageId="2" pageNumber="53">
(
<figureCitation id="13595498FF8C4B6FC7B8992C175F5F7E" box="[159,265,769,796]" captionStart="Figures 18" captionStartId="3.[151,236,1965,1989]" captionTargetBox="[175,1427,226,1921]" captionTargetId="figure-13@3.[151,1427,223,1946]" captionTargetPageId="3" captionText="Figures 18. Chelis spp.: adults. Depositories of the specimens: 1, 3 in ASV; 2 in WIGJ; 4 in SDM; 58 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232634" httpUri="https://zenodo.org/record/13232634/files/figure.png" pageId="2" pageNumber="53">Figs 14</figureCitation>
,
<figureCitation id="13595498FF8C4B6FC632992C17705F7E" box="[277,294,769,796]" captionStart="Figure 9" captionStartId="4.[151,226,1050,1074]" captionTargetBox="[163,1425,212,1032]" captionTargetId="figure-18@4.[151,1436,189,1032]" captionTargetPageId="4" captionText="Figure 9. Chelis gratiosa kartashovi, paratypes. The specimens are deposited in VKV (photo by V. Kartashov)." figureDoi="http://doi.org/10.5281/zenodo.13232636" httpUri="https://zenodo.org/record/13232636/files/figure.png" pageId="2" pageNumber="53">9</figureCitation>
,
<figureCitation id="13595498FF8C4B6FC614992C17045F7E" box="[307,338,769,796]" captionStart="Figures 1720" captionStartId="6.[151,236,2002,2026]" captionTargetBox="[163,1426,222,1936]" captionTargetId="figure-18@6.[151,1436,222,1961]" captionTargetPageId="6" captionText="Figures 1720. Chelis gratiosa sspp.: adults. Depositories of the specimens: 17 in ASV/WIGJ; 1820 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232642" httpUri="https://zenodo.org/record/13232642/files/figure.png" pageId="2" pageNumber="53">17</figureCitation>
,
<figureCitation id="13595498FF8C4B6FC678992C17285F7E" box="[351,382,769,796]" captionStart="Figures 2428" captionStartId="8.[151,236,1909,1933]" captionTargetBox="[155,1436,204,1841]" captionTargetId="figure-18@8.[151,1436,189,1872]" captionTargetPageId="8" captionText="Figures 2428. Chelis spp.: adults. Depositories of the specimens: 24 in ASV; 2528 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232648" httpUri="https://zenodo.org/record/13232648/files/figure.png" pageId="2" pageNumber="53">24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3781B96FF8C4B6FC7B099681307582E" pageId="2" pageNumber="53" type="materials_examined">
<paragraph id="8BDD481DFF8C4B6FC7B0996817C75FA7" blockId="2.[151,1436,836,965]" pageId="2" pageNumber="53">
<emphasis id="B916940FFF8C4B6FC7B09968171E5F3D" bold="true" box="[151,328,837,863]" pageId="2" pageNumber="53">Type material</emphasis>
.
<materialsCitation id="3B0A4240FF8C4B6FC67F996817C75FA7" collectingDate="2018-07-15" collectingDateMax="2019-05-18" collectingDateMin="2018-07-15" collectionCode="V, ASV, WIGJ" country="Kyrgyzstan" elevation="3600" location="Transalai Range" pageId="2" pageNumber="53" specimenCount="1" typeStatus="holotype">
<emphasis id="B916940FFF8C4B6FC67F9968179C5F3D" bold="true" box="[344,458,837,863]" pageId="2" pageNumber="53">
<typeStatus id="54D9F6BFFF8C4B6FC67F9968179C5F3D" box="[344,458,837,863]" pageId="2" pageNumber="53" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="13595498FF8C4B6FC6FD996914725F3D" box="[474,548,836,863]" captionStart="Figures 18" captionStartId="3.[151,236,1965,1989]" captionTargetBox="[175,1427,226,1921]" captionTargetId="figure-13@3.[151,1427,223,1946]" captionTargetPageId="3" captionText="Figures 18. Chelis spp.: adults. Depositories of the specimens: 1, 3 in ASV; 2 in WIGJ; 4 in SDM; 58 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232634" httpUri="https://zenodo.org/record/13232634/files/figure.png" pageId="2" pageNumber="53">Figs 1</figureCitation>
,
<figureCitation id="13595498FF8C4B6FC513996914035F3D" box="[564,597,836,863]" captionStart="Figures 1720" captionStartId="6.[151,236,2002,2026]" captionTargetBox="[163,1426,222,1936]" captionTargetId="figure-18@6.[151,1436,222,1961]" captionTargetPageId="6" captionText="Figures 1720. Chelis gratiosa sspp.: adults. Depositories of the specimens: 17 in ASV/WIGJ; 1820 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232642" httpUri="https://zenodo.org/record/13232642/files/figure.png" pageId="2" pageNumber="53">17</figureCitation>
): male, [
<collectingCountry id="F375088DFF8C4B6FC5999968151E5F3D" box="[702,840,837,863]" name="Kyrgyzstan" pageId="2" pageNumber="53">Kyrgyzstan</collectingCountry>
,
<location id="8EBD1EC6FF8C4B6FC473996912415F3D" LSID="urn:lsid:plazi:treatment:03CBF90BFF8C4B68C7B098CD14265CBC:8EBD1EC6FF8C4B6FC473996912415F3D" box="[852,1047,836,863]" country="Kyrgyzstan" name="Transalai Range" pageId="2" pageNumber="53">Transalai Range</location>
] “KIRGIZIA S., Chon- Alai reg., Zaalaisky mountains range, Doroot-Korgon vill., Altyn-dara river, Aram-Kungei valley, H~
<quantity id="4C9AE5F8FF8C4B6FC3F9994B13655FE3" box="[1246,1331,870,897]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="53" unit="m" value="3600.0">
<elevation id="004FAF2EFF8C4B6FC3F9994B13655FE3" box="[1246,1331,870,897]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="53" unit="m" value="3600.0">3600m</elevation>
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, original female
<date id="FFDC6EDDFF8C4B6FC7D099A517295FC1" box="[247,383,904,931]" pageId="2" pageNumber="53" value="2018-07-15">
<collectingDate id="EF989735FF8C4B6FC7D099A517295FC1" box="[247,383,904,931]" pageId="2" pageNumber="53" value="2018-07-15">15.vii.2018</collectingDate>
</date>
, ex. ovo
<date id="FFDC6EDDFF8C4B6FC52399A514FE5FC1" box="[516,680,904,931]" pageId="2" pageNumber="53" value="2019-05-14" valueMax="2019-05-18" valueMin="2019-05-14">
<collectingDate id="EF989735FF8C4B6FC52399A514FE5FC1" box="[516,680,904,931]" pageId="2" pageNumber="53" value="2019-05-14" valueMax="2019-05-18" valueMin="2019-05-14">1418.v.2019</collectingDate>
</date>
, leg.
<collectionCode id="ED73D0D8FF8C4B6FC5DE99A415445FC1" box="[761,786,905,931]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="53" type="Museum">V</collectionCode>
. Kartashov”, gen. slide No.: AV5839 (Coll.
<collectionCode id="ED73D0D8FF8C4B6FC27D99A513CE5FC1" box="[1370,1432,904,931]" pageId="2" pageNumber="53">ASV</collectionCode>
, designated to
<collectionCode id="ED73D0D8FF8C4B6FC61D998717D35FA7" box="[314,389,938,965]" pageId="2" pageNumber="53">WIGJ</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BDD481DFF8C4B6FC7B099C01307582E" blockId="2.[151,1436,1005,1100]" pageId="2" pageNumber="53">
<materialsCitation id="3B0A4240FF8C4B6FC7B099C017E15848" collectionCode="ASV, VKV, WIGJ" location="Colls" pageId="2" pageNumber="53" specimenCount="22" specimenCount-female="13" specimenCount-male="9" typeStatus="paratype">
<emphasis id="B916940FFF8C4B6FC7B099C017425865" bold="true" box="[151,276,1005,1031]" pageId="2" pageNumber="53">
<typeStatus id="54D9F6BFFF8C4B6FC7B099C017425865" box="[151,276,1005,1031]" pageId="2" pageNumber="53" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D648394FF8C4B6FC60199C017D3586A" box="[294,389,1005,1032]" count="9" pageId="2" pageNumber="53" type="male">9 males</specimenCount>
,
<specimenCount id="9D648394FF8C4B6FC6B399C0144F586A" box="[404,537,1005,1032]" count="13" pageId="2" pageNumber="53" type="female">13 females</specimenCount>
, the same data as in the
<typeStatus id="54D9F6BFFF8C4B6FC46199C015E6586A" box="[838,944,1005,1032]" pageId="2" pageNumber="53" type="holotype">holotype</typeStatus>
, gen. slide No.: AV5840 (female) (
<location id="8EBD1EC6FF8C4B6FC27C99C013CC586A" LSID="urn:lsid:plazi:treatment:03CBF90BFF8C4B68C7B098CD14265CBC:8EBD1EC6FF8C4B6FC27C99C013CC586A" box="[1371,1434,1005,1032]" name="Colls" pageId="2" pageNumber="53">Colls</location>
<collectionCode id="ED73D0D8FF8C4B6FC7B09E2216825848" box="[151,212,1039,1066]" pageId="2" pageNumber="53">ASV</collectionCode>
,
<collectionCode id="ED73D0D8FF8C4B6FC7C39E3D17725848" box="[228,292,1040,1066]" pageId="2" pageNumber="53">VKV</collectionCode>
and
<collectionCode id="ED73D0D8FF8C4B6FC6439E2217F95848" box="[356,431,1039,1066]" pageId="2" pageNumber="53">WIGJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B0A4240FF8C4B6FC6E09E2212F8582E" collectingDate="2018-07-15" collectingDateMax="2018-07-21" collectingDateMin="2018-07-15" collectionCode="V" collectorName="Kartashov" country="Kyrgyzstan" elevation="3600" location="Kirgizia" municipality="Chon-Alaisky Distr." pageId="2" pageNumber="53" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D648394FF8C4B6FC6E09E2214625848" box="[455,564,1039,1066]" count="1" pageId="2" pageNumber="53" type="female">1 female</specimenCount>
, [
<collectingCountry id="F375088DFF8C4B6FC56E9E3D14805848" box="[585,726,1040,1066]" name="Kyrgyzstan" pageId="2" pageNumber="53">Kyrgyzstan</collectingCountry>
, Transalai Range]
<location id="8EBD1EC6FF8C4B6FC4999E2212775848" LSID="urn:lsid:plazi:treatment:03CBF90BFF8C4B68C7B098CD14265CBC:8EBD1EC6FF8C4B6FC4999E2212775848" box="[958,1057,1039,1066]" country="Kyrgyzstan" municipality="Chon-Alaisky Distr." name="Kirgizia" pageId="2" pageNumber="53">Kirgizia</location>
, Zaalaisky Ran.,
<collectingMunicipality id="6BB9D267FF8C4B6FC3D19E22168A582E" pageId="2" pageNumber="53">Chon-Alaisky Distr.</collectingMunicipality>
, Altyn-Dara Riv., Aram-Kungei Val.,
<date id="FFDC6EDDFF8C4B6FC5B89E1C1502582E" box="[671,852,1073,1100]" pageId="2" pageNumber="53" value="2018-07-15" valueMax="2018-07-21" valueMin="2018-07-15">
<collectingDate id="EF989735FF8C4B6FC5B89E1C1502582E" box="[671,852,1073,1100]" pageId="2" pageNumber="53" value="2018-07-15" valueMax="2018-07-21" valueMin="2018-07-15">1521.vii.2018</collectingDate>
</date>
, H-
<quantity id="4C9AE5F8FF8C4B6FC4A79E1C1582582E" box="[896,980,1073,1100]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="53" unit="m" value="3600.0">
<elevation id="004FAF2EFF8C4B6FC4A79E1C1582582E" box="[896,980,1073,1100]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="53" unit="m" value="3600.0">3600m</elevation>
</quantity>
,
<collectionCode id="ED73D0D8FF8C4B6FC4C79E1F15AF582E" box="[992,1017,1074,1100]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="53" type="Museum">V</collectionCode>
.
<collectorName id="26972DCBFF8C4B6FC3249E1C122D582E" box="[1027,1147,1073,1100]" pageId="2" pageNumber="53">Kartashov</collectorName>
leg.
</materialsCitation>
(Coll.
<collectionCode id="ED73D0D8FF8C4B6FC2279E1C1312582E" box="[1280,1348,1073,1100]" country="United Kingdom" httpUri="http://grbio.org/cool/gw1v-bpc3" name="Stroud and District Museum" pageId="2" pageNumber="53">SDM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3781B96FF8C4B6FC7B09E59174158D3" pageId="2" pageNumber="53" type="etymology">
<paragraph id="8BDD481DFF8C4B6FC7B09E59174158D3" blockId="2.[151,1434,1140,1201]" pageId="2" pageNumber="53">
<emphasis id="B916940FFF8C4B6FC7B09E59177058EC" bold="true" box="[151,294,1140,1166]" pageId="2" pageNumber="53">Etymology.</emphasis>
The new subspecies is dedicated to prominent Russian
<taxonomicName id="4C62339EFF8C4B6FC3309E5912F358ED" authorityName="Linnaeus" authorityYear="1758" box="[1047,1189,1140,1167]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
collector
<collectingRegion id="49A686FFFF8C4B6FC2179E5913CC58ED" box="[1328,1434,1140,1167]" country="Russia" name="Vladimir" pageId="2" pageNumber="53">Vladimir</collectingRegion>
Kartashov.
</paragraph>
</subSubSection>
<subSubSection id="C3781B96FF8C4B6FC7B09EF714E35A8D" pageId="2" pageNumber="53" type="diagnosis">
<paragraph id="8BDD481DFF8C4B6FC7B09EF714E35A8D" blockId="2.[151,1437,1241,1775]" pageId="2" pageNumber="53">
<emphasis id="B916940FFF8C4B6FC7B09EF717465896" bold="true" box="[151,272,1242,1268]" pageId="2" pageNumber="53">Diagnosis</emphasis>
. The new subspecies (
<figureCitation id="13595498FF8C4B6FC5049EF414C65896" box="[547,656,1241,1268]" captionStart="Figures 18" captionStartId="3.[151,236,1965,1989]" captionTargetBox="[175,1427,226,1921]" captionTargetId="figure-13@3.[151,1427,223,1946]" captionTargetPageId="3" captionText="Figures 18. Chelis spp.: adults. Depositories of the specimens: 1, 3 in ASV; 2 in WIGJ; 4 in SDM; 58 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232634" httpUri="https://zenodo.org/record/13232634/files/figure.png" pageId="2" pageNumber="53">Figs 14</figureCitation>
) is most similar externally to
<taxonomicName id="4C62339EFF8C4B6FC4D89EF413555896" baseAuthorityName="Grum-Grshimailo" baseAuthorityYear="1890" box="[1023,1283,1241,1268]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="gratiosa" subSpecies="rupicola">
<emphasis id="B916940FFF8C4B6FC4D89EF413555896" box="[1023,1283,1241,1268]" italics="true" pageId="2" pageNumber="53">Ch. gratiosa rupicola</emphasis>
</taxonomicName>
(
<figureCitation id="13595498FF8C4B6FC2319EF413085896" box="[1302,1374,1241,1268]" captionStart="Figures 18" captionStartId="3.[151,236,1965,1989]" captionTargetBox="[175,1427,226,1921]" captionTargetId="figure-13@3.[151,1427,223,1946]" captionTargetPageId="3" captionText="Figures 18. Chelis spp.: adults. Depositories of the specimens: 1, 3 in ASV; 2 in WIGJ; 4 in SDM; 58 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232634" httpUri="https://zenodo.org/record/13232634/files/figure.png" pageId="2" pageNumber="53">Fig. 5</figureCitation>
) and also
<taxonomicName id="4C62339EFF8C4B6FC7F59ED6176D5977" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis, Ivinskis &amp; Churkin" baseAuthorityYear="2000" box="[210,315,1275,1302]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="rasa">
<emphasis id="B916940FFF8C4B6FC7F59ED6176D5977" box="[210,315,1275,1302]" italics="true" pageId="2" pageNumber="53">Ch. rasa</emphasis>
</taxonomicName>
(
<figureCitation id="13595498FF8C4B6FC6779ED617895974" box="[336,479,1275,1302]" captionStart="Figures 1016" captionStartId="5.[151,236,1989,2013]" captionTargetBox="[160,1422,480,1948]" captionTargetId="figure-51@5.[151,1434,480,1971]" captionTargetPageId="5" captionText="Figures 1016. Chelis spp.: adults. The specimens are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232640" httpUri="https://zenodo.org/record/13232640/files/figure.png" pageId="2" pageNumber="53">Figs 13, 14</figureCitation>
,
<figureCitation id="13595498FF8C4B6FC6D79ED614465974" box="[496,528,1275,1302]" captionStart="Figures 2123" captionStartId="7.[151,236,1994,2018]" captionTargetBox="[237,1328,196,1963]" captionTargetId="figure-13@7.[151,1406,189,1975]" captionTargetPageId="7" captionText="Figures 2123. Chelis spp.: adults. The specimens are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232646" httpUri="https://zenodo.org/record/13232646/files/figure.png" pageId="2" pageNumber="53">22</figureCitation>
,
<figureCitation id="13595498FF8C4B6FC5079ED614695974" box="[544,575,1275,1302]" captionStart="Figures 2428" captionStartId="8.[151,236,1909,1933]" captionTargetBox="[155,1436,204,1841]" captionTargetId="figure-18@8.[151,1436,189,1872]" captionTargetPageId="8" captionText="Figures 2428. Chelis spp.: adults. Depositories of the specimens: 24 in ASV; 2528 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232648" httpUri="https://zenodo.org/record/13232648/files/figure.png" pageId="2" pageNumber="53">28</figureCitation>
) and
<taxonomicName id="4C62339EFF8C4B6FC5AD9ED615745974" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Dubatolov" baseAuthorityYear="1996" box="[650,802,1275,1302]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="golbecki">
<emphasis id="B916940FFF8C4B6FC5AD9ED615745974" box="[650,802,1275,1302]" italics="true" pageId="2" pageNumber="53">Ch. golbecki</emphasis>
</taxonomicName>
(
<figureCitation id="13595498FF8C4B6FC4119ED615905974" box="[822,966,1275,1302]" captionStart="Figures 1016" captionStartId="5.[151,236,1989,2013]" captionTargetBox="[160,1422,480,1948]" captionTargetId="figure-51@5.[151,1434,480,1971]" captionTargetPageId="5" captionText="Figures 1016. Chelis spp.: adults. The specimens are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232640" httpUri="https://zenodo.org/record/13232640/files/figure.png" pageId="2" pageNumber="53">Figs 15, 16</figureCitation>
,
<figureCitation id="13595498FF8C4B6FC4F19ED615A05974" box="[982,1014,1275,1302]" captionStart="Figures 2123" captionStartId="7.[151,236,1994,2018]" captionTargetBox="[237,1328,196,1963]" captionTargetId="figure-13@7.[151,1406,189,1975]" captionTargetPageId="7" captionText="Figures 2123. Chelis spp.: adults. The specimens are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232646" httpUri="https://zenodo.org/record/13232646/files/figure.png" pageId="2" pageNumber="53">23</figureCitation>
,
<figureCitation id="13595498FF8C4B6FC3209ED6127E5974" box="[1031,1064,1275,1302]" captionStart="Figures 2428" captionStartId="8.[151,236,1909,1933]" captionTargetBox="[155,1436,204,1841]" captionTargetId="figure-18@8.[151,1436,189,1872]" captionTargetPageId="8" captionText="Figures 2428. Chelis spp.: adults. Depositories of the specimens: 24 in ASV; 2528 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232648" httpUri="https://zenodo.org/record/13232648/files/figure.png" pageId="2" pageNumber="53">27</figureCitation>
), but is distinguishable by the pinkish suffusion on the thorax and the forewing (it is absent in the three similar taxa), and the intense blackish suffusion on hindwing making it almost uniformly dark with only a small pale spot in the subterminal or terminal area medially (whereas in the three similar taxa, the hindwing has intense blackish or dark brown suffusion in the subterminal area only, while the medial and basal areas are weakly suffused with blackish scales and the discal spot is present). Additionally, the new subspecies differs from
<taxonomicName id="4C62339EFF8C4B6FC2279F89133E59DC" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis, Ivinskis &amp; Churkin" baseAuthorityYear="2000" box="[1280,1384,1444,1471]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="rasa">
<emphasis id="B916940FFF8C4B6FC2279F89133E59DC" box="[1280,1384,1444,1471]" italics="true" pageId="2" pageNumber="53">Ch. rasa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C62339EFF8C4B6FC7B09FEB177B5983" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Dubatolov" baseAuthorityYear="1996" box="[151,301,1478,1505]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="golbecki">
<emphasis id="B916940FFF8C4B6FC7B09FEB177B5983" box="[151,301,1478,1505]" italics="true" pageId="2" pageNumber="53">Ch. golbecki</emphasis>
</taxonomicName>
by the longer rami of the male antenna, and from
<taxonomicName id="4C62339EFF8C4B6FC4A89FEB15A35982" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis, Ivinskis &amp; Churkin" baseAuthorityYear="2000" box="[911,1013,1478,1505]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="rasa">
<emphasis id="B916940FFF8C4B6FC4A89FEB15A35982" box="[911,1013,1478,1505]" italics="true" pageId="2" pageNumber="53">Ch. rasa</emphasis>
</taxonomicName>
by its less elongate forewing apex. The male genital capsule of
<emphasis id="B916940FFF8C4B6FC6C09FCA14E55A60" box="[487,691,1511,1538]" italics="true" pageId="2" pageNumber="53">Ch. g. kartashovi</emphasis>
(
<figureCitation id="13595498FF8C4B6FC5E39FCA154F5A60" box="[708,793,1511,1538]" captionStart="Figures 1720" captionStartId="6.[151,236,2002,2026]" captionTargetBox="[163,1426,222,1936]" captionTargetId="figure-18@6.[151,1436,222,1961]" captionTargetPageId="6" captionText="Figures 1720. Chelis gratiosa sspp.: adults. Depositories of the specimens: 17 in ASV/WIGJ; 1820 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232642" httpUri="https://zenodo.org/record/13232642/files/figure.png" pageId="2" pageNumber="53">Fig. 17</figureCitation>
) is nearly identical to that of
<emphasis id="B916940FFF8C4B6FC35E9FCA137A5A60" box="[1145,1324,1511,1538]" italics="true" pageId="2" pageNumber="53">Ch. g. rupicola</emphasis>
(
<figureCitation id="13595498FF8C4B6FC2199FCA13C55A60" box="[1342,1427,1511,1538]" captionStart="Figures 1720" captionStartId="6.[151,236,2002,2026]" captionTargetBox="[163,1426,222,1936]" captionTargetId="figure-18@6.[151,1436,222,1961]" captionTargetPageId="6" captionText="Figures 1720. Chelis gratiosa sspp.: adults. Depositories of the specimens: 17 in ASV/WIGJ; 1820 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232642" httpUri="https://zenodo.org/record/13232642/files/figure.png" pageId="2" pageNumber="53">Fig. 18</figureCitation>
) and is also similar to those of
<taxonomicName id="4C62339EFF8C4B6FC6D89C2414E75A41" authorityName="gratiosa (Grum-Grshimailo" authorityYear="1890" baseAuthorityName="gratiosa (Grum-Grshimailo" baseAuthorityYear="1890" box="[511,689,1545,1572]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="gratiosa" subSpecies="gratiosa">
<emphasis id="B916940FFF8C4B6FC6D89C2414E75A41" box="[511,689,1545,1572]" italics="true" pageId="2" pageNumber="53">Ch. g. gratiosa</emphasis>
</taxonomicName>
(
<figureCitation id="13595498FF8C4B6FC5E59C24154F5A46" box="[706,793,1545,1572]" captionStart="Figures 1720" captionStartId="6.[151,236,2002,2026]" captionTargetBox="[163,1426,222,1936]" captionTargetId="figure-18@6.[151,1436,222,1961]" captionTargetPageId="6" captionText="Figures 1720. Chelis gratiosa sspp.: adults. Depositories of the specimens: 17 in ASV/WIGJ; 1820 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232642" httpUri="https://zenodo.org/record/13232642/files/figure.png" pageId="2" pageNumber="53">Fig. 20</figureCitation>
) and
<emphasis id="B916940FFF8C4B6FC47A9C2415A95A46" box="[861,1023,1545,1572]" italics="true" pageId="2" pageNumber="53">Ch. g. flavala</emphasis>
(
<figureCitation id="13595498FF8C4B6FC3369C24123E5A46" box="[1041,1128,1545,1572]" captionStart="Figures 1720" captionStartId="6.[151,236,2002,2026]" captionTargetBox="[163,1426,222,1936]" captionTargetId="figure-18@6.[151,1436,222,1961]" captionTargetPageId="6" captionText="Figures 1720. Chelis gratiosa sspp.: adults. Depositories of the specimens: 17 in ASV/WIGJ; 1820 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232642" httpUri="https://zenodo.org/record/13232642/files/figure.png" pageId="2" pageNumber="53">Fig. 19</figureCitation>
), but differs from the two latter subspecies by its more rectangular cucullus. The aedeagus of
<emphasis id="B916940FFF8C4B6FC4EE9C0612CE5A24" box="[969,1176,1579,1606]" italics="true" pageId="2" pageNumber="53">Ch. g. kartashovi</emphasis>
differs from those of the three similar subspecies by its markedly narrower dentate carinal plate. The vesicae of the four subspecies display no remarkable differences. As the female genitalia of
<emphasis id="B916940FFF8C4B6FC3229C4312ED5AEB" box="[1029,1211,1646,1673]" italics="true" pageId="2" pageNumber="53">Ch. g. rupicola</emphasis>
and
<emphasis id="B916940FFF8C4B6FC3DF9C4313CA5AEB" box="[1272,1436,1646,1673]" italics="true" pageId="2" pageNumber="53">Ch. g. flavala</emphasis>
are unknown therefore the female genitalia morphology of the new species (
<figureCitation id="13595498FF8C4B6FC30E9CBD12D75AC9" box="[1065,1153,1680,1707]" captionStart="Figures 2428" captionStartId="8.[151,236,1909,1933]" captionTargetBox="[155,1436,204,1841]" captionTargetId="figure-18@8.[151,1436,189,1872]" captionTargetPageId="8" captionText="Figures 2428. Chelis spp.: adults. Depositories of the specimens: 24 in ASV; 2528 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232648" httpUri="https://zenodo.org/record/13232648/files/figure.png" pageId="2" pageNumber="53">Fig. 24</figureCitation>
) is compared to that of
<taxonomicName id="4C62339EFF8C4B6FC7B09C9F171C5AAE" authorityName="gratiosa (Grum-Grshimailo" authorityYear="1890" baseAuthorityName="gratiosa (Grum-Grshimailo" baseAuthorityYear="1890" box="[151,330,1714,1741]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="gratiosa" subSpecies="gratiosa">
<emphasis id="B916940FFF8C4B6FC7B09C9F171C5AAE" box="[151,330,1714,1741]" italics="true" pageId="2" pageNumber="53">Ch. g. gratiosa</emphasis>
</taxonomicName>
(
<figureCitation id="13595498FF8C4B6FC67C9C9F17E35AAF" box="[347,437,1714,1741]" captionStart="Figures 2428" captionStartId="8.[151,236,1909,1933]" captionTargetBox="[155,1436,204,1841]" captionTargetId="figure-18@8.[151,1436,189,1872]" captionTargetPageId="8" captionText="Figures 2428. Chelis spp.: adults. Depositories of the specimens: 24 in ASV; 2528 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232648" httpUri="https://zenodo.org/record/13232648/files/figure.png" pageId="2" pageNumber="53">Fig. 25</figureCitation>
):
<emphasis id="B916940FFF8C4B6FC6EC9C9F14CF5AAF" box="[459,665,1714,1741]" italics="true" pageId="2" pageNumber="53">Ch. g. kartashovi</emphasis>
has a somewhat narrower ductus bursae and its appendix bursae is longer than that of the nominate subspecies.
</paragraph>
</subSubSection>
<subSubSection id="C3781B96FF8C4B69C7B09D3A14AF5B1C" lastPageId="4" lastPageNumber="55" pageId="2" pageNumber="53" type="description">
<paragraph id="8BDD481DFF8C4B69C7B09D3A14AF5B1C" blockId="2.[151,1436,1815,2044]" lastBlockId="4.[151,1436,1149,1918]" lastPageId="4" lastPageNumber="55" pageId="2" pageNumber="53">
<emphasis id="B916940FFF8C4B6FC7B09D3A177E5B53" bold="true" box="[151,296,1815,1841]" pageId="2" pageNumber="53">Description</emphasis>
.
<emphasis id="B916940FFF8C4B6FC61F9D3A14EC5B53" bold="true" box="[312,698,1815,1841]" pageId="2" pageNumber="53">External morphology of adults</emphasis>
(
<figureCitation id="13595498FF8C4B6FC5EC9D3A15605B50" box="[715,822,1815,1842]" captionStart="Figures 18" captionStartId="3.[151,236,1965,1989]" captionTargetBox="[175,1427,226,1921]" captionTargetId="figure-13@3.[151,1427,223,1946]" captionTargetPageId="3" captionText="Figures 18. Chelis spp.: adults. Depositories of the specimens: 1, 3 in ASV; 2 in WIGJ; 4 in SDM; 58 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232634" httpUri="https://zenodo.org/record/13232634/files/figure.png" pageId="2" pageNumber="53">Figs 14</figureCitation>
,
<figureCitation id="13595498FF8C4B6FC4649D3A15025B50" box="[835,852,1815,1842]" captionStart="Figure 9" captionStartId="4.[151,226,1050,1074]" captionTargetBox="[163,1425,212,1032]" captionTargetId="figure-18@4.[151,1436,189,1032]" captionTargetPageId="4" captionText="Figure 9. Chelis gratiosa kartashovi, paratypes. The specimens are deposited in VKV (photo by V. Kartashov)." figureDoi="http://doi.org/10.5281/zenodo.13232636" httpUri="https://zenodo.org/record/13232636/files/figure.png" pageId="2" pageNumber="53">9</figureCitation>
). Forewing length
<quantity id="4C9AE5F8FF8C4B6FC3129D3A12E65B50" box="[1077,1200,1815,1842]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="2" pageNumber="53" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11-12 mm</quantity>
in males and
<quantity id="4C9AE5F8FF8C4B6FC27F9D3A16935B31" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="2" pageNumber="53" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">11-13 mm</quantity>
in females. Male antenna shortly bipectinate, female antenna serrate. Sexual dimorphism limited: compared to males, females have forewing with more rounded apex and more reduced markings and dark suffusion on hindwing less intense. Head dark brown with admixture of pinkish scales. Collar dark brown with pinkish margin. Thorax blackish brown with two pinkish longitudinal stripes and pinkish margins of tegulae. Upperside of abdomen blackish brown with ochreous brown triangular spots laterally, underside ochreous brown; tip of abdomen densely covered in ochreous brown scales. Forewing ground color creamy with intense pinkish suffusion. Markings blackish brown. Subbasal spot asymmetrically V-shaped. Subbasal area with large elongate triangular spot. Antemedial line strongly angled medially, consisting of two large elliptical spots connected to each other by an oblique stroke and a costal dash. Postmedial line angled on the Cu vein dilated in the cell and at costa and fused with antemedial line at anal wing margin. Subterminal line interrupted, represented by four large triangular spots. Terminal area and anal margin blackish brown, inner edge of terminal area strongly zigzagged. Cilia pinkish brown. Hindwing with intense dark suffusion making it nearly uniformly blackish brown. Subterminal area with a small pale medial spot which may be protruding to the terminal area. Cilia brown.
<emphasis id="B916940FFF8A4B69C5039F47148159E6" bold="true" box="[548,727,1386,1412]" pageId="4" pageNumber="55">Male genitalia</emphasis>
(
<figureCitation id="13595498FF8A4B69C5C09F44156959E6" box="[743,831,1385,1412]" captionStart="Figures 1720" captionStartId="6.[151,236,2002,2026]" captionTargetBox="[163,1426,222,1936]" captionTargetId="figure-18@6.[151,1436,222,1961]" captionTargetPageId="6" captionText="Figures 1720. Chelis gratiosa sspp.: adults. Depositories of the specimens: 17 in ASV/WIGJ; 1820 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232642" httpUri="https://zenodo.org/record/13232642/files/figure.png" pageId="4" pageNumber="55">Fig. 17</figureCitation>
). Uncus large, broadly triangular with elongate and tapered distal part pointed apically. Tuba analis broad, membranous. Tegumen short, with thick and heavily sclerotized arms. Juxta large, X-shaped with wide arms, moderately sclerotized. Vinculum large, heavily sclerotized, U-shaped. Valva short, its basal and medial sections wide, cucullus ca. 3 times narrower than base of valva, rectangular with rounded corners. Sacculus narrow, heavily sclerotized. Aedeagus large, elongate, its distal half somewhat curved dorsad; dorsal carinal plate narrow, triangular, covered in tiny denticles. Vesica broad, C-like curved dorsad and anteriorly, with a small membranous subbasal diverticulum dorsally, four short granulated diverticula outwardly and globular membranous subapical diverticulum inwardly.
<emphasis id="B916940FFF8A4B69C6979C5514D65AF0" bold="true" box="[432,640,1656,1682]" pageId="4" pageNumber="55">Female genitalia</emphasis>
(
<figureCitation id="13595498FF8A4B69C5B49C5A14B85AF0" box="[659,750,1655,1682]" captionStart="Figures 2428" captionStartId="8.[151,236,1909,1933]" captionTargetBox="[155,1436,204,1841]" captionTargetId="figure-18@8.[151,1436,189,1872]" captionTargetPageId="8" captionText="Figures 2428. Chelis spp.: adults. Depositories of the specimens: 24 in ASV; 2528 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232648" httpUri="https://zenodo.org/record/13232648/files/figure.png" pageId="4" pageNumber="55">Fig. 24</figureCitation>
). Papillae anales wide, rectangular with rounded corners, setose. Apophyses posteriores elongate, thin, with rounded tips. Apophyses anteriores fully reduced. Postvaginal area with narrow pocket-like incision medially. Ostium bursae broad. Ductus bursae long, dorso-ventrally flattened, somewhat tapered anteriorly, its posterior three fifth heavily sclerotized, while anterior two fifth weakly sclerotized. Corpus bursae broad, elliptical, membranous, with area of weak granulation posteriorly. Medial section of corpus bursae armed with two round signa bursae covered with tiny denticles. Appendix bursae elongate with broad basal section, scobinated basally and membranous distally, situated postero-ventrally, directed laterally to the right side.
</paragraph>
</subSubSection>
<caption id="DF1D1895FF8D4B6EC7B09D8017735B81" ID-DOI="http://doi.org/10.5281/zenodo.13232634" ID-Zenodo-Dep="13232634" httpUri="https://zenodo.org/record/13232634/files/figure.png" pageId="3" pageNumber="54" startId="3.[151,236,1965,1989]" targetBox="[175,1427,226,1921]" targetPageId="3" targetType="figure">
<paragraph id="8BDD481DFF8D4B6EC7B09D8017735B81" blockId="3.[151,1436,1964,2019]" pageId="3" pageNumber="54">
<emphasis id="B916940FFF8D4B6EC7B09D80177F5BA7" bold="true" box="[151,297,1965,1989]" pageId="3" pageNumber="54">Figures 18.</emphasis>
<taxonomicName id="4C62339EFF8D4B6EC6109D8117EE5BA6" box="[311,440,1964,1988]" class="Insecta" family="Erebidae" genus="Chelis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B916940FFF8D4B6EC6109D81172D5BA6" box="[311,379,1964,1988]" italics="true" pageId="3" pageNumber="54">Chelis</emphasis>
spp.
</taxonomicName>
: adults. Depositories of the specimens: 1, 3 in ASV; 2 in WIGJ; 4 in SDM; 58 in MWM/ZSM.
</paragraph>
</caption>
<caption id="DF1D1895FF8A4B69C7B09E3713175850" ID-DOI="http://doi.org/10.5281/zenodo.13232636" ID-Zenodo-Dep="13232636" box="[151,1345,1050,1075]" httpUri="https://zenodo.org/record/13232636/files/figure.png" pageId="4" pageNumber="55" startId="4.[151,226,1050,1074]" targetBox="[163,1425,212,1032]" targetPageId="4" targetType="figure">
<paragraph id="8BDD481DFF8A4B69C7B09E3713175850" blockId="4.[151,1345,1050,1075]" box="[151,1345,1050,1075]" pageId="4" pageNumber="55">
<emphasis id="B916940FFF8A4B69C7B09E3716A85850" bold="true" box="[151,254,1050,1074]" pageId="4" pageNumber="55">Figure 9.</emphasis>
<taxonomicName id="4C62339EFF8A4B69C6239E3714485850" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" box="[260,542,1050,1074]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="gratiosa" subSpecies="kartashovi">
<emphasis id="B916940FFF8A4B69C6239E3714485850" box="[260,542,1050,1074]" italics="true" pageId="4" pageNumber="55">Chelis gratiosa kartashovi</emphasis>
</taxonomicName>
, paratypes. The specimens are deposited in VKV (photo by V. Kartashov).
</paragraph>
</caption>
<subSubSection id="C3781B96FF8A4B68C7B09D8A14265CBC" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55" type="distribution">
<paragraph id="8BDD481DFF8A4B69C7B09D8A14645467" blockId="4.[151,1435,1959,2053]" pageId="4" pageNumber="55">
<emphasis id="B916940FFF8A4B69C7B09D8A17A75BA3" bold="true" box="[151,497,1959,1985]" pageId="4" pageNumber="55">Distribution and bionomics</emphasis>
. The new subspecies is known from Transalai Range in southern
<collectingCountry id="F375088DFF8A4B69C22B9D8513C15BA0" box="[1292,1431,1960,1986]" name="Kyrgyzstan" pageId="4" pageNumber="55">Kyrgyzstan</collectingCountry>
. Single female was found siting on ground in an alpine grassland (at ca
<quantity id="4C9AE5F8FF8A4B69C4FB9DE512635B81" box="[988,1077,1992,2019]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="55" unit="m" value="3600.0">3600 m</quantity>
altitude) where the vegetation was damaged by grazing livestock.
</paragraph>
<paragraph id="8BDD481DFF8B4B68C7B09AE914265CBC" blockId="5.[151,624,196,222]" box="[151,624,196,222]" pageId="5" pageNumber="56">
<emphasis id="B916940FFF8B4B68C7B09AE914265CBC" bold="true" box="[151,624,196,222]" pageId="5" pageNumber="56">Descriptions of brachipterous females</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>