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30 KiB
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246 lines
30 KiB
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<document id="3D22DE5D6AACEC41A6256489EDB230CA" ID-DOI="10.37520/aemnp.2021.009" ID-ISSN="1804-6487" ID-Zenodo-Dep="5037330" ID-ZooBank="27EA8466-6B29-40F4-8236-DD49061C523B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624895377019" checkinUser="carolina" docAuthor="Liu, Tian-tian, Ono, Hiroki & Maruyama, Munetoshi" docDate="2021" docId="03CB023BAC07FF8F4FEFF8CF13F1FA6C" docLanguage="en" docName="ActaEntMusNatPra.61.1.163-201.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2021.009" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Bryothinusa japonica Liu, Ono & Maruyama 2020" docType="treatment" docVersion="3" lastPageNumber="176" masterDocId="FFF27A43AC0AFF824F48FFFB175FFFCD" masterDocTitle="Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="201" masterPageNumber="163" pageNumber="176" updateTime="1698977316832" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="D8761C411CA436C82FBC939A24AFB28F">Revision of the intertidal rove beetle genus Bryothinusa from Japan (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="9748B3D3682761A0C28818171653282E">Liu, Tian-tian</mods:namePart>
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<mods:affiliation id="0D21F442F154101AE0F442E78E0EEFC4">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395, Japan & The Kyushu University Museum, Hakozaki</mods:affiliation>
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<mods:namePart id="64E5B044DEC9FC4A5C11931054A4EADD">Ono, Hiroki</mods:namePart>
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<mods:affiliation id="6884972F3FC6E7608C7A27ACCE6A03E7">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan</mods:affiliation>
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<mods:namePart id="A8A63B31715207C536DEC88F771A853C">Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation id="16E80073D72BDFD88C927BA684299CAE">The Kyushu University Museum, Hakozaki</mods:affiliation>
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<mods:nameIdentifier id="58C0E1C5DF6C80A7768FA27754B28D8E" type="email">dendrolasius@gmail.com</mods:nameIdentifier>
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<mods:title id="398070C68BAA940C3873220BB8711310">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="C9361D4BD2FCA0F3A535AB58B0BBDB17">2021</mods:date>
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<subSubSection id="C378E0A6AC07FF8F4FEFF8CF15B8F881" box="[167,743,1844,1868]" pageId="13" pageNumber="176" type="nomenclature">
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<paragraph id="8BDDB32DAC07FF8F4FEFF8CF15B8F881" blockId="13.[167,743,1844,1868]" box="[167,743,1844,1868]" pageId="13" pageNumber="176">
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<heading id="D0950441AC07FF8F4FEFF8CF15B8F881" box="[167,743,1844,1868]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="176" reason="3">
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<taxonomicName id="4C62C8AEAC07FF8F4FEFF8CF15B8F881" ID-CoL="8XW4L" authority="Liu, Ono & Maruyama, 2020" authorityName="Liu, Ono & Maruyama" authorityYear="2020" box="[167,743,1844,1868]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="japonica">
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<emphasis id="B9166F3FAC07FF8F4FEFF8CF15B8F881" bold="true" box="[167,743,1844,1868]" pageId="13" pageNumber="176">
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<emphasis id="B9166F3FAC07FF8F4FEFF8CF16CAF881" bold="true" box="[167,405,1844,1868]" italics="true" pageId="13" pageNumber="176">Bryothinusa japonica</emphasis>
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<bibRefCitation id="EFF3CEDCAC07FF8F4ED4F8CE15B8F881" author="LIU T. T. & ONO H. & MARUYAMA M." box="[412,743,1845,1868]" pageId="13" pageNumber="176" pagination="585 - 593" refId="ref25551" refString="LIU T. T., ONO H. & MARUYAMA M. 2020: A new species and new record of the intertidal rove beetle genus Bryothinusa Casey (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 4743 (4): 585 - 593." type="journal article" year="2020">Liu, Ono & Maruyama, 2020</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C378E0A6AC07FF8F4E0AF8A815D3F84C" pageId="13" pageNumber="176" type="description">
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<paragraph id="8BDDB32DAC07FF8F4E0AF8A81513F8AB" blockId="13.[151,761,1875,2077]" box="[322,588,1875,1894]" pageId="13" pageNumber="176">
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(
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<figureCitation id="1359AFA8AC07FF8F4E00F8A816D3F8AB" box="[328,396,1875,1894]" captionStart="Fig" captionStartId="2.[151,182,1972,1991]" captionTargetBox="[189,1405,205,1944]" captionTargetId="figure-13@2.[175,1411,189,1947]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Bryothinusa spp. A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.5037332" httpUri="https://zenodo.org/record/5037332/files/figure.png" pageId="13" pageNumber="176">Figs 1F</figureCitation>
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;
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<figureCitation id="1359AFA8AC07FF8F4EDDF8A816F0F8AB" box="[405,431,1875,1894]" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="13" pageNumber="176">2F</figureCitation>
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;
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<figureCitation id="1359AFA8AC07FF8F4EFFF8A8169CF8AB" box="[439,451,1875,1894]" captionStart="Fig" captionStartId="14.[151,182,2000,2019]" captionTargetBox="[159,1400,189,1927]" captionTargetId="figure-13@14.[155,1433,189,1977]" captionTargetPageId="14" captionText="Fig. 8. Bryothinusa japonica Liu, Ono & Maruyama, 2020. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm. Modified from Lංඎ et al. (2020)." figureDoi="http://doi.org/10.5281/zenodo.5037348" httpUri="https://zenodo.org/record/5037348/files/figure.png" pageId="13" pageNumber="176">8</figureCitation>
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;
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<figureCitation id="1359AFA8AC07FF8F4E85F8A81578F8AB" box="[461,551,1875,1894]" captionStart="Fig" captionStartId="34.[151,182,1955,1974]" captionTargetBox="[167,1408,193,1935]" captionTargetId="figure-13@34.[161,1426,189,1938]" captionTargetPageId="34" captionText="Fig. 21. Collection localities of Bryothinusa species in Japan: A – B. koreana Ahn & Jeon, 2004 and B. japonica Liu, Ono & Maruyama, 2020 in Shima-nogita, Fukuoka; B – B. aikoae sp. nov., B. fulvipennis sp. nov., B. japonica and B. nigra sp. nov. in Kuroto, Chiba; C – B. hokkaidensis sp. nov. in Shunkunitai, Hokkaidô; D – B. nigra sp. nov. and B. tsutsuii (Sawada, 1955) in Edateku-jima; E – B. nakanei (Sawada, 1955) in Nagura-anparu, Ishigaki-jima; F – B. gangjinensis Ahn & Jeon, 2004 in Iki-no-matsubara, Fukuoka; G – B. algarum Sawada, 1971, B. japonica and B. minuta (Sawada, 1955) in Sunosaki, Fukuoka." figureDoi="http://doi.org/10.5281/zenodo.5037374" httpUri="https://zenodo.org/record/5037374/files/figure.png" pageId="13" pageNumber="176">21A–B, G</figureCitation>
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;
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<figureCitation id="1359AFA8AC07FF8F4D78F8A81519F8AB" box="[560,582,1875,1894]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="13" pageNumber="176">22</figureCitation>
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)
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</paragraph>
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<paragraph id="8BDDB32DAC07FF8F4E4BF89515D3F84C" blockId="13.[151,761,1875,2077]" box="[259,652,1902,1921]" pageId="13" pageNumber="176">(Japanese name: Nippon-nagisa-hanekakushi)</paragraph>
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</subSubSection>
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<subSubSection id="C378E0A6AC07FF8F4FDFF8741529F870" pageId="13" pageNumber="176" type="reference_group">
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<paragraph id="8BDDB32DAC07FF8F4FDFF8741529F870" blockId="13.[151,761,1875,2077]" pageId="13" pageNumber="176">
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<treatmentCitationGroup id="AB729403AC07FF8F4FDFF8741529F870" pageId="13" pageNumber="176">
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<treatmentCitation id="0AC3953CAC07FF8F4FDFF87415FDF86E" ID-DOI="http://doi.org/10.5281/zenodo.3691439" ID-Zenodo-Dep="3691439" author="LIU T. T. & ONO H. & MARUYAMA M." box="[151,674,1935,1955]" httpUri="http://treatment.plazi.org/id/D5082F0FFFC4B54AFF223EE2FD78FF57" page="586" pageId="13" pageNumber="176" year="2020">
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<taxonomicName id="4C62C8AEAC07FF8F4FDFF87415FDF86E" ID-CoL="8XW4L" authority="Liu, Ono & Maruyama, 2020: 586" authorityName="Liu, Ono & Maruyama" authorityPageNumber="586" authorityYear="2020" box="[151,674,1935,1955]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="japonica">
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<emphasis id="B9166F3FAC07FF8F4FDFF8741605F86F" box="[151,346,1935,1954]" italics="true" pageId="13" pageNumber="176">Bryothinusa japonica</emphasis>
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<bibRefCitation id="EFF3CEDCAC07FF8F4E2BF86B15FDF86E" author="LIU T. T. & ONO H. & MARUYAMA M." box="[355,674,1936,1955]" pageId="13" pageNumber="176" pagination="585 - 593" refId="ref25551" refString="LIU T. T., ONO H. & MARUYAMA M. 2020: A new species and new record of the intertidal rove beetle genus Bryothinusa Casey (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 4743 (4): 585 - 593." type="journal article" year="2020">Liu, Ono & Maruyama, 2020: 586</bibRefCitation>
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(original description;
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||
<typeStatus id="54D90D8FAC07FF8F4E57F851161BF870" box="[287,324,1962,1981]" pageId="13" pageNumber="176">type</typeStatus>
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locality:
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<collectingCountry id="F375F3BDAC07FF8F4EDDF8511697F870" box="[405,456,1962,1981]" name="Japan" pageId="13" pageNumber="176">Japan</collectingCountry>
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, Kyûshû,
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<collectingRegion id="49A67DCFAC07FF8F4D55F8511533F870" box="[541,620,1962,1981]" country="Japan" name="Fukuoka" pageId="13" pageNumber="176">Fukuoka</collectingRegion>
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).
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C378E0A6AC07FF8F4FDFF82F130FFCEC" pageId="13" pageNumber="176" type="materials_examined">
|
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<paragraph id="8BDDB32DAC07FF8F4FDFF82F15A7F7D0" blockId="13.[151,761,1875,2077]" pageId="13" pageNumber="176">
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<emphasis id="B9166F3FAC07FF8F4FDFF82F16A8F82A" bold="true" box="[151,503,2004,2023]" pageId="13" pageNumber="176">
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Material examined.
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<collectingCountry id="F375F3BDAC07FF8F4E18F82F16C7F82A" box="[336,408,2004,2023]" name="Japan" pageId="13" pageNumber="176">JAPAN</collectingCountry>
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: HඈඇඌΗǕ:
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</emphasis>
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||
7 JJ
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<specimenCount id="9D6478A4AC07FF8F4D7BF82F150CF82A" box="[563,595,1998,2023]" pageId="13" pageNumber="176" type="female">1 ♀</specimenCount>
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||
, 22 unsexed spec., Matsumi-ga-ura,Ôchiba, Kosai-shi, Shizuoka-ken,
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<date id="FFDC95EDAC07FF8F4D0EF81415FEF7CF" box="[582,673,2031,2050]" pageId="13" pageNumber="176" value="2004-04-11">11.iv.2004</date>
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, Y.Tahira; 8 unsexed spec., Miya-gawa, Iso-chô,Ise-shi, Mie-ken,
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<date id="FFDC95EDAC07FF8F4D2AF7F115ECF7D0" box="[610,691,2058,2077]" pageId="13" pageNumber="176" value="2008-06-08">8.vi.2008</date>
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, K.Aki-
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</paragraph>
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<paragraph id="8BDDB32DAC07FF8F4C74FF3F1300FDFC" blockId="13.[828,1437,189,561]" pageId="13" pageNumber="176">
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ta; 4 JJ
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<specimenCount id="9D6478A4AC07FF8F4CC1FF3F14E9FF1B" box="[905,950,189,215]" pageId="13" pageNumber="176" type="female">2 ♀♀</specimenCount>
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, 67 unsexed spec., Kumozu-gawa, Karasu-chô, Mie-ken,
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<date id="FFDC95EDAC07FF8F4C74FF2514CEFF3C" box="[828,913,222,241]" pageId="13" pageNumber="176" value="2004-05-15">15.v.2004</date>
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, K.
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<collectingRegion id="49A67DCFAC07FF8F4CF8FF2514BDFF3C" box="[944,994,222,241]" country="Japan" name="Akita" pageId="13" pageNumber="176">Akita</collectingRegion>
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; 2 JJ
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<specimenCount id="9D6478A4AC07FF8F4B54FF251366FF3C" box="[1052,1081,216,241]" pageId="13" pageNumber="176" type="female">1♀</specimenCount>
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, 26 unsexed spec., Mouth of Kansa-gawa, Nansei-chô, Mie-ken,
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<date id="FFDC95EDAC07FF8F4CB2FF02131AFEC1" box="[1018,1093,249,268]" pageId="13" pageNumber="176" value="2008-05-05">5.v.2008</date>
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, K.
|
||
<collectingRegion id="49A67DCFAC07FF8F4B2DFF0213C8FEC1" box="[1125,1175,249,268]" country="Japan" name="Akita" pageId="13" pageNumber="176">Akita</collectingRegion>
|
||
; 4 JJ 4 unsexed spec., Mouth of Yoshii-gawa, Nishikôzai, Okayama-shi, Okayama-ken,
|
||
<date id="FFDC95EDAC07FF8F4A61FEEF1220FEEA" box="[1321,1407,276,295]" pageId="13" pageNumber="176" value="2004-10-24">24.x.2004</date>
|
||
, O. Yamaji; 2 JJ
|
||
<specimenCount id="9D6478A4AC07FF8F4CFFFED514B9FE8C" box="[951,998,296,321]" pageId="13" pageNumber="176" type="female">4 ♀♀</specimenCount>
|
||
, 6 unsexed spec., Mitsuguchi-kaigan, Yasuura-chô, Kure, Hiroshima-ken,
|
||
<date id="FFDC95EDAC07FF8F4CB2FEB21302FE91" box="[1018,1117,329,348]" pageId="13" pageNumber="176" value="2019-07-28">28.vii.2019</date>
|
||
, H. Yoshigou.
|
||
<emphasis id="B9166F3FAC07FF8F4B90FEB2126DFE91" bold="true" box="[1240,1330,329,348]" pageId="13" pageNumber="176">SΗංκඈκඎ:</emphasis>
|
||
2 JJ
|
||
<specimenCount id="9D6478A4AC07FF8F4A20FEB212C9FE96" box="[1384,1430,322,348]" pageId="13" pageNumber="176" type="female">5 ♀♀</specimenCount>
|
||
, 112 unsexed spec., Shimoda, Shimanto-shi, Kôchi-ken,
|
||
<date id="FFDC95EDAC07FF8F4A56FE9F1227FEBA" box="[1310,1400,356,375]" pageId="13" pageNumber="176" value="2012-04-21">21.iv.2012</date>
|
||
, M. Maruyama; 1 J
|
||
<specimenCount id="9D6478A4AC07FF8F4C84FE8514A1FE5C" box="[972,1022,376,401]" pageId="13" pageNumber="176" type="female">4 ♀♀</specimenCount>
|
||
, 2 unsexed spec., Kagami-gawa, Takajô-machi, Kôchi-shi, Kôchi-ken,
|
||
<date id="FFDC95EDAC07FF8F4B48FE621302FE61" box="[1024,1117,409,428]" pageId="13" pageNumber="176" value="2008-06-20">20.vi.2008</date>
|
||
, H. Fujimoto.
|
||
<emphasis id="B9166F3FAC07FF8F4B92FE621272FE61" bold="true" box="[1242,1325,409,428]" pageId="13" pageNumber="176">KඒǕඌΗǕ:</emphasis>
|
||
3 JJ
|
||
<specimenCount id="9D6478A4AC07FF8F4A2EFE6212D9FE66" box="[1382,1414,402,428]" pageId="13" pageNumber="176" type="female">1 ♀</specimenCount>
|
||
, 4 unsexed spec., Sunosaki, Imazu, Nishi-ku, Fukuoka-shi, Fukuoka-ken,
|
||
<date id="FFDC95EDAC07FF8F4C74FE3514CCFE2C" box="[828,915,462,481]" pageId="13" pageNumber="176" value="2020-10-16">16.x.2020</date>
|
||
,
|
||
<collectionCode id="ED732BE8AC07FF8F4CD2FE3514F7FE2C" box="[922,936,462,481]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="176">T</collectionCode>
|
||
. Liu &
|
||
<collectionCode id="ED732BE8AC07FF8F4CADFE3514ACFE2C" box="[997,1011,462,481]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="176">T</collectionCode>
|
||
. Nozaki; 8 unsexed spec., Mouth of Sakurai-gawa, Shima-Nogita, Itoshima-shi, Fukuoka-ken,
|
||
<date id="FFDC95EDAC07FF8F4BE6FE121255FE31" box="[1198,1290,489,508]" pageId="13" pageNumber="176" value="2020-04-21">21.iv.2020</date>
|
||
, Y. Hisasue;
|
||
<specimenCount id="9D6478A4AC07FF8F4A3FFE1212C8FE36" box="[1399,1431,482,508]" pageId="13" pageNumber="176" type="female">1 ♀</specimenCount>
|
||
, same locality,
|
||
<date id="FFDC95EDAC07FF8F4CFEFDFF1352FDDA" box="[950,1037,516,535]" pageId="13" pageNumber="176" value="2018-10-11">11.x.2018</date>
|
||
, Liu et al. Deposited at
|
||
<collectionCode id="ED732BE8AC07FF8F4B90FDFF1255FDDA" box="[1240,1290,516,535]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="13" pageNumber="176" type="Herbarium">KUM</collectionCode>
|
||
but some will be distributed to
|
||
<collectionCode id="ED732BE8AC07FF8F4CFAFDE514B1FDFC" box="[946,1006,542,561]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="13" pageNumber="176">NMPC</collectionCode>
|
||
and
|
||
<collectionCode id="ED732BE8AC07FF8F4B50FDE51304FDFC" box="[1048,1115,542,561]" pageId="13" pageNumber="176">BMNH</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDDB32DAC07FF8F4C74FDB1130FFCEC" blockId="13.[828,1437,586,1441]" pageId="13" pageNumber="176">
|
||
<emphasis id="B9166F3FAC07FF8F4C74FDB1131CFDAC" bold="true" box="[828,1091,586,609]" pageId="13" pageNumber="176">Additional description.</emphasis>
|
||
Number of facets 23–25 as
|
||
<figureCitation id="1359AFA8AC07FF8F4A3AFDB11406FD4C" captionStart="Fig" captionStartId="4.[151,182,1978,1997]" captionTargetBox="[196,1391,190,1955]" captionTargetId="figure-13@4.[193,1394,189,1958]" captionTargetPageId="4" captionText="Fig. 2. Compound eyes of Bryothinusa spp.A – B. algarum Sawada, 1971; B – B. minuta (Sawada, 1955); C – B. tsutsuii (Sawada, 1955); D – B. koreana Ahn & Jeon, 2004; E – B. yoshigoui sp. nov.; F – B. japonica Liu, Ono & Maruyama, 2020; G – B. sakishimana Sawada, 1991; H – B. constricta sp. nov.; I – B. hokkaidensis sp. nov.; J – B. fulvipennis sp. nov.; K – B. itsuroi sp. nov.; L – B. aikoae sp. nov.; M – B. okinawana sp. nov.; N – B. nigra sp. nov.; O – B. nakanei (Sawada, 1955); P – B. moriguchii sp. nov.; Q – B. gangjinensis Ahn & Jeon, 2004. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5037334" httpUri="https://zenodo.org/record/5037334/files/figure.png" pageId="13" pageNumber="176">
|
||
Fig.
|
||
<specimenCount id="9D6478A4AC07FF8F4C74FD911406FD4C" box="[828,857,618,641]" pageId="13" pageNumber="176" type="generic">2F</specimenCount>
|
||
</figureCitation>
|
||
. Antennal segment I 1.2 times longer than II, almost 2.4 times longer than III, and longer than XI. Mouthparts with median tooth longer than 1/4 of mandibles; 6 setae on inner margin of lacinia; male and female abdominal sternite VIII, spermatheca and aedeagus as in
|
||
<figureCitation id="1359AFA8AC07FF8F4A71FD1112C7FCCC" box="[1337,1432,746,769]" captionStart="Fig" captionStartId="14.[151,182,2000,2019]" captionTargetBox="[159,1400,189,1927]" captionTargetId="figure-13@14.[155,1433,189,1977]" captionTargetPageId="14" captionText="Fig. 8. Bryothinusa japonica Liu, Ono & Maruyama, 2020. A – labrum; B – mandible, dorsal view; C – lacinia, dorsal view; D – labium; E – mentum; F – male sternite VIII; G – female sternite VIII; H – median lobe, ventral view; I – median lobe, lateral view; J – paramere; K – spermatheca. Scale bars: 0.1 mm. Modified from Lංඎ et al. (2020)." figureDoi="http://doi.org/10.5281/zenodo.5037348" httpUri="https://zenodo.org/record/5037348/files/figure.png" pageId="13" pageNumber="176">Figure 8</figureCitation>
|
||
. See also Lංඎ et al. (2020).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC07FF8F4C74FCD114E5FC0C" pageId="13" pageNumber="176" type="diagnosis">
|
||
<paragraph id="8BDDB32DAC07FF8F4C74FCD114E5FC0C" blockId="13.[828,1437,586,1441]" pageId="13" pageNumber="176">
|
||
<emphasis id="B9166F3FAC07FF8F4C74FCD1136DFC8C" bold="true" box="[828,1074,810,833]" pageId="13" pageNumber="176">Differential diagnosis.</emphasis>
|
||
In general appearance, the species is closely allied to
|
||
<taxonomicName id="4C62C8AEAC07FF8F4B49FCB11334FCAC" authorityName="" baseAuthorityName="Sawada" baseAuthorityYear="1955" box="[1025,1131,842,865]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="tsutsuii">
|
||
<emphasis id="B9166F3FAC07FF8F4B49FCB11334FCAC" box="[1025,1131,842,865]" italics="true" pageId="13" pageNumber="176">B. tsutsuii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C62C8AEAC07FF8F4B30FCB113B0FCAC" authorityName="Sawada" authorityYear="1971" box="[1144,1263,841,865]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="algarum">
|
||
<emphasis id="B9166F3FAC07FF8F4B30FCB113B0FCAC" box="[1144,1263,841,865]" italics="true" pageId="13" pageNumber="176">B. algarum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C62C8AEAC07FF8F4BB4FCB11230FCAC" authorityName="Ahn & Jeon" authorityYear="2004" box="[1276,1391,841,865]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="koreana">
|
||
<emphasis id="B9166F3FAC07FF8F4BB4FCB11230FCAC" box="[1276,1391,841,865]" italics="true" pageId="13" pageNumber="176">B. koreana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C62C8AEAC07FF8F4C74FC91149DFC4C" authority="Liu & Ono & Maruyama, 2021" authorityName="Liu & Ono & Maruyama" authorityYear="2021" box="[828,962,873,897]" class="Insecta" family="Staphylinidae" genus="Bryothinusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="176" phylum="Arthropoda" rank="species" species="yoshigoui" status="sp. nov.">
|
||
<emphasis id="B9166F3FAC07FF8F4C74FC91149DFC4C" box="[828,962,873,897]" italics="true" pageId="13" pageNumber="176">B. yoshigoui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A225D244AC07FF8F4C80FC911341FC4C" box="[968,1054,874,897]" pageId="13" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, but easily distinguished from them by the apical part of spermatheca without granule-like projections.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC07FF8F4C74FC3112C2FB8C" pageId="13" pageNumber="176" type="distribution">
|
||
<paragraph id="8BDDB32DAC07FF8F4C74FC3112C2FB8C" blockId="13.[828,1437,586,1441]" pageId="13" pageNumber="176">
|
||
<emphasis id="B9166F3FAC07FF8F4C74FC311491FC2C" bold="true" box="[828,974,970,993]" pageId="13" pageNumber="176">Distribution.</emphasis>
|
||
<collectingCountry id="F375F3BDAC07FF8F4C9CFC31134EFC2C" box="[980,1041,970,993]" name="Japan" pageId="13" pageNumber="176">Japan</collectingCountry>
|
||
(
|
||
<emphasis id="B9166F3FAC07FF8F4B57FC3113DEFC2C" bold="true" box="[1055,1153,970,993]" pageId="13" pageNumber="176">Honshû:</emphasis>
|
||
Chiba-ken, Shizuoka-ken, Mie-ken, Ôsaka-fu, Hyôgo-ken, Wakayama-ken, Okayama-ken, Hiroshima-ken, Yamaguchi-ken;
|
||
<emphasis id="B9166F3FAC07FF8F4BB8FBF11208FBEC" bold="true" box="[1264,1367,1034,1057]" pageId="13" pageNumber="176">Shikoku:</emphasis>
|
||
Kagawa-ken, Ehime-ken, Kôchi-ken;
|
||
<emphasis id="B9166F3FAC07FF8F4BD0FBD113A5FB8C" bold="true" box="[1176,1274,1066,1089]" pageId="13" pageNumber="176">Kyûshû:</emphasis>
|
||
Fukuoka-ken).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C378E0A6AC07FF8F4C74FBB113F1FA6C" pageId="13" pageNumber="176" type="discussion">
|
||
<paragraph id="8BDDB32DAC07FF8F4C74FBB11395FAEC" blockId="13.[828,1437,586,1441]" pageId="13" pageNumber="176">
|
||
<emphasis id="B9166F3FAC07FF8F4C74FBB114F4FBAC" bold="true" box="[828,939,1098,1121]" pageId="13" pageNumber="176">Remarks.</emphasis>
|
||
About
|
||
<specimenCount id="9D6478A4AC07FF8F4B4AFBB113FBFBAC" box="[1026,1188,1098,1121]" pageId="13" pageNumber="176" type="generic">200 specimens</specimenCount>
|
||
were collected before the original description but they were not included in the
|
||
<typeStatus id="54D90D8FAC07FF8F4C74FB711435FB6C" box="[828,874,1162,1185]" pageId="13" pageNumber="176">type</typeStatus>
|
||
series due to the difficulty of identification. There was considerable variation in the colouration of the specimens, ranging from light brown to brown, but it is unclear whether this is due to geographical variation or to the methods in which the specimens were preserved.
|
||
</paragraph>
|
||
<paragraph id="8BDDB32DAC07FF8F4C12FAD113F1FA6C" blockId="13.[828,1437,586,1441]" pageId="13" pageNumber="176">
|
||
This species is quite common in sandy to mud flats in estuaries in
|
||
<collectingCountry id="F375F3BDAC07FF8F4CA9FAB1137EFAAC" box="[993,1057,1354,1377]" name="Japan" pageId="13" pageNumber="176">Japan</collectingCountry>
|
||
, mainly west of Kantô District. A distribution map of examined specimens of this species in
|
||
<collectingCountry id="F375F3BDAC07FF8F4C1FFA7114CBFA6C" box="[855,916,1418,1441]" name="Japan" pageId="13" pageNumber="176">Japan</collectingCountry>
|
||
is illustrated in
|
||
<figureCitation id="1359AFA8AC07FF8F4B08FA7113F5FA6C" box="[1088,1194,1418,1441]" captionStart="Fig" captionStartId="35.[150,181,1388,1407]" captionTargetBox="[150,1434,191,1370]" captionTargetId="figure-236@35.[150,1435,191,1371]" captionTargetPageId="35" captionText="Fig. 22. Map of examined specimens of Bryothinusa spp. from Japan: white inverted triangle – B. algarum Sawada, 1971; black inverted triangle – B. minuta (Sawada, 1955); white star – B. tsutsuii (Sawada, 1955); black star – B. koreana Ahn & Jeon, 2004; black square – B. yoshigoui sp. nov.; white triangle – B. japonica Liu, Ono & Maruyama, 2020; black diamond – B. sakishimana Sawada, 1991; white diamond – B. constricta sp. nov.; white square – B. hokkaidensis sp. nov.; black triangle – B. fulvipennis sp. nov.; double circle – B. itsuroi sp. nov.; arrow – B. aikoae sp. nov.; black hart – B. okinawana sp. nov.; circled dot – B. nigra sp. nov.; white circle – B. nakanei (Sawada, 1955); white hart – B. moriguchii sp. nov.; black circle – B. gangjinensis Ahn & Jeon, 2004." figureDoi="http://doi.org/10.5281/zenodo.5037376" httpUri="https://zenodo.org/record/5037376/files/figure.png" pageId="13" pageNumber="176">Figure 22</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |