190 lines
23 KiB
XML
190 lines
23 KiB
XML
<document id="03FE4B5FDD125C8666EEB5B947F3AF0A" ID-DOI="10.11646/zootaxa.4093.2.7" ID-GBIF-Dataset="4ece7a98-bd32-4fb5-acc4-ecb7128b8382" ID-ISSN="1175-5326" ID-Zenodo-Dep="267656" ID-ZooBank="9EBEA915-88BA-473F-9CC5-A944240D7095" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458649234170" checkinUser="plazi" docAuthor="Cumming, Jeffrey M., Brooks, Scott E. & Sinclair, Bradley J." docDate="2016" docId="03C84D7AFF8CFFC3FF25FDB3FCA5F834" docLanguage="en" docName="zootaxa.4093.2.7.pdf.imf" docOrigin="Zootaxa 4093 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Philetus Melander" docType="treatment" docVersion="9" lastPageNumber="263" masterDocId="FFF13502FF8EFFC1FFB2FF95FFF3FF99" masterDocTitle="Review of the little-known western Nearctic fly genus Philetus Melander (Diptera: Empididae), with a discussion of its phylogenetic assignment" masterLastPageNumber="274" masterPageNumber="261" pageNumber="263" updateTime="1698185882827" updateUser="plazi">
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<mods:mods id="C3323928C4EB30994F83061C75316867" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5EDB9962D18882797FFF8963C2F89469">
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<mods:title id="3B4AB0BB89E690C7C6C603D8D4D4430F">Review of the little-known western Nearctic fly genus Philetus Melander (Diptera: Empididae), with a discussion of its phylogenetic assignment</mods:title>
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<mods:name id="260B4137415069003595DC10C60D7A9A" type="personal">
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<mods:namePart id="0A1BB06C3922EBDB57CF6A9AE8C6DD8E">Cumming, Jeffrey M.</mods:namePart>
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<mods:namePart id="4468DA0FA767AE370194C15B5B86B35B">Brooks, Scott E.</mods:namePart>
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<mods:roleTerm id="410D2AACB7BD8096202A2AC104DF1F6C">Author</mods:roleTerm>
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<mods:namePart id="B369A1058AF5B0425AD91A794EB66AAB">Sinclair, Bradley J.</mods:namePart>
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<mods:title id="0568490AECDE724FD24A2195D17CA1C1">Zootaxa</mods:title>
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<mods:part id="49C99032EEFFD7B4116E94EA71814C91">
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<mods:date id="019292C68D79E3E36B75198654C78E6C">2016</mods:date>
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<mods:detail id="255398860AF08276082996CBB78ACF58" type="volume">
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<mods:number id="9661ABCF7438D98026AE6842744D57E8">4093</mods:number>
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<mods:identifier id="2B6C9A98116AA9E1C2F4F44EB0A01529" type="DOI">10.11646/zootaxa.4093.2.7</mods:identifier>
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<treatment id="03C84D7AFF8CFFC3FF25FDB3FCA5F834" ID-DOI="http://doi.org/10.5281/zenodo.5696400" ID-GBIF-Taxon="119209038" ID-Zenodo-Dep="5696400" LSID="urn:lsid:plazi:treatment:03C84D7AFF8CFFC3FF25FDB3FCA5F834" httpUri="http://treatment.plazi.org/id/03C84D7AFF8CFFC3FF25FDB3FCA5F834" lastPageNumber="263" pageId="2" pageNumber="263">
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<subSubSection id="C37BAFE7FF8CFFC3FF25FDB3FE20FDD9" box="[151,467,550,576]" pageId="2" pageNumber="263" type="nomenclature">
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<paragraph id="8BDEFC6CFF8CFFC3FF25FDB3FE20FDD9" blockId="2.[151,467,550,576]" box="[151,467,550,576]" pageId="2" pageNumber="263">
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<heading id="D0964B00FF8CFFC3FF25FDB3FE20FDD9" bold="true" box="[151,467,550,576]" fontSize="11" level="1" pageId="2" pageNumber="263" reason="1">
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<emphasis id="B915207EFF8CFFC3FF25FDB3FE20FDD9" bold="true" box="[151,467,550,576]" pageId="2" pageNumber="263">
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Genus
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<taxonomicName id="4C6187EFFF8CFFC3FF5CFDB3FE20FDD9" ID-CoL="6ND5" ID-ENA="1713971" authority="Melander" authorityName="Melander" box="[238,467,550,576]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="B915207EFF8CFFC3FF5CFDB3FEA2FDD9" bold="true" box="[238,337,550,576]" italics="true" pageId="2" pageNumber="263">Philetus</emphasis>
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Melander
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37BAFE7FF8CFFC3FF25FDF8FDE2FD58" pageId="2" pageNumber="263" type="reference_group">
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<paragraph id="8BDEFC6CFF8CFFC3FF25FDF8FDE2FD58" blockId="2.[151,1436,621,705]" pageId="2" pageNumber="263">
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<taxonomicName id="4C6187EFFF8CFFC3FF25FDF8FE3DFD1A" ID-CoL="6ND5" authority="Melander, 1928: 110" authorityName="Melander" authorityPageNumber="110" authorityYear="1928" box="[151,462,621,643]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="B915207EFF8CFFC3FF25FDF8FF19FD1A" box="[151,234,621,643]" italics="true" pageId="2" pageNumber="263">Philetus</emphasis>
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Melander, 1928: 110
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</taxonomicName>
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.
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<typeStatus id="54DA42CEFF8CFFC3FE69FDFBFDFEFD1A" box="[475,525,622,643]" pageId="2" pageNumber="263">Type</typeStatus>
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species,
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<taxonomicName id="4C6187EFFF8CFFC3FDC2FDF8FC0FFD1A" ID-CoL="779YZ" authority="Melander, 1928" authorityName="Melander" authorityYear="1928" box="[624,1020,621,643]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="memorandus">
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<emphasis id="B915207EFF8CFFC3FDC2FDF8FCBCFD1A" box="[624,847,621,643]" italics="true" pageId="2" pageNumber="263">Philetus memorandus</emphasis>
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Melander, 1928
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</taxonomicName>
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(original designation). Melander, 1965: 455; Poole, 1996: 158, 368; Sinclair & Cumming, 2006: 74, 102–105, 107; Yang
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<emphasis id="B915207EFF8CFFC3FBBDFD18FBCEFD3B" box="[1039,1085,652,674]" italics="true" pageId="2" pageNumber="263">et al</emphasis>
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., 2007: 350;
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<collectingCountry id="F376BCFCFF8CFFC3FB74FD19FAD0FD3B" box="[1222,1315,652,674]" name="Marshall Islands" pageId="2" pageNumber="263">Marshall</collectingCountry>
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, 2012: 288; Cumming
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<emphasis id="B915207EFF8CFFC3FE80FD39FEACFD58" box="[306,351,683,705]" italics="true" pageId="2" pageNumber="263">et al</emphasis>
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., 2014: 171, 205.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37BAFE7FF8CFFC3FF25FD7DFB95FA4D" pageId="2" pageNumber="263" type="diagnosis">
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<paragraph id="8BDEFC6CFF8CFFC3FF25FD7DFDA2FAFE" blockId="2.[151,1437,744,1965]" pageId="2" pageNumber="263">
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<emphasis id="B915207EFF8CFFC3FF25FD7DFEF9FC98" bold="true" box="[151,266,744,769]" pageId="2" pageNumber="263">Diagnosis</emphasis>
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. The genus is characterized by the following features: small size (body length 3.0–5.0 mm) brownish grey flies with silvery grey tomentum (
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<figureCitation id="135AE0E9FF8CFFC3FDE1FC99FD69FCBD" box="[595,666,780,805]" captionStart="FIGURES 1 − 3" captionStartId="3.[151,263,1628,1650]" captionTargetBox="[151,1436,195,1603]" captionTargetId="figure@3.[151,1436,195,1603]" captionTargetPageId="3" captionText="FIGURES 1 − 3. Philetus habitus photograph and habitat. 1. Philetus sp., live female; photograph by S. A. Marshall. 2. Sitka spruce canopy, Upper Carmanah Valley, Vancouver Island, British Columbia. 3. Coastal wet temperate rainforest, Upper Carmanah Valley." httpUri="https://zenodo.org/record/267657/files/figure.png" pageId="2" pageNumber="263">Figs 1</figureCitation>
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,
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<figureCitation id="135AE0E9FF8CFFC3FD1BFC99FD49FCBD" box="[681,698,780,804]" captionStart="FIGURES 4 − 8" captionStartId="5.[151,263,1379,1401]" captionTargetBox="[172,1436,177,1357]" captionTargetId="figure@5.[151,1436,177,1358]" captionTargetPageId="5" captionText="FIGURES 4 − 8. Philetus morphology. 4. P. memorandus, pinned male. 5. P. schizophorus, male head, lateral view. 6. P. schizophorus, male head, anterior view. 7. P. memorandus, female antenna. 8. P. schizophorus, female antenna. Abbreviation: pped—postpedicel." httpUri="https://zenodo.org/record/267658/files/figure.png" pageId="2" pageNumber="263">4</figureCitation>
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); head produced obliquely downwards; ocelli positioned forward on broad V-shaped frons; eyes bare, dichoptic in male and female; inner eye incision present; antenna inserted just below middle of eye (
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<figureCitation id="135AE0E9FF8CFFC3FE24FCC1FDF3FCF4" box="[406,512,852,877]" captionStart="FIGURES 4 − 8" captionStartId="5.[151,263,1379,1401]" captionTargetBox="[172,1436,177,1357]" captionTargetId="figure@5.[151,1436,177,1358]" captionTargetPageId="5" captionText="FIGURES 4 − 8. Philetus morphology. 4. P. memorandus, pinned male. 5. P. schizophorus, male head, lateral view. 6. P. schizophorus, male head, anterior view. 7. P. memorandus, female antenna. 8. P. schizophorus, female antenna. Abbreviation: pped—postpedicel." httpUri="https://zenodo.org/record/267658/files/figure.png" pageId="2" pageNumber="263">Figs 5, 6</figureCitation>
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); apical stylus slightly tapered with 3 articles including minute terminal hyaline process, subequal to or slightly shorter than length of pyriform postpedicel (
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<figureCitation id="135AE0E9FF8CFFC3FC5CFCEDFBA0FC09" box="[1006,1107,888,913]" captionStart="FIGURES 4 − 8" captionStartId="5.[151,263,1379,1401]" captionTargetBox="[172,1436,177,1357]" captionTargetId="figure@5.[151,1436,177,1358]" captionTargetPageId="5" captionText="FIGURES 4 − 8. Philetus morphology. 4. P. memorandus, pinned male. 5. P. schizophorus, male head, lateral view. 6. P. schizophorus, male head, anterior view. 7. P. memorandus, female antenna. 8. P. schizophorus, female antenna. Abbreviation: pped—postpedicel." httpUri="https://zenodo.org/record/267658/files/figure.png" pageId="2" pageNumber="263">Figs 7, 8</figureCitation>
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); proboscis moderately long; epipharyngeal blades movable; palpus straight, directed forward (
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<figureCitation id="135AE0E9FF8CFFC3FC3AFC09FC01FC2C" box="[904,1010,924,949]" captionStart="FIGURES 4 − 8" captionStartId="5.[151,263,1379,1401]" captionTargetBox="[172,1436,177,1357]" captionTargetId="figure@5.[151,1436,177,1358]" captionTargetPageId="5" captionText="FIGURES 4 − 8. Philetus morphology. 4. P. memorandus, pinned male. 5. P. schizophorus, male head, lateral view. 6. P. schizophorus, male head, anterior view. 7. P. memorandus, female antenna. 8. P. schizophorus, female antenna. Abbreviation: pped—postpedicel." httpUri="https://zenodo.org/record/267658/files/figure.png" pageId="2" pageNumber="263">Figs 5, 6</figureCitation>
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); head and thoracic chaetotaxy dark; mesonotal setae on each side with a few short irregular presutural acrostichal setae, row of 4–6 long dorsocentral and 3–4 intra-alar setae, 1–2 postpronotal setae, 1–2 presutural supra-alar setae, 2 notopleural setae, 1 postalar seta, and 2 (pairs) of long scutellar setae; mesopleuron and laterotergite bare; legs without prominent bristles; wing (Fig. 9) with venation complete (for Eremoneura), including nearly straight Sc reaching C, forked R4+5, cell cua angled apically with CuA curved back towards short evanescent CuA+CuP vein, basal costal bristle present,
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<taxonomicName id="4C6187EFFF8CFFC3FAAAFBC6FA6FFBF5" box="[1304,1436,1107,1132]" pageId="2" pageNumber="263" rank="species" species="pterostigma">pterostigma</taxonomicName>
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faint, anal lobe developed; male terminalia (Figs 11–20) with median apical hypandrial process, gonocoxal apodeme projecting anteriorly as process from anterodorsal margin of hypandrium, postgonite broad, ejaculatory apodeme lever-like and articulated to base of phallus, phallus tubular with desclerotized apex, epandrium deeply emarginate with left and right lamella connected basally, epandrial lobe present apically, proctiger differentiated into a long ventral and short medial subepandrial lobe attached to prolonged dorsal cercus; female terminalia (Fig. 10) with terminal segments partially telescopic, tergite 10 undivided without acanthophorites, cercus narrow, spermatheca unpigmented and sac-like.
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</paragraph>
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<paragraph id="8BDEFC6CFF8CFFC3FF75FAE6FB95FA4D" blockId="2.[151,1437,744,1965]" pageId="2" pageNumber="263">
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<emphasis id="B915207EFF8CFFC3FF75FAE6FE0DFA15" bold="true" box="[199,510,1395,1420]" pageId="2" pageNumber="263">Geographical distribution</emphasis>
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(
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<figureCitation id="135AE0E9FF8CFFC3FDBCFAE6FD91FA12" box="[526,610,1395,1420]" captionStart="FIGURES 21 − 23" captionStartId="10.[151,263,1620,1642]" captionTargetBox="[152,1433,632,1590]" captionTargetId="figure@10.[151,1436,628,1599]" captionTargetPageId="10" captionText="FIGURES 21 − 23. Distribution of Philetus. 21. Overall distribution of the genus Philetus, including undetermined female specimens. 22. Known distribution of P. memorandus. 23. Known distribution of P. schizophorus." httpUri="https://zenodo.org/record/267661/files/figure.png" pageId="2" pageNumber="263">Fig. 21</figureCitation>
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). Known from Alaska and the Yukon Territory in the north to California, Nevada and Arizona in the south, including records from British
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<collectingCountry id="F376BCFCFF8CFFC3FC1CFA03FBECFA36" box="[942,1055,1430,1455]" name="United States of America" pageId="2" pageNumber="263">Columbia</collectingCountry>
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, Washington, Oregon, Montana, Wyoming and Colorado (with undetermined female specimens listed in the Appendix).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37BAFE7FF8CFFC3FF75FA4BFCA5F834" pageId="2" pageNumber="263" type="discussion">
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<paragraph id="8BDEFC6CFF8CFFC3FF75FA4BFB3BF880" blockId="2.[151,1437,744,1965]" pageId="2" pageNumber="263">
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<emphasis id="B915207EFF8CFFC3FF75FA4BFEC7FA6E" bold="true" box="[199,308,1502,1527]" pageId="2" pageNumber="263">Remarks</emphasis>
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. In the Nearctic Region, this genus is most similar to
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<taxonomicName id="4C6187EFFF8CFFC3FC00FA75FB4AFA6E" authority="Melander" authorityName="Melander" box="[946,1209,1502,1527]" class="Insecta" family="Empididae" genus="Hesperempis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="B915207EFF8CFFC3FC00FA75FBB0FA6E" box="[946,1091,1504,1527]" italics="true" pageId="2" pageNumber="263">Hesperempis</emphasis>
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Melander
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</taxonomicName>
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and somewhat like
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<taxonomicName id="4C6187EFFF8CFFC3FF25F996FEB0F985" authority="Meigen" authorityName="Meigen" box="[151,323,1539,1564]" class="Insecta" family="Empididae" genus="Hilara" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="B915207EFF8CFFC3FF25F996FF11F982" box="[151,226,1539,1563]" italics="true" pageId="2" pageNumber="263">Hilara</emphasis>
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Meigen
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</taxonomicName>
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, but can be most easily distinguished from these empidid genera by the oblique form of the head with forward positioned ocelli and low inserted antennae.
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<taxonomicName id="4C6187EFFF8CFFC3FCAFF9B2FC8AF9A6" box="[797,889,1575,1599]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="B915207EFF8CFFC3FCAFF9B2FC8AF9A6" box="[797,889,1575,1599]" italics="true" pageId="2" pageNumber="263">Philetus</emphasis>
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</taxonomicName>
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also has straight forward-directed palps and vein Sc nearly straight and joining C at an acute angle unlike
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<taxonomicName id="4C6187EFFF8CFFC3FC86F9DEFC8CF9FA" box="[820,895,1611,1635]" class="Insecta" family="Empididae" genus="Hilara" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="B915207EFF8CFFC3FC86F9DEFC8CF9FA" box="[820,895,1611,1635]" italics="true" pageId="2" pageNumber="263">Hilara</emphasis>
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</taxonomicName>
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. In addition,
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<taxonomicName id="4C6187EFFF8CFFC3FBADF9DEFB99F9FA" box="[1055,1130,1611,1635]" class="Insecta" family="Empididae" genus="Hilara" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="B915207EFF8CFFC3FBADF9DEFB99F9FA" box="[1055,1130,1611,1635]" italics="true" pageId="2" pageNumber="263">Hilara</emphasis>
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</taxonomicName>
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and most members of the Hilarini have vein R1 swollen prior to reaching C and males usually have the first tarsomere of the foreleg expanded (Sinclair & Cumming 2006). In their key to Nearctic genera of
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<taxonomicName id="4C6187EFFF8CFFC3FC78F900FAC2F934" authority="Steyskal & Knutson" authorityName="Steyskal & Knutson" box="[970,1329,1685,1710]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="family">Empididae, Steyskal & Knutson</taxonomicName>
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(1981, p. 617) refer to a complete lack of thoracic setae in
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<taxonomicName id="4C6187EFFF8CFFC3FD07F92FFCB5F948" box="[693,838,1722,1745]" class="Insecta" family="Empididae" genus="Hesperempis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="B915207EFF8CFFC3FD07F92FFCB5F948" box="[693,838,1722,1745]" italics="true" pageId="2" pageNumber="263">Hesperempis</emphasis>
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</taxonomicName>
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compared with
|
||
<taxonomicName id="4C6187EFFF8CFFC3FC4BF92FFBA6F94B" box="[1017,1109,1722,1746]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915207EFF8CFFC3FC4BF92FFBA6F94B" box="[1017,1109,1722,1746]" italics="true" pageId="2" pageNumber="263">Philetus</emphasis>
|
||
</taxonomicName>
|
||
, but this does not apply to all the included species of
|
||
<taxonomicName id="4C6187EFFF8CFFC3FE14F94BFCB8F96F" authority="Cumming et al. 2014" authorityName="Cumming et al." authorityYear="2014" box="[422,843,1757,1782]" class="Insecta" family="Empididae" genus="Hesperempis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915207EFF8CFFC3FE14F94BFDC4F96C" box="[422,567,1758,1781]" italics="true" pageId="2" pageNumber="263">Hesperempis</emphasis>
|
||
(Cumming
|
||
<emphasis id="B915207EFF8CFFC3FD71F94BFD05F96C" box="[707,758,1757,1781]" italics="true" pageId="2" pageNumber="263">et al</emphasis>
|
||
. 2014)
|
||
</taxonomicName>
|
||
. However, the dark setae on the head and thorax of
|
||
<taxonomicName id="4C6187EFFF8CFFC3FF25F897FF00F883" box="[151,243,1794,1818]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915207EFF8CFFC3FF25F897FF00F883" box="[151,243,1794,1818]" italics="true" pageId="2" pageNumber="263">Philetus</emphasis>
|
||
</taxonomicName>
|
||
are diagnostic compared with the pale more inconspicuous chaetotaxy of
|
||
<taxonomicName id="4C6187EFFF8CFFC3FB82F897FB32F880" box="[1072,1217,1794,1817]" class="Insecta" family="Empididae" genus="Hesperempis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915207EFF8CFFC3FB82F897FB32F880" box="[1072,1217,1794,1817]" italics="true" pageId="2" pageNumber="263">Hesperempis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDEFC6CFF8CFFC3FF75F8B3FCA5F834" blockId="2.[151,1437,744,1965]" pageId="2" pageNumber="263">
|
||
Except for differences in the male and female terminalia, specimens of
|
||
<taxonomicName id="4C6187EFFF8CFFC3FBE5F8B0FB40F8A4" box="[1111,1203,1829,1853]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915207EFF8CFFC3FBE5F8B0FB40F8A4" box="[1111,1203,1829,1853]" italics="true" pageId="2" pageNumber="263">Philetus</emphasis>
|
||
</taxonomicName>
|
||
also resemble the brachystomatid genus
|
||
<taxonomicName id="4C6187EFFF8CFFC3FE21F8DFFD75F8FB" authority="Haliday" authorityName="Haliday" box="[403,646,1865,1890]" class="Insecta" family="Brachystomatidae" genus="Heleodromia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915207EFF8CFFC3FE21F8DFFDD5F8FB" box="[403,550,1866,1890]" italics="true" pageId="2" pageNumber="263">Heleodromia</emphasis>
|
||
Haliday
|
||
</taxonomicName>
|
||
, although they can be easily separated by the presence of a forked R4+5 wing vein in
|
||
<taxonomicName id="4C6187EFFF8CFFC3FE9EF8E5FE7BF811" box="[300,392,1904,1928]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915207EFF8CFFC3FE9EF8E5FE7BF811" box="[300,392,1904,1928]" italics="true" pageId="2" pageNumber="263">Philetus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<collectingCountry id="F376BCFCFF8CFFC3FE25F8E5FE08F810" box="[407,507,1904,1929]" name="Marshall Islands" pageId="2" pageNumber="263">Marshall</collectingCountry>
|
||
(2012: 288, fig. 5) provided a good-quality photograph of a live female specimen of
|
||
<taxonomicName id="4C6187EFFF8CFFC3FF07F800FEE2F834" box="[181,273,1941,1965]" class="Insecta" family="Empididae" genus="Philetus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B915207EFF8CFFC3FF07F800FEE2F834" box="[181,273,1941,1965]" italics="true" pageId="2" pageNumber="263">Philetus</emphasis>
|
||
</taxonomicName>
|
||
, which is reproduced here with permission (
|
||
<figureCitation id="135AE0E9FF8CFFC3FCB1F801FCB7F834" box="[771,836,1940,1965]" captionStart="FIGURES 1 − 3" captionStartId="3.[151,263,1628,1650]" captionTargetBox="[151,1436,195,1603]" captionTargetId="figure@3.[151,1436,195,1603]" captionTargetPageId="3" captionText="FIGURES 1 − 3. Philetus habitus photograph and habitat. 1. Philetus sp., live female; photograph by S. A. Marshall. 2. Sitka spruce canopy, Upper Carmanah Valley, Vancouver Island, British Columbia. 3. Coastal wet temperate rainforest, Upper Carmanah Valley." httpUri="https://zenodo.org/record/267657/files/figure.png" pageId="2" pageNumber="263">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |