421 lines
52 KiB
XML
421 lines
52 KiB
XML
<document id="FC24D60D95CF3FEEF10B4C1D6C477206" ID-CLB-Dataset="27727" ID-DOI="10.11646/zootaxa.4535.1.1" ID-GBIF-Dataset="a047bcd9-ab65-4e3f-b07a-c830c7af1072" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615773" ID-ZooBank="ABE9DB1F-7378-4571-90C4-A3FDB66527F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553788891023" checkinUser="plazi" docAuthor="Freyhof, Jörg, Bayçelebi, Esra & Geiger, Matthias" docDate="2018" docId="03BB0D181F78FFB3718BFE4BFCB8FCCF" docLanguage="en" docName="zootaxa.4535.1.1.pdf" docOrigin="Zootaxa 4535 (1)" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Cobitis troasensis Freyhof & Bayçelebi & Geiger 2018, new species" docType="treatment" docVersion="11" lastPageNumber="67" masterDocId="FF8275601F38FFF1711CFFFCFFABFF93" masterDocTitle="Review of the genus Cobitis in the Middle East, with the description of eight new species (Teleostei: Cobitidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="65" updateTime="1698715239854" updateUser="ExternalLinkService">
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<mods:mods id="0FEF39D1A5242CB422696599DF7A282A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="434A707F3A43D6A821DC982E9754F863">
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<mods:title id="D3C6D6DDA9FBCD9D862E891568831847">Review of the genus Cobitis in the Middle East, with the description of eight new species (Teleostei: Cobitidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="8A535252566045AEC883A187BE538F66" type="personal">
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<mods:roleTerm id="CC1E2C663C00991A9BE7A964C5E6DE46">Author</mods:roleTerm>
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<mods:namePart id="1573945DCEA61E5578843FAAAB1217DD">Freyhof, Jörg</mods:namePart>
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<mods:roleTerm id="7D47C03BF87AD1FAA3F8E672AD8A4642">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FD91DB346DEE579E307AC2A5B01F1B4C">Bayçelebi, Esra</mods:namePart>
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</mods:name>
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<mods:name id="2914F4D43364DFA1CAE46B8B315465AF" type="personal">
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<mods:roleTerm id="5140944708608356D5193C6858EDA434">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4E0B314B3CEF4475210937E02D462D7A">Geiger, Matthias</mods:namePart>
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<mods:typeOfResource id="4962C1815F8792FF10A06D3631A58269">text</mods:typeOfResource>
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<mods:titleInfo id="3889E0F8A5C107132E8492B32AB73294">
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<mods:title id="0E58064247AC28F5D68011AC2788C0F1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B96A754297BB318B6F0CBAE1805AA32A">
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<mods:date id="91EBD516C96F5440888AF67B9E402F1B">2018</mods:date>
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<mods:number id="179D1D4F12460407DB4FF4CDAE291AB3">2018-12-21</mods:number>
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<mods:detail id="CB7C15DD8D200073BA17E2DD1C9EAE6F" type="volume">
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<mods:number id="C9B8387EC41FE668B8DD05330FC061AD">4535</mods:number>
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<mods:detail id="23A65D8E44DDE623CA376950A287DBE3" type="issue">
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<mods:classification id="D622EAD19A9190BB0B1E6988E4EF095B">journal article</mods:classification>
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<mods:identifier id="99D0396497EA34ACBBCB467B51528C0C" type="CLB-Dataset">27727</mods:identifier>
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<mods:identifier id="09D08BB100B3C5BF5B343DE1ADDF01A8" type="DOI">10.11646/zootaxa.4535.1.1</mods:identifier>
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<mods:identifier id="22D30F6EE5C12321ABD2ACA55365B480" type="GBIF-Dataset">a047bcd9-ab65-4e3f-b07a-c830c7af1072</mods:identifier>
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<mods:identifier id="59A5E09FEA2DC37BC2C1950FDE8673F9" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="56C467EB01E8CDCE677715EC863E4923" type="Zenodo-Dep">2615773</mods:identifier>
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</mods:mods>
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<treatment id="03BB0D181F78FFB3718BFE4BFCB8FCCF" ID-DOI="http://doi.org/10.5281/zenodo.5699591" ID-GBIF-Taxon="154521876" ID-Zenodo-Dep="5699591" LSID="urn:lsid:plazi:treatment:03BB0D181F78FFB3718BFE4BFCB8FCCF" httpUri="http://treatment.plazi.org/id/03BB0D181F78FFB3718BFE4BFCB8FCCF" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65">
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<subSubSection id="C308EF851F78FFB1718BFE4BFDA1FE41" box="[151,522,439,466]" pageId="64" pageNumber="65" type="nomenclature">
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<paragraph id="8BADBC0E1F78FFB1718BFE4BFDA1FE41" blockId="64.[151,522,439,499]" box="[151,522,439,466]" pageId="64" pageNumber="65">
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<heading id="D0E50B621F78FFB1718BFE4BFDA1FE41" bold="true" box="[151,522,439,466]" fontSize="11" level="1" pageId="64" pageNumber="65" reason="1">
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<emphasis id="B966601C1F78FFB1718BFE4BFDA1FE41" bold="true" box="[151,522,439,466]" pageId="64" pageNumber="65">
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<taxonomicName id="4C12C78D1F78FFB1718BFE4BFEC0FE42" authority="Freyhof & Bayçelebi & Geiger, 2018" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[151,363,439,465]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="troasensis" status="sp. nov.">
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||
<emphasis id="B966601C1F78FFB1718BFE4BFEC0FE42" bold="true" box="[151,363,439,465]" italics="true" pageId="64" pageNumber="65">Cobitis troasensis</emphasis>
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||
</taxonomicName>
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||
,
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<taxonomicNameLabel id="A255DD671F78FFB17066FE44FDA1FE41" box="[378,522,440,466]" pageId="64" pageNumber="65" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C308EF851F78FFB1718BFE26FE88FE60" box="[151,291,474,499]" pageId="64" pageNumber="65" type="description">
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<paragraph id="8BADBC0E1F78FFB1718BFE26FE88FE60" blockId="64.[151,522,439,499]" box="[151,291,474,499]" pageId="64" pageNumber="65">
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(
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<figureCitation id="1329A08B1F78FFB17183FE26FEB1FE60" box="[159,282,474,499]" captionStart-0="FIGURE 51" captionStart-1="FIGURE 52" captionStart-2="FIGURE 53" captionStartId-0="65.[151,250,802,824]" captionStartId-1="65.[151,250,1251,1273]" captionStartId-2="65.[151,250,1818,1840]" captionTargetBox-0="[151,1409,372,746]" captionTargetBox-1="[186,1410,926,1206]" captionTargetBox-2="[151,1435,1354,1797]" captionTargetId-0="figure@65.[151,1436,337,781]" captionTargetId-1="figure@65.[151,1436,904,1229]" captionTargetId-2="figure@65.[151,1435,1354,1797]" captionTargetPageId-0="65" captionTargetPageId-1="65" captionTargetPageId-2="65" captionText-0="FIGURE 51. Cobitis troasensis, left column from the top: males, ZFMK ICH-098603, holotype, 50 mm SL; paratypes, FSJF 2383, 47 mm SL, 43 mm SL; right column from the top: females, 76 mm SL, 72 mm SL, 55 mm SL; Turkey: stream Tuzla." captionText-1="FIGURE 52. Cobitis troasensis, left column from the top: males, ZFMK ICH-098603, holotype, 50 mm SL; paratypes, FSJF 2383, 47 mm SL, 43 mm SL; right column from the top: females, 76 mm SL, 72 mm SL, 55 mm SL; Turkey: stream Tuzla." captionText-2="FIGURE 53. Cobitis troasensis, FSJF 2317, male, 60 mm SL; Turkey: stream Tuzla." httpUri-0="https://zenodo.org/record/2615877/files/figure.png" httpUri-1="https://zenodo.org/record/2615880/files/figure.png" httpUri-2="https://zenodo.org/record/2615883/files/figure.png" pageId="64" pageNumber="65">Fig. 51–53</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C308EF851F78FFB1718BFDDEFCBEFD7D" pageId="64" pageNumber="65" type="materials_examined">
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<paragraph id="8BADBC0E1F78FFB1718BFDDEFE94FDCC" blockId="64.[151,1437,546,2011]" pageId="64" pageNumber="65">
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<materialsCitation id="3B7AB6531F78FFB1718BFDDEFE94FDCC" ID-GBIF-Occurrence="2120487510" collectionCode="ZFMK" country="Turkey" location="Ayvacik" municipality="Tuzla" pageId="64" pageNumber="65" specimenCode="ICH-098603, 50" specimenCount="1" stateProvince="Canakkale" typeStatus="holotype">
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<emphasis id="B966601C1F78FFB1718BFDDEFEA0FDA8" bold="true" box="[151,267,546,571]" pageId="64" pageNumber="65">
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<typeStatus id="54A902AC1F78FFB1718BFDDEFEADFDA8" box="[151,262,546,571]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
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||
.
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</emphasis>
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||
<collectionCode id="ED0324CB1F78FFB1700FFDDFFECAFDA9" box="[275,353,547,570]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="64" pageNumber="65" type="Museum">ZFMK</collectionCode>
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<specimenCode id="DBB414751F78FFB17074FDDEFE5DFDA8" box="[360,502,546,571]" collectionCode="ICH-" pageId="64" pageNumber="65">ICH-098603</specimenCode>
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,
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<quantity id="4CEA11EB1F78FFB17319FDDEFDFEFDA9" box="[517,597,546,571]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="64" pageNumber="65" unit="mm" value="50.0">
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<specimenCode id="DBB414751F78FFB17319FDDEFD8AFDA8" box="[517,545,546,571]" collectionCode="ICH-" pageId="64" pageNumber="65">50</specimenCode>
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||
mm
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||
</quantity>
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||
SL;
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||
<collectingCountry id="F305FC9E1F78FFB17391FDDEFD4AFDA8" box="[653,737,546,571]" name="Turkey" pageId="64" pageNumber="65">Turkey</collectingCountry>
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:
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<collectingRegion id="49D672EC1F78FFB173F1FDDEFCC8FDA8" box="[749,867,546,571]" country="Turkey" name="Canakkale" pageId="64" pageNumber="65">Çanakkale</collectingRegion>
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||
prov.: stream
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||
<collectingMunicipality id="6BC926741F78FFB1751AFDDEFBEEFDA8" box="[1030,1093,546,571]" pageId="64" pageNumber="65">Tuzla</collectingMunicipality>
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about
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||
<quantity id="4CEA11EB1F78FFB1758FFDDFFB66FDA8" box="[1171,1229,546,571]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="64" pageNumber="65" unit="km" value="1.0">1 km</quantity>
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||
south of
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||
<location id="8ECDEAD51F78FFB17425FDDEFA33FDA8" LSID="urn:lsid:plazi:treatment:03BB0D181F78FFB3718BFE4BFCB8FCCF:8ECDEAD51F78FFB17425FDDEFA33FDA8" box="[1337,1432,546,571]" country="Turkey" municipality="Tuzla" name="Ayvacik" pageId="64" pageNumber="65" stateProvince="Canakkale">Ayvacık</location>
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||
, 39.596 26.438.
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||
</materialsCitation>
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</paragraph>
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||
<paragraph id="8BADBC0E1F78FFB171DBFD96FBE0FD58" blockId="64.[151,1437,546,2011]" pageId="64" pageNumber="65">
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||
<materialsCitation id="3B7AB6531F78FFB171DBFD96FBE3FD58" ID-GBIF-Occurrence="2120484900" collectionCode="FSJF" country="Turkey" county="Tuzla" location="Tuzla" municipality="Behramkale" pageId="64" pageNumber="65" specimenCode="FSJF 2383, 8, 43-76" specimenCount="1" stateProvince="Canakkale" typeStatus="paratype">
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||
<emphasis id="B966601C1F78FFB171DBFD96FEEDFD10" bold="true" box="[199,326,618,643]" pageId="64" pageNumber="65">
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<typeStatus id="54A902AC1F78FFB171DBFD96FEEAFD10" box="[199,321,618,643]" pageId="64" pageNumber="65" type="paratype">Paratypes</typeStatus>
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||
.
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||
</emphasis>
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||
<specimenCode id="DBB414751F78FFB1704DFD97FE7BFD10" box="[337,464,619,643]" collectionCode="FSJF" pageId="64" pageNumber="65">FSJF 2383</specimenCode>
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||
,
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||
<specimenCode id="DBB414751F78FFB170C3FD97FE44FD10" box="[479,495,619,643]" collectionCode="FSJF" pageId="64" pageNumber="65">8</specimenCode>
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,
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||
<quantity id="4CEA11EB1F78FFB1731CFD97FDD6FD11" box="[512,637,618,643]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" metricValueMax="7.6" metricValueMin="4.3" pageId="64" pageNumber="65" unit="mm" value="59.5" valueMax="76.0" valueMin="43.0">
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||
<specimenCode id="DBB414751F78FFB1731CFD97FDEDFD10" box="[512,582,618,643]" collectionCode="FSJF" pageId="64" pageNumber="65">43–76</specimenCode>
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||
mm
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||
</quantity>
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||
SL; same data as holotype—FSJF 2317, 2,
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||
<quantity id="4CEA11EB1F78FFB17595FD96FAADFD11" box="[1161,1286,618,643]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" metricValueMax="6.0" metricValueMin="5.9" pageId="64" pageNumber="65" unit="mm" value="59.5" valueMax="60.0" valueMin="59.0">59–60 mm</quantity>
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||
SL;
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||
<collectingCountry id="F305FC9E1F78FFB17458FD96FA33FD10" box="[1348,1432,618,643]" name="Turkey" pageId="64" pageNumber="65">Turkey</collectingCountry>
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:
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||
<collectingRegion id="49D672EC1F78FFB1718BFD72FEA6FD34" box="[151,269,654,679]" country="Turkey" name="Canakkale" pageId="64" pageNumber="65">Çanakkale</collectingRegion>
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||
prov.: stream
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||
<collectingCounty id="62CCC4821F78FFB170A6FD72FE52FD34" box="[442,505,654,679]" pageId="64" pageNumber="65">Tuzla</collectingCounty>
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about
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||
<quantity id="4CEA11EB1F78FFB17351FD73FD20FD34" box="[589,651,654,679]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="64" pageNumber="65" unit="km" value="1.0">1 km</quantity>
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north of
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||
<collectingMunicipality id="6BC926741F78FFB173E2FD72FC23FD34" box="[766,904,654,679]" pageId="64" pageNumber="65">Behramkale</collectingMunicipality>
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||
, 39.498 26.333.—FFR 0 5529, 1,
|
||
<quantity id="4CEA11EB1F78FFB1740AFD72FAC2FD35" box="[1302,1385,654,679]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="64" pageNumber="65" unit="mm" value="65.0">65 mm</quantity>
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||
SL;
|
||
<collectingCountry id="F305FC9E1F78FFB1718BFD4EFF40FD58" box="[151,235,690,715]" name="Turkey" pageId="64" pageNumber="65">Turkey</collectingCountry>
|
||
:
|
||
<collectingRegion id="49D672EC1F78FFB171E9FD4EFEC0FD58" box="[245,363,690,715]" country="Turkey" name="Canakkale" pageId="64" pageNumber="65">Çanakkale</collectingRegion>
|
||
prov.: stream
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||
<location id="8ECDEAD51F78FFB17310FD4EFDE0FD58" LSID="urn:lsid:plazi:treatment:03BB0D181F78FFB3718BFE4BFCB8FCCF:8ECDEAD51F78FFB17310FD4EFDE0FD58" box="[524,587,690,715]" country="Turkey" county="Tuzla" municipality="Behramkale" name="Tuzla" pageId="64" pageNumber="65" stateProvince="Canakkale">Tuzla</location>
|
||
about
|
||
<quantity id="4CEA11EB1F78FFB1738BFD4FFD7BFD58" box="[663,720,690,715]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="64" pageNumber="65" unit="km" value="1.0">1 km</quantity>
|
||
south of
|
||
<location id="8ECDEAD51F78FFB17224FD4EFC3CFD58" LSID="urn:lsid:plazi:treatment:03BB0D181F78FFB3718BFE4BFCB8FCCF:8ECDEAD51F78FFB17224FD4EFC3CFD58" box="[824,919,690,715]" country="Turkey" county="Tuzla" municipality="Behramkale" name="Ayvacik" pageId="64" pageNumber="65" stateProvince="Canakkale">Ayvacık</location>
|
||
, 39.586 26.417
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BADBC0E1F78FFB171DBFD2AFCBEFD7D" blockId="64.[151,1437,546,2011]" box="[199,789,726,751]" pageId="64" pageNumber="65">
|
||
<emphasis id="B966601C1F78FFB171DBFD2AFD7AFD7C" bold="true" box="[199,721,726,751]" pageId="64" pageNumber="65">Material used in molecular genetic analysis.</emphasis>
|
||
None
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308EF851F78FFB171DBFD06FEA0FB50" pageId="64" pageNumber="65" type="diagnosis">
|
||
<paragraph id="8BADBC0E1F78FFB171DBFD06FC36FC54" blockId="64.[151,1437,546,2011]" pageId="64" pageNumber="65">
|
||
<emphasis id="B966601C1F78FFB171DBFD06FEEAFC80" bold="true" box="[199,321,762,787]" pageId="64" pageNumber="65">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C12C78D1F78FFB17057FD07FDBCFC81" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[331,535,763,787]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F78FFB17057FD07FDBCFC81" box="[331,535,763,787]" italics="true" pageId="64" pageNumber="65">Cobitis troasensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C12C78D1F78FFB17201FD07FC37FC80" box="[797,924,763,787]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="fahireae">
|
||
<emphasis id="B966601C1F78FFB17201FD07FC37FC80" box="[797,924,763,787]" italics="true" pageId="64" pageNumber="65">C. fahireae</emphasis>
|
||
</taxonomicName>
|
||
by having 11–15 small, irregularly commashaped blotches in Z4 anterior to the dorsal-fin origin (vs. 6–8 large, roundish, triangular, squarish, or horizontally elongate, often very densely set and fused to each other). In
|
||
<taxonomicName id="4C12C78D1F78FFB17227FCBFFC67FCC9" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[827,972,835,858]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F78FFB17227FCBFFC67FCC9" box="[827,972,835,858]" italics="true" pageId="64" pageNumber="65">C. troasensis</emphasis>
|
||
</taxonomicName>
|
||
, the blotches in Z2 are usually fused into a stripe reaching to the vertical or the anus, fused with the pigmentation in Z1 and Z3 on the caudal peduncle into a mottled pattern (vs. blotches in Z2 usually well separate) and the black spot at the upper caudal-fin base is small and usually roundish (vs. absent in many populations of
|
||
<taxonomicName id="4C12C78D1F78FFB1720CFC53FC27FC55" box="[784,908,942,966]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="fahireae">
|
||
<emphasis id="B966601C1F78FFB1720CFC53FC27FC55" box="[784,908,942,966]" italics="true" pageId="64" pageNumber="65">C. fahireae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BADBC0E1F78FFB171DBFC2FFEA0FB50" blockId="64.[151,1437,546,2011]" pageId="64" pageNumber="65">
|
||
<taxonomicName id="4C12C78D1F78FFB171DBFC2FFE24FC79" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[199,399,979,1003]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F78FFB171DBFC2FFE24FC79" box="[199,399,979,1003]" italics="true" pageId="64" pageNumber="65">Cobitis troasensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C12C78D1F78FFB1739BFC2FFD4EFC79" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[647,741,979,1002]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F78FFB1739BFC2FFD31FC79" box="[647,666,979,1002]" italics="true" pageId="64" pageNumber="65">C</emphasis>
|
||
.
|
||
<emphasis id="B966601C1F78FFB173BBFC2FFD4EFC79" box="[679,741,979,1002]" italics="true" pageId="64" pageNumber="65">emrei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C12C78D1F78FFB173EFFC2FFC09FC78" box="[755,930,979,1003]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="elongatoides">
|
||
<emphasis id="B966601C1F78FFB173EFFC2FFC09FC78" box="[755,930,979,1003]" italics="true" pageId="64" pageNumber="65">C. elongatoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C12C78D1F78FFB172ACFC2FFB8FFC79" authorityName="Vasil'eva & Vasil'ev" authorityYear="2006" box="[944,1060,979,1002]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="pontica">
|
||
<emphasis id="B966601C1F78FFB172ACFC2FFB8FFC79" box="[944,1060,979,1002]" italics="true" pageId="64" pageNumber="65">C. pontica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C12C78D1F78FFB1752EFC2FFB6AFC78" authorityName="Erk'akan, Atalay-Ekmekçi & Nalbant" authorityYear="1998" box="[1074,1217,979,1003]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="splendens">
|
||
<emphasis id="B966601C1F78FFB1752EFC2FFB6AFC78" box="[1074,1217,979,1003]" italics="true" pageId="64" pageNumber="65">C. splendens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C12C78D1F78FFB175EAFC2FFAD1FC79" box="[1270,1402,979,1002]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="tanaitica">
|
||
<emphasis id="B966601C1F78FFB175EAFC2FFAD1FC79" box="[1270,1402,979,1002]" italics="true" pageId="64" pageNumber="65">C. tanaitica</emphasis>
|
||
</taxonomicName>
|
||
by having 11–15 small, comma-shaped blotches in Z4 along the flank anterior to the dorsal-fin origin (vs. 5–7 horizontally elongate, often very densely set and fused into short or longer stripes in
|
||
<taxonomicName id="4C12C78D1F78FFB17579FBE7FB6DFBA1" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[1125,1222,1051,1074]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F78FFB17579FBE7FBD3FBA1" box="[1125,1144,1051,1074]" italics="true" pageId="64" pageNumber="65">C</emphasis>
|
||
.
|
||
<emphasis id="B966601C1F78FFB1759BFBE7FB6DFBA1" box="[1159,1222,1051,1074]" italics="true" pageId="64" pageNumber="65">emrei</emphasis>
|
||
</taxonomicName>
|
||
, well separate and irregularly roundish or squarish in
|
||
<taxonomicName id="4C12C78D1F78FFB17358FBC3FD71FBC5" authorityName="Erk'akan, Atalay-Ekmekçi & Nalbant" authorityYear="1998" box="[580,730,1086,1110]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="splendens">
|
||
<emphasis id="B966601C1F78FFB17358FBC3FD71FBC5" box="[580,730,1086,1110]" italics="true" pageId="64" pageNumber="65">C. splendens</emphasis>
|
||
</taxonomicName>
|
||
, vertically elongate or roundish and well separated in
|
||
<taxonomicName id="4C12C78D1F78FFB1749EFBC3FE8DFBE8" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="elongatoides">
|
||
<emphasis id="B966601C1F78FFB1749EFBC3FE8DFBE8" italics="true" pageId="64" pageNumber="65">C. elongatoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C12C78D1F78FFB17029FB9FFE07FBE9" authorityName="Vasil'eva & Vasil'ev" authorityYear="2006" box="[309,428,1123,1146]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="pontica">
|
||
<emphasis id="B966601C1F78FFB17029FB9FFE07FBE9" box="[309,428,1123,1146]" italics="true" pageId="64" pageNumber="65">C. pontica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C12C78D1F78FFB170F9FB9FFDC7FBE9" box="[485,620,1123,1146]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="tanaitica">
|
||
<emphasis id="B966601C1F78FFB170F9FB9FFDC7FBE9" box="[485,620,1123,1146]" italics="true" pageId="64" pageNumber="65">C. tanaitica</emphasis>
|
||
</taxonomicName>
|
||
) and the black spot at the upper caudal-fin base smaller or equal to the eye diameter (vs. larger than the eye diameter in
|
||
<taxonomicName id="4C12C78D1F78FFB173F8FB7BFC32FB0D" box="[740,921,1158,1182]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="elongatoides">
|
||
<emphasis id="B966601C1F78FFB173F8FB7BFC32FB0D" box="[740,921,1158,1182]" italics="true" pageId="64" pageNumber="65">C. elongatoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C12C78D1F78FFB172B0FB7BFB8EFB0D" authorityName="Vasil'eva & Vasil'ev" authorityYear="2006" box="[940,1061,1159,1182]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="pontica">
|
||
<emphasis id="B966601C1F78FFB172B0FB7BFB8EFB0D" box="[940,1061,1159,1182]" italics="true" pageId="64" pageNumber="65">C. pontica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C12C78D1F78FFB17524FB7BFB30FB0D" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[1080,1179,1159,1182]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F78FFB17524FB7BFB30FB0D" box="[1080,1179,1159,1182]" italics="true" pageId="64" pageNumber="65">C. emrei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C12C78D1F78FFB175B2FB7BFAE9FB0D" authorityName="Erk'akan, Atalay-Ekmekçi & Nalbant" authorityYear="1998" box="[1198,1346,1158,1182]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="splendens">
|
||
<emphasis id="B966601C1F78FFB175B2FB7BFAE9FB0D" box="[1198,1346,1158,1182]" italics="true" pageId="64" pageNumber="65">C. splendens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C12C78D1F78FFB1749EFB7BFF50FB51" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="tanaitica">
|
||
<emphasis id="B966601C1F78FFB1749EFB7BFF50FB51" italics="true" pageId="64" pageNumber="65">C. tanaitica</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308EF851F78FFB371DBFB32FF5AFE43" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65" type="description">
|
||
<paragraph id="8BADBC0E1F78FFB171DBFB32FCD2FA2C" blockId="64.[151,1437,546,2011]" pageId="64" pageNumber="65">
|
||
<taxonomicName id="4C12C78D1F78FFB171DBFB32FE24FB75" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[199,399,1230,1254]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F78FFB171DBFB32FE24FB75" box="[199,399,1230,1254]" italics="true" pageId="64" pageNumber="65">Cobitis troasensis</emphasis>
|
||
</taxonomicName>
|
||
is further distinguished from
|
||
<taxonomicName id="4C12C78D1F78FFB173C0FB33FC97FB75" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[732,828,1231,1254]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F78FFB173C0FB33FD44FB75" box="[732,751,1231,1254]" italics="true" pageId="64" pageNumber="65">C</emphasis>
|
||
.
|
||
<emphasis id="B966601C1F78FFB173E1FB33FC97FB75" box="[765,828,1231,1254]" italics="true" pageId="64" pageNumber="65">emrei</emphasis>
|
||
</taxonomicName>
|
||
by having a more slender body (body width 8–9% SL vs.
|
||
<quantity id="4CEA11EB1F78FFB171A1FB0FFEBAFA98" box="[189,273,1266,1291]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="64" pageNumber="65" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">9–11 in</quantity>
|
||
the male; 7–10% SL vs.
|
||
<quantity id="4CEA11EB1F78FFB17334FB0FFD21FA98" box="[552,650,1266,1291]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="64" pageNumber="65" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">10–12 in</quantity>
|
||
the female) and a greater head length in the male (21% SL vs. 21–24). It is further distinguished from
|
||
<taxonomicName id="4C12C78D1F78FFB17318FAEBFD3DFABD" authorityName="Erk'akan, Atalay-Ekmekçi & Nalbant" authorityYear="1998" box="[516,662,1302,1326]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="splendens">
|
||
<emphasis id="B966601C1F78FFB17318FAEBFD3DFABD" box="[516,662,1302,1326]" italics="true" pageId="64" pageNumber="65">C. splendens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C12C78D1F78FFB173CFFAEBFCF1FABD" box="[723,858,1303,1326]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="tanaitica">
|
||
<emphasis id="B966601C1F78FFB173CFFAEBFCF1FABD" box="[723,858,1303,1326]" italics="true" pageId="64" pageNumber="65">C. tanaitica</emphasis>
|
||
</taxonomicName>
|
||
by the pigmentation in Z3 reaching to the caudal peduncle in the male (vs. Z3 reduced to about head length in
|
||
<taxonomicName id="4C12C78D1F78FFB17246FAC7FC41FAC0" authorityName="Erk'akan, Atalay-Ekmekçi & Nalbant" authorityYear="1998" box="[858,1002,1339,1363]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="splendens">
|
||
<emphasis id="B966601C1F78FFB17246FAC7FCC6FAC1" box="[858,877,1339,1362]" italics="true" pageId="64" pageNumber="65">C</emphasis>
|
||
.
|
||
<emphasis id="B966601C1F78FFB17260FAC7FC41FAC0" box="[892,1002,1339,1363]" italics="true" pageId="64" pageNumber="65">splendens</emphasis>
|
||
</taxonomicName>
|
||
, Z3 usually reaching to the dorsal-fin base in
|
||
<taxonomicName id="4C12C78D1F78FFB171EEFAA3FED2FAE5" box="[242,377,1375,1398]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="tanaitica">
|
||
<emphasis id="B966601C1F78FFB171EEFAA3FED2FAE5" box="[242,377,1375,1398]" italics="true" pageId="64" pageNumber="65">C. tanaitica</emphasis>
|
||
</taxonomicName>
|
||
) and Z3 wider than Z
|
||
<quantity id="4CEA11EB1F78FFB17367FAA3FD03FAE4" box="[635,680,1374,1399]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="64" pageNumber="65" unit="in" value="2.0">2 in</quantity>
|
||
the male (vs. usually Z3 narrower than Z2).
|
||
<taxonomicName id="4C12C78D1F78FFB175A8FAA2FA2BFAE5" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[1204,1408,1374,1398]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F78FFB175A8FAA2FA2BFAE5" box="[1204,1408,1374,1398]" italics="true" pageId="64" pageNumber="65">Cobitis troasensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C12C78D1F78FFB1706DFA7FFDB4FA08" authorityName="Erk'akan, Atalay-Ekmekçi & Nalbant" authorityYear="1998" box="[369,543,1411,1435]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="puncticulata">
|
||
<emphasis id="B966601C1F78FFB1706DFA7FFDB4FA08" box="[369,543,1411,1435]" italics="true" pageId="64" pageNumber="65">C. puncticulata</emphasis>
|
||
</taxonomicName>
|
||
by lacking the pigmentation on the flank below Z4 (vs. pigmented) and having a pale yellowish or whitish background coloration (vs. brownish).
|
||
</paragraph>
|
||
<paragraph id="8BADBC0E1F78FFB171DBFA37FDF4F994" blockId="64.[151,1437,546,2011]" pageId="64" pageNumber="65">
|
||
<taxonomicName id="4C12C78D1F78FFB171DBFA37FE25FA71" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[199,398,1483,1507]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F78FFB171DBFA37FE25FA71" box="[199,398,1483,1507]" italics="true" pageId="64" pageNumber="65">Cobitis troasensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C12C78D1F78FFB17394FA37FD5FFA70" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[648,756,1483,1507]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="afifeae">
|
||
<emphasis id="B966601C1F78FFB17394FA37FD5FFA70" box="[648,756,1483,1507]" italics="true" pageId="64" pageNumber="65">C. afifeae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C12C78D1F78FFB1721EFA37FC0BFA70" authorityName="Erkakan, Ozdemir & Ozeren" authorityYear="2017" box="[770,928,1483,1507]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="dorademiri">
|
||
<emphasis id="B966601C1F78FFB1721EFA37FC0BFA70" box="[770,928,1483,1507]" italics="true" pageId="64" pageNumber="65">C. dorademiri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B966601C1F78FFB172B2FA37FB5CFA71" box="[942,1271,1483,1507]" italics="true" pageId="64" pageNumber="65">
|
||
<taxonomicName id="4C12C78D1F78FFB172B2FA37FB99FA70" authorityName="Battalgazi" authorityYear="1944" box="[942,1074,1483,1507]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="phrygica">C. phrygica</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C12C78D1F78FFB17575FA37FB5CFA71" authorityName="Karaman" authorityYear="1955" box="[1129,1271,1483,1506]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="strumicae">C. strumicae</taxonomicName>
|
||
</emphasis>
|
||
by having one laminae circularis in the males (vs. two).
|
||
</paragraph>
|
||
<paragraph id="8BADBC0E1F78FFB171DBF9EEFBA8F890" blockId="64.[151,1437,546,2011]" pageId="64" pageNumber="65">
|
||
<emphasis id="B966601C1F78FFB171DBF9EEFEF2F9B8" bold="true" box="[199,345,1554,1579]" pageId="64" pageNumber="65">Description.</emphasis>
|
||
See
|
||
<figureCitation id="1329A08B1F78FFB17091F9EEFD85F9B8" box="[397,558,1554,1579]" captionStart-0="FIGURE 51" captionStart-1="FIGURE 52" captionStart-2="FIGURE 53" captionStartId-0="65.[151,250,802,824]" captionStartId-1="65.[151,250,1251,1273]" captionStartId-2="65.[151,250,1818,1840]" captionTargetBox-0="[151,1409,372,746]" captionTargetBox-1="[186,1410,926,1206]" captionTargetBox-2="[151,1435,1354,1797]" captionTargetId-0="figure@65.[151,1436,337,781]" captionTargetId-1="figure@65.[151,1436,904,1229]" captionTargetId-2="figure@65.[151,1435,1354,1797]" captionTargetPageId-0="65" captionTargetPageId-1="65" captionTargetPageId-2="65" captionText-0="FIGURE 51. Cobitis troasensis, left column from the top: males, ZFMK ICH-098603, holotype, 50 mm SL; paratypes, FSJF 2383, 47 mm SL, 43 mm SL; right column from the top: females, 76 mm SL, 72 mm SL, 55 mm SL; Turkey: stream Tuzla." captionText-1="FIGURE 52. Cobitis troasensis, left column from the top: males, ZFMK ICH-098603, holotype, 50 mm SL; paratypes, FSJF 2383, 47 mm SL, 43 mm SL; right column from the top: females, 76 mm SL, 72 mm SL, 55 mm SL; Turkey: stream Tuzla." captionText-2="FIGURE 53. Cobitis troasensis, FSJF 2317, male, 60 mm SL; Turkey: stream Tuzla." httpUri-0="https://zenodo.org/record/2615877/files/figure.png" httpUri-1="https://zenodo.org/record/2615880/files/figure.png" httpUri-2="https://zenodo.org/record/2615883/files/figure.png" pageId="64" pageNumber="65">Figures 51–53</figureCitation>
|
||
for general appearance and
|
||
<tableCitation id="C69089B51F78FFB17275F9EEFC10F9B8" box="[873,955,1554,1579]" captionStart="TABLE 9" captionStartId="66.[151,239,904,927]" captionText="TABLE 9. Morphometric data of Cobitis troasensis (holotype ZFMK ICH-098603 and paratypes FSJF 2383, n = 8; FSJF 2317, n=2; FFR 05529, n=1). The calculations include the holotype." pageId="64" pageNumber="65">Table 9</tableCitation>
|
||
for morphometric data of the
|
||
<typeStatus id="54A902AC1F78FFB17417F9EEFAC6F9B8" box="[1291,1389,1554,1579]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D1477871F78FFB1718BF9CBFE81F9DC" box="[151,298,1591,1615]" pageId="64" pageNumber="65" type="generic" typeStatus="paratypes">11 paratypes</specimenCount>
|
||
. Greatest body depth at or slightly anterior to dorsal-fin origin, decreasing towards caudal-fin base. Head profile slightly convex, head length 1.0–1.4 times in body depth. Snout pointed, its length 0.6–0.9 times in postorbital length. Eye diameter 0.2–0.4 times in head depth at eye, 1.3–2.7 times in interorbital width. Caudal peduncle 1.2–1.7 times longer than deep. Pelvic axillary lobe present. Margin of dorsal and anal fins convex. Caudal fin truncate. Very shallow ventral keel on caudal peduncle. External part of the suborbital spine bifurcate, reaching slightly beyond centre of eye. Largest recorded specimen
|
||
<quantity id="4CEA11EB1F78FFB1729AF916FC7FF891" box="[902,980,1770,1795]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="64" pageNumber="65" unit="mm" value="76.0">76 mm</quantity>
|
||
SL.
|
||
</paragraph>
|
||
<paragraph id="8BADBC0E1F78FFB071DBF8F2FBFDFF46" blockId="64.[151,1437,546,2011]" lastBlockId="65.[151,1436,151,285]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
|
||
Dorsal fin with 3 unbranched and 6½ (12) branched rays. Anal fin with 3 unbranched and 5½ (11) and 6½ (1) branched rays. Caudal fin with 7+7 branched rays in the
|
||
<specimenCount id="9D1477871F78FFB17212F8CEFCC8F8D8" box="[782,867,1842,1867]" pageId="64" pageNumber="65" type="male">5 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D1477871F78FFB17285F8CFFBA3F8D8" box="[921,1032,1842,1867]" pageId="64" pageNumber="65" type="female">4 females</specimenCount>
|
||
, 7+
|
||
<quantity id="4CEA11EB1F78FFB1752DF8CFFBF0F8D8" box="[1073,1115,1842,1867]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="64" pageNumber="65" unit="in" value="8.0">8 in</quantity>
|
||
the
|
||
<specimenCount id="9D1477871F78FFB17596F8CFFB5DF8D8" box="[1162,1270,1842,1867]" pageId="64" pageNumber="65" type="female">2 females</specimenCount>
|
||
and 8+
|
||
<quantity id="4CEA11EB1F78FFB17456F8CFFADFF8D8" box="[1354,1396,1842,1867]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="64" pageNumber="65" unit="in" value="8.0">8 in</quantity>
|
||
the one female. Pectoral fin with 7 (4) and 8 (8) branched rays and pelvic fin with 5 (1) and 6 (11) branched rays. Body completely covered by embedded scales, except on belly and breast. Scales small. Focal area in subdorsal scales about 1/2 or 1/3 of vertical scale diameter. Lateral line short, with 4–6, pores reaching to anterior part of pectoral fin. Lips (
|
||
<figureCitation id="1329A08B1F78FFB17017F83EFECBF848" box="[267,352,1986,2011]" captionStart="FIGURE 54" captionStartId="67.[151,250,1575,1597]" captionTargetBox="[151,1436,220,1518]" captionTargetId="figure@67.[151,1436,220,1518]" captionTargetPageId="67" captionText="FIGURE 54. Upper row: Cobitis afifeae, FSJF 3022, paratype, 76 mm SL; Turkey: Büyük Menderes River; C. aliyeae, FSJF 2427, paratype, female, 65 mm SL; Turkey: Seyhan River; C. anabelae, ZFMK ICH-98633, holotype, male, 53 mm SL; Turkey: Karasu River; C. emrei, FSJF 1960, paratype, female, 77 mm SL; Turkey: Lake Sapanca basin; Lower row: C. erkakanae, FSJF 2510, paratype, female, 68 mm SL; Turkey: canal at Lake Gölbaşı; C. joergbohleni, ZFMK ICH-97624, holotype, female, 60 mm SL; Turkey: spring Soysallı; C. pirii, FSJF 2474, paratype, female, 95 mm SL; Turkey: streams Çayköy; C. troasensis, FSJF 2383, paratype, female, 72 mm SL; Turkey: stream Tuzla." httpUri="https://zenodo.org/record/2615885/files/figure.png" pageId="64" pageNumber="65">Fig. 54</figureCitation>
|
||
) thin, mental lobes of lower lip short, usually well separated, not produced into a barbel-like process. Rostral barbel reaching base of mandibular barbel. Mandibular barbel reaching to or slightly beyond vertical of nostril. Maxillary barbel reaching vertical of front border or middle of eye.
|
||
</paragraph>
|
||
<paragraph id="8BADBC0E1F79FFB071DBFF23FF44FE8E" blockId="65.[151,1436,151,285]" pageId="65" pageNumber="66">
|
||
<emphasis id="B966601C1F79FFB071DBFF23FE13FF6B" bold="true" box="[199,440,223,248]" pageId="65" pageNumber="66">Sexual dimorphism.</emphasis>
|
||
Males have a longer pectoral fin (18–24% SL vs. 14–16) and one lamina circularis (vs. absent).
|
||
</paragraph>
|
||
<caption id="DF6DEC861F79FFB0718BFCDEFAD0FCC4" httpUri="https://zenodo.org/record/2615877/files/figure.png" pageId="65" pageNumber="66" startId="65.[151,250,802,824]" targetBox="[151,1409,372,746]" targetPageId="65">
|
||
<paragraph id="8BADBC0E1F79FFB0718BFCDEFAD0FCC4" blockId="65.[151,1435,802,855]" pageId="65" pageNumber="66">
|
||
<emphasis id="B966601C1F79FFB0718BFCDEFE8BFCAB" bold="true" box="[151,288,802,824]" pageId="65" pageNumber="66">FIGURE 51.</emphasis>
|
||
<taxonomicName id="4C12C78D1F79FFB0703BFCDEFE75FCAB" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[295,478,802,824]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F79FFB0703BFCDEFE75FCAB" box="[295,478,802,824]" italics="true" pageId="65" pageNumber="66">Cobitis troasensis</emphasis>
|
||
</taxonomicName>
|
||
, left column from the top: males, ZFMK ICH-098603, holotype, 50 mm SL; paratypes, FSJF 2383, 47 mm SL, 43 mm SL; right column from the top: females, 76 mm SL, 72 mm SL, 55 mm SL; Turkey: stream Tuzla.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6DEC861F79FFB0718BFB1FFAD0FA8B" httpUri="https://zenodo.org/record/2615880/files/figure.png" pageId="65" pageNumber="66" startId="65.[151,250,1251,1273]" targetBox="[186,1410,926,1206]" targetPageId="65">
|
||
<paragraph id="8BADBC0E1F79FFB0718BFB1FFAD0FA8B" blockId="65.[151,1435,1251,1304]" pageId="65" pageNumber="66">
|
||
<emphasis id="B966601C1F79FFB0718BFB1FFE8BFB6A" bold="true" box="[151,288,1251,1273]" pageId="65" pageNumber="66">FIGURE 52.</emphasis>
|
||
<taxonomicName id="4C12C78D1F79FFB0703BFB1FFE75FB6A" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[295,478,1251,1273]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F79FFB0703BFB1FFE75FB6A" box="[295,478,1251,1273]" italics="true" pageId="65" pageNumber="66">Cobitis troasensis</emphasis>
|
||
</taxonomicName>
|
||
, left column from the top: males, ZFMK ICH-098603, holotype, 50 mm SL; paratypes, FSJF 2383, 47 mm SL, 43 mm SL; right column from the top: females, 76 mm SL, 72 mm SL, 55 mm SL; Turkey: stream Tuzla.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6DEC861F79FFB0718BF8E6FC5CF8A3" box="[151,1015,1818,1840]" httpUri="https://zenodo.org/record/2615883/files/figure.png" pageId="65" pageNumber="66" startId="65.[151,250,1818,1840]" targetBox="[151,1435,1354,1797]" targetPageId="65">
|
||
<paragraph id="8BADBC0E1F79FFB0718BF8E6FC5CF8A3" blockId="65.[151,1015,1818,1840]" box="[151,1015,1818,1840]" pageId="65" pageNumber="66">
|
||
<emphasis id="B966601C1F79FFB0718BF8E6FE8BF8A3" bold="true" box="[151,288,1818,1840]" pageId="65" pageNumber="66">FIGURE 53.</emphasis>
|
||
<taxonomicName id="4C12C78D1F79FFB07039F8E6FE71F8A3" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[293,474,1818,1840]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F79FFB07039F8E6FE71F8A3" box="[293,474,1818,1840]" italics="true" pageId="65" pageNumber="66">Cobitis troasensis</emphasis>
|
||
</taxonomicName>
|
||
, FSJF 2317, male, 60 mm SL; Turkey: stream Tuzla.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BADBC0E1F79FFB371DBF8A1FF5AFE43" blockId="65.[151,1437,1885,2018]" lastBlockId="66.[151,1436,151,861]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
|
||
<emphasis id="B966601C1F79FFB071DBF8A1FEF4F8E5" bold="true" box="[199,351,1885,1910]" pageId="65" pageNumber="66">Colouration.</emphasis>
|
||
Body whitish with a dark-brown pigmentation pattern organised in one mid-dorsal and four lateral zones. Mid-dorsal pigmentation consists of a series of 16–20 roundish blotches in few and a marbled pattern on mid-dorsal back in most individuals. Many, very small spots in Z1, Z1 as wide as or wider than Z2, pigmentation not reaching dorsally to interspaces of mid-dorsal blotches, reaching to dorsal-fin base, pigmentation in Z1 fused with pigmentation in Z2 and Z3 on postdorsal flank. Spots in Z2 of about ½–1 pupil size, very densely set, usually fused into a stripe anterior to dorsal-fin base. Pigmentation in Z2 reaching to anus in some individuals, reaching to caudal peduncle in other individuals. Few, very small spots in Z3 along predorsal flank in most male, many small spots in female, Z3 much wider than Z
|
||
<date id="FFAC9ACE1F7AFFB373D2FEF8FC87FE8E" box="[718,812,260,285]" pageId="66" pageNumber="67" value="2015-11-02">2. 11–15</date>
|
||
predorsal, 3–4 subdorsal and 10–17 postdorsal blotches in Z4, blotches small, irregularly comma-shaped anterior to anal-fin origin, larger, more squarish or roundish on caudal peduncle. A single black spot at upper caudal-fin base, distinct, smaller than eye size. Upper part of head, opercle and snout covered by small spots. A dark-brown stripe between eye and snout. Fins hyaline. Caudal fin with 4–8 and dorsal fin with 4–5 dark, sometimes irregular shaped bars. No pigmentation on paired fins. Barbels whitish.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308EF851F7AFFB371DBFE20FD11FD8B" pageId="66" pageNumber="67" type="etymology">
|
||
<paragraph id="8BADBC0E1F7AFFB371DBFE20FD11FD8B" blockId="66.[151,1436,151,861]" pageId="66" pageNumber="67">
|
||
<emphasis id="B966601C1F7AFFB371DBFE20FEE5FE66" bold="true" box="[199,334,476,501]" pageId="66" pageNumber="67">Etymology.</emphasis>
|
||
Named for Alexandria Troas, an ancient Greek city located southeast of the modern village Dalyan in the
|
||
<collectingRegion id="49D672EC1F7AFFB37029FE03FDB2FD8B" box="[309,537,511,536]" country="Turkey" name="Canakkale" pageId="66" pageNumber="67">Çanakkale Province</collectingRegion>
|
||
. An adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308EF851F7AFFB371DBFDD8FD15FDF3" pageId="66" pageNumber="67" type="distribution">
|
||
<paragraph id="8BADBC0E1F7AFFB371DBFDD8FD15FDF3" blockId="66.[151,1436,151,861]" pageId="66" pageNumber="67">
|
||
<emphasis id="B966601C1F7AFFB371DBFDD8FECBFDAE" bold="true" box="[199,352,548,573]" pageId="66" pageNumber="67">Distribution.</emphasis>
|
||
Known only from the Tuzla River. The Tuzla River is flowing to the northernmost Aegean Sea in
|
||
<collectingCountry id="F305FC9E1F7AFFB371A8FDBBFEAEFDF3" box="[180,261,583,608]" name="Turkey" pageId="66" pageNumber="67">Turkey</collectingCountry>
|
||
, just at the entrance of the Dardanelles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308EF851F7AFFB371DBFD90FCB8FCCF" pageId="66" pageNumber="67" type="discussion">
|
||
<paragraph id="8BADBC0E1F7AFFB371DBFD90FCB8FCCF" blockId="66.[151,1436,151,861]" pageId="66" pageNumber="67">
|
||
<emphasis id="B966601C1F7AFFB371DBFD90FE90FD16" bold="true" box="[199,315,620,645]" pageId="66" pageNumber="67">Remarks.</emphasis>
|
||
No molecular data are available from
|
||
<taxonomicName id="4C12C78D1F7AFFB373EAFD90FC21FD17" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[758,906,620,644]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F7AFFB373EAFD90FC21FD17" box="[758,906,620,644]" italics="true" pageId="66" pageNumber="67">C. troasensis</emphasis>
|
||
</taxonomicName>
|
||
but we expect this species to belong to the
|
||
<taxonomicName id="4C12C78D1F7AFFB3749EFD90FF76FD34" authorityName="Linnaeus" authorityYear="1758" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="taenia">
|
||
<emphasis id="B966601C1F7AFFB3749EFD90FF76FD34" italics="true" pageId="66" pageNumber="67">C. taenia</emphasis>
|
||
</taxonomicName>
|
||
species group, as it has only one scale-shaped lamina canestini in the male (vs. two in other species groups in the area).
|
||
<taxonomicName id="4C12C78D1F7AFFB37030FD48FE53FD5F" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[300,504,692,716]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F7AFFB37030FD48FE53FD5F" box="[300,504,692,716]" italics="true" pageId="66" pageNumber="67">Cobitis troasensis</emphasis>
|
||
</taxonomicName>
|
||
might be endemic to the Tuzla River. Tuzla River is also inhabited by
|
||
<taxonomicName id="4C12C78D1F7AFFB37420FD48FEACFD63" authorityName="Özulug & Freyhof" authorityYear="2011" class="Actinopterygii" family="Cyprinidae" genus="Squalius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="aristotelis">
|
||
<emphasis id="B966601C1F7AFFB37420FD48FEACFD63" italics="true" pageId="66" pageNumber="67">Squalius aristotelis</emphasis>
|
||
</taxonomicName>
|
||
, a second species described as being endemic to the Tuzla River by
|
||
<bibRefCitation id="EF83C1FF1F7AFFB37502FD2BFAEDFD63" author="Ozulug, M. & Freyhof, J." box="[1054,1350,727,752]" pageId="66" pageNumber="67" pagination="107 - 148" refId="ref48692" refString="Ozulug, M. & Freyhof, J. (2011) Revision of the genus Squalius in western and central Anatolia, with description of four new species (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 22, 107 - 148." type="journal article" year="2011">Özuluğ & Freyhof (2011)</bibRefCitation>
|
||
. Future studies should investigate the fish fauna of Tuzla River and adjacent rivers more in detail not only to better understand the biogeographic relationships of fish species inhabiting this river, but also to find out, if
|
||
<taxonomicName id="4C12C78D1F7AFFB37417FCDCFA37FCA4" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[1291,1436,800,823]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="troasensis">
|
||
<emphasis id="B966601C1F7AFFB37417FCDCFA37FCA4" box="[1291,1436,800,823]" italics="true" pageId="66" pageNumber="67">C. troasensis</emphasis>
|
||
</taxonomicName>
|
||
(and
|
||
<taxonomicName id="4C12C78D1F7AFFB371CCFCB8FEF7FCCF" authorityName="Özulug & Freyhof" authorityYear="2011" box="[208,348,836,860]" class="Actinopterygii" family="Cyprinidae" genus="Squalius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="aristotelis">
|
||
<emphasis id="B966601C1F7AFFB371CCFCB8FEF7FCCF" box="[208,348,836,860]" italics="true" pageId="66" pageNumber="67">S. aristotelis</emphasis>
|
||
</taxonomicName>
|
||
) are indeed endemic to this small area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |