591 lines
71 KiB
XML
591 lines
71 KiB
XML
<document id="1B2FDE6E9CEDFA4D0803904A016164F7" ID-CLB-Dataset="27727" ID-DOI="10.11646/zootaxa.4535.1.1" ID-GBIF-Dataset="a047bcd9-ab65-4e3f-b07a-c830c7af1072" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615773" ID-ZooBank="ABE9DB1F-7378-4571-90C4-A3FDB66527F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553788891023" checkinUser="plazi" docAuthor="Freyhof, Jörg, Bayçelebi, Esra & Geiger, Matthias" docDate="2018" docId="03BB0D181F0BFFC7718BFF6BFC28FEF6" docLanguage="en" docName="zootaxa.4535.1.1.pdf" docOrigin="Zootaxa 4535 (1)" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Cobitis emrei Freyhof & Bayçelebi & Geiger 2018, new species" docType="treatment" docVersion="11" lastPageNumber="54" masterDocId="FF8275601F38FFF1711CFFFCFFABFF93" masterDocTitle="Review of the genus Cobitis in the Middle East, with the description of eight new species (Teleostei: Cobitidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="52" updateTime="1698715239854" updateUser="ExternalLinkService">
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<mods:mods id="A08A67BE05CC1F851039BB5EBE909024" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4D7181F4A7A3D08DCA3533B2631443AE">
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<mods:title id="D69DD74E1FD7FAE76E8081B0AC3BCF38">Review of the genus Cobitis in the Middle East, with the description of eight new species (Teleostei: Cobitidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="FC6B437C7DF238DB0425E84DE718FAD4" type="personal">
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<mods:role id="29CD6B20989674601F74DF9CE0B3281C">
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<mods:roleTerm id="54EC337E84F1DC34F430642EFAB4B8DB">Author</mods:roleTerm>
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<mods:namePart id="92AA052C3EF455E2D4E7923ADC629F0A">Freyhof, Jörg</mods:namePart>
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</mods:name>
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<mods:roleTerm id="E85C3BCBE14FCF6428A96CF96D513B1D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E49EDA6D827761904AACC3187D40B0A5">Bayçelebi, Esra</mods:namePart>
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</mods:name>
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<mods:name id="4B210477D4B7F5605B743FA022C80C40" type="personal">
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<mods:roleTerm id="A487C1EEA325373CFF10615EFF9F9C31">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7B9D6720FB58613A4662D33CAAF22254">Geiger, Matthias</mods:namePart>
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<mods:typeOfResource id="B1EE46C5AA5631C1C77078C927A095BD">text</mods:typeOfResource>
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<mods:titleInfo id="15299B62E6B0A627F89900CD23AA34A9">
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<mods:title id="8118DAD94E3848C7065DC06583A75555">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5BD81CA30BE97E66119052898B0C51AE">
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<mods:date id="F0C1C987B19877D9F640FC39F2D9C2DE">2018</mods:date>
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<mods:detail id="B921CDE7573ECD905D0A34DDEBABA6BD" type="pubDate">
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<mods:number id="3831A0D639DE9AB37CBD43635BC078EA">2018-12-21</mods:number>
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</mods:detail>
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<mods:detail id="F6EF7F7AA84AD52C3CC1A2994E01E47E" type="volume">
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<mods:number id="2399A819E27821E18CAC0C34887FCE8C">4535</mods:number>
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</mods:detail>
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<mods:detail id="639AAA69AD6458934487B84DF1EA721E" type="issue">
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<mods:number id="9A2B351DE2B12701087A955506E40669">1</mods:number>
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<mods:start id="73CEE9F693CA8A0F78229EEBBCD8D0C4">1</mods:start>
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<mods:classification id="6FD132A563342B297B69FDEBB620AE51">journal article</mods:classification>
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<mods:identifier id="A453640DBDF8E9CE25E9FD492CEDD282" type="CLB-Dataset">27727</mods:identifier>
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<mods:identifier id="7187C3A7FDD969E16D3BDA832879ECCE" type="DOI">10.11646/zootaxa.4535.1.1</mods:identifier>
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<mods:identifier id="9031ED36279A8D7284AEB71E6C8894DC" type="GBIF-Dataset">a047bcd9-ab65-4e3f-b07a-c830c7af1072</mods:identifier>
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<mods:identifier id="67E11A6E217A530CED4CAAAFEAFC3659" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F12D150D9C42423D32821A938C6D719F" type="Zenodo-Dep">2615773</mods:identifier>
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<mods:identifier id="84F440B2C121261944E139F5DDE4EC95" type="ZooBank">ABE9DB1F-7378-4571-90C4-A3FDB66527F3</mods:identifier>
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</mods:mods>
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<treatment id="03BB0D181F0BFFC7718BFF6BFC28FEF6" ID-DOI="http://doi.org/10.5281/zenodo.5699583" ID-GBIF-Taxon="154521904" ID-Zenodo-Dep="5699583" LSID="urn:lsid:plazi:treatment:03BB0D181F0BFFC7718BFF6BFC28FEF6" httpUri="http://treatment.plazi.org/id/03BB0D181F0BFFC7718BFF6BFC28FEF6" lastPageId="54" lastPageNumber="54" pageId="51" pageNumber="52">
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<subSubSection id="C308EF851F0BFFC2718BFF6BFE7DFF21" box="[151,470,151,178]" pageId="51" pageNumber="52" type="nomenclature">
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<paragraph id="8BADBC0E1F0BFFC2718BFF6BFE7DFF21" blockId="51.[151,470,151,211]" box="[151,470,151,178]" pageId="51" pageNumber="52">
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<heading id="D0E50B621F0BFFC2718BFF6BFE7DFF21" bold="true" box="[151,470,151,178]" fontSize="11" level="1" pageId="51" pageNumber="52" reason="1">
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<emphasis id="B966601C1F0BFFC2718BFF6BFE7DFF21" bold="true" box="[151,470,151,178]" pageId="51" pageNumber="52">
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<taxonomicName id="4C12C78D1F0BFFC2718BFF6BFE9CFF22" authority="Freyhof & Bayçelebi & Geiger, 2018" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[151,311,151,177]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="emrei" status="sp. nov.">
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<emphasis id="B966601C1F0BFFC2718BFF6BFE9CFF22" bold="true" box="[151,311,151,177]" italics="true" pageId="51" pageNumber="52">Cobitis emrei</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A255DD671F0BFFC2705AFF64FE7DFF21" box="[326,470,152,178]" pageId="51" pageNumber="52" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C308EF851F0BFFC2718BFF46FE88FF40" box="[151,291,186,211]" pageId="51" pageNumber="52" type="description">
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||
<paragraph id="8BADBC0E1F0BFFC2718BFF46FE88FF40" blockId="51.[151,470,151,211]" box="[151,291,186,211]" pageId="51" pageNumber="52">
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||
(
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<figureCitation id="1329A08B1F0BFFC27183FF46FEB0FF40" box="[159,283,186,211]" captionStart-0="FIGURE 39" captionStart-1="FIGURE 40" captionStart-2="FIGURE 41" captionStartId-0="51.[151,250,1599,1621]" captionStartId-1="52.[151,250,1032,1054]" captionStartId-2="52.[151,250,1555,1577]" captionTargetBox-0="[151,1436,1092,1578]" captionTargetBox-1="[191,1408,616,979]" captionTargetBox-2="[151,1435,1135,1544]" captionTargetId-0="figure@51.[151,1436,1092,1578]" captionTargetId-1="figure@52.[151,1436,589,1011]" captionTargetId-2="figure@52.[151,1435,1135,1544]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="52" captionText-0="FIGURE 39. Cobitis emrei, left column from the top: males, ZFMK ICH-099187, holotype, 56 mm SL; paratypes, FSJF 1960, 54 mm SL, 48 mm SL; right column from the top: females, 77 mm SL, 72 mm SL, 69 mm SL female; Turkey: Lake Sapanca basin." captionText-1="FIGURE 40. Cobitis emrei, left column from the top: males, ZFMK ICH-099187, holotype, 56 mm SL; paratypes, FSJF 1960, 54 mm SL, 48 mm SL; right column from the top: females, 77 mm SL, 72 mm SL, 69 mm SL; Turkey: Lake Sapanca basin." captionText-2="FIGURE 41. Cobitis emrei, FSJF 1960, 55 mm SL, male; Turkey: stream Kurtköy flowing to Lake Sapanca." httpUri-0="https://zenodo.org/record/2615853/files/figure.png" httpUri-1="https://zenodo.org/record/2615855/files/figure.png" httpUri-2="https://zenodo.org/record/2615857/files/figure.png" pageId="51" pageNumber="52">Fig. 39–41</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C308EF851F0BFFC2718BFEFEFEB1FE5C" pageId="51" pageNumber="52" type="materials_examined">
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<paragraph id="8BADBC0E1F0BFFC2718BFEFEFD83FEAC" blockId="51.[151,1437,258,1039]" pageId="51" pageNumber="52">
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<materialsCitation id="3B7AB6531F0BFFC2718BFEFEFD83FEAC" ID-GBIF-Occurrence="2120485956" collectionCode="ZFMK" country="Turkey" county="Adapazari" location="Lake Sapanca" municipality="Kurtkoy" pageId="51" pageNumber="52" specimenCode="ICH-099187, 56" specimenCount="1" stateProvince="Sakarya" typeStatus="holotype">
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<emphasis id="B966601C1F0BFFC2718BFEFEFEA0FE88" bold="true" box="[151,267,258,283]" pageId="51" pageNumber="52">
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||
<typeStatus id="54A902AC1F0BFFC2718BFEFEFEADFE88" box="[151,262,258,283]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
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||
.
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</emphasis>
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<collectionCode id="ED0324CB1F0BFFC2700FFEFFFECAFE89" box="[275,353,259,282]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="51" pageNumber="52" type="Museum">ZFMK</collectionCode>
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<specimenCode id="DBB414751F0BFFC27076FEFFFE51FE88" box="[362,506,259,283]" collectionCode="ICH-" pageId="51" pageNumber="52">ICH-099187</specimenCode>
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||
,
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||
<quantity id="4CEA11EB1F0BFFC2731BFEFEFDF3FE89" box="[519,600,258,283]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="51" pageNumber="52" unit="mm" value="56.0">
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<specimenCode id="DBB414751F0BFFC2731BFEFEFD88FE88" box="[519,547,258,283]" collectionCode="ICH-" pageId="51" pageNumber="52">56</specimenCode>
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mm
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||
</quantity>
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||
SL;
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<collectingCountry id="F305FC9E1F0BFFC2738DFEFEFD4EFE88" box="[657,741,258,283]" name="Turkey" pageId="51" pageNumber="52">Turkey</collectingCountry>
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||
:
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||
<collectingCounty id="62CCC4821F0BFFC273EEFEFEFCCEFE88" box="[754,869,258,283]" pageId="51" pageNumber="52">Adapazarı</collectingCounty>
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||
(
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||
<collectingRegion id="49D672EC1F0BFFC27269FEFEFC79FE88" box="[885,978,258,283]" country="Turkey" name="Sakarya" pageId="51" pageNumber="52">Sakarya</collectingRegion>
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||
) prov.: stream
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<collectingMunicipality id="6BC926741F0BFFC27563FEFEFB77FE88" box="[1151,1244,258,283]" pageId="51" pageNumber="52">Kurtköy</collectingMunicipality>
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||
flowing to
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||
<location id="8ECDEAD51F0BFFC27478FEFEFF5FFEAC" LSID="urn:lsid:plazi:treatment:03BB0D181F0BFFC7718BFF6BFC28FEF6:8ECDEAD51F0BFFC27478FEFEFF5FFEAC" country="Turkey" county="Adapazari" municipality="Kurtkoy" name="Lake Sapanca" pageId="51" pageNumber="52" stateProvince="Sakarya">Lake Sapanca</location>
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||
at
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<location id="8ECDEAD51F0BFFC2700AFEDAFEDFFEAC" LSID="urn:lsid:plazi:treatment:03BB0D181F0BFFC7718BFF6BFC28FEF6:8ECDEAD51F0BFFC2700AFEDAFEDFFEAC" box="[278,372,294,319]" country="Turkey" county="Adapazari" municipality="Kurtkoy" name="Kurtkoy" pageId="51" pageNumber="52" stateProvince="Sakarya">Kurtköy</location>
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, 40.713 30.175.
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BADBC0E1F0BFFC271DBFEB6FC00FEF0" blockId="51.[151,1437,258,1039]" box="[199,939,330,355]" pageId="51" pageNumber="52">
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<materialsCitation id="3B7AB6531F0BFFC271DBFEB6FC0CFEF0" ID-GBIF-Occurrence="2120485957" box="[199,935,330,355]" collectionCode="FSJF" country="Turkey" county="Adapazari" location="Lake Sapanca" municipality="Kurtkoy" pageId="51" pageNumber="52" specimenCode="FSJF 1960, 10, 48-77" specimenCount="1" stateProvince="Sakarya" typeStatus="paratype">
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<emphasis id="B966601C1F0BFFC271DBFEB6FEEDFEF0" bold="true" box="[199,326,330,355]" pageId="51" pageNumber="52">
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<typeStatus id="54A902AC1F0BFFC271DBFEB6FEEAFEF0" box="[199,321,330,355]" pageId="51" pageNumber="52" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<specimenCode id="DBB414751F0BFFC27050FEB7FE63FEF0" box="[332,456,330,355]" collectionCode="FSJF" pageId="51" pageNumber="52">FSJF 1960</specimenCode>
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,
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<specimenCode id="DBB414751F0BFFC270CFFEB7FE5AFEF0" box="[467,497,331,355]" collectionCode="FSJF" pageId="51" pageNumber="52">10</specimenCode>
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||
,
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<quantity id="4CEA11EB1F0BFFC270E1FEB7FDDDFEF1" box="[509,630,331,355]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="7.7" metricValueMin="4.8" pageId="51" pageNumber="52" unit="mm" value="62.5" valueMax="77.0" valueMin="48.0">
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<specimenCode id="DBB414751F0BFFC270E1FEB7FDE8FEF1" box="[509,579,331,355]" collectionCode="FSJF" pageId="51" pageNumber="52">48–77</specimenCode>
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mm
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</quantity>
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SL; same data as holotype
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BADBC0E1F0BFFC271DBFE92FEB1FE5C" blockId="51.[151,1437,258,1039]" pageId="51" pageNumber="52">
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||
<emphasis id="B966601C1F0BFFC271DBFE92FD4AFE14" bold="true" box="[199,737,366,391]" pageId="51" pageNumber="52">Material used in molecular genetic analysis.</emphasis>
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FSJF DNA-173;
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<collectingCountry id="F305FC9E1F0BFFC272ADFE92FBAEFE14" box="[945,1029,366,391]" name="Turkey" pageId="51" pageNumber="52">Turkey</collectingCountry>
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: Adapazarı (
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<collectingRegion id="49D672EC1F0BFFC27584FE92FB5EFE14" box="[1176,1269,366,391]" country="Turkey" name="Sakarya" pageId="51" pageNumber="52">Sakarya</collectingRegion>
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||
) prov.: stream Kurtköy flowing to Lake Sapanca at Kurtköy, 40.712 30.175. (GenBank accession numbers:
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<accessionNumber id="944121ED1F0BFFC2743EFE6EFA33FE38" box="[1314,1432,402,427]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ553154" pageId="51" pageNumber="52">KJ553154</accessionNumber>
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||
,
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<accessionNumber id="944121ED1F0BFFC2718BFE4AFEA4FE5C" box="[151,271,438,463]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KJ553118" pageId="51" pageNumber="52">KJ553118</accessionNumber>
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).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C308EF851F0BFFC271DBFE26FA26FB9C" pageId="51" pageNumber="52" type="diagnosis">
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||
<paragraph id="8BADBC0E1F0BFFC271DBFE26FED8FC80" blockId="51.[151,1437,258,1039]" pageId="51" pageNumber="52">
|
||
<emphasis id="B966601C1F0BFFC271DBFE26FEEAFE60" bold="true" box="[199,321,474,499]" pageId="51" pageNumber="52">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C12C78D1F0BFFC27054FE27FE75FE61" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[328,478,475,499]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="emrei">
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<emphasis id="B966601C1F0BFFC27054FE27FE75FE61" box="[328,478,475,499]" italics="true" pageId="51" pageNumber="52">Cobitis emrei</emphasis>
|
||
</taxonomicName>
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||
is distinguished from other
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<taxonomicName id="4C12C78D1F0BFFC27200FE27FCC6FE60" box="[796,877,475,499]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B966601C1F0BFFC27200FE27FCC6FE60" box="[796,877,475,499]" italics="true" pageId="51" pageNumber="52">Cobitis</emphasis>
|
||
</taxonomicName>
|
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species in the Black Sea basins by a combination of characters, none of them unique. It is distinguished from
|
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<taxonomicName id="4C12C78D1F0BFFC27250FE03FC76FD85" authorityName="Erk'akan, Atalay-Ekmekçi & Nalbant" authorityYear="1998" box="[844,989,510,534]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="splendens">
|
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<emphasis id="B966601C1F0BFFC27250FE03FC76FD85" box="[844,989,510,534]" italics="true" pageId="51" pageNumber="52">C. splendens</emphasis>
|
||
</taxonomicName>
|
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by the pigmentation in Z3 reaching to between the nape and the vertical of the dorsal-fin origin or even to the caudal peduncle in the males (vs. Z3 reduced to about head length), the pigmentation in Z3 usually wider than zone Z2 (vs. narrower or of equal width), the blotches in Z4 being horizontally elongate, often very densely set and fused into short or longer stripes in the male and some female (vs. Z4 anterior to dorsal-fin origin, irregularly roundish or squarish, well separate, very rarely fused into a stripe) and the back with 7–11 dark-brown large, roundish or squarish blotches, in one individual examined, blotches fused (vs. a marmorate pattern on the back or the blotches very indistinct from each other and irregularly shaped).
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||
</paragraph>
|
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<paragraph id="8BADBC0E1F0BFFC271DBFCE2FA26FB9C" blockId="51.[151,1437,258,1039]" pageId="51" pageNumber="52">
|
||
<taxonomicName id="4C12C78D1F0BFFC271DBFCE2FEF5FCA5" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[199,350,798,822]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="emrei">
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||
<emphasis id="B966601C1F0BFFC271DBFCE2FEF5FCA5" box="[199,350,798,822]" italics="true" pageId="51" pageNumber="52">Cobitis emrei</emphasis>
|
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</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C12C78D1F0BFFC27347FCE3FD69FCA5" authorityName="Linnaeus" authorityYear="1758" box="[603,706,799,822]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="taenia">
|
||
<emphasis id="B966601C1F0BFFC27347FCE3FD69FCA5" box="[603,706,799,822]" italics="true" pageId="51" pageNumber="52">C. taenia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C12C78D1F0BFFC273CDFCE3FCECFCA5" authorityName="Vasil'eva & Vasil'ev" authorityYear="2006" box="[721,839,799,822]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="pontica">
|
||
<emphasis id="B966601C1F0BFFC273CDFCE3FCECFCA5" box="[721,839,799,822]" italics="true" pageId="51" pageNumber="52">C. pontica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C12C78D1F0BFFC27262FCE3FB84FCA5" box="[894,1071,798,822]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="elongatoides">
|
||
<emphasis id="B966601C1F0BFFC27262FCE3FB84FCA5" box="[894,1071,798,822]" italics="true" pageId="51" pageNumber="52">C. elongatoides</emphasis>
|
||
</taxonomicName>
|
||
by having horizontally elongate blotches in Z4 often very densely set and fused into short or longer stripes in the male and some female (vs. blotches vertically elongate, roundish, triangular or squarish, not fused), the focal zone in the subdorsal scales small, about ¼ or less than 1/3 of the maximum scale diameter (vs. large about 1/3 or more in
|
||
<taxonomicName id="4C12C78D1F0BFFC275D3FC77FA9CFC31" authorityName="Linnaeus" authorityYear="1758" box="[1231,1335,907,930]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="taenia">
|
||
<emphasis id="B966601C1F0BFFC275D3FC77FA9CFC31" box="[1231,1335,907,930]" italics="true" pageId="51" pageNumber="52">C. taenia</emphasis>
|
||
</taxonomicName>
|
||
) and the black spot at the upper caudal-fin base roundish or ovoid, larger than the eye diameter in the male (vs. usually comma-shaped, smaller than the eye diameter in the male and female
|
||
<taxonomicName id="4C12C78D1F0BFFC27283FC2FFBADFC79" authorityName="Linnaeus" authorityYear="1758" box="[927,1030,979,1002]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="taenia">
|
||
<emphasis id="B966601C1F0BFFC27283FC2FFBADFC79" box="[927,1030,979,1002]" italics="true" pageId="51" pageNumber="52">C. taenia</emphasis>
|
||
</taxonomicName>
|
||
) and Z3 fully covered by very small spots forming a sand-like pattern in the female (vs. pigmentation in Z3 not sand-like in
|
||
<taxonomicName id="4C12C78D1F0BFFC27577FC0BFB7AFB9D" authorityName="Linnaeus" authorityYear="1758" box="[1131,1233,1015,1038]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="taenia">
|
||
<emphasis id="B966601C1F0BFFC27577FC0BFB7AFB9D" box="[1131,1233,1015,1038]" italics="true" pageId="51" pageNumber="52">C. taenia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C12C78D1F0BFFC27414FC0BFAD7FB9D" authorityName="Vasil'eva & Vasil'ev" authorityYear="2006" box="[1288,1404,1015,1038]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="pontica">
|
||
<emphasis id="B966601C1F0BFFC27414FC0BFAD7FB9D" box="[1288,1404,1015,1038]" italics="true" pageId="51" pageNumber="52">C. pontica</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6DEC861F0BFFC2718BF9C3FF78F901" httpUri="https://zenodo.org/record/2615853/files/figure.png" pageId="51" pageNumber="52" startId="51.[151,250,1599,1621]" targetBox="[151,1436,1092,1578]" targetPageId="51">
|
||
<paragraph id="8BADBC0E1F0BFFC2718BF9C3FF78F901" blockId="51.[151,1436,1599,1682]" pageId="51" pageNumber="52">
|
||
<emphasis id="B966601C1F0BFFC2718BF9C3FEB4F9C6" bold="true" box="[151,287,1599,1621]" pageId="51" pageNumber="52">FIGURE 39.</emphasis>
|
||
<taxonomicName id="4C12C78D1F0BFFC27039F9C3FE06F9C6" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[293,429,1599,1621]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F0BFFC27039F9C3FE06F9C6" box="[293,429,1599,1621]" italics="true" pageId="51" pageNumber="52">Cobitis emrei</emphasis>
|
||
</taxonomicName>
|
||
, left column from the top: males, ZFMK ICH-099187, holotype, 56 mm SL; paratypes, FSJF 1960, 54 mm SL, 48 mm SL; right column from the top: females, 77 mm SL, 72 mm SL, 69 mm SL female; Turkey: Lake Sapanca basin.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C308EF851F0BFFC471DBF946FA30F95F" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52" type="description">
|
||
<paragraph id="8BADBC0E1F0BFFC271DBF946FE99F8F1" blockId="51.[151,1436,1722,2035]" pageId="51" pageNumber="52">
|
||
<taxonomicName id="4C12C78D1F0BFFC271DBF946FEF5F941" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[199,350,1722,1746]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F0BFFC271DBF946FEF5F941" box="[199,350,1722,1746]" italics="true" pageId="51" pageNumber="52">Cobitis emrei</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C12C78D1F0BFFC27343F946FD4EF941" box="[607,741,1722,1746]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="tanaitica">
|
||
<emphasis id="B966601C1F0BFFC27343F946FD4EF941" box="[607,741,1722,1746]" italics="true" pageId="51" pageNumber="52">C. tanaitica</emphasis>
|
||
</taxonomicName>
|
||
by the pigmentation in Z3 reaching to between the nape and the vertical of the dorsal-fin origin or even to the caudal peduncle in the males (vs. Z3 usually reaching to a point anterior to the dorsal-fin base), the blotches in Z4 being horizontally elongate, often very densely set and fused into short or longer stripes in the males (vs. vertically elongate, not fused) and Z3 wider than Z2 (vs. usually Z3 narrower Z2).
|
||
</paragraph>
|
||
<paragraph id="8BADBC0E1F0BFFC571DBF892FE45FF46" blockId="51.[151,1436,1722,2035]" lastBlockId="52.[151,1437,151,536]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
|
||
<taxonomicName id="4C12C78D1F0BFFC271DBF892FEF6F815" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[199,349,1902,1926]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F0BFFC271DBF892FEF6F815" box="[199,349,1902,1926]" italics="true" pageId="51" pageNumber="52">Cobitis emrei</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C12C78D1F0BFFC27349F893FD79F815" box="[597,722,1902,1926]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="fahireae">
|
||
<emphasis id="B966601C1F0BFFC27349F893FD79F815" box="[597,722,1902,1926]" italics="true" pageId="51" pageNumber="52">C. fahireae</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<collectingRegion id="49D672EC1F0BFFC27221F893FC39F814" box="[829,914,1903,1927]" country="Greece" name="Voreio Aigaio" pageId="51" pageNumber="52">Aegean</collectingRegion>
|
||
Sea basin by having a large, roundish or ovoid black spot at the upper caudal-fin base, larger than eye diameter in the males (vs. usually comma-shaped, smaller than eye diameter in the male and female or spot absent). It is distinguished from
|
||
<taxonomicName id="4C12C78D1F0BFFC2754CF84BFB6BF85D" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[1104,1216,1974,1998]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="afifeae">
|
||
<emphasis id="B966601C1F0BFFC2754CF84BFB6BF85D" box="[1104,1216,1974,1998]" italics="true" pageId="51" pageNumber="52">C. afifeae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C12C78D1F0BFFC275CCF84BFAD9F85D" authorityName="Erkakan, Ozdemir & Ozeren" authorityYear="2017" box="[1232,1394,1974,1998]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="dorademiri">
|
||
<emphasis id="B966601C1F0BFFC275CCF84BFAD9F85D" box="[1232,1394,1974,1998]" italics="true" pageId="51" pageNumber="52">C. dorademiri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B966601C1F0BFFC2749EF84BFE10F861" italics="true" pageId="51" pageNumber="52">
|
||
<taxonomicName id="4C12C78D1F0BFFC2749EF84BFF56F861" authorityName="Battalgazi" authorityYear="1944" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="phrygica">C. phrygica</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C12C78D1F0BFFC27017F826FE10F861" authorityName="Erk'akan, Atalay-Ekmekçi & Nalbant" authorityYear="1998" box="[267,443,2010,2034]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="puncticulata">C. puncticulata</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C12C78D1F0BFFC270EBF826FD1BF861" authorityName="Hanko" authorityYear="1924" box="[503,688,2010,2034]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="simplicispina">
|
||
<emphasis id="B966601C1F0BFFC270EBF826FD1BF861" box="[503,688,2010,2034]" italics="true" pageId="51" pageNumber="52">C. simplicispina</emphasis>
|
||
</taxonomicName>
|
||
by having one laminae circularis in the males (vs. two).
|
||
<taxonomicName id="4C12C78D1F0BFFC57457F826FF7DFF3C" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="52" lastPageNumber="53" order="Cypriniformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F0BFFC57457F826FF7DFF3C" italics="true" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">Cobitis emrei</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C12C78D1F0CFFC570CCFF64FD92FF3C" authorityName="Eagderi, Jouladeh-Roudbar, Jalili, Sayyadzadeh & Esmaeili" authorityYear="2017" box="[464,569,152,175]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="saniae">
|
||
<emphasis id="B966601C1F0CFFC570CCFF64FD92FF3C" box="[464,569,152,175]" italics="true" pageId="52" pageNumber="53">C. saniae</emphasis>
|
||
</taxonomicName>
|
||
by having the base of the lamina circularis narrowly attached to the pectoralfin ray (vs. widely connected).
|
||
</paragraph>
|
||
<paragraph id="8BADBC0E1F0CFFC571DBFF23FCA3FE1B" blockId="52.[151,1437,151,536]" pageId="52" pageNumber="53">
|
||
<emphasis id="B966601C1F0CFFC571DBFF23FEF2FF6B" bold="true" box="[199,345,223,248]" pageId="52" pageNumber="53">Description.</emphasis>
|
||
See
|
||
<figureCitation id="1329A08B1F0CFFC57091FF23FD85FF6B" box="[397,558,223,248]" captionStart-0="FIGURE 49" captionStart-1="FIGURE 51" captionStartId-0="61.[151,250,1505,1527]" captionStartId-1="65.[151,250,802,824]" captionTargetBox-0="[171,1421,1094,1442]" captionTargetBox-1="[151,1409,372,746]" captionTargetId-0="figure@61.[151,1436,1058,1483]" captionTargetId-1="figure@65.[151,1436,337,781]" captionTargetPageId-0="61" captionTargetPageId-1="65" captionText-0="FIGURE 49. Cobitis pirii, left column from the top: males, ZFMK ICH-99635, holotype, 67 mm SL; paratypes, FSJF 2469, 61 mm SL, 59 mm SL; right column from the top: females, FSJF 2474, 95 mm SL, FSJF 2469 74 mm SL, 72 mm SL; Turkey: streams Aksu and Çayköy." captionText-1="FIGURE 51. Cobitis troasensis, left column from the top: males, ZFMK ICH-098603, holotype, 50 mm SL; paratypes, FSJF 2383, 47 mm SL, 43 mm SL; right column from the top: females, 76 mm SL, 72 mm SL, 55 mm SL; Turkey: stream Tuzla." httpUri-0="https://zenodo.org/record/2615875/files/figure.png" httpUri-1="https://zenodo.org/record/2615877/files/figure.png" pageId="52" pageNumber="53">Figures 49–51</figureCitation>
|
||
for general appearance and
|
||
<tableCitation id="C69089B51F0CFFC57275FF23FC10FF6B" box="[873,955,223,248]" captionStart="TABLE 5" captionStartId="53.[151,239,152,175]" captionText="TABLE 5. Morphometric data of Cobitis emrei (holotype ZFMK ICH 099187 and paratypes FSJF 1960; n = 10. The calculations include the holotype." pageId="52" pageNumber="53">Table 5</tableCitation>
|
||
for morphometric data of the
|
||
<typeStatus id="54A902AC1F0CFFC57417FF23FAC6FF6B" box="[1291,1389,223,248]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
|
||
and the
|
||
<specimenCount id="9D1477871F0CFFC571D8FEF8FEF2FE8E" box="[196,345,260,285]" pageId="52" pageNumber="53" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
|
||
. Greatest body depth at or slightly anterior to dorsal-fin origin, decreasing towards caudal-fin base. Head profile slightly convex, head length 1.1
|
||
<emphasis id="B966601C1F0CFFC573C2FED5FD47FED3" box="[734,748,297,320]" italics="true" pageId="52" pageNumber="53">–</emphasis>
|
||
1.4 times in body depth. Snout oblong or pointed, its length 0.6
|
||
<emphasis id="B966601C1F0CFFC571A6FEB1FF63FEF7" box="[186,200,333,356]" italics="true" pageId="52" pageNumber="53">–</emphasis>
|
||
0.8 times in postorbital length. Eye diameter 0 2
|
||
<emphasis id="B966601C1F0CFFC573F0FEB1FD51FEF7" box="[748,762,333,356]" italics="true" pageId="52" pageNumber="53">–</emphasis>
|
||
0 4 times in head depth at eye, 1.0
|
||
<emphasis id="B966601C1F0CFFC5759CFEB1FB25FEF7" box="[1152,1166,333,356]" italics="true" pageId="52" pageNumber="53">–</emphasis>
|
||
2.0 times in interorbital width. Caudal peduncle 1.0
|
||
<emphasis id="B966601C1F0CFFC570D6FE8DFE73FE1B" box="[458,472,369,392]" italics="true" pageId="52" pageNumber="53">–</emphasis>
|
||
1.6 times longer than deep.
|
||
</paragraph>
|
||
<paragraph id="8BADBC0E1F0CFFC571DBFE68FE2DFD8B" blockId="52.[151,1437,151,536]" pageId="52" pageNumber="53">
|
||
Pelvic axillary lobe present or absent. Margin of dorsal fin convex and margin of anal fin convex. Caudal fin truncate or slightly rounded. No dorsal keel on caudal peduncle and a shallow ventral keel on caudal peduncle. External part of the suborbital spine bifurcate, reaching to or slightly beyond centre of eye. Largest recorded specimen
|
||
<quantity id="4CEA11EB1F0CFFC57014FDFCFEFCFD84" box="[264,343,512,535]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="52" pageNumber="53" unit="mm" value="77.0">77 mm</quantity>
|
||
SL.
|
||
</paragraph>
|
||
<caption id="DF6DEC861F0CFFC5718BFBF4FA29FBAE" httpUri="https://zenodo.org/record/2615855/files/figure.png" pageId="52" pageNumber="53" startId="52.[151,250,1032,1054]" targetBox="[191,1408,616,979]" targetPageId="52">
|
||
<paragraph id="8BADBC0E1F0CFFC5718BFBF4FA29FBAE" blockId="52.[151,1436,1032,1085]" pageId="52" pageNumber="53">
|
||
<emphasis id="B966601C1F0CFFC5718BFBF4FEB4FB8D" bold="true" box="[151,287,1032,1054]" pageId="52" pageNumber="53">FIGURE 40.</emphasis>
|
||
<taxonomicName id="4C12C78D1F0CFFC57039FBF4FE06FB8D" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[293,429,1032,1054]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F0CFFC57039FBF4FE06FB8D" box="[293,429,1032,1054]" italics="true" pageId="52" pageNumber="53">Cobitis emrei</emphasis>
|
||
</taxonomicName>
|
||
, left column from the top: males, ZFMK ICH-099187, holotype, 56 mm SL; paratypes, FSJF 1960, 54 mm SL, 48 mm SL; right column from the top: females, 77 mm SL, 72 mm SL, 69 mm SL; Turkey: Lake Sapanca basin.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6DEC861F0CFFC5718BF9EFFB43F9BA" box="[151,1256,1555,1577]" httpUri="https://zenodo.org/record/2615857/files/figure.png" pageId="52" pageNumber="53" startId="52.[151,250,1555,1577]" targetBox="[151,1435,1135,1544]" targetPageId="52">
|
||
<paragraph id="8BADBC0E1F0CFFC5718BF9EFFB43F9BA" blockId="52.[151,1256,1555,1577]" box="[151,1256,1555,1577]" pageId="52" pageNumber="53">
|
||
<emphasis id="B966601C1F0CFFC5718BF9EFFEB4F9BA" bold="true" box="[151,287,1555,1577]" pageId="52" pageNumber="53">FIGURE 41.</emphasis>
|
||
<taxonomicName id="4C12C78D1F0CFFC57039F9EFFE05F9BA" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[293,430,1555,1577]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F0CFFC57039F9EFFE05F9BA" box="[293,430,1555,1577]" italics="true" pageId="52" pageNumber="53">Cobitis emrei</emphasis>
|
||
</taxonomicName>
|
||
, FSJF 1960, 55 mm SL, male; Turkey: stream Kurtköy flowing to Lake Sapanca.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BADBC0E1F0CFFC571DBF9AAFD97F81C" blockId="52.[151,1437,1622,1935]" pageId="52" pageNumber="53">
|
||
Dorsal fin with 3 unbranched and 6½ (4) and 7½ (7) branched rays. Anal fin with 3 unbranched and 5½ branched rays. Caudal fin with 6+5 branched rays in the male, 7+7 branched rays in the
|
||
<specimenCount id="9D1477871F0CFFC5756FF986FB67F900" box="[1139,1228,1658,1683]" pageId="52" pageNumber="53" type="male">5 males</specimenCount>
|
||
, 7+
|
||
<quantity id="4CEA11EB1F0CFFC575E9F986FA8BF900" box="[1269,1312,1658,1683]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="52" pageNumber="53" unit="in" value="6.0">6 in</quantity>
|
||
the female and 7+
|
||
<quantity id="4CEA11EB1F0CFFC571FBF963FEBEF924" box="[231,277,1694,1719]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="52" pageNumber="53" unit="in" value="7.0">7 in</quantity>
|
||
the
|
||
<specimenCount id="9D1477871F0CFFC57050F963FE14F924" box="[332,447,1694,1719]" pageId="52" pageNumber="53" type="female">4 females</specimenCount>
|
||
. Pectoral fin with 8 (9) and 9 (2) branched rays and pelvic fin with 6 (10) and 5 (1) branched rays. Body completely covered by embedded scales, except on belly and breast. Scales small. Focal zone in subdorsal scales about 1/4 or less than 1/3 of vertical scale diameter. Lateral line short, with 3–6 pores or an open slit in skin along lateral line. Lips (
|
||
<figureCitation id="1329A08B1F0CFFC57335F8F6FDD7F8B0" box="[553,636,1802,1827]" captionStart="FIGURE 54" captionStartId="67.[151,250,1575,1597]" captionTargetBox="[151,1436,220,1518]" captionTargetId="figure@67.[151,1436,220,1518]" captionTargetPageId="67" captionText="FIGURE 54. Upper row: Cobitis afifeae, FSJF 3022, paratype, 76 mm SL; Turkey: Büyük Menderes River; C. aliyeae, FSJF 2427, paratype, female, 65 mm SL; Turkey: Seyhan River; C. anabelae, ZFMK ICH-98633, holotype, male, 53 mm SL; Turkey: Karasu River; C. emrei, FSJF 1960, paratype, female, 77 mm SL; Turkey: Lake Sapanca basin; Lower row: C. erkakanae, FSJF 2510, paratype, female, 68 mm SL; Turkey: canal at Lake Gölbaşı; C. joergbohleni, ZFMK ICH-97624, holotype, female, 60 mm SL; Turkey: spring Soysallı; C. pirii, FSJF 2474, paratype, female, 95 mm SL; Turkey: streams Çayköy; C. troasensis, FSJF 2383, paratype, female, 72 mm SL; Turkey: stream Tuzla." httpUri="https://zenodo.org/record/2615885/files/figure.png" pageId="52" pageNumber="53">Fig. 54</figureCitation>
|
||
) thin and mental lobes of lower lip short, usually well separated from lower lip, rarely produced into a short barbel-like process. Rostral barbel reaching base of mandibular barbel. Mandibular barbel reaching to beyond vertical of nostril or corner of eye. Maxillary barbel reaching vertical of front border or usually middle of eye.
|
||
</paragraph>
|
||
<caption id="DF6DEC861F0DFFC4718BFF64FE55FF5C" pageId="53" pageNumber="54">
|
||
<paragraph id="8BADBC0E1F0DFFC4718BFF64FE55FF5C" blockId="53.[151,1436,151,207]" pageId="53" pageNumber="54">
|
||
<emphasis id="B966601C1F0DFFC4718BFF64FEA2FF3C" bold="true" box="[151,265,152,175]" pageId="53" pageNumber="54">TABLE 5.</emphasis>
|
||
Morphometric data of
|
||
<taxonomicName id="4C12C78D1F0DFFC47317FF6BFD37FF3C" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[523,668,151,175]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F0DFFC47317FF6BFD37FF3C" box="[523,668,151,175]" italics="true" pageId="53" pageNumber="54">Cobitis emrei</emphasis>
|
||
</taxonomicName>
|
||
(holotype ZFMK ICH 0 99187 and paratypes FSJF 1960; n = 10. The calculations include the holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BADBC0E1F0DFFC473B0FF16FA98FB1D" pageId="53" pageNumber="54">
|
||
<table id="F9124EAE1F0D000E7183FF16FAEBFB1D" box="[159,1344,234,1166]" colsContinueFrom="50.[159,1346,234,1166]" colsContinueIn="56.[159,1346,234,1166]" gridcols="6" gridrows="23" pageId="53" pageNumber="54">
|
||
<tr id="3522BE4C1F0D000E7183FF16FAEBFE93" box="[159,1344,234,256]" gridrow="0" pageId="53" pageNumber="54" rowspan-0="1" rowspan-4="1" rowspan-5="1">
|
||
<th id="76F3D7301F0D000E73B0FF16FCAFFE93" box="[684,772,234,256]" gridcol="1" gridrow="0" pageId="53" pageNumber="54">holotype</th>
|
||
<th id="76F3D7301F0D000E7258FF16FBB5FE93" box="[836,1054,234,256]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="53" pageNumber="54">holotype & paratypes</th>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FEE8FAEBFEB9" box="[159,1344,276,298]" gridrow="1" pageId="53" pageNumber="54" rowspan-0="1" rowspan-3="1" rowspan-5="1">
|
||
<td id="76F3D7301F0D000E73B0FEE8FCAFFEB9" box="[684,772,276,298]" gridcol="1" gridrow="1" pageId="53" pageNumber="54">male</td>
|
||
<td id="76F3D7301F0D000E7258FEE8FC01FEB9" box="[836,938,276,298]" gridcol="2" gridrow="1" pageId="53" pageNumber="54">male</td>
|
||
<td id="76F3D7301F0D000E754FFEE8FB13FEB9" box="[1107,1208,276,298]" gridcol="4" gridrow="1" pageId="53" pageNumber="54">female</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FEC2FAEBFEC0" box="[159,1344,318,339]" gridrow="2" pageId="53" pageNumber="54" rowspan-0="1" rowspan-1="1">
|
||
<td id="76F3D7301F0D000E7258FEC2FC01FEC0" box="[836,938,318,339]" gridcol="2" gridrow="2" pageId="53" pageNumber="54">range</td>
|
||
<td id="76F3D7301F0D000E72FEFEC2FBB5FEC0" box="[994,1054,318,339]" gridcol="3" gridrow="2" pageId="53" pageNumber="54">mean</td>
|
||
<td id="76F3D7301F0D000E754FFEC2FB13FEC0" box="[1107,1208,318,339]" gridcol="4" gridrow="2" pageId="53" pageNumber="54">range</td>
|
||
<td id="76F3D7301F0D000E741BFEC2FAEBFEC0" box="[1287,1344,318,339]" gridcol="5" gridrow="2" pageId="53" pageNumber="54">mean</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FE9AFAEBFEEF" box="[159,1344,358,380]" gridrow="3" pageId="53" pageNumber="54" rowspan-3="1" rowspan-5="1">
|
||
<th id="76F3D7301F0D000E7183FE9AFE4AFEEF" box="[159,481,358,380]" gridcol="0" gridrow="3" pageId="53" pageNumber="54">SL (mm)</th>
|
||
<td id="76F3D7301F0D000E73B0FE9AFCAFFEEF" box="[684,772,358,380]" gridcol="1" gridrow="3" pageId="53" pageNumber="54">56</td>
|
||
<td id="76F3D7301F0D000E7258FE9AFC01FEEF" box="[836,938,358,380]" gridcol="2" gridrow="3" pageId="53" pageNumber="54">48–56</td>
|
||
<td id="76F3D7301F0D000E754FFE9AFB13FEEF" box="[1107,1208,358,380]" gridcol="4" gridrow="3" pageId="53" pageNumber="54">69-77</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FE6CFAEBFE35" box="[159,1344,400,422]" gridrow="4" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FE6CFAEBFE35" box="[159,1344,400,422]" colspan="6" colspanRight="5" gridcol="0" gridrow="4" pageId="53" pageNumber="54">In percent of standard length</th>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FE45FAEBFE5C" box="[159,1344,441,463]" gridrow="5" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FE45FE4AFE5C" box="[159,481,441,463]" gridcol="0" gridrow="5" pageId="53" pageNumber="54">Head length</th>
|
||
<td id="76F3D7301F0D000E73B0FE45FCAFFE5C" box="[684,772,441,463]" gridcol="1" gridrow="5" pageId="53" pageNumber="54">21.9</td>
|
||
<td id="76F3D7301F0D000E7258FE45FC01FE5C" box="[836,938,441,463]" gridcol="2" gridrow="5" pageId="53" pageNumber="54">21.4–23.8</td>
|
||
<td id="76F3D7301F0D000E72FEFE45FBB5FE5C" box="[994,1054,441,463]" gridcol="3" gridrow="5" pageId="53" pageNumber="54">22.2</td>
|
||
<td id="76F3D7301F0D000E754FFE45FB13FE5C" box="[1107,1208,441,463]" gridcol="4" gridrow="5" pageId="53" pageNumber="54">19.8–21.2</td>
|
||
<td id="76F3D7301F0D000E741BFE45FAEBFE5C" box="[1287,1344,441,463]" gridcol="5" gridrow="5" pageId="53" pageNumber="54">20.5</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FE1EFAEBFE6B" box="[159,1344,482,504]" gridrow="6" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FE1EFE4AFE6B" box="[159,481,482,504]" gridcol="0" gridrow="6" pageId="53" pageNumber="54">Predorsal length</th>
|
||
<td id="76F3D7301F0D000E73B0FE1EFCAFFE6B" box="[684,772,482,504]" gridcol="1" gridrow="6" pageId="53" pageNumber="54">51.9</td>
|
||
<td id="76F3D7301F0D000E7258FE1EFC01FE6B" box="[836,938,482,504]" gridcol="2" gridrow="6" pageId="53" pageNumber="54">51.9–58.9</td>
|
||
<td id="76F3D7301F0D000E72FEFE1EFBB5FE6B" box="[994,1054,482,504]" gridcol="3" gridrow="6" pageId="53" pageNumber="54">54.1</td>
|
||
<td id="76F3D7301F0D000E754FFE1EFB13FE6B" box="[1107,1208,482,504]" gridcol="4" gridrow="6" pageId="53" pageNumber="54">53.0–54.9</td>
|
||
<td id="76F3D7301F0D000E741BFE1EFAEBFE6B" box="[1287,1344,482,504]" gridcol="5" gridrow="6" pageId="53" pageNumber="54">53.8</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FDF0FAEBFDB1" box="[159,1344,524,546]" gridrow="7" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FDF0FE4AFDB1" box="[159,481,524,546]" gridcol="0" gridrow="7" pageId="53" pageNumber="54">Prepelvic length</th>
|
||
<td id="76F3D7301F0D000E73B0FDF0FCAFFDB1" box="[684,772,524,546]" gridcol="1" gridrow="7" pageId="53" pageNumber="54">53.2</td>
|
||
<td id="76F3D7301F0D000E7258FDF0FC01FDB1" box="[836,938,524,546]" gridcol="2" gridrow="7" pageId="53" pageNumber="54">52.0–57.5</td>
|
||
<td id="76F3D7301F0D000E72FEFDF0FBB5FDB1" box="[994,1054,524,546]" gridcol="3" gridrow="7" pageId="53" pageNumber="54">54.8</td>
|
||
<td id="76F3D7301F0D000E754FFDF0FB13FDB1" box="[1107,1208,524,546]" gridcol="4" gridrow="7" pageId="53" pageNumber="54">52.3–55.2</td>
|
||
<td id="76F3D7301F0D000E741BFDF0FAEBFDB1" box="[1287,1344,524,546]" gridcol="5" gridrow="7" pageId="53" pageNumber="54">53.5</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FDC9FAEBFDD8" box="[159,1344,565,587]" gridrow="8" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FDC9FE4AFDD8" box="[159,481,565,587]" gridcol="0" gridrow="8" pageId="53" pageNumber="54">Preanal length</th>
|
||
<td id="76F3D7301F0D000E73B0FDC9FCAFFDD8" box="[684,772,565,587]" gridcol="1" gridrow="8" pageId="53" pageNumber="54">77.8</td>
|
||
<td id="76F3D7301F0D000E7258FDC9FC01FDD8" box="[836,938,565,587]" gridcol="2" gridrow="8" pageId="53" pageNumber="54">77.8–80.7</td>
|
||
<td id="76F3D7301F0D000E72FEFDC9FBB5FDD8" box="[994,1054,565,587]" gridcol="3" gridrow="8" pageId="53" pageNumber="54">79.6</td>
|
||
<td id="76F3D7301F0D000E754FFDC9FB13FDD8" box="[1107,1208,565,587]" gridcol="4" gridrow="8" pageId="53" pageNumber="54">79.8–81.2</td>
|
||
<td id="76F3D7301F0D000E741BFDC9FAEBFDD8" box="[1287,1344,565,587]" gridcol="5" gridrow="8" pageId="53" pageNumber="54">80.5</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FDA2FAEBFDE7" box="[159,1344,606,628]" gridrow="9" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FDA2FE4AFDE7" box="[159,481,606,628]" gridcol="0" gridrow="9" pageId="53" pageNumber="54">Body depth at dorsal-fin origin</th>
|
||
<td id="76F3D7301F0D000E73B0FDA2FCAFFDE7" box="[684,772,606,628]" gridcol="1" gridrow="9" pageId="53" pageNumber="54">18.3</td>
|
||
<td id="76F3D7301F0D000E7258FDA2FC01FDE7" box="[836,938,606,628]" gridcol="2" gridrow="9" pageId="53" pageNumber="54">16.2–18.3</td>
|
||
<td id="76F3D7301F0D000E72FEFDA2FBB5FDE7" box="[994,1054,606,628]" gridcol="3" gridrow="9" pageId="53" pageNumber="54">17.4</td>
|
||
<td id="76F3D7301F0D000E754FFDA2FB13FDE7" box="[1107,1208,606,628]" gridcol="4" gridrow="9" pageId="53" pageNumber="54">17.0–18.4</td>
|
||
<td id="76F3D7301F0D000E741BFDA2FAEBFDE7" box="[1287,1344,606,628]" gridcol="5" gridrow="9" pageId="53" pageNumber="54">17.9</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FD74FAEBFD0D" box="[159,1344,648,670]" gridrow="10" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FD74FE4AFD0D" box="[159,481,648,670]" gridcol="0" gridrow="10" pageId="53" pageNumber="54">Body width at dorsal-fin origin</th>
|
||
<td id="76F3D7301F0D000E73B0FD74FCAFFD0D" box="[684,772,648,670]" gridcol="1" gridrow="10" pageId="53" pageNumber="54">9.1</td>
|
||
<td id="76F3D7301F0D000E7258FD74FC01FD0D" box="[836,938,648,670]" gridcol="2" gridrow="10" pageId="53" pageNumber="54">8.9–10.8</td>
|
||
<td id="76F3D7301F0D000E72FEFD74FBB5FD0D" box="[994,1054,648,670]" gridcol="3" gridrow="10" pageId="53" pageNumber="54">9.6</td>
|
||
<td id="76F3D7301F0D000E754FFD74FB13FD0D" box="[1107,1208,648,670]" gridcol="4" gridrow="10" pageId="53" pageNumber="54">10.1–11.8</td>
|
||
<td id="76F3D7301F0D000E741BFD74FAEBFD0D" box="[1287,1344,648,670]" gridcol="5" gridrow="10" pageId="53" pageNumber="54">10.9</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FD4DFAEBFD54" box="[159,1344,689,711]" gridrow="11" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FD4DFE4AFD54" box="[159,481,689,711]" gridcol="0" gridrow="11" pageId="53" pageNumber="54">Depth of caudal peduncle</th>
|
||
<td id="76F3D7301F0D000E73B0FD4DFCAFFD54" box="[684,772,689,711]" gridcol="1" gridrow="11" pageId="53" pageNumber="54">9.3</td>
|
||
<td id="76F3D7301F0D000E7258FD4DFC01FD54" box="[836,938,689,711]" gridcol="2" gridrow="11" pageId="53" pageNumber="54">10.7–12.0</td>
|
||
<td id="76F3D7301F0D000E72FEFD4DFBB5FD54" box="[994,1054,689,711]" gridcol="3" gridrow="11" pageId="53" pageNumber="54">11.3</td>
|
||
<td id="76F3D7301F0D000E754FFD4DFB13FD54" box="[1107,1208,689,711]" gridcol="4" gridrow="11" pageId="53" pageNumber="54">9.4–10.2</td>
|
||
<td id="76F3D7301F0D000E741BFD4DFAEBFD54" box="[1287,1344,689,711]" gridcol="5" gridrow="11" pageId="53" pageNumber="54">9.8</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FD26FAEBFD63" box="[159,1344,730,752]" gridrow="12" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FD26FE4AFD63" box="[159,481,730,752]" gridcol="0" gridrow="12" pageId="53" pageNumber="54">Length of caudal peduncle</th>
|
||
<td id="76F3D7301F0D000E73B0FD26FCAFFD63" box="[684,772,730,752]" gridcol="1" gridrow="12" pageId="53" pageNumber="54">12.0</td>
|
||
<td id="76F3D7301F0D000E7258FD26FC01FD63" box="[836,938,730,752]" gridcol="2" gridrow="12" pageId="53" pageNumber="54">11.5–16.6</td>
|
||
<td id="76F3D7301F0D000E72FEFD26FBB5FD63" box="[994,1054,730,752]" gridcol="3" gridrow="12" pageId="53" pageNumber="54">13.8</td>
|
||
<td id="76F3D7301F0D000E754FFD26FB13FD63" box="[1107,1208,730,752]" gridcol="4" gridrow="12" pageId="53" pageNumber="54">12.4–15.2</td>
|
||
<td id="76F3D7301F0D000E741BFD26FAEBFD63" box="[1287,1344,730,752]" gridcol="5" gridrow="12" pageId="53" pageNumber="54">13.9</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FCF8FAEBFC89" box="[159,1344,772,794]" gridrow="13" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FCF8FE4AFC89" box="[159,481,772,794]" gridcol="0" gridrow="13" pageId="53" pageNumber="54">Dorsal-fin depth</th>
|
||
<td id="76F3D7301F0D000E73B0FCF8FCAFFC89" box="[684,772,772,794]" gridcol="1" gridrow="13" pageId="53" pageNumber="54">18.7</td>
|
||
<td id="76F3D7301F0D000E7258FCF8FC01FC89" box="[836,938,772,794]" gridcol="2" gridrow="13" pageId="53" pageNumber="54">20.1–21.9</td>
|
||
<td id="76F3D7301F0D000E72FEFCF8FBB5FC89" box="[994,1054,772,794]" gridcol="3" gridrow="13" pageId="53" pageNumber="54">20.8</td>
|
||
<td id="76F3D7301F0D000E754FFCF8FB13FC89" box="[1107,1208,772,794]" gridcol="4" gridrow="13" pageId="53" pageNumber="54">17.5–19.2</td>
|
||
<td id="76F3D7301F0D000E741BFCF8FAEBFC89" box="[1287,1344,772,794]" gridcol="5" gridrow="13" pageId="53" pageNumber="54">18.2</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FCD1FAEBFCD0" box="[159,1344,813,835]" gridrow="14" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FCD1FE4AFCD0" box="[159,481,813,835]" gridcol="0" gridrow="14" pageId="53" pageNumber="54">Caudal-fin length</th>
|
||
<td id="76F3D7301F0D000E73B0FCD1FCAFFCD0" box="[684,772,813,835]" gridcol="1" gridrow="14" pageId="53" pageNumber="54">16.6</td>
|
||
<td id="76F3D7301F0D000E7258FCD1FC01FCD0" box="[836,938,813,835]" gridcol="2" gridrow="14" pageId="53" pageNumber="54">18.1–20.9</td>
|
||
<td id="76F3D7301F0D000E72FEFCD1FBB5FCD0" box="[994,1054,813,835]" gridcol="3" gridrow="14" pageId="53" pageNumber="54">18.7</td>
|
||
<td id="76F3D7301F0D000E754FFCD1FB13FCD0" box="[1107,1208,813,835]" gridcol="4" gridrow="14" pageId="53" pageNumber="54">15.4–19.0</td>
|
||
<td id="76F3D7301F0D000E741BFCD1FAEBFCD0" box="[1287,1344,813,835]" gridcol="5" gridrow="14" pageId="53" pageNumber="54">17.2</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FCAAFAEBFCFF" box="[159,1344,854,876]" gridrow="15" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FCAAFE4AFCFF" box="[159,481,854,876]" gridcol="0" gridrow="15" pageId="53" pageNumber="54">Pectoral-fin length</th>
|
||
<td id="76F3D7301F0D000E73B0FCAAFCAFFCFF" box="[684,772,854,876]" gridcol="1" gridrow="15" pageId="53" pageNumber="54">20.1</td>
|
||
<td id="76F3D7301F0D000E7258FCAAFC01FCFF" box="[836,938,854,876]" gridcol="2" gridrow="15" pageId="53" pageNumber="54">21.3–22.6</td>
|
||
<td id="76F3D7301F0D000E72FEFCAAFBB5FCFF" box="[994,1054,854,876]" gridcol="3" gridrow="15" pageId="53" pageNumber="54">21.7</td>
|
||
<td id="76F3D7301F0D000E754FFCAAFB13FCFF" box="[1107,1208,854,876]" gridcol="4" gridrow="15" pageId="53" pageNumber="54">13.4–16.4</td>
|
||
<td id="76F3D7301F0D000E741BFCAAFAEBFCFF" box="[1287,1344,854,876]" gridcol="5" gridrow="15" pageId="53" pageNumber="54">14.4</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FC7CFAEBFC05" box="[159,1344,896,918]" gridrow="16" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FC7CFE4AFC05" box="[159,481,896,918]" gridcol="0" gridrow="16" pageId="53" pageNumber="54">Pelvic-fin length</th>
|
||
<td id="76F3D7301F0D000E73B0FC7CFCAFFC05" box="[684,772,896,918]" gridcol="1" gridrow="16" pageId="53" pageNumber="54">18.1</td>
|
||
<td id="76F3D7301F0D000E7258FC7CFC01FC05" box="[836,938,896,918]" gridcol="2" gridrow="16" pageId="53" pageNumber="54">15.2–16.9</td>
|
||
<td id="76F3D7301F0D000E72FEFC7CFBB5FC05" box="[994,1054,896,918]" gridcol="3" gridrow="16" pageId="53" pageNumber="54">16.0</td>
|
||
<td id="76F3D7301F0D000E754FFC7CFB13FC05" box="[1107,1208,896,918]" gridcol="4" gridrow="16" pageId="53" pageNumber="54">12.9–14.5</td>
|
||
<td id="76F3D7301F0D000E741BFC7CFAEBFC05" box="[1287,1344,896,918]" gridcol="5" gridrow="16" pageId="53" pageNumber="54">13.4</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FC55FAEBFC2C" box="[159,1344,937,959]" gridrow="17" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FC55FAEBFC2C" box="[159,1344,937,959]" colspan="6" colspanRight="5" gridcol="0" gridrow="17" pageId="53" pageNumber="54">In percent of head length</th>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FC2EFAEBFC7B" box="[159,1344,978,1000]" gridrow="18" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FC2EFE4AFC7B" box="[159,481,978,1000]" gridcol="0" gridrow="18" pageId="53" pageNumber="54">Snout length</th>
|
||
<td id="76F3D7301F0D000E73B0FC2EFCAFFC7B" box="[684,772,978,1000]" gridcol="1" gridrow="18" pageId="53" pageNumber="54">41</td>
|
||
<td id="76F3D7301F0D000E7258FC2EFC01FC7B" box="[836,938,978,1000]" gridcol="2" gridrow="18" pageId="53" pageNumber="54">35–41</td>
|
||
<td id="76F3D7301F0D000E72FEFC2EFBB5FC7B" box="[994,1054,978,1000]" gridcol="3" gridrow="18" pageId="53" pageNumber="54">38.1</td>
|
||
<td id="76F3D7301F0D000E754FFC2EFB13FC7B" box="[1107,1208,978,1000]" gridcol="4" gridrow="18" pageId="53" pageNumber="54">38–42</td>
|
||
<td id="76F3D7301F0D000E741BFC2EFAEBFC7B" box="[1287,1344,978,1000]" gridcol="5" gridrow="18" pageId="53" pageNumber="54">40.7</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FC00FAEBFB81" box="[159,1344,1020,1042]" gridrow="19" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FC00FE4AFB81" box="[159,481,1020,1042]" gridcol="0" gridrow="19" pageId="53" pageNumber="54">Postorbital distance</th>
|
||
<td id="76F3D7301F0D000E73B0FC00FCAFFB81" box="[684,772,1020,1042]" gridcol="1" gridrow="19" pageId="53" pageNumber="54">52</td>
|
||
<td id="76F3D7301F0D000E7258FC00FC01FB81" box="[836,938,1020,1042]" gridcol="2" gridrow="19" pageId="53" pageNumber="54">51–58</td>
|
||
<td id="76F3D7301F0D000E72FEFC00FBB5FB81" box="[994,1054,1020,1042]" gridcol="3" gridrow="19" pageId="53" pageNumber="54">54.0</td>
|
||
<td id="76F3D7301F0D000E754FFC00FB13FB81" box="[1107,1208,1020,1042]" gridcol="4" gridrow="19" pageId="53" pageNumber="54">50–57</td>
|
||
<td id="76F3D7301F0D000E741BFC00FAEBFB81" box="[1287,1344,1020,1042]" gridcol="5" gridrow="19" pageId="53" pageNumber="54">53.4</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FBD9FAEBFBA8" box="[159,1344,1061,1083]" gridrow="20" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FBD9FE4AFBA8" box="[159,481,1061,1083]" gridcol="0" gridrow="20" pageId="53" pageNumber="54">Head depth at eye</th>
|
||
<td id="76F3D7301F0D000E73B0FBD9FCAFFBA8" box="[684,772,1061,1083]" gridcol="1" gridrow="20" pageId="53" pageNumber="54">51</td>
|
||
<td id="76F3D7301F0D000E7258FBD9FC01FBA8" box="[836,938,1061,1083]" gridcol="2" gridrow="20" pageId="53" pageNumber="54">47–55</td>
|
||
<td id="76F3D7301F0D000E72FEFBD9FBB5FBA8" box="[994,1054,1061,1083]" gridcol="3" gridrow="20" pageId="53" pageNumber="54">51.3</td>
|
||
<td id="76F3D7301F0D000E754FFBD9FB13FBA8" box="[1107,1208,1061,1083]" gridcol="4" gridrow="20" pageId="53" pageNumber="54">52–55</td>
|
||
<td id="76F3D7301F0D000E741BFBD9FAEBFBA8" box="[1287,1344,1061,1083]" gridcol="5" gridrow="20" pageId="53" pageNumber="54">53.8</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FBB2FAEBFBF7" box="[159,1344,1102,1124]" gridrow="21" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FBB2FE4AFBF7" box="[159,481,1102,1124]" gridcol="0" gridrow="21" pageId="53" pageNumber="54">Eye diameter</th>
|
||
<td id="76F3D7301F0D000E73B0FBB2FCAFFBF7" box="[684,772,1102,1124]" gridcol="1" gridrow="21" pageId="53" pageNumber="54">16</td>
|
||
<td id="76F3D7301F0D000E7258FBB2FC01FBF7" box="[836,938,1102,1124]" gridcol="2" gridrow="21" pageId="53" pageNumber="54">13–18</td>
|
||
<td id="76F3D7301F0D000E72FEFBB2FBB5FBF7" box="[994,1054,1102,1124]" gridcol="3" gridrow="21" pageId="53" pageNumber="54">15.8</td>
|
||
<td id="76F3D7301F0D000E754FFBB2FB13FBF7" box="[1107,1208,1102,1124]" gridcol="4" gridrow="21" pageId="53" pageNumber="54">17–19</td>
|
||
<td id="76F3D7301F0D000E741BFBB2FAEBFBF7" box="[1287,1344,1102,1124]" gridcol="5" gridrow="21" pageId="53" pageNumber="54">17.8</td>
|
||
</tr>
|
||
<tr id="3522BE4C1F0D000E7183FB84FAEBFB1D" box="[159,1344,1144,1166]" gridrow="22" pageId="53" pageNumber="54">
|
||
<th id="76F3D7301F0D000E7183FB84FE4AFB1D" box="[159,481,1144,1166]" gridcol="0" gridrow="22" pageId="53" pageNumber="54">Interorbital width</th>
|
||
<td id="76F3D7301F0D000E73B0FB84FCAFFB1D" box="[684,772,1144,1166]" gridcol="1" gridrow="22" pageId="53" pageNumber="54">12</td>
|
||
<td id="76F3D7301F0D000E7258FB84FC01FB1D" box="[836,938,1144,1166]" gridcol="2" gridrow="22" pageId="53" pageNumber="54">9–14</td>
|
||
<td id="76F3D7301F0D000E72FEFB84FBB5FB1D" box="[994,1054,1144,1166]" gridcol="3" gridrow="22" pageId="53" pageNumber="54">12.3</td>
|
||
<td id="76F3D7301F0D000E754FFB84FB13FB1D" box="[1107,1208,1144,1166]" gridcol="4" gridrow="22" pageId="53" pageNumber="54">17–19</td>
|
||
<td id="76F3D7301F0D000E741BFB84FAEBFB1D" box="[1287,1344,1144,1166]" gridcol="5" gridrow="22" pageId="53" pageNumber="54">17.8</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BADBC0E1F0DFFC471DBFB2EFE24FA9C" blockId="53.[151,1437,1234,2022]" pageId="53" pageNumber="54">
|
||
<emphasis id="B966601C1F0DFFC471DBFB2EFE13FB78" bold="true" box="[199,440,1234,1259]" pageId="53" pageNumber="54">Sexual dimorphism.</emphasis>
|
||
Males have a longer pectoral fin than females (21
|
||
<emphasis id="B966601C1F0DFFC472E5FB2FFBACFB79" box="[1017,1031,1235,1258]" italics="true" pageId="53" pageNumber="54">–</emphasis>
|
||
23% SL vs. 13
|
||
<emphasis id="B966601C1F0DFFC475ADFB2FFB14FB79" box="[1201,1215,1235,1258]" italics="true" pageId="53" pageNumber="54">–</emphasis>
|
||
16) and one lamina circularis (vs. absent).
|
||
</paragraph>
|
||
<paragraph id="8BADBC0E1F0DFFC471D1FADCFA30F95F" blockId="53.[151,1437,1234,2022]" pageId="53" pageNumber="54">
|
||
<emphasis id="B966601C1F0DFFC471D1FADCFECEFAAA" bold="true" box="[205,357,1312,1337]" pageId="53" pageNumber="54">Colouration.</emphasis>
|
||
Background colour whitish with dark-brown pigmentation pattern organised in one mid-dorsal and four lateral zones. Mid-dorsal pigmentation consist of a series of 14–20, roundish blotches, often irregularlyshaped blotches, forming a marbled pattern, fused into short or long dark-brown stripes in few individuals. Zone Z1 fully covered by very small spots forming a sand-like pattern, as wide as or wider than Z2, reaching on predorsal body to interspaces of mid-dorsal blotches, fused with Z2 and Z3 on postdorsal flank. Zone Z2 with elongate blotches larger than eye, usually fused on anterior part of flank in male, especially on flank anterior to dorsal-fin origin. Zone Z3 fully covered by very small spots forming a sand-like pattern, confluent with zone Z1 and Z2 on postdorsal flank, much wider than Z2. Pigmentation in Zone Z4 with 5–7 predorsal, 2–3 subdorsal and 5–8 postdorsal blotches, blotches elongate, usually much longer than wide, often confluent and forming short stripes. A single, distinct, eye sized, black spot at upper caudal-fin base. Upper part of head, opercle and snout covered by large spots or short vermiculation. A dark-brown stripe from eye to snout. Fins hyaline. Caudal fin with 5–7 and dorsal fin with 5–8 dark, sometimes irregularly-shaped bars. Few dark-brown spots in paired fins. Barbels whitish.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308EF851F0DFFC471DBF92DFC1CF89D" pageId="53" pageNumber="54" type="etymology">
|
||
<paragraph id="8BADBC0E1F0DFFC471DBF92DFC1CF89D" blockId="53.[151,1437,1234,2022]" pageId="53" pageNumber="54">
|
||
<emphasis id="B966601C1F0DFFC471DBF92DFEE5F979" bold="true" box="[199,334,1745,1770]" pageId="53" pageNumber="54">Etymology.</emphasis>
|
||
Named for Yunus Emre (about
|
||
<date id="FFAC9ACE1F0DFFC473AAF92EFC9DF979" box="[694,822,1746,1770]" pageId="53" pageNumber="54" value="1238" valueMax="1320">1238–1320</date>
|
||
) the folk poet, philosopher and Sufi mystic who is the pioneer of Turkish poetry in
|
||
<collectingCountry id="F305FC9E1F0DFFC470C5F909FD96F89D" box="[473,573,1781,1806]" name="Turkey" pageId="53" pageNumber="54">Anatolia</collectingCountry>
|
||
. A noun in genitive, indeclinable.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308EF851F0DFFC471DBF8E5FB7DF8A1" box="[199,1238,1817,1842]" pageId="53" pageNumber="54" type="distribution">
|
||
<paragraph id="8BADBC0E1F0DFFC471DBF8E5FB7DF8A1" blockId="53.[151,1437,1234,2022]" box="[199,1238,1817,1842]" pageId="53" pageNumber="54">
|
||
<emphasis id="B966601C1F0DFFC471DBF8E5FECBF8A1" bold="true" box="[199,352,1817,1842]" pageId="53" pageNumber="54">Distribution.</emphasis>
|
||
<taxonomicName id="4C12C78D1F0DFFC4707BF8E5FE56F8A2" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[359,509,1817,1841]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F0DFFC4707BF8E5FE56F8A2" box="[359,509,1817,1841]" italics="true" pageId="53" pageNumber="54">Cobitis emrei</emphasis>
|
||
</taxonomicName>
|
||
has been only identified from the Lake Sapanca basin in
|
||
<collectingCountry id="F305FC9E1F0DFFC4759CF8E5FB79F8A1" box="[1152,1234,1817,1842]" name="Turkey" pageId="53" pageNumber="54">Turkey</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C308EF851F0DFFC771DBF8C1FC28FEF6" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="discussion">
|
||
<paragraph id="8BADBC0E1F0DFFC771DBF8C1FC28FEF6" blockId="53.[151,1437,1234,2022]" lastBlockId="54.[151,1436,151,357]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
|
||
<emphasis id="B966601C1F0DFFC471DBF8C1FE90F8C5" bold="true" box="[199,315,1853,1878]" pageId="53" pageNumber="54">Remarks.</emphasis>
|
||
Erk'akan
|
||
<emphasis id="B966601C1F0DFFC470B1F8C2FE74F8C6" box="[429,479,1853,1877]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
|
||
. (1999) identified this species as
|
||
<taxonomicName id="4C12C78D1F0DFFC47241F8C2FBAAF8C6" authorityName="Karaman" authorityYear="1928" box="[861,1025,1853,1877]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="vardarensis">
|
||
<emphasis id="B966601C1F0DFFC47241F8C2FBAAF8C6" box="[861,1025,1853,1877]" italics="true" pageId="53" pageNumber="54">C. vardarensis</emphasis>
|
||
</taxonomicName>
|
||
, which is widespread in the river of the European Aegean Sea basin. Our molecular data clearly reject this hypothesis. While
|
||
<taxonomicName id="4C12C78D1F0DFFC475BDF89EFAEDF8EA" authorityName="Karaman" authorityYear="1928" box="[1185,1350,1889,1913]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="vardarensis">
|
||
<emphasis id="B966601C1F0DFFC475BDF89EFAEDF8EA" box="[1185,1350,1889,1913]" italics="true" pageId="53" pageNumber="54">C. vardarensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C12C78D1F0DFFC4749EF89EFF7DF80E" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="53" pageNumber="68" phylum="Chordata" rank="species" species="emrei">
|
||
C.
|
||
<emphasis id="B966601C1F0DFFC4718BF87AFF7DF80E" box="[151,214,1926,1949]" italics="true" pageId="53" pageNumber="54">emrei</emphasis>
|
||
</taxonomicName>
|
||
, both belong to the
|
||
<taxonomicName id="4C12C78D1F0DFFC470D8F87AFD85F80E" authorityName="Linnaeus" authorityYear="1758" box="[452,558,1926,1949]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="taenia">
|
||
<emphasis id="B966601C1F0DFFC470D8F87AFD85F80E" box="[452,558,1926,1949]" italics="true" pageId="53" pageNumber="54">C. taenia</emphasis>
|
||
</taxonomicName>
|
||
species group, they are not closely related. Erk'akan
|
||
<emphasis id="B966601C1F0DFFC47580F87AFB7BF80E" box="[1180,1232,1925,1949]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
|
||
. (1998) report
|
||
<taxonomicName id="4C12C78D1F0DFFC4749EF87AFEB2F852" authorityName="Karaman" authorityYear="1928" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="vardarensis">
|
||
<emphasis id="B966601C1F0DFFC4749EF87AFEB2F852" italics="true" pageId="53" pageNumber="54">C. vardarensis</emphasis>
|
||
</taxonomicName>
|
||
also from the lower Sakarya River drainage, but we did not find
|
||
<taxonomicName id="4C12C78D1F0DFFC47504F855FBC2F852" box="[1048,1129,1961,1985]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="53" pageNumber="54" phylum="Chordata" rank="genus">
|
||
<emphasis id="B966601C1F0DFFC47504F855FBC2F852" box="[1048,1129,1961,1985]" italics="true" pageId="53" pageNumber="54">Cobitis</emphasis>
|
||
</taxonomicName>
|
||
in the lower Sakarya, and suspect that these populations might be identified as
|
||
<emphasis id="B966601C1F0DFFC473F7F832FC5BF876" box="[747,1008,1997,2021]" italics="true" pageId="53" pageNumber="54">
|
||
<taxonomicName id="4C12C78D1F0DFFC473F7F832FCE5F876" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[747,846,1998,2021]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="emrei">C. emrei</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C12C78D1F0DFFC47245F831FC5BF876" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[857,1008,1997,2021]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="emrei">Cobitis emrei</taxonomicName>
|
||
</emphasis>
|
||
seems to be closer to
|
||
<taxonomicName id="4C12C78D1F0DFFC475F0F832FA37F876" box="[1260,1436,1997,2021]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="elongatoides">
|
||
<emphasis id="B966601C1F0DFFC475F0F832FB54F876" box="[1260,1279,1998,2021]" italics="true" pageId="53" pageNumber="54">C</emphasis>
|
||
.
|
||
<emphasis id="B966601C1F0DFFC47411F831FA37F876" box="[1293,1436,1997,2021]" italics="true" pageId="53" pageNumber="54">elongatoides</emphasis>
|
||
</taxonomicName>
|
||
from the Danube and Elbe drainages,
|
||
<taxonomicName id="4C12C78D1F0EFFC77320FF64FD1AFF3C" authorityName="Vasil'eva & Vasil'ev" authorityYear="2006" box="[572,689,152,175]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="pontica">
|
||
<emphasis id="B966601C1F0EFFC77320FF64FD1AFF3C" box="[572,689,152,175]" italics="true" pageId="54" pageNumber="55">C. pontica</emphasis>
|
||
</taxonomicName>
|
||
from the rivers south of the Danube in
|
||
<collectingCountry id="F305FC9E1F0EFFC77575FF6BFB62FF23" box="[1129,1225,151,176]" name="Bulgaria" pageId="54" pageNumber="55">Bulgaria</collectingCountry>
|
||
and Thrace and
|
||
<taxonomicName id="4C12C78D1F0EFFC7749EFF64FF50FF47" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="tanaitica">
|
||
<emphasis id="B966601C1F0EFFC7749EFF64FF50FF47" italics="true" pageId="54" pageNumber="55">C. tanaitica</emphasis>
|
||
</taxonomicName>
|
||
from the northern Black Sea basin.
|
||
<taxonomicName id="4C12C78D1F0EFFC77380FF40FC9FFF47" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[668,820,188,212]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F0EFFC77380FF40FC9FFF47" box="[668,820,188,212]" italics="true" pageId="54" pageNumber="55">Cobitis emrei</emphasis>
|
||
</taxonomicName>
|
||
is superficially very similar to
|
||
<taxonomicName id="4C12C78D1F0EFFC77581FF40FAE4FF47" box="[1181,1359,188,212]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="elongatoides">
|
||
<emphasis id="B966601C1F0EFFC77581FF40FAE4FF47" box="[1181,1359,188,212]" italics="true" pageId="54" pageNumber="55">C. elongatoides</emphasis>
|
||
</taxonomicName>
|
||
and is distinguished by small details in the colour pattern only. Both species are well distinguished by molecular characters and might represent almost real cryptic species. Based on DNA barcoding
|
||
<taxonomicName id="4C12C78D1F0EFFC7756DFEF8FB78FE8F" authorityName="Freyhof & Bayçelebi & Geiger" authorityYear="2018" box="[1137,1235,260,284]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="emrei">
|
||
<emphasis id="B966601C1F0EFFC7756DFEF8FB78FE8F" box="[1137,1235,260,284]" italics="true" pageId="54" pageNumber="55">C. emrei</emphasis>
|
||
</taxonomicName>
|
||
is well separated from all other included
|
||
<taxonomicName id="4C12C78D1F0EFFC770B2FED4FE54FED3" box="[430,511,296,320]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="B966601C1F0EFFC770B2FED4FE54FED3" box="[430,511,296,320]" italics="true" pageId="54" pageNumber="55">Cobitis</emphasis>
|
||
</taxonomicName>
|
||
, and by a minimum K2P distance of 2.5% to
|
||
<taxonomicName id="4C12C78D1F0EFFC77538FED4FB06FEAC" box="[1060,1197,296,319]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="tanaitica">
|
||
<emphasis id="B966601C1F0EFFC77538FED4FB06FEAC" box="[1060,1197,296,319]" italics="true" pageId="54" pageNumber="55">C. tanaitica</emphasis>
|
||
</taxonomicName>
|
||
, which is its closest relative. It is also supported by the PTP approach as distinct entity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |