treatments-xml/data/03/B5/C9/03B5C9112C4EFFCD90C4D8AEFE8D28E2.xml
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<document id="D3AE58B4AA336F3876F8BFF70B7C399D" ID-DOI="10.11646/zootaxa.4743.3.9" ID-GBIF-Dataset="b2dbeccc-0f3e-424a-882b-85f810bf6301" ID-ISSN="1175-5326" ID-Zenodo-Dep="3688346" ID-ZooBank="C4B81BE0-7A4C-4EC4-8DDE-AAA565B2ED00" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582721007500" checkinUser="plazi" docAuthor="Hendrich, Lars &amp; Balke, Michael" docDate="2020" docId="03B5C9112C4EFFCD90C4D8AEFE8D28E2" docLanguage="en" docName="zootaxa.4743.3.9.pdf" docOrigin="Zootaxa 4743 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydaticus (Prodaticus) hauthi Hendrich &amp; Balke 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="421" masterDocId="FF8CB1692C4FFFCF9053D900FF892969" masterDocTitle="Hydaticus (Prodaticus) hauthi sp. nov., a new diving beetle from the cloud forest in the Cordillera El Sira, Peru (Coleoptera: Dytiscidae)" masterLastPageNumber="426" masterPageNumber="419" pageNumber="420" updateTime="1698722592272" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="6D36186C770878BFA06D73D8142E0189">Hydaticus (Prodaticus) hauthi sp. nov., a new diving beetle from the cloud forest in the Cordillera El Sira, Peru (Coleoptera: Dytiscidae)</mods:title>
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<mods:name id="928906672E1579A69400BA17FA7B5C05" type="personal">
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<mods:namePart id="17B9962CA1280FA413DBF75BFE615B28">Hendrich, Lars</mods:namePart>
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<mods:namePart id="979FBF63202D0673C3954F89FF43D2B1">Balke, Michael</mods:namePart>
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<mods:typeOfResource id="A682F06DA4670CF17FAB04B4DCD67486">text</mods:typeOfResource>
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<mods:title id="ED34C3D67096D2825BF7EB3719852B77">Zootaxa</mods:title>
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<mods:part id="7F783310D4C21B1EB4653E033E53B9D9">
<mods:date id="B5559302D19F12A758729C0A71834833">2020</mods:date>
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<mods:number id="C51ACC76A9D27CE8E3FCE88B2EF684F8">2020-02-26</mods:number>
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<mods:number id="0BDC969BCC59873DF7AD97E110885713">4743</mods:number>
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<treatment id="03B5C9112C4EFFCD90C4D8AEFE8D28E2" ID-DOI="http://doi.org/10.5281/zenodo.3691604" ID-GBIF-Taxon="162268587" ID-Zenodo-Dep="3691604" LSID="urn:lsid:plazi:treatment:03B5C9112C4EFFCD90C4D8AEFE8D28E2" httpUri="http://treatment.plazi.org/id/03B5C9112C4EFFCD90C4D8AEFE8D28E2" lastPageId="2" lastPageNumber="421" pageId="1" pageNumber="420">
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<paragraph id="8BA378072C4EFFCE90C4D8AEFDE228A0" blockId="1.[151,619,430,493]" box="[151,619,430,457]" pageId="1" pageNumber="420">
<heading id="D0EBCF6B2C4EFFCE90C4D8AEFDE228A0" bold="true" box="[151,619,430,457]" fontSize="11" level="1" pageId="1" pageNumber="420" reason="1">
<emphasis id="B968A4152C4EFFCE90C4D8AEFDE228A0" bold="true" box="[151,619,430,457]" pageId="1" pageNumber="420">
<taxonomicName id="4C1C03842C4EFFCE90C4D8AEFD8F28A0" ID-CoL="8LZF9" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[151,518,430,457]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="hauthi" status="sp. nov." subGenus="Prodaticus">
<emphasis id="B968A4152C4EFFCE90C4D8AEFE9828A0" bold="true" box="[151,273,430,457]" italics="true" pageId="1" pageNumber="420">Hydaticus</emphasis>
(
<emphasis id="B968A4152C4EFFCE9170D8AEFE2C28A0" bold="true" box="[291,421,430,457]" italics="true" pageId="1" pageNumber="420">Prodaticus</emphasis>
)
<emphasis id="B968A4152C4EFFCE91E5D8AEFD8F28A0" bold="true" box="[438,518,430,457]" italics="true" pageId="1" pageNumber="420">hauthi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B196E2C4EFFCE925ED8AFFDE228A0" box="[525,619,431,457]" pageId="1" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3062B8C2C4EFFCE90C4D8D3FEE52884" box="[151,364,466,493]" pageId="1" pageNumber="420" type="description">
<paragraph id="8BA378072C4EFFCE90C4D8D3FEE52884" blockId="1.[151,619,430,493]" box="[151,364,466,493]" pageId="1" pageNumber="420">
(
<figureCitation id="132764822C4EFFCE90CCD8D3FF7A2884" box="[159,243,467,493]" captionStart="FIGURE 1" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[278,1310,179,1918]" captionTargetId="figure@4.[278,1310,179,1918]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of H. hauthi sp. nov., holotype, male (a); H. lateralis (b) and H. panguana (c), both from Panguana Research Station, Huanuco Department, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3688348" httpUri="https://zenodo.org/record/3688348/files/figure.png" pageId="1" pageNumber="420">Figs 1a</figureCitation>
,
<figureCitation id="132764822C4EFFCE9153D8D3FE992884" box="[256,272,467,493]" captionStart="FIGURES 23" captionStartId="5.[151,264,1939,1964]" captionTargetBox="[185,1403,181,1920]" captionTargetId="figure@5.[185,1403,181,1920]" captionTargetPageId="5" captionText="FIGURES 23. Median lobe of aedeagus in dorsal (a) and lateral view (b): H. hauthi sp. nov. (2), H. lateralis (3) and H. panguana (4). The anterior margin of the membranous lateral “wings” of the median lobe in 3b and 4 b have been enhanced digitally for more contrast and better visibility. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3688350" httpUri="https://zenodo.org/record/3688350/files/figure.png" pageId="1" pageNumber="420">2</figureCitation>
,
<figureCitation id="132764822C4EFFCE914FD8D2FEC12884" box="[284,328,466,493]" captionStart="FIGURES 56" captionStartId="6.[151,264,1946,1971]" captionTargetBox="[183,1404,519,1920]" captionTargetId="figure@6.[183,1404,181,1920]" captionTargetPageId="6" captionText="FIGURES 56. Cloud forest at 1500 m in the Cordillera El Sira, Peru (5) and habitat of Hydaticus hauthi sp. nov. in Peru (6) (Photos by Armin Niessner)." figureDoi="http://doi.org/10.5281/zenodo.3688352" httpUri="https://zenodo.org/record/3688352/files/figure.png" pageId="1" pageNumber="420">5, 6</figureCitation>
,
<figureCitation id="132764822C4EFFCE9107D8D3FEEA2884" box="[340,355,467,493]" captionStart="FIGURE 7" captionStartId="7.[151,250,1268,1293]" captionTargetId="figure@7.[319,1267,198,1239]" captionTargetPageId="7" captionText="FIGURE 7. Map, type locality of H. hauthi sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.3689101" httpUri="https://zenodo.org/record/3689101/files/figure.png" pageId="1" pageNumber="420">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3062B8C2C4EFFCE90C4DB1BFCBD2BAC" pageId="1" pageNumber="420" type="materials_examined">
<paragraph id="8BA378072C4EFFCE90C4DB1BFB6C2B5C" blockId="1.[151,1437,538,2041]" box="[151,1253,538,565]" pageId="1" pageNumber="420">
<emphasis id="B968A4152C4EFFCE90C4DB1BFEBF2B5C" bold="true" box="[151,310,538,565]" pageId="1" pageNumber="420">Type locality.</emphasis>
<materialsCitation id="3B74725A2C4EFFCE916FDB1BFB682B5C" ID-GBIF-Occurrence="2571181075" box="[316,1249,538,565]" country="Peru" elevation="1550" latitude="-9.426126" location="Cerros del Sira" longLatPrecision="1" longitude="-74.73566" municipality="Department" pageId="1" pageNumber="420" specimenCount="1" stateProvince="Huanuco">
<collectingCountry id="F30B38972C4EFFCE916FDB1BFEFB2B5C" box="[316,370,539,565]" name="Peru" pageId="1" pageNumber="420">Peru</collectingCountry>
,
<collectingRegion id="49D8B6E52C4EFFCE912DDB1BFE6A2B5C" box="[382,483,539,565]" country="Peru" name="Huanuco" pageId="1" pageNumber="420">Huanuco</collectingRegion>
<collectingMunicipality id="6BC7E27D2C4EFFCE91B9DB1BFDF82B5C" box="[490,625,539,565]" pageId="1" pageNumber="420">Department</collectingMunicipality>
,
<location id="8EC32EDC2C4EFFCE922FDB1BFCAE2B5C" LSID="urn:lsid:plazi:treatment:03B5C9112C4EFFCD90C4D8AEFE8D28E2:8EC32EDC2C4EFFCE922FDB1BFCAE2B5C" box="[636,807,539,565]" country="Peru" latitude="-9.426126" longLatPrecision="1" longitude="-74.73566" municipality="Department" name="Cerros del Sira" pageId="1" pageNumber="420" stateProvince="Huanuco">Cerros del Sira</location>
,
<quantity id="4CE4D5E22C4EFFCE9361DB1AFC192B5C" box="[818,912,538,565]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="420" unit="m" value="1550.0">
<elevation id="00319F342C4EFFCE9361DB1AFC192B5C" box="[818,912,538,565]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="420" unit="m" value="1550.0">1,550 m</elevation>
</quantity>
,
<geoCoordinate id="EE281EC02C4EFFCE93CFDB1AFBBC2B5C" box="[924,1077,538,565]" degrees="9" direction="south" minutes="25" orientation="latitude" pageId="1" pageNumber="420" precision="1" seconds="34.05" value="-9.426126">9°2534.05”S</geoCoordinate>
<geoCoordinate id="EE281EC02C4EFFCE946FDB1BFB682B5C" box="[1084,1249,539,565]" degrees="74" direction="west" minutes="44" orientation="longitude" pageId="1" pageNumber="420" precision="1" seconds="8.38" value="-74.73566">74°448.38”W</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA378072C4EFFCE9094DB3FFCBD2BAC" blockId="1.[151,1437,538,2041]" pageId="1" pageNumber="420">
<materialsCitation id="3B74725A2C4EFFCE9094DB3FFEEB2BC8" ID-GBIF-Occurrence="2571181076" collectingDate="2012-06-10" collectionCode="MHN-UNMSM" country="Peru" determinerName="Balke" elevation="1550" latitude="-9.426126" location="Cerros del Sira" longLatPrecision="1" longitude="-74.73566" pageId="1" pageNumber="420" specimenCount="2" stateProvince="Huanuco" typeStatus="holotype">
<emphasis id="B968A4152C4EFFCE9094DB3FFEF12B30" bold="true" box="[199,376,575,601]" pageId="1" pageNumber="420">
<typeStatus id="54A7C6A52C4EFFCE9094DB3FFEBF2B30" box="[199,310,575,601]" pageId="1" pageNumber="420" type="holotype">Holotype</typeStatus>
,male
</emphasis>
(
<collectionCode id="ED0DE0C22C4EFFCE91D1DB3FFDB72B30" box="[386,574,575,601]" pageId="1" pageNumber="420">MHN-UNMSM</collectionCode>
).“
<collectingCountry id="F30B38972C4EFFCE9205DB3FFD2A2B30" box="[598,675,575,601]" name="Peru" pageId="1" pageNumber="420">PERU</collectingCountry>
,
<collectingRegion id="49D8B6E52C4EFFCE92F9DB3FFC862B30" box="[682,783,575,601]" country="Peru" name="Huanuco" pageId="1" pageNumber="420">Huanuco</collectingRegion>
Dep.,
<location id="8EC32EDC2C4EFFCE9301DB3FFC7C2B30" LSID="urn:lsid:plazi:treatment:03B5C9112C4EFFCD90C4D8AEFE8D28E2:8EC32EDC2C4EFFCE9301DB3FFC7C2B30" box="[850,1013,575,601]" country="Peru" latitude="-9.426126" longLatPrecision="1" longitude="-74.73566" name="Cerros del Sira" pageId="1" pageNumber="420" stateProvince="Huanuco">Cerros del Sira</location>
,
<quantity id="4CE4D5E22C4EFFCE93AFDB3EFBC62B30" box="[1020,1103,574,601]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="420" unit="m" value="1550.0">
<elevation id="00319F342C4EFFCE93AFDB3EFBC62B30" box="[1020,1103,574,601]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="420" unit="m" value="1550.0">1550m</elevation>
</quantity>
,
<geoCoordinate id="EE281EC02C4EFFCE9404DB3EFB782B30" box="[1111,1265,574,601]" degrees="9" direction="south" minutes="25" orientation="latitude" pageId="1" pageNumber="420" precision="1" seconds="34.05" value="-9.426126">9°2534.05”S</geoCoordinate>
<geoCoordinate id="EE281EC02C4EFFCE94A0DB3FFA112B30" box="[1267,1432,575,601]" degrees="74" direction="west" minutes="44" orientation="longitude" pageId="1" pageNumber="420" precision="1" seconds="8.38" value="-74.73566">74°448.38”W</geoCoordinate>
,
<date id="FFA25EC72C4EFFCE90C4DB63FE832B14" box="[151,266,611,637]" pageId="1" pageNumber="420" value="2012-06-10">
<collectingDate id="EFE6A72F2C4EFFCE90C4DB63FE832B14" box="[151,266,611,637]" pageId="1" pageNumber="420" value="2012-06-10">6.10.2012</collectingDate>
</date>
, local collector”, “
<typeStatus id="54A7C6A52C4EFFCE9189DB63FDCC2B14" box="[474,581,611,637]" pageId="1" pageNumber="420" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C1C03842C4EFFCE921EDB63FC872B14" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[589,782,611,637]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="hauthi" status="sp. nov.">
<emphasis id="B968A4152C4EFFCE921EDB63FC872B14" box="[589,782,611,637]" italics="true" pageId="1" pageNumber="420">Hydaticus hauthi</emphasis>
</taxonomicName>
<emphasis id="B968A4152C4EFFCE9346DB63FCE62B14" bold="true" box="[789,879,611,637]" pageId="1" pageNumber="420">
<taxonomicNameLabel id="A25B196E2C4EFFCE9346DB63FCE62B14" box="[789,879,611,637]" pageId="1" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Hendrich &amp;
<determinerName id="7E7F94C32C4EFFCE9458DB63FBC52B14" box="[1035,1100,611,637]" pageId="1" pageNumber="420">Balke</determinerName>
det. 2019” [red printed label] (
<collectionCode id="ED0DE0C22C4EFFCE90CCDB87FED22BC8" box="[159,347,647,673]" pageId="1" pageNumber="420">MHN-UNMSM</collectionCode>
)
</materialsCitation>
.
<emphasis id="B968A4152C4EFFCE913FDB87FD052BC8" bold="true" box="[364,652,646,673]" pageId="1" pageNumber="420">
<typeStatus id="54A7C6A52C4EFFCE913FDB87FE6D2BC8" box="[364,484,647,673]" pageId="1" pageNumber="420" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D1AB38E2C4EFFCE91A7DB86FD0C2BC8" box="[500,645,646,673]" pageId="1" pageNumber="420" type="generic">5 specimens</specimenCount>
)
</emphasis>
:
<materialsCitation id="3B74725A2C4EFFCE92C8DB87FCB92BAC" ID-GBIF-Occurrence="2571181074" collectionCode="ZSM" country="Peru" pageId="1" pageNumber="420" specimenCount="4" specimenCount-male="4" stateProvince="Huanuco" typeStatus="holotype">
<specimenCount id="9D1AB38E2C4EFFCE92C8DB87FD782BC8" box="[667,753,647,673]" pageId="1" pageNumber="420" type="male">4 males</specimenCount>
and one female with the same label data as
<typeStatus id="54A7C6A52C4EFFCE94B3DB87FACB2BC8" box="[1248,1346,647,673]" pageId="1" pageNumber="420" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED0DE0C22C4EFFCE9502DB87FA192BC8" box="[1361,1424,647,673]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="420" type="Museum">ZSM</collectionCode>
). Each
<typeStatus id="54A7C6A52C4EFFCE9085DBABFEBF2BAC" box="[214,310,683,709]" pageId="1" pageNumber="420" type="paratype">paratype</typeStatus>
is provided with a red printed
<typeStatus id="54A7C6A52C4EFFCE92DCDBABFD662BAC" box="[655,751,683,709]" pageId="1" pageNumber="420" type="paratype">paratype</typeStatus>
label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3062B8C2C4EFFCE9094DBCEFE312F94" pageId="1" pageNumber="420" type="description">
<paragraph id="8BA378072C4EFFCE9094DBCEFE2F2A58" blockId="1.[151,1437,538,2041]" pageId="1" pageNumber="420">
<emphasis id="B968A4152C4EFFCE9094DBCEFE6E2B80" bold="true" box="[199,487,718,745]" pageId="1" pageNumber="420">
Description of
<typeStatus id="54A7C6A52C4EFFCE912ADBCFFE6B2B80" box="[377,482,719,745]" pageId="1" pageNumber="420" type="holotype">holotype</typeStatus>
.
</emphasis>
TL =
<quantity id="4CE4D5E22C4EFFCE9263DBCFFD112B80" box="[560,664,719,745]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="1" pageNumber="420" unit="mm" value="14.1">14.1 mm</quantity>
; TL-H =
<quantity id="4CE4D5E22C4EFFCE9350DBCEFCE22B80" box="[771,875,718,745]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="1" pageNumber="420" unit="mm" value="13.5">13.5 mm</quantity>
; MW =
<quantity id="4CE4D5E22C4EFFCE9398DBCEFBAC2B80" box="[971,1061,718,745]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="420" unit="mm" value="7.5">7.5 mm</quantity>
. Body oval, elongate, slightly attenuated posteriorly, dorsoventrally depressed; subparallel medially, widest just behind middle, lateral outline continuous in dorsal aspect.
</paragraph>
<paragraph id="8BA378072C4EFFCE9094DA3AFEE82AA8" blockId="1.[151,1437,538,2041]" pageId="1" pageNumber="420">
<emphasis id="B968A4152C4EFFCE9094DA3AFEA92A3C" bold="true" box="[199,288,826,853]" pageId="1" pageNumber="420">Colour.</emphasis>
Dorsal surface predominantly black. Head capsule black with clypeus and frons slightly testaceous (
<figureCitation id="132764822C4EFFCE90F3DA5FFF792A10" box="[160,240,863,889]" captionStart="FIGURE 1" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[278,1310,179,1918]" captionTargetId="figure@4.[278,1310,179,1918]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of H. hauthi sp. nov., holotype, male (a); H. lateralis (b) and H. panguana (c), both from Panguana Research Station, Huanuco Department, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3688348" httpUri="https://zenodo.org/record/3688348/files/figure.png" pageId="1" pageNumber="420">Fig. 1a</figureCitation>
). Pronotum black with lateral margins testaceous. Elytron black, lateral margin of apex sparsely irrorated. Ventral surface predominantly black, antenna, prosternum, mouthparts and fore legs testaceous, middle and hind legs almost black.
</paragraph>
<paragraph id="8BA378072C4EFFCE9094DACAFE702CB4" blockId="1.[151,1437,538,2041]" pageId="1" pageNumber="420">
<emphasis id="B968A4152C4EFFCE9094DACAFE652A8C" bold="true" box="[199,492,970,997]" pageId="1" pageNumber="420">Structure and sculpture.</emphasis>
Head with fine dense punctation and scattered stronger punctures; additionally with strong punctures concentrated in two spots in anterior third, and with longitudinal rows of strong punctures along postero-medial margins of eyes. Pronotum with posterolateral angle produced and acute, posterior margin broadly sinuate and recurved lateral to basomedial lobe. Pronotum with fine, dense punctation, and row of strong punctures along anterior margin; scattered strong punctures and traces of wrinkles near lateral margins, and row of stronger punctures near posterior margin on both sides of middle. Elytron with very fine microreticulation and scattered stronger punctures. Two rows of well-developed and deep punctures visible. Ventral surface with fine microreticulation; metacoxae and ventrites with fine scattered punctation and with stronger punctures along margin of last ventrite. Prosternal process almost flat. Metaventral wing relatively broad medially, narrow laterally with anterolateral margin straight. Dorsal surface of protarsomeres I and II with strong and dense reticulations and a series of spines. Protibia with stout pegs on apicodorsal surface. Mesotarsomeres ventrally with a small basal brush of setae. Posterior surface of metafemur with few large punctures on anterior surface and without interspersed small punctures. Metatibia without punctures on anterior surface; posterior surface with dorsal series of seven bifid setae curved ventro-basally, not parallel to dorsal margin; metatibial spurs acuminate apically. Metatarsal claws unequal, anterior claw shorter than posterior one.
</paragraph>
<paragraph id="8BA378072C4EFFCE9094DCE7FDCF2F20" blockId="1.[151,1437,538,2041]" pageId="1" pageNumber="420">
<emphasis id="B968A4152C4EFFCE9094DCE7FEF32F68" bold="true" box="[199,378,1510,1537]" pageId="1" pageNumber="420">Male genitalia.</emphasis>
Median lobe of aedeagus narrow in ventral view; in lateral view enlarged in apical third and with apex rounded (
<figureCitation id="132764822C4EFFCE9124DF0BFE342F4C" box="[375,445,1547,1573]" captionStart="FIGURES 23" captionStartId="5.[151,264,1939,1964]" captionTargetBox="[185,1403,181,1920]" captionTargetId="figure@5.[185,1403,181,1920]" captionTargetPageId="5" captionText="FIGURES 23. Median lobe of aedeagus in dorsal (a) and lateral view (b): H. hauthi sp. nov. (2), H. lateralis (3) and H. panguana (4). The anterior margin of the membranous lateral “wings” of the median lobe in 3b and 4 b have been enhanced digitally for more contrast and better visibility. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3688350" httpUri="https://zenodo.org/record/3688350/files/figure.png" pageId="1" pageNumber="420">Fig. 2</figureCitation>
). Parameres are not illustrated because by no means useful in identification of most species of
<taxonomicName id="4C1C03842C4EFFCE90BBDF2FFDCB2F20" authority="(Megna et al. 2019)" baseAuthorityName="Megna" baseAuthorityYear="2019" box="[232,578,1583,1609]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="genus">
<emphasis id="B968A4152C4EFFCE90BBDF2FFED32F20" box="[232,346,1583,1609]" italics="true" pageId="1" pageNumber="420">Hydaticus</emphasis>
(
<bibRefCitation id="EF8D05F62C4EFFCE913BDF2FFDB22F20" author="Megna, Y. S. &amp; Balke, M. &amp; Apenborn, R. &amp; Hendrich, L." box="[360,571,1583,1609]" pageId="1" pageNumber="420" pagination="113 - 130" refId="ref3615" refString="Megna, Y. S., Balke, M., Apenborn, R. &amp; Hendrich, L. (2019) A review of Peruvian diving beetles of the genus Hydaticus Leach, 1817, with description of Hydaticus (Prodaticus) panguana sp. nov. and notes on other Neotropical species (Coleoptera: Dytiscidae). Zootaxa, 4615 (1), 113 - 130. https: // doi. org / 10.11646 / zootaxa. 4615.1.5" type="journal article" year="2019">
Megna
<emphasis id="B968A4152C4EFFCE91ECDF30FE7E2F20" box="[447,503,1583,1609]" italics="true" pageId="1" pageNumber="420">et al.</emphasis>
2019
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BA378072C4EFFCE9094DF52FE312F94" blockId="1.[151,1437,538,2041]" pageId="1" pageNumber="420">
<emphasis id="B968A4152C4EFFCE9094DF52FE312F04" bold="true" box="[199,440,1618,1645]" pageId="1" pageNumber="420">Sexual dimorphism.</emphasis>
Male with protarsomeres I-III enlarged to form a large oval palette ventrally bearing 22 rounded suckers of different sizes; male protarsomeres I and II with a series of spines across dorsal surface; mesotarsomeres IIII moderately enlarged, ventrally with 14 rounded suckers. Wrinkles on pronotum similar to those in males. Female with TL =
<quantity id="4CE4D5E22C4EFFCE91E7DFBFFD932FB0" box="[436,538,1727,1753]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="1" pageNumber="420" unit="mm" value="13.6">13.6 mm</quantity>
; TL-H =
<quantity id="4CE4D5E22C4EFFCE92D2DFBFFD6E2FB0" box="[641,743,1727,1753]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="1" pageNumber="420" unit="mm" value="12.9">12.9 mm</quantity>
; MW =
<quantity id="4CE4D5E22C4EFFCE9312DFBFFC102FB0" box="[833,921,1727,1753]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="420" unit="mm" value="7.4">7.4 mm</quantity>
; metatarsomeres of female with natatory setae on posteroventral margin.
</paragraph>
</subSubSection>
<subSubSection id="C3062B8C2C4EFFCE9094DE06FE4A2E90" pageId="1" pageNumber="420" type="diagnosis">
<paragraph id="8BA378072C4EFFCE9094DE06FE4A2E90" blockId="1.[151,1437,538,2041]" pageId="1" pageNumber="420">
<emphasis id="B968A4152C4EFFCE9094DE06FE472E48" bold="true" box="[199,462,1798,1825]" pageId="1" pageNumber="420">Differential diagnosis.</emphasis>
The habitus of
<taxonomicName id="4C1C03842C4EFFCE922FDE07FCB42E48" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[636,829,1799,1825]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="hauthi" status="sp. nov.">
<emphasis id="B968A4152C4EFFCE922FDE07FCB42E48" box="[636,829,1799,1825]" italics="true" pageId="1" pageNumber="420">Hydaticus hauthi</emphasis>
</taxonomicName>
<emphasis id="B968A4152C4EFFCE9310DE07FC142E48" bold="true" box="[835,925,1799,1825]" pageId="1" pageNumber="420">
<taxonomicNameLabel id="A25B196E2C4EFFCE9310DE07FC142E48" box="[835,925,1799,1825]" pageId="1" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is near to that of
<taxonomicName id="4C1C03842C4EFFCE940CDE08FE562E2C" authority="Megna, Balke, Apenborn &amp; Hendrich, 2019" authorityName="Megna, Balke, Apenborn &amp; Hendrich" authorityYear="2019" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="panguana">
<emphasis id="B968A4152C4EFFCE940CDE08FB782E48" box="[1119,1265,1800,1825]" italics="true" pageId="1" pageNumber="420">H. panguana</emphasis>
<bibRefCitation id="EF8D05F62C4EFFCE94ABDE07FE562E2C" author="Megna, Y. S. &amp; Balke, M. &amp; Apenborn, R. &amp; Hendrich, L." pageId="1" pageNumber="420" pagination="113 - 130" refId="ref3615" refString="Megna, Y. S., Balke, M., Apenborn, R. &amp; Hendrich, L. (2019) A review of Peruvian diving beetles of the genus Hydaticus Leach, 1817, with description of Hydaticus (Prodaticus) panguana sp. nov. and notes on other Neotropical species (Coleoptera: Dytiscidae). Zootaxa, 4615 (1), 113 - 130. https: // doi. org / 10.11646 / zootaxa. 4615.1.5" type="journal article" year="2019">Megna, Balke, Apenborn &amp; Hendrich, 2019</bibRefCitation>
</taxonomicName>
(
<figureCitation id="132764822C4EFFCE91A3DE2BFDCB2E2C" box="[496,578,1835,1861]" captionStart="FIGURE 1" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[278,1310,179,1918]" captionTargetId="figure@4.[278,1310,179,1918]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of H. hauthi sp. nov., holotype, male (a); H. lateralis (b) and H. panguana (c), both from Panguana Research Station, Huanuco Department, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3688348" httpUri="https://zenodo.org/record/3688348/files/figure.png" pageId="1" pageNumber="420">Fig. 1c</figureCitation>
), but both species can be separated by their different size (TL =
<quantity id="4CE4D5E22C4EFFCE957FDE2BFF4A2E00" metricMagnitude="-2" metricUnit="m" metricValue="1.41" metricValueMax="1.46" metricValueMin="1.3599999999999999" pageId="1" pageNumber="420" unit="mm" value="14.1" valueMax="14.6" valueMin="13.6">13.614.6 mm</quantity>
in
<taxonomicName id="4C1C03842C4EFFCE90BBDE50FEDD2E00" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[232,340,1871,1897]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="hauthi" status="sp. nov.">
<emphasis id="B968A4152C4EFFCE90BBDE50FEDD2E00" box="[232,340,1871,1897]" italics="true" pageId="1" pageNumber="420">H. hauthi</emphasis>
</taxonomicName>
<emphasis id="B968A4152C4EFFCE9108DE4FFE3F2E00" bold="true" box="[347,438,1871,1897]" pageId="1" pageNumber="420">
<taxonomicNameLabel id="A25B196E2C4EFFCE9108DE4FFE3F2E00" box="[347,438,1871,1897]" pageId="1" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and TL = 13.014.0 mm in
<taxonomicName id="4C1C03842C4EFFCE92A4DE50FC022E00" authorityName="Megna, Balke, Apenborn &amp; Hendrich" authorityYear="2019" box="[759,907,1872,1897]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="panguana">
<emphasis id="B968A4152C4EFFCE92A4DE50FC022E00" box="[759,907,1872,1897]" italics="true" pageId="1" pageNumber="420">H. panguana</emphasis>
</taxonomicName>
), colour patterns of the elytra (
<figureCitation id="132764822C4EFFCE94BFDE4FFAD42E00" box="[1260,1373,1871,1897]" captionStart="FIGURE 1" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[278,1310,179,1918]" captionTargetId="figure@4.[278,1310,179,1918]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of H. hauthi sp. nov., holotype, male (a); H. lateralis (b) and H. panguana (c), both from Panguana Research Station, Huanuco Department, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3688348" httpUri="https://zenodo.org/record/3688348/files/figure.png" pageId="1" pageNumber="420">Figs 1a, c</figureCitation>
), and length and shape of the median lobes (
<figureCitation id="132764822C4EFFCE9216DE73FD072EE4" box="[581,654,1907,1933]" captionStart="FIGURES 23" captionStartId="5.[151,264,1939,1964]" captionTargetBox="[185,1403,181,1920]" captionTargetId="figure@5.[185,1403,181,1920]" captionTargetPageId="5" captionText="FIGURES 23. Median lobe of aedeagus in dorsal (a) and lateral view (b): H. hauthi sp. nov. (2), H. lateralis (3) and H. panguana (4). The anterior margin of the membranous lateral “wings” of the median lobe in 3b and 4 b have been enhanced digitally for more contrast and better visibility. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3688350" httpUri="https://zenodo.org/record/3688350/files/figure.png" pageId="1" pageNumber="420">Figs 2</figureCitation>
, 4). The other larger species in
<collectingCountry id="F30B38972C4EFFCE93BCDE73FBAC2EE4" box="[1007,1061,1907,1933]" name="Peru" pageId="1" pageNumber="420">Peru</collectingCountry>
,
<taxonomicName id="4C1C03842C4EFFCE9463DE74FADD2EE4" authority="Laporte, 1835" authorityName="Laporte" authorityYear="1835" box="[1072,1364,1906,1933]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B968A4152C4EFFCE9463DE74FB262EE4" box="[1072,1199,1907,1933]" italics="true" pageId="1" pageNumber="420">H. lateralis</emphasis>
Laporte, 1835
</taxonomicName>
(TL =
<quantity id="4CE4D5E22C4EFFCE90C4DE97FEB42ED8" box="[151,317,1943,1969]" metricMagnitude="-2" metricUnit="m" metricValue="1.335" metricValueMax="1.38" metricValueMin="1.29" pageId="1" pageNumber="420" unit="mm" value="13.35" valueMax="13.8" valueMin="12.9">12.913.8 mm</quantity>
), has yellow longitudinal stripes and a subtriangular, postmedial and broad spot on each elytron (
<figureCitation id="132764822C4EFFCE953CDE97FF3D2EBF" captionStart="FIGURE 1" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[278,1310,179,1918]" captionTargetId="figure@4.[278,1310,179,1918]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of H. hauthi sp. nov., holotype, male (a); H. lateralis (b) and H. panguana (c), both from Panguana Research Station, Huanuco Department, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3688348" httpUri="https://zenodo.org/record/3688348/files/figure.png" pageId="1" pageNumber="420">Fig. 1b</figureCitation>
). Furthermore, the median lobe of
<taxonomicName id="4C1C03842C4EFFCE9268DEBCFD332EBC" authorityName="Laporte" authorityYear="1835" box="[571,698,1979,2005]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B968A4152C4EFFCE9268DEBCFD332EBC" box="[571,698,1979,2005]" italics="true" pageId="1" pageNumber="420">H. lateralis</emphasis>
</taxonomicName>
is much smaller and has a different shape than the median lobe of
<taxonomicName id="4C1C03842C4EFFCE90C4DEE0FE882E90" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[151,257,2015,2041]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="hauthi" status="sp. nov.">
<emphasis id="B968A4152C4EFFCE90C4DEE0FE882E90" box="[151,257,2015,2041]" italics="true" pageId="1" pageNumber="420">H. hauthi</emphasis>
</taxonomicName>
<emphasis id="B968A4152C4EFFCE915BDEDFFEEB2E90" bold="true" box="[264,354,2015,2041]" pageId="1" pageNumber="420">
<taxonomicNameLabel id="A25B196E2C4EFFCE915BDEDFFEEB2E90" box="[264,354,2015,2041]" pageId="1" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132764822C4EFFCE9122DEDFFE3C2E90" box="[369,437,2015,2041]" captionStart="FIGURES 23" captionStartId="5.[151,264,1939,1964]" captionTargetBox="[185,1403,181,1920]" captionTargetId="figure@5.[185,1403,181,1920]" captionTargetPageId="5" captionText="FIGURES 23. Median lobe of aedeagus in dorsal (a) and lateral view (b): H. hauthi sp. nov. (2), H. lateralis (3) and H. panguana (4). The anterior margin of the membranous lateral “wings” of the median lobe in 3b and 4 b have been enhanced digitally for more contrast and better visibility. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3688350" httpUri="https://zenodo.org/record/3688350/files/figure.png" pageId="1" pageNumber="420">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3062B8C2C4DFFCD9094D999FAD829BE" pageId="2" pageNumber="421" type="etymology">
<paragraph id="8BA378072C4DFFCD9094D999FAD829BE" blockId="2.[151,1436,153,395]" pageId="2" pageNumber="421">
<emphasis id="B968A4152C4DFFCD9094D999FEC729DA" bold="true" box="[199,334,153,179]" pageId="2" pageNumber="421">Etymology.</emphasis>
The new species is dedicated to our young and keen colleague David Hauth (Marburg,
<collectingCountry id="F30B38972C4DFFCD9577D999FA1829DA" box="[1316,1425,153,179]" name="Germany" pageId="2" pageNumber="421">Germany</collectingCountry>
), specialist for different xylophagous beetle families. The specific epithet is a substantive in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3062B8C2C4DFFCD9094D9E0FE9B2849" pageId="2" pageNumber="421" type="distribution">
<paragraph id="8BA378072C4DFFCD9094D9E0FE9B2849" blockId="2.[151,1436,153,395]" pageId="2" pageNumber="421">
<emphasis id="B968A4152C4DFFCD9094D9E0FEE82992" bold="true" box="[199,353,224,251]" pageId="2" pageNumber="421">Distribution.</emphasis>
So far
<taxonomicName id="4C1C03842C4DFFCD91E6D9E1FDFE2992" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[437,631,225,251]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="hauthi" status="sp. nov.">
<emphasis id="B968A4152C4DFFCD91E6D9E1FDFE2992" box="[437,631,225,251]" italics="true" pageId="2" pageNumber="421">Hydaticus hauthi</emphasis>
</taxonomicName>
<emphasis id="B968A4152C4DFFCD922DD9E1FD502992" bold="true" box="[638,729,225,251]" pageId="2" pageNumber="421">
<taxonomicNameLabel id="A25B196E2C4DFFCD922DD9E1FD502992" box="[638,729,225,251]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from the
<typeStatus id="54A7C6A52C4DFFCD93A1D9E1FBAB2992" box="[1010,1058,225,251]" pageId="2" pageNumber="421">type</typeStatus>
locality in the El Sira Mountains (
<figureCitation id="132764822C4DFFCD90CCD805FE8D2849" box="[159,260,260,288]" captionStart="FIGURES 56" captionStartId="6.[151,264,1946,1971]" captionTargetBox="[183,1404,519,1920]" captionTargetId="figure@6.[183,1404,181,1920]" captionTargetPageId="6" captionText="FIGURES 56. Cloud forest at 1500 m in the Cordillera El Sira, Peru (5) and habitat of Hydaticus hauthi sp. nov. in Peru (6) (Photos by Armin Niessner)." figureDoi="http://doi.org/10.5281/zenodo.3688352" httpUri="https://zenodo.org/record/3688352/files/figure.png" pageId="2" pageNumber="421">Figs 5, 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3062B8C2C4DFFCD9094D829FE8D28E2" pageId="2" pageNumber="421" type="biology_ecology">
<paragraph id="8BA378072C4DFFCD9094D829FE8D28E2" blockId="2.[151,1436,153,395]" pageId="2" pageNumber="421">
<emphasis id="B968A4152C4DFFCD9094D829FEA5282A" bold="true" box="[199,300,297,323]" pageId="2" pageNumber="421">Ecology.</emphasis>
All specimens of
<taxonomicName id="4C1C03842C4DFFCD9251D829FD4F282A" authority="Hendrich &amp; Balke, 2020" authorityName="Hendrich &amp; Balke" authorityYear="2020" box="[514,710,297,323]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="hauthi" status="sp. nov.">
<emphasis id="B968A4152C4DFFCD9251D829FD4F282A" box="[514,710,297,323]" italics="true" pageId="2" pageNumber="421">Hydaticus hauthi</emphasis>
</taxonomicName>
<emphasis id="B968A4152C4DFFCD929CD829FCA2282A" bold="true" box="[719,811,297,323]" pageId="2" pageNumber="421">
<taxonomicNameLabel id="A25B196E2C4DFFCD929CD829FCA2282A" box="[719,811,297,323]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected in a partly shaded, shallow, temporary forest pool, rich in decaying leaves (
<figureCitation id="132764822C4DFFCD926AD84DFDF62801" box="[569,639,333,360]" captionStart="FIGURES 56" captionStartId="6.[151,264,1946,1971]" captionTargetBox="[183,1404,519,1920]" captionTargetId="figure@6.[183,1404,181,1920]" captionTargetPageId="6" captionText="FIGURES 56. Cloud forest at 1500 m in the Cordillera El Sira, Peru (5) and habitat of Hydaticus hauthi sp. nov. in Peru (6) (Photos by Armin Niessner)." figureDoi="http://doi.org/10.5281/zenodo.3688352" httpUri="https://zenodo.org/record/3688352/files/figure.png" pageId="2" pageNumber="421">Fig. 6</figureCitation>
). No other
<taxonomicName id="4C1C03842C4DFFCD9350D84DFCF2280E" box="[771,891,333,359]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
were collected, just a few
<taxonomicName id="4C1C03842C4DFFCD94E2D84DFAA8280E" box="[1201,1313,333,359]" class="Insecta" family="Gyrinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="family">Gyrinidae</taxonomicName>
have been observed.
</paragraph>
</subSubSection>
</treatment>
</document>