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401 lines
40 KiB
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<document ID-DOI="http://doi.org/10.5281/zenodo.3728741" ID-GBIF-Dataset="1cd9553c-1ca8-45ec-9a0e-2ba5bcdb5987" ID-GBIF-Taxon="162716766" ID-Zenodo-Dep="3728741" approvalRequired="13" approvalRequired_for_document="2" approvalRequired_for_textStreams="11" checkinTime="1584986461704" checkinUser="valdenar" docAuthor="P Érez Santa- Rita, Jose V., Valdivia, Alejandra & Baixeras, Joaquín" docDate="2020" docId="03AC545B12650840BB86FB5BFF0A3A52" docLanguage="en" docName="PROC.ENTOMOL.SOC.WASH.122.1.127.pdf.imf" docOrigin="Proceedings of the Entomological Society of Washington 122 (1)" docStyle="DocumentStyle{}" docStyleName="procentomolsocwash.2010.journal_article" docTitle="Aspinipogon samaypatensis P Érez Santa- Rita & Valdivia & Baixeras 2020, sp. n." docType="treatment" docVersion="6" lastPageId="5" lastPageNumber="132" masterDocId="FF952C2312640845B920FFF1FFED3927" masterDocTitle="Description Of A New Species Of Aspinipogon Razowski And Becker, 2010, With The First Reported Female Of The Genus (Lepidoptera: Tortricidae)" masterLastPageNumber="134" masterPageNumber="127" pageId="1" pageNumber="128" updateTime="1643447700084" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Description Of A New Species Of Aspinipogon Razowski And Becker, 2010, With The First Reported Female Of The Genus (Lepidoptera: Tortricidae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>P Érez Santa- Rita, Jose V.</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Valdivia, Alejandra</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Baixeras, Joaquín</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Proceedings of the Entomological Society of Washington</mods:title>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-02-29</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>122</mods:number>
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<mods:number>1</mods:number>
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<mods:start>127</mods:start>
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<mods:end>134</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.4289/0013-8797.122.1.127</mods:identifier>
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<mods:identifier type="GBIF-Dataset">1cd9553c-1ca8-45ec-9a0e-2ba5bcdb5987</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3724729</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3728741" ID-GBIF-Taxon="162716766" ID-Zenodo-Dep="3728741" LSID="urn:lsid:plazi:treatment:03AC545B12650840BB86FB5BFF0A3A52" httpUri="http://treatment.plazi.org/id/03AC545B12650840BB86FB5BFF0A3A52" lastPageId="5" lastPageNumber="132" pageId="1" pageNumber="128">
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<subSubSection pageId="1" pageNumber="128" type="nomenclature">
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<paragraph blockId="1.[678,1133,1194,1218]" box="[678,1133,1194,1218]" pageId="1" pageNumber="128">
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<heading box="[678,1133,1194,1218]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="128" reason="2">
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<emphasis bold="true" box="[678,1133,1194,1218]" pageId="1" pageNumber="128">
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<taxonomicName authority="P Érez Santa- Rita & Valdivia & Baixeras, 2020" authorityName="P Érez Santa- Rita & Valdivia & Baixeras" authorityYear="2020" box="[678,1043,1194,1218]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="samaypatensis" status="sp. n.">
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<emphasis bold="true" box="[678,1043,1194,1218]" italics="true" pageId="1" pageNumber="128">Aspinipogon samaypatensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[1060,1133,1194,1218]" pageId="1" pageNumber="128" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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<paragraph blockId="1.[668,1143,1243,1334]" pageId="1" pageNumber="128">
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http://zoobank.org/
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<uuid pageId="1" pageNumber="128">A2B500B8-C9FF- 46F9-A50F-0206D92AA0C8</uuid>
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(
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<figureCitation box="[845,966,1309,1334]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="3.[140,186,1332,1352]" captionStartId-1="4.[191,237,1588,1608]" captionTargetBox-0="[113,1157,198,1297]" captionTargetBox-1="[164,666,955,1554]" captionTargetId-0="figure@3.[111,1159,196,1299]" captionTargetId-1="figure@4.[162,667,953,1555]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figs. 2–6. Morphological characters of Aspinipogon samaypatensis sp. n. 2, Habitus, holotype, male, Bolivia, Santa Cruz, Pampagrande, 25 January 2011, MNKM. 3, Male genitalia, phallus removed, slide JBA20799. 4, Phallus uneverted, slide JBA20714. 5, Phallus with everted vesical, slide JBA20798. 6, Female genitalia, slide JBA20721. Scale bars: 5 mm (Fig. 1); 500 µm (Figs. 3‒6). Abbreviations. ab: accessory bursa. d: diverticulum. ds: ductus seminalis." captionText-1="Figs. 7–8. Everted phallus of Aspinipogon samaypatensis sp. n. 7, Left aspect. 8, Right aspect, slide JBA20893. Abbreviations. c: cornutus. dd: dorsal diverticulum. ld: left diverticulum. rd: right diverticulum. sd: small diverticula. Scale bars: 500 µm." figureDoi-0="http://doi.org/10.5281/zenodo.3724735" figureDoi-1="http://doi.org/10.5281/zenodo.3724737" httpUri-0="https://zenodo.org/record/3724735/files/figure.png" httpUri-1="https://zenodo.org/record/3724737/files/figure.png" pageId="1" pageNumber="128">Figs. 2–8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="132" pageId="1" pageNumber="128" type="diagnosis">
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<paragraph blockId="1.[651,1161,1358,1749]" lastBlockId="2.[159,670,827,1749]" lastPageId="2" lastPageNumber="129" pageId="1" pageNumber="128">
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Diagnosis.—The forewing pattern in both species of
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<taxonomicName authorityName="RAZOWSKI AND BECKER" authorityYear="2010" box="[843,1002,1392,1416]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="genus">
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<emphasis box="[843,1002,1392,1416]" italics="true" pageId="1" pageNumber="128">Aspinipogon</emphasis>
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</taxonomicName>
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consists of a system of parallel fasciae between the costa and the dorsum obliquely arranged with respect to the termen. The median fascia in
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<taxonomicName authorityName="P Érez Santa- Rita & Valdivia & Baixeras" authorityYear="2020" box="[775,998,1525,1549]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="samaypatensis">
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<emphasis box="[775,998,1525,1549]" italics="true" pageId="1" pageNumber="128">A. samaypatensis</emphasis>
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</taxonomicName>
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extends approximately on one-third of the costa whereas in
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<taxonomicName authorityName="Razowski and Becker" authorityYear="2010" box="[801,914,1591,1615]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="bellus">
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<emphasis box="[801,914,1591,1615]" italics="true" pageId="1" pageNumber="128">A. bellus</emphasis>
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</taxonomicName>
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it extends on onesixth of costa. In
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<taxonomicName authorityName="P Érez Santa- Rita & Valdivia & Baixeras" authorityYear="2020" box="[887,1109,1624,1648]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="samaypatensis">
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<emphasis box="[887,1109,1624,1648]" italics="true" pageId="1" pageNumber="128">A. samaypatensis</emphasis>
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</taxonomicName>
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the colour of the markings is darker than in
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<taxonomicName authorityName="Razowski and Becker" authorityYear="2010" box="[651,757,1691,1715]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="bellus">
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<emphasis box="[651,757,1691,1715]" italics="true" pageId="1" pageNumber="128">A. bellus</emphasis>
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</taxonomicName>
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, especially in the basal-subbasal blotch. The distal markings (postmedian + preterminal) are more contrasted with respect to the interfascial areas in
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<taxonomicName authorityName="Razowski and Becker" authorityYear="2010" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="bellus">
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<emphasis italics="true" pageId="2" pageNumber="129">A. bellus</emphasis>
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</taxonomicName>
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than in
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<taxonomicName authorityName="P Érez Santa- Rita & Valdivia & Baixeras" authorityYear="2020" box="[367,595,894,918]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="samaypatensis">
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<emphasis box="[367,595,894,918]" italics="true" pageId="2" pageNumber="129">A. samaypatensis</emphasis>
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</taxonomicName>
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. The male genitalia of
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<taxonomicName authorityName="P Érez Santa- Rita & Valdivia & Baixeras" authorityYear="2020" box="[395,618,927,951]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="samaypatensis">
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<emphasis box="[395,618,927,951]" italics="true" pageId="2" pageNumber="129">A. samaypatensis</emphasis>
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</taxonomicName>
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are easily distinguished from those of
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<taxonomicName authorityName="Razowski and Becker" authorityYear="2010" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="bellus">
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<emphasis italics="true" pageId="2" pageNumber="129">A. bellus</emphasis>
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</taxonomicName>
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. In
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<taxonomicName authorityName="P Érez Santa- Rita & Valdivia & Baixeras" authorityYear="2020" box="[287,507,993,1017]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="samaypatensis">
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<emphasis box="[287,507,993,1017]" italics="true" pageId="2" pageNumber="129">A. samaypatensis</emphasis>
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</taxonomicName>
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the cucullus is smoothly elongate and distally fingerlike, whereas in
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<taxonomicName authorityName="Razowski and Becker" authorityYear="2010" box="[403,512,1060,1084]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="bellus">
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<emphasis box="[403,512,1060,1084]" italics="true" pageId="2" pageNumber="129">A. bellus</emphasis>
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</taxonomicName>
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the cucullus is subtriangular, ventrally spined, and with a distinct neck. The sacculus is pointed in both species but relatively longer and more sinuous in
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<taxonomicName authorityName="P Érez Santa- Rita & Valdivia & Baixeras" authorityYear="2020" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="samaypatensis">
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<emphasis box="[538,557,1193,1217]" italics="true" pageId="2" pageNumber="129">A</emphasis>
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.
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<emphasis italics="true" pageId="2" pageNumber="129">samaypatensis</emphasis>
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</taxonomicName>
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<emphasis box="[266,270,1227,1250]" italics="true" pageId="2" pageNumber="129">.</emphasis>
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The cornutus in both species is well sclerotized; the apical portion is curved downward in
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<taxonomicName authorityName="Razowski and Becker" authorityYear="2010" box="[423,533,1292,1316]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="bellus">
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<emphasis box="[423,533,1292,1316]" italics="true" pageId="2" pageNumber="129">A. bellus</emphasis>
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</taxonomicName>
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, whereas it is porrect in
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<taxonomicName authorityName="P Érez Santa- Rita & Valdivia & Baixeras" authorityYear="2020" box="[313,520,1325,1349]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="samaypatensis">
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<emphasis box="[313,520,1325,1349]" italics="true" pageId="2" pageNumber="129">A. samaypatensis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3724733" ID-Zenodo-Dep="3724733" httpUri="https://zenodo.org/record/3724733/files/figure.png" pageId="2" pageNumber="129" startId="2.[191,228,708,728]" targetBox="[163,666,197,675]" targetPageId="2">
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<paragraph blockId="2.[159,670,708,784]" pageId="2" pageNumber="129">
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Fig. 1. Venation in
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<taxonomicName authority="P Érez Santa- Rita & Valdivia & Baixeras, 2020" authorityName="P Érez Santa- Rita & Valdivia & Baixeras" authorityYear="2020" box="[397,670,709,729]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="samaypatensis" status="sp. n.">
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<emphasis box="[397,670,709,729]" italics="true" pageId="2" pageNumber="129">Aspinipogon samaypatensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[159,216,737,756]" pageId="2" pageNumber="129" rank="species">sp. n.</taxonomicNameLabel>
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Based on slide JBA20816. Dashed lines indicate obsolescent veins. Scale bars: 2 mm.
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</paragraph>
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</caption>
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<paragraph blockId="2.[159,670,827,1749]" lastBlockId="2.[701,1212,196,1749]" pageId="2" pageNumber="129">
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Description.—
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<emphasis box="[377,445,1359,1383]" italics="true" pageId="2" pageNumber="129">Head</emphasis>
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: Typically tortricoid. Vertex with light brown scales. Frons light brown, slightly convex, nearly naked. Antenna brownish ochre, length ca. 0.5 as long as forewing costa, dorsally scaled with two rows of scales per flagellomere, ventrally ciliated, cilia less than 0.2 times width of flagellomere. Maxillary palpus inconspicuous. Labial palpus porrect, length (all three segments combined) ca. 1.3 times diameter of compound eye, uniformly and densely scaled; first segment short, second segment long, straight, and rough scaled, third segment short and straight; opening of organ of vom Rath in lateral position. Haustellum well developed, naked. Ocellus well developed; chaetosomata reduced.
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</paragraph>
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<paragraph blockId="2.[701,1212,196,1749]" pageId="2" pageNumber="129">
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<emphasis box="[733,823,396,420]" italics="true" pageId="2" pageNumber="129">Thorax</emphasis>
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: Pronotum, anterior half of mesoscutum, and tegulae covered by brown scales, posterior half of mesoscutum covered by dark brown scales, metanotum covered by ochre scales; smooth-scaled including tegulae, without tufts of scales. Sterna and legs pale whitish brown, male foreleg hairpencil absent. Forewing length
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(mean = 6.9; n = 7) in males,
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<quantity box="[1061,1155,694,719]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="2" pageNumber="129" unit="mm" value="8.7">8.7 mm</quantity>
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(n = 1) in females. Forewing pattern (
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<figureCitation box="[1124,1202,727,752]" captionStart="Figs" captionStartId="3.[140,186,1332,1352]" captionTargetBox="[113,1157,198,1297]" captionTargetId="figure@3.[111,1159,196,1299]" captionTargetPageId="3" captionText="Figs. 2–6. Morphological characters of Aspinipogon samaypatensis sp. n. 2, Habitus, holotype, male, Bolivia, Santa Cruz, Pampagrande, 25 January 2011, MNKM. 3, Male genitalia, phallus removed, slide JBA20799. 4, Phallus uneverted, slide JBA20714. 5, Phallus with everted vesical, slide JBA20798. 6, Female genitalia, slide JBA20721. Scale bars: 5 mm (Fig. 1); 500 µm (Figs. 3‒6). Abbreviations. ab: accessory bursa. d: diverticulum. ds: ductus seminalis." figureDoi="http://doi.org/10.5281/zenodo.3724735" httpUri="https://zenodo.org/record/3724735/files/figure.png" pageId="2" pageNumber="129">Fig. 2</figureCitation>
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) not sexually dimorphic. Forewing upperside light brown with scattered dark yellowish-green marks on costa; costal strigulae weakly expressed; basal and subbasal fascia fused, dark brown in posterior half as a dark brown dorsal subtriangular blotch delimited by whitish scales, anterior half subfused with interfascial area as a costal pale band; interfascial area as a dorsal spot concolorous with median fascia but delimited by whitish scales; median fascia broad, greenish olivaceous brown proximally, dorsally darker; an interfascial area between median and postmedian fasciae light brown delimited by darker scales, extending between Sc-R1 on costa and tornus; postmedian and preterminal fasciae fused and rather concolorous with median fascia; fringe with a mixture of dark brown and cream ochre scales. Forewing underside light brown, pale on the base, pale interradial marks on costa. Hindwing upperside uniformly light brown with brownish paler scales in area overlapping with forewing; fringe pale. Hindwing underside whitish.
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||
</paragraph>
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<paragraph blockId="2.[701,1212,196,1749]" lastBlockId="4.[701,1212,196,1749]" lastPageId="4" lastPageNumber="131" pageId="2" pageNumber="129">
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<emphasis box="[733,848,1657,1681]" italics="true" pageId="2" pageNumber="129">Abdomen</emphasis>
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: Dorsad brown grey, pale cream ochreous cephalad and caudad. Segment
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||
<quantity box="[820,873,1724,1749]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="2" pageNumber="129" unit="in" value="8.0">8 in</quantity>
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||
males with long modified scales projecting caudad covering genital segments. Male genitalia (
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<figureCitation box="[477,609,1558,1583]" captionStart="Figs" captionStartId="3.[140,186,1332,1352]" captionTargetBox="[113,1157,198,1297]" captionTargetId="figure@3.[111,1159,196,1299]" captionTargetPageId="3" captionText="Figs. 2–6. Morphological characters of Aspinipogon samaypatensis sp. n. 2, Habitus, holotype, male, Bolivia, Santa Cruz, Pampagrande, 25 January 2011, MNKM. 3, Male genitalia, phallus removed, slide JBA20799. 4, Phallus uneverted, slide JBA20714. 5, Phallus with everted vesical, slide JBA20798. 6, Female genitalia, slide JBA20721. Scale bars: 5 mm (Fig. 1); 500 µm (Figs. 3‒6). Abbreviations. ab: accessory bursa. d: diverticulum. ds: ductus seminalis." figureDoi="http://doi.org/10.5281/zenodo.3724735" httpUri="https://zenodo.org/record/3724735/files/figure.png" pageId="3" pageNumber="130">Figs. 3‒5</figureCitation>
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) (based on three preparations) with tegumen simple; uncus as a small slight protuberance; socius short, hairy, appressed to tegumen; gnathos reduced to a small sclerotized lateral curved plate; anal cone slightly sclerotized; transtilla simple, continuous, slender but evident; valva well sclerotized, length
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<quantity box="[939,1094,1590,1616]" metricMagnitude="-1" metricUnit="m" metricValue="9.55" metricValueMax="9.7" metricValueMin="9.4" pageId="3" pageNumber="130" unit="mm" value="955.0" valueMax="970.0" valueMin="940.0">940–970 µm</quantity>
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from the base of sacculus to apex of cucullus, costa fairly straight, length
|
||
<quantity box="[1001,1160,1657,1682]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" metricValueMax="6.2" metricValueMin="5.8" pageId="3" pageNumber="130" unit="mm" value="600.0" valueMax="620.0" valueMin="580.0">580–620 µm</quantity>
|
||
from apex to base; cucullus simple, fingerlike, hairy, especially distally; sacculus long, sinuous, same length as costa of valva, strongly sclerotized, projected distally in a upcurved point; pulvinus absent; basal process simple; vinculum strong; basal shield of the juxta simple, semicircular, projected caudad and dorsad as a sclerotized plate, medially membranous; phallus (
|
||
<figureCitation box="[263,376,428,453]" captionStart="Figs" captionStartId="3.[140,186,1332,1352]" captionTargetBox="[113,1157,198,1297]" captionTargetId="figure@3.[111,1159,196,1299]" captionTargetPageId="3" captionText="Figs. 2–6. Morphological characters of Aspinipogon samaypatensis sp. n. 2, Habitus, holotype, male, Bolivia, Santa Cruz, Pampagrande, 25 January 2011, MNKM. 3, Male genitalia, phallus removed, slide JBA20799. 4, Phallus uneverted, slide JBA20714. 5, Phallus with everted vesical, slide JBA20798. 6, Female genitalia, slide JBA20721. Scale bars: 5 mm (Fig. 1); 500 µm (Figs. 3‒6). Abbreviations. ab: accessory bursa. d: diverticulum. ds: ductus seminalis." figureDoi="http://doi.org/10.5281/zenodo.3724735" httpUri="https://zenodo.org/record/3724735/files/figure.png" pageId="4" pageNumber="131">Figs. 4, 5</figureCitation>
|
||
,
|
||
<figureCitation box="[389,433,428,453]" captionStart="Figs" captionStartId="4.[191,237,1588,1608]" captionTargetBox="[164,666,955,1554]" captionTargetId="figure@4.[162,667,953,1555]" captionTargetPageId="4" captionText="Figs. 7–8. Everted phallus of Aspinipogon samaypatensis sp. n. 7, Left aspect. 8, Right aspect, slide JBA20893. Abbreviations. c: cornutus. dd: dorsal diverticulum. ld: left diverticulum. rd: right diverticulum. sd: small diverticula. Scale bars: 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3724737" httpUri="https://zenodo.org/record/3724737/files/figure.png" pageId="4" pageNumber="131">7, 8</figureCitation>
|
||
) ca. same length as valva, with coecum penis simple, caulis absent, vesica complex, asymmetrical, membranous (no acanthae), with three main diverticula when fully everted, two basal, laterally arranged, left one bearing a single long saberlike, non-deciduous cornutus attached laterobasally, inwardly directed, and longitudinally striated (no deciduous cornuti detected in female genitalia corpus bursae), right one smaller, without sclerotization, a third diverticulum dorsal subconical, slightly directed to right, bearing three small diverticula and distally taking nonsclerotized gonopore at tip of a funnellike delicately membranous extension. Segment
|
||
<quantity box="[824,879,296,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="131" unit="in" value="7.0">7 in</quantity>
|
||
female without modified scaling. Female genitalia (
|
||
<figureCitation box="[1034,1110,329,354]" captionStart="Figs" captionStartId="3.[140,186,1332,1352]" captionTargetBox="[113,1157,198,1297]" captionTargetId="figure@3.[111,1159,196,1299]" captionTargetPageId="3" captionText="Figs. 2–6. Morphological characters of Aspinipogon samaypatensis sp. n. 2, Habitus, holotype, male, Bolivia, Santa Cruz, Pampagrande, 25 January 2011, MNKM. 3, Male genitalia, phallus removed, slide JBA20799. 4, Phallus uneverted, slide JBA20714. 5, Phallus with everted vesical, slide JBA20798. 6, Female genitalia, slide JBA20721. Scale bars: 5 mm (Fig. 1); 500 µm (Figs. 3‒6). Abbreviations. ab: accessory bursa. d: diverticulum. ds: ductus seminalis." figureDoi="http://doi.org/10.5281/zenodo.3724735" httpUri="https://zenodo.org/record/3724735/files/figure.png" pageId="4" pageNumber="131">Fig. 6</figureCitation>
|
||
) (based on one preparation) with ostium broad, groovelike; sterigma slightly sclerotized; antrum extremely short, broad, unsclerotized; ductus bursae wide, strongly sclerotized, relatively short and continuous with corpus bursae, dorsoventrally flattened, slightly narrowed before cervix, left side posteriorly expanded in a sclerotized diverticulum; corpus bursae sub-spherical, covered with numerous well-developed acanthae individually arranged and in bands or clusters, ventrally with an accessory bursa membranous, but with complex folds (stained dark with chlorazol black); no distinct signum. Ductus seminalis ventrally connected to anterior membranous part of corpus bursae; no bulla seminalis detected. A single globular spermatophore without collum extracted. Anal papillae uniformly covered by strong and long hairs; posterior apophysis short; ovipositor short, stout.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3724735" ID-Zenodo-Dep="3724735" httpUri="https://zenodo.org/record/3724735/files/figure.png" pageId="3" pageNumber="130" startId="3.[140,186,1332,1352]" targetBox="[113,1157,198,1297]" targetPageId="3">
|
||
<paragraph blockId="3.[109,1162,1332,1463]" pageId="3" pageNumber="130">
|
||
Figs. 2–6. Morphological characters of
|
||
<taxonomicName authority="P Érez Santa- Rita & Valdivia & Baixeras, 2020" authorityName="P Érez Santa- Rita & Valdivia & Baixeras" authorityYear="2020" box="[575,861,1332,1352]" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="samaypatensis" status="sp. n.">
|
||
<emphasis box="[575,861,1332,1352]" italics="true" pageId="3" pageNumber="130">Aspinipogon samaypatensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[871,930,1333,1352]" pageId="3" pageNumber="130" rank="species">sp. n.</taxonomicNameLabel>
|
||
2, Habitus, holotype, male, Bolivia, Santa Cruz, Pampagrande, 25 January 2011, MNKM. 3, Male genitalia, phallus removed, slide JBA20799. 4, Phallus uneverted, slide JBA20714. 5, Phallus with everted vesical, slide JBA20798. 6, Female genitalia, slide JBA20721. Scale bars: 5 mm (Fig. 1); 500 µm (Figs. 3‒6). Abbreviations. ab: accessory bursa. d: diverticulum. ds: ductus seminalis.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="4.[701,1212,196,1749]" lastBlockId="5.[109,619,196,885]" lastPageId="5" lastPageNumber="132" pageId="4" pageNumber="131">
|
||
Type material.—
|
||
<materialsCitation ID-GBIF-Occurrence="2579594354" collectingDate="2011-01-25" collectionCode="MNKM" collectorName="J. Baixeras & A. Valdivia & G. Fernandez" country="Bolivia" county="Florida Province" elevation="1575" location="Hueco de la Pascana" municipality="Pampa Grande Municipality" pageId="4" pageNumber="131" specimenCount="1" stateProvince="Santa Cruz Department" typeStatus="holotype">
|
||
<emphasis bold="true" box="[939,1065,1093,1117]" pageId="4" pageNumber="131">
|
||
<typeStatus box="[939,1060,1093,1117]" pageId="4" pageNumber="131">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Ơ,
|
||
<collectingCountry box="[1117,1207,1093,1118]" name="Bolivia" pageId="4" pageNumber="131">Bolivia</collectingCountry>
|
||
,
|
||
<collectingRegion box="[701,990,1126,1151]" country="Bolivia" name="Santa Cruz" pageId="4" pageNumber="131">Santa Cruz Department</collectingRegion>
|
||
,
|
||
<collectingCounty box="[1003,1206,1126,1151]" pageId="4" pageNumber="131">Florida Province</collectingCounty>
|
||
,
|
||
<collectingMunicipality box="[701,1060,1159,1184]" pageId="4" pageNumber="131">Pampa Grande Municipality</collectingMunicipality>
|
||
, locality of
|
||
<location LSID="urn:lsid:plazi:treatment:03AC545B12650840BB86FB5BFF0A3A52:8EDAB39612600841BB9DFB58FC533DE7" box="[701,958,1192,1217]" country="Bolivia" county="Florida Province" municipality="Pampa Grande Municipality" name="Hueco de la Pascana" pageId="4" pageNumber="131" stateProvince="Santa Cruz Department">Hueco de la Pascana</location>
|
||
,
|
||
<quantity box="[972,1069,1192,1217]" metricMagnitude="3" metricUnit="m" metricValue="1.575" pageId="4" pageNumber="131" unit="m" value="1575.0">
|
||
<elevation box="[972,1069,1192,1217]" metricMagnitude="3" metricUnit="m" metricValue="1.575" pageId="4" pageNumber="131" unit="m" value="1575.0">1575 m</elevation>
|
||
</quantity>
|
||
, 18
|
||
<emphasis box="[1112,1124,1192,1217]" italics="true" pageId="4" pageNumber="131">°</emphasis>
|
||
7.09
|
||
<emphasis bold="true" box="[1176,1186,1192,1217]" italics="true" pageId="4" pageNumber="131">̍</emphasis>
|
||
S, 64
|
||
<emphasis box="[731,743,1225,1250]" italics="true" pageId="4" pageNumber="131">°</emphasis>
|
||
3.58
|
||
<emphasis bold="true" box="[796,806,1225,1250]" italics="true" pageId="4" pageNumber="131">̍</emphasis>
|
||
W,
|
||
<date box="[852,1010,1225,1250]" pageId="4" pageNumber="131" value="2011-01-25">
|
||
<collectingDate box="[852,1010,1225,1250]" pageId="4" pageNumber="131" value="2011-01-25">25 Jan 2011</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1027,1165,1225,1250]" pageId="4" pageNumber="131">J. Baixeras</collectorName>
|
||
,
|
||
<collectorName pageId="4" pageNumber="131">A. Valdivia</collectorName>
|
||
and
|
||
<collectorName box="[888,1065,1259,1284]" pageId="4" pageNumber="131">G. Fernández</collectorName>
|
||
(
|
||
<collectionCode box="[1095,1199,1259,1284]" pageId="4" pageNumber="131">MNKM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation ID-GBIF-Occurrence="2579594356" collectingDate="1997-10-23" collectingDateMax="2011-02-04" collectingDateMin="1997-10-23" collectionCode="O, JBA" collectorName="J. Baixeras & A. Valdivia & G. Fernandez & G. Fernandez. Locality & Samaipata & Samaipata Municipality & Achira" country="Bolivia" county="Florida Province" elevation="1554" lastPageId="5" lastPageNumber="132" location="Hueco de la Pascana" municipality="Pampa Grande Municipality" pageId="4" pageNumber="131" specimenCode="GS 20721" specimenCount="2" specimenCount-female="2" stateProvince="Santa Cruz Department" typeStatus="paratype">
|
||
<emphasis bold="true" box="[701,829,1292,1316]" pageId="4" pageNumber="131">
|
||
<typeStatus box="[701,829,1292,1316]" pageId="4" pageNumber="131">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(6
|
||
<collectionCode box="[864,886,1291,1316]" pageId="4" pageNumber="131">Ơ</collectionCode>
|
||
,
|
||
<specimenCount box="[904,942,1291,1317]" pageId="4" pageNumber="131" type="female">1♀</specimenCount>
|
||
):
|
||
<collectingCountry box="[970,1062,1292,1317]" name="Bolivia" pageId="4" pageNumber="131">Bolivia</collectingCountry>
|
||
:
|
||
<collectingRegion country="Bolivia" name="Santa Cruz" pageId="4" pageNumber="131">Santa Cruz Department</collectingRegion>
|
||
,
|
||
<collectingCounty box="[877,1099,1325,1350]" pageId="4" pageNumber="131">Florida Province</collectingCounty>
|
||
,
|
||
<collectingMunicipality pageId="4" pageNumber="131">Pampa Grande Municipality</collectingMunicipality>
|
||
, locality of
|
||
<location LSID="urn:lsid:plazi:treatment:03AC545B12650840BB86FB5BFF0A3A52:8EDAB39612600841BD68FABEFCD23CAF" country="Bolivia" county="Florida Province" municipality="Pampa Grande Municipality" name="Hueco de la Pascana" pageId="4" pageNumber="131" stateProvince="Santa Cruz Department">Hueco de la Pascana</location>
|
||
, 18
|
||
<emphasis box="[875,887,1391,1416]" italics="true" pageId="4" pageNumber="131">°</emphasis>
|
||
7.09
|
||
<emphasis bold="true" box="[939,949,1391,1416]" italics="true" pageId="4" pageNumber="131">̍</emphasis>
|
||
S, 64
|
||
<emphasis box="[1013,1025,1391,1416]" italics="true" pageId="4" pageNumber="131">°</emphasis>
|
||
3.58
|
||
<emphasis bold="true" box="[1078,1088,1391,1416]" italics="true" pageId="4" pageNumber="131">̍</emphasis>
|
||
W,
|
||
<date pageId="4" pageNumber="131" value="2011-01-30">
|
||
<collectingDate pageId="4" pageNumber="131" value="2011-01-30">30 Jan 2011</collectingDate>
|
||
</date>
|
||
(1
|
||
<collectionCode box="[794,817,1424,1449]" pageId="4" pageNumber="131">Ơ</collectionCode>
|
||
),
|
||
<date box="[842,994,1425,1450]" pageId="4" pageNumber="131" value="2011-02-04">
|
||
<collectingDate box="[842,994,1425,1450]" pageId="4" pageNumber="131" value="2011-02-04">04 Feb 2011</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount box="[1012,1048,1424,1450]" pageId="4" pageNumber="131" type="female">1♀</specimenCount>
|
||
),
|
||
<collectorName box="[1073,1206,1425,1450]" pageId="4" pageNumber="131">J. Baixeras</collectorName>
|
||
,
|
||
<collectorName box="[701,837,1458,1483]" pageId="4" pageNumber="131">A. Valdivia</collectorName>
|
||
and
|
||
<collectorName box="[895,1056,1458,1483]" pageId="4" pageNumber="131">G. Fernández</collectorName>
|
||
(
|
||
<specimenCode box="[1072,1198,1458,1483]" collectionCode="GS" pageId="4" pageNumber="131">GS 20721</specimenCode>
|
||
).
|
||
<location LSID="urn:lsid:plazi:treatment:03AC545B12650840BB86FB5BFF0A3A52:8EDAB39612600841BB9DFA22FBCE3CCB" box="[701,1059,1491,1516]" country="Bolivia" county="Florida Province" municipality="Pampa Grande Municipality" name="Locality of Agua Clarita" pageId="4" pageNumber="131" stateProvince="Santa Cruz Department">Locality of Agua Clarita</location>
|
||
,
|
||
<quantity box="[1092,1206,1491,1516]" metricMagnitude="3" metricUnit="m" metricValue="1.554" pageId="4" pageNumber="131" unit="m" value="1554.0">
|
||
<elevation box="[1092,1206,1491,1516]" metricMagnitude="3" metricUnit="m" metricValue="1.554" pageId="4" pageNumber="131" unit="m" value="1554.0">1554 m</elevation>
|
||
</quantity>
|
||
, 17
|
||
<emphasis box="[731,743,1524,1549]" italics="true" pageId="4" pageNumber="131">°</emphasis>
|
||
56.74
|
||
<emphasis bold="true" box="[811,821,1524,1549]" italics="true" pageId="4" pageNumber="131">̍</emphasis>
|
||
S, 64
|
||
<emphasis box="[882,894,1524,1549]" italics="true" pageId="4" pageNumber="131">°</emphasis>
|
||
7.97
|
||
<emphasis bold="true" box="[946,956,1524,1549]" italics="true" pageId="4" pageNumber="131">̍</emphasis>
|
||
W,
|
||
<date box="[997,1141,1524,1549]" pageId="4" pageNumber="131" value="2011-01-27">
|
||
<collectingDate box="[997,1141,1524,1549]" pageId="4" pageNumber="131" value="2011-01-27">27 Jan 2011</collectingDate>
|
||
</date>
|
||
(1
|
||
<collectionCode box="[1171,1194,1524,1549]" pageId="4" pageNumber="131">Ơ</collectionCode>
|
||
),
|
||
<collectorName box="[701,835,1558,1583]" pageId="4" pageNumber="131">J. Baixeras</collectorName>
|
||
,
|
||
<collectorName box="[847,983,1558,1583]" pageId="4" pageNumber="131">A. Valdivia</collectorName>
|
||
and
|
||
<collectorName pageId="4" pageNumber="131">G. Fernández. Locality</collectorName>
|
||
of Antiguo camino a
|
||
<collectorName box="[1079,1206,1591,1616]" pageId="4" pageNumber="131">Samaipata</collectorName>
|
||
,
|
||
<quantity box="[701,799,1624,1649]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="131" unit="m" value="1300.0">
|
||
<elevation box="[701,799,1624,1649]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="131" unit="m" value="1300.0">1300 m</elevation>
|
||
</quantity>
|
||
, 18
|
||
<emphasis box="[843,855,1623,1648]" italics="true" pageId="4" pageNumber="131">°</emphasis>
|
||
03
|
||
<emphasis bold="true" box="[885,895,1623,1648]" italics="true" pageId="4" pageNumber="131">̍</emphasis>
|
||
S, 64
|
||
<emphasis box="[960,972,1623,1648]" italics="true" pageId="4" pageNumber="131">°</emphasis>
|
||
05
|
||
<emphasis bold="true" box="[1002,1012,1623,1648]" italics="true" pageId="4" pageNumber="131">̍</emphasis>
|
||
W,
|
||
<date box="[1057,1211,1624,1649]" pageId="4" pageNumber="131" value="1997-10-23">
|
||
<collectingDate box="[1057,1211,1624,1649]" pageId="4" pageNumber="131" value="1997-10-23">23 Oct 1997</collectingDate>
|
||
</date>
|
||
(3
|
||
<collectionCode box="[726,749,1657,1682]" pageId="4" pageNumber="131">Ơ</collectionCode>
|
||
),
|
||
<collectorName box="[793,943,1657,1682]" pageId="4" pageNumber="131">J. Baixeras</collectorName>
|
||
(microscope slides
|
||
<accessionNumber isEnumeration="true" pageId="4" pageNumber="131">
|
||
<collectionCode box="[701,750,1690,1715]" pageId="4" pageNumber="131">JBA</collectionCode>
|
||
20714,
|
||
<collectionCode box="[830,880,1690,1715]" pageId="4" pageNumber="131">JBA</collectionCode>
|
||
20721,
|
||
<collectionCode box="[959,1009,1690,1715]" pageId="4" pageNumber="131">JBA</collectionCode>
|
||
20798,
|
||
<collectionCode box="[1088,1137,1690,1715]" pageId="4" pageNumber="131">JBA</collectionCode>
|
||
20799,
|
||
<collectionCode box="[701,751,1724,1749]" pageId="4" pageNumber="131">JBA</collectionCode>
|
||
20801,
|
||
<collectionCode box="[830,879,1724,1749]" pageId="4" pageNumber="131">JBA</collectionCode>
|
||
20816,
|
||
<collectionCode box="[960,1011,1724,1749]" pageId="4" pageNumber="131">JBA</collectionCode>
|
||
20893
|
||
</accessionNumber>
|
||
).
|
||
<collectorName lastPageId="5" lastPageNumber="132" pageId="4" pageNumber="131">Samaipata Municipality</collectorName>
|
||
, locality of
|
||
<collectorName box="[416,502,196,221]" pageId="5" pageNumber="132">Achira</collectorName>
|
||
,
|
||
<quantity box="[516,613,196,221]" metricMagnitude="3" metricUnit="m" metricValue="1.456" pageId="5" pageNumber="132" unit="m" value="1456.0">
|
||
<elevation box="[516,613,196,221]" metricMagnitude="3" metricUnit="m" metricValue="1.456" pageId="5" pageNumber="132" unit="m" value="1456.0">1456 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date box="[109,259,229,254]" pageId="5" pageNumber="132" value="2011-02-01">
|
||
<collectingDate box="[109,259,229,254]" pageId="5" pageNumber="132" value="2011-02-01">01 Feb 2011</collectingDate>
|
||
</date>
|
||
(1
|
||
<collectionCode box="[290,313,229,254]" pageId="5" pageNumber="132">Ơ</collectionCode>
|
||
),
|
||
<collectorName box="[337,471,229,254]" pageId="5" pageNumber="132">J. Baixeras</collectorName>
|
||
,
|
||
<collectorName box="[481,618,229,254]" pageId="5" pageNumber="132">A. Valdivia</collectorName>
|
||
and
|
||
<collectorName box="[162,331,262,287]" pageId="5" pageNumber="132">G. Fernández</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation ID-GBIF-Occurrence="2579594355" collectionCode="MNKM" country="Bolivia" pageId="5" pageNumber="132" specimenCount="1" stateProvince="Santa Cruz" typeStatus="paratype">
|
||
<typeStatus box="[343,461,263,287]" pageId="5" pageNumber="132">Paratypes</typeStatus>
|
||
deposited in
|
||
<collectionCode box="[109,208,297,320]" pageId="5" pageNumber="132">MNKM</collectionCode>
|
||
and ICBiBE
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3724737" ID-Zenodo-Dep="3724737" httpUri="https://zenodo.org/record/3724737/files/figure.png" pageId="4" pageNumber="131" startId="4.[191,237,1588,1608]" targetBox="[164,666,955,1554]" targetPageId="4">
|
||
<paragraph blockId="4.[159,670,1588,1747]" pageId="4" pageNumber="131">
|
||
Figs. 7–8. Everted phallus of
|
||
<taxonomicName authority="P Érez Santa- Rita & Valdivia & Baixeras, 2020" authorityName="P Érez Santa- Rita & Valdivia & Baixeras" authorityYear="2020" class="Insecta" family="Tortricidae" genus="Aspinipogon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="131" phylum="Arthropoda" rank="species" species="samaypatensis" status="sp. n.">
|
||
<emphasis italics="true" pageId="4" pageNumber="131">Aspinipogon samaypatensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[317,374,1616,1635]" pageId="4" pageNumber="131" rank="species">sp. n.</taxonomicNameLabel>
|
||
7, Left aspect. 8, Right aspect, slide JBA20893. Abbreviations. c: cornutus. dd: dorsal diverticulum. ld: left diverticulum. rd: right diverticulum. sd: small diverticula. Scale bars: 500 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="5.[109,619,196,885]" pageId="5" pageNumber="132">
|
||
Biology and distribution.—The immature stages are unknown. Adults were collected in January (n = 3), February (n = 2) and October (n = 3) at middle elevations (
|
||
<quantity box="[262,450,462,487]" metricMagnitude="3" metricUnit="m" metricValue="1.4375" metricValueMax="1.575" metricValueMin="1.3" pageId="5" pageNumber="132" unit="m" value="1437.5" valueMax="1575.0" valueMin="1300.0">1300‒1575 m</quantity>
|
||
) in
|
||
<collectingCountry box="[519,614,462,487]" name="Bolivia" pageId="5" pageNumber="132">Bolivia</collectingCountry>
|
||
,
|
||
<collectingRegion box="[109,397,495,520]" country="Bolivia" name="Santa Cruz" pageId="5" pageNumber="132">Santa Cruz Department</collectingRegion>
|
||
, Florida Province, in the municipalities of Pampa Grande and Samaipata, localities of Achira, Agua Clarita, Hueco de la Pascana. The collecting sites are mainly transitional areas to Chaco forest.
|
||
</paragraph>
|
||
<paragraph blockId="5.[109,619,196,885]" pageId="5" pageNumber="132">
|
||
Etymology.—The specific epithet refers to the Quechuan word “Samaypata,” which translates as “height of rest.” Samaipata is the locality in
|
||
<collectingCountry box="[433,528,794,819]" name="Bolivia" pageId="5" pageNumber="132">Bolivia</collectingCountry>
|
||
, where specimens of this new species were collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |