176 lines
21 KiB
XML
176 lines
21 KiB
XML
<document id="BCF87176822DA04AF92D01029A0608BF" ID-DOI="10.1206/0003-0090.455.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="7161371" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665202843977" checkinUser="felipe" docAuthor="Voss, Robert S." docDate="2022" docId="03A487D6FFC4FFD6ADE73C92FB6CFEF1" docLanguage="en" docName="BAmMusNatHist.2022.455.1-74.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (455)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-455/issue-1/0003-0090.455.1.1/An-Annotated-Checklist-of-Recent-Opossums-Mammalia-Didelphidae/10.1206/0003-0090.455.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Marmosa (Exulomarmosa) robinsoni Bangs 1898" docType="treatment" docVersion="4" lastPageNumber="12" masterDocId="FF9DFFAEFFC8FFDAAD4A3C70FFEBFFEA" masterDocTitle="An Annotated Checklist Of Recent Opossums (Mammalia: Didelphidae)" masterLastPageNumber="77" masterPageNumber="1" pageNumber="12" updateTime="1699366015932" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="E444EA1CC057A4C4985C59267216EBEE">
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<mods:title id="01E8DBDA8C6079DD3D99D89B8F1302DF">An Annotated Checklist Of Recent Opossums (Mammalia: Didelphidae)</mods:title>
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<mods:namePart id="4CC8F00140893E65CF3F9027BA8ABEC6">Voss, Robert S.</mods:namePart>
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<mods:typeOfResource id="BF6C748EFE6C7D3B203B52F85E028182">text</mods:typeOfResource>
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<mods:title id="EEDB9CDE6B1FED0D121689323DEED4C7">Bulletin of the American Museum of Natural History</mods:title>
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<mods:part id="CC8FE58952D16D55AEB373EB14B27313">
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<mods:date id="EEEA4C633957C04C0D9B1515DCDD9D04">2022</mods:date>
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<mods:number id="3DC5F8EC2780260C3A5083FF5434C13E">2022-04-04</mods:number>
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<mods:detail id="4BBD4B829B4FCC58D03D1235351C6184" type="volume">
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<mods:number id="76E29E40074656DF5D1936833567E495">2022</mods:number>
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<mods:detail id="4F1954D0EA26AFDD70A96753F4D05F89" type="issue">
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<mods:number id="D945E7608D6331E0F19A9876832EE89B">455</mods:number>
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<mods:url id="80B9946889B6EF301C6850C2390A8EB3">https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-455/issue-1/0003-0090.455.1.1/An-Annotated-Checklist-of-Recent-Opossums-Mammalia-Didelphidae/10.1206/0003-0090.455.1.1.full</mods:url>
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</mods:location>
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<mods:classification id="12134229FD28C95C5AF10A5EFB9E2883">journal article</mods:classification>
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<mods:identifier id="7E9C8C452EFC85341FF12F240F6CE2AF" type="DOI">10.1206/0003-0090.455.1.1</mods:identifier>
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<mods:identifier id="CF1F667612B1136553A250984402CBFB" type="ISSN">0003-0090</mods:identifier>
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<mods:identifier id="17521FB0BC89B80713E82C61A1D00D98" type="Zenodo-Dep">7161371</mods:identifier>
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<treatment id="03A487D6FFC4FFD6ADE73C92FB6CFEF1" ID-DOI="http://doi.org/10.5281/zenodo.7161413" ID-Zenodo-Dep="7161413" LSID="urn:lsid:plazi:treatment:03A487D6FFC4FFD6ADE73C92FB6CFEF1" httpUri="http://treatment.plazi.org/id/03A487D6FFC4FFD6ADE73C92FB6CFEF1" lastPageNumber="12" pageId="12" pageNumber="12">
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<subSubSection id="C317654BFFC4FFD6ADE73C92FE5EFEF1" pageId="12" pageNumber="12" type="nomenclature">
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<paragraph id="8BB236C0FFC4FFD6ADE73C92FE5EFEF1" blockId="12.[173,572,226,283]" pageId="12" pageNumber="12">
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<heading id="D0FA81ACFFC4FFD6ADE73C92FE5EFEF1" centered="true" fontSize="9" level="2" pageId="12" pageNumber="12" reason="2">
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<taxonomicName id="4C0D4D43FFC4FFD6ADE73C92FE5EFEF1" ID-CoL="3Y83J" authority="Bangs, 1898" authorityName="Bangs" authorityYear="1898" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="12" phylum="Chordata" rank="species" species="robinsoni" subGenus="Exulomarmosa">
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<emphasis id="B979EAD2FFC4FFD6ADE73C92FEFFFF13" box="[173,276,226,249]" italics="true" pageId="12" pageNumber="12">Marmosa</emphasis>
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(
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<emphasis id="B979EAD2FFC4FFD6AC6E3C92FE2CFF10" box="[292,455,226,250]" italics="true" pageId="12" pageNumber="12">Exulomarmosa</emphasis>
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)
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<emphasis id="B979EAD2FFC4FFD6AC923C92FDD7FF10" box="[472,572,226,250]" italics="true" pageId="12" pageNumber="12">robinsoni</emphasis>
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Bangs, 1898
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C317654BFFC4FFD6ADC63D34FEE1FD4D" pageId="12" pageNumber="12" type="materials_examined">
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<paragraph id="8BB236C0FFC4FFD6ADC63D34FDA7FDEB" blockId="12.[108,636,324,1737]" pageId="12" pageNumber="12">
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<materialsCitation id="3B653C9DFFC4FFD6ADC63D34FDA3FDEB" collectionCode="MCZ" country="Venezuela" elevation="200" latitude="10.98" location="El Valle de Espirito Santo" longLatPrecision="778" longitude="-63.87" municipality="Margarita" pageId="12" pageNumber="12" specimenCode="B7749" specimenCount="1" stateProvince="Nueva Esparta" typeStatus="holotype">
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TYPE MATERIAL AND TYPE LOCALITY:
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<collectionCode id="ED1CAE05FFC4FFD6AF0B3D34FD97FEB6" box="[577,636,324,348]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="12" pageNumber="12" type="Museum">MCZ</collectionCode>
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<specimenCode id="DBAB9EBBFFC4FFD6AD263D15FF59FE97" box="[108,178,357,381]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="12" pageNumber="12" type="Museum">B7749</specimenCode>
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, the
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<typeStatus id="54B68862FFC4FFD6ADAE3D15FEA8FE97" box="[228,323,357,381]" pageId="12" pageNumber="12" type="holotype">holotype</typeStatus>
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by original designation, consists of the skin and skull of an adult male collected at
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<location id="8ED2601BFFC4FFD6AD9D3DD7FE15FE55" LSID="urn:lsid:plazi:treatment:03A487D6FFC4FFD6ADE73C92FB6CFEF1:8ED2601BFFC4FFD6AD9D3DD7FE15FE55" box="[215,510,423,447]" country="Venezuela" latitude="10.98" longLatPrecision="778" longitude="-63.87" municipality="Margarita" name="El Valle de Espirito Santo" pageId="12" pageNumber="12" stateProvince="Nueva Esparta">El Valle de Espírito Santo</location>
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(
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<geoCoordinate id="EE395007FFC4FFD6AF5A3DD7FD9CFE54" box="[528,631,423,447]" degrees="10.98" direction="north" orientation="latitude" pageId="12" pageNumber="12" precision="555" value="10.98">10.98° N</geoCoordinate>
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,
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<geoCoordinate id="EE395007FFC4FFD6AD263DB8FF3EFE0A" box="[108,213,456,480]" degrees="63.87" direction="west" orientation="longitude" pageId="12" pageNumber="12" precision="555" value="-63.87">63.87° W</geoCoordinate>
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; ca.
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<quantity id="4CF59B25FFC4FFD6AC403DB8FEB8FE0A" box="[266,339,456,480]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="12" unit="m" value="200.0">
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<elevation id="0020D1F3FFC4FFD6AC403DB8FEB8FE0A" box="[266,339,456,480]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="12" unit="m" value="200.0">200 m</elevation>
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</quantity>
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) on Isla
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<collectingMunicipality id="6BD6ACBAFFC4FFD6ACF33DB8FDC3FE0A" box="[441,552,456,480]" pageId="12" pageNumber="12">Margarita</collectingMunicipality>
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,
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<collectingRegion id="49C9F822FFC4FFD6AF7F3DB9FF12FDEB" country="Venezuela" name="Nueva Esparta" pageId="12" pageNumber="12">Nueva Esparta state</collectingRegion>
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,
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<collectingCountry id="F31A7650FFC4FFD6AC4F3D99FE9FFDEB" box="[261,372,489,513]" name="Venezuela" pageId="12" pageNumber="12">Venezuela</collectingCountry>
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(
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<bibRefCitation id="EF9C4B31FFC4FFD6ACC83D99FDABFDEB" author="Rossi, R. V. & R. S. Voss & D. P. Lunde" box="[386,576,489,513]" pageId="12" pageNumber="12" pagination="1 - 83" refId="ref50085" refString="Rossi, R. V., R. S. Voss, and D. P. Lunde. 2010. A revision of the didelphid marsupial genus Marmosa. Part 1. The species in Tate's " Mexicana " and " Mitis " sections and other closely related forms. Bulletin of the American Museum of Natural History 334: 1 - 83." type="journal article" year="2010">Rossi et al., 2010</bibRefCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BB236C0FFC4FFD6ADC63E7AFEE1FD4D" blockId="12.[108,636,324,1737]" pageId="12" pageNumber="12">
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SYNONYMS:
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<emphasis id="B979EAD2FFC4FFD6AC5D3E7BFEA5FDC8" box="[279,334,523,546]" italics="true" pageId="12" pageNumber="12">casta</emphasis>
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<bibRefCitation id="EF9C4B31FFC4FFD6AC163E7AFE15FDC8" author="Thomas, O." box="[348,510,522,546]" pageId="12" pageNumber="12" pagination="120 - 158" refId="ref51643" refString="Thomas, O. 1911. The mammals of the tenth edition of Linnaeus; an attempt to fix the types of the genera and the exact bases and localities of the species. Proceedings of the Zoological Society of London 1910: 120 - 158." type="journal article" year="1911">Thomas, 1911</bibRefCitation>
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;
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<emphasis id="B979EAD2FFC4FFD6AF5A3E7AFD97FDC8" box="[528,636,522,546]" italics="true" pageId="12" pageNumber="12">chapmani</emphasis>
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Allen, 1900;
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<emphasis id="B979EAD2FFC4FFD6AC4A3E5BFE9FFDA9" box="[256,372,555,579]" italics="true" pageId="12" pageNumber="12">fulviventer</emphasis>
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Bangs, 1901;
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<emphasis id="B979EAD2FFC4FFD6AF533E5BFD97FDA9" box="[537,636,555,579]" italics="true" pageId="12" pageNumber="12">grenadae</emphasis>
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<bibRefCitation id="EF9C4B31FFC4FFD6AD263E3CFEE3FD8E" author="Thomas, O." box="[108,264,588,612]" pageId="12" pageNumber="12" pagination="120 - 158" refId="ref51643" refString="Thomas, O. 1911. The mammals of the tenth edition of Linnaeus; an attempt to fix the types of the genera and the exact bases and localities of the species. Proceedings of the Zoological Society of London 1910: 120 - 158." type="journal article" year="1911">Thomas, 1911</bibRefCitation>
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;
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<emphasis id="B979EAD2FFC4FFD6AC5F3E3CFE6FFD8E" box="[277,388,588,612]" italics="true" pageId="12" pageNumber="12">luridivolta</emphasis>
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Goodwin, 1961;
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<taxonomicName id="4C0D4D43FFC4FFD6AF0D3E3DFF1AFD6C" authority="Bangs, 1898" authorityName="Bangs" authorityYear="1898" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="12" phylum="Chordata" rank="species" species="mitis">
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<emphasis id="B979EAD2FFC4FFD6AF0D3E3DFD97FD8E" box="[583,636,589,612]" italics="true" pageId="12" pageNumber="12">mitis</emphasis>
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Bangs, 1898
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</taxonomicName>
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;
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<emphasis id="B979EAD2FFC4FFD6ADB73E1EFEAFFD6F" box="[253,324,622,645]" italics="true" pageId="12" pageNumber="12">nesaea</emphasis>
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<bibRefCitation id="EF9C4B31FFC4FFD6AC013E1DFE0EFD6C" author="Thomas, O." box="[331,485,621,646]" pageId="12" pageNumber="12" pagination="120 - 158" refId="ref51643" refString="Thomas, O. 1911. The mammals of the tenth edition of Linnaeus; an attempt to fix the types of the genera and the exact bases and localities of the species. Proceedings of the Zoological Society of London 1910: 120 - 158." type="journal article" year="1911">Thomas, 1911</bibRefCitation>
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;
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<emphasis id="B979EAD2FFC4FFD6ACBB3E1DFD97FD6F" box="[497,636,621,645]" italics="true" pageId="12" pageNumber="12">pallidiventris</emphasis>
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Osgood, 1912.
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</paragraph>
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</subSubSection>
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<subSubSection id="C317654BFFC4FFD6ADC63EC0FEEFFC3A" pageId="12" pageNumber="12" type="distribution">
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<paragraph id="8BB236C0FFC4FFD6ADC63EC0FEEFFC3A" blockId="12.[108,636,324,1737]" pageId="12" pageNumber="12">
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DISTRIBUTION:
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<taxonomicName id="4C0D4D43FFC4FFD6AC0F3EC0FDCFFD22" authorityName="Bangs" authorityYear="1898" box="[325,548,688,712]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="12" phylum="Chordata" rank="species" species="robinsoni">
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<emphasis id="B979EAD2FFC4FFD6AC0F3EC0FDCFFD22" box="[325,548,688,712]" italics="true" pageId="12" pageNumber="12">Marmosa robinsoni</emphasis>
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</taxonomicName>
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occurs primarily in dry forests, but also occasionally in other habitats from western
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<collectingCountry id="F31A7650FFC4FFD6ACE33E82FDEAFCE3" box="[425,513,754,777]" name="Panama" pageId="12" pageNumber="12">Panama</collectingCountry>
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to
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<collectingCountry id="F31A7650FFC4FFD6AF6C3E82FF7BFCC1" name="Colombia" pageId="12" pageNumber="12">Colombia</collectingCountry>
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, northern
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<collectingCountry id="F31A7650FFC4FFD6AC453F63FE6FFCC1" box="[271,388,787,811]" name="Venezuela" pageId="12" pageNumber="12">Venezuela</collectingCountry>
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, and several adjacent continental-shelf islands (including Isla Margarita,
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<collectingCountry id="F31A7650FFC4FFD6ADF23F25FEF1FC87" box="[184,282,853,877]" name="Trinidad and Tobago" pageId="12" pageNumber="12">Trinidad</collectingCountry>
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, and
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<collectingCountry id="F31A7650FFC4FFD6AC1E3F25FE42FC87" box="[340,425,853,877]" name="Trinidad and Tobago" pageId="12" pageNumber="12">Tobago</collectingCountry>
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); the species is also known from
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<collectingCountry id="F31A7650FFC4FFD6AC483F06FE8FFC64" box="[258,356,886,910]" name="Grenada" pageId="12" pageNumber="12">Grenada</collectingCountry>
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, a Caribbean island that is not on the continental shelf (
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<bibRefCitation id="EF9C4B31FFC4FFD6ACA53FE7FF4FFC3A" author="Rossi, R. V. & R. S. Voss & D. P. Lunde" pageId="12" pageNumber="12" pagination="1 - 83" refId="ref50085" refString="Rossi, R. V., R. S. Voss, and D. P. Lunde. 2010. A revision of the didelphid marsupial genus Marmosa. Part 1. The species in Tate's " Mexicana " and " Mitis " sections and other closely related forms. Bulletin of the American Museum of Natural History 334: 1 - 83." type="journal article" year="2010">Rossi et al., 2010</bibRefCitation>
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: fig. 25).
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</paragraph>
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</subSubSection>
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<subSubSection id="C317654BFFC4FFD6ADC63FA9FB6CFEF1" pageId="12" pageNumber="12" type="discussion">
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<paragraph id="8BB236C0FFC4FFD6ADC63FA9FB6CFEF1" blockId="12.[108,636,324,1737]" lastBlockId="12.[684,1211,226,283]" pageId="12" pageNumber="12">
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REMARKS: See
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<bibRefCitation id="EF9C4B31FFC4FFD6AC093FA9FDCFFC1B" author="Rossi, R. V. & R. S. Voss & D. P. Lunde" box="[323,548,985,1009]" pageId="12" pageNumber="12" pagination="1 - 83" refId="ref50085" refString="Rossi, R. V., R. S. Voss, and D. P. Lunde. 2010. A revision of the didelphid marsupial genus Marmosa. Part 1. The species in Tate's " Mexicana " and " Mitis " sections and other closely related forms. Bulletin of the American Museum of Natural History 334: 1 - 83." type="journal article" year="2010">Rossi et al. (2010)</bibRefCitation>
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for an emended description, tabulated measurement data, and morphological comparisons with congeneric species. Several taxa that were formerly treated as subspecies or synonyms of
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<taxonomicName id="4C0D4D43FFC4FFD6AD26380FFEA8FB7D" authorityName="Bangs" authorityYear="1898" box="[108,323,1151,1175]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="12" phylum="Chordata" rank="species" species="robinsoni">
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<emphasis id="B979EAD2FFC4FFD6AD26380FFEA8FB7D" box="[108,323,1151,1175]" italics="true" pageId="12" pageNumber="12">Marmosa robinsoni</emphasis>
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</taxonomicName>
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(e.g., by
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<bibRefCitation id="EF9C4B31FFC4FFD6ACE3380FFD9DFB7C" author="Hershkovitz, P." box="[425,630,1150,1175]" pageId="12" pageNumber="12" pagination="547 - 569" refId="ref47601" refString="Hershkovitz, P. 1951. Mammals from British Honduras, Mexico, Jamaica, and Haiti. Fieldiana Zoology 31: 547 - 569." type="journal article" year="1951">Hershkovitz, 1951</bibRefCitation>
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;
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<bibRefCitation id="EF9C4B31FFC4FFD6AD2638D0FF09FB52" author="Hall, E. R." box="[108,226,1184,1208]" pageId="12" pageNumber="12" refId="ref47452" refString="Hall, E. R. 1981. The mammals of North America (2 nd ed., 2 vols.). New York: John Wiley and Sons." type="book" year="1981">Hall, 1981</bibRefCitation>
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; Creighton and Gardner, 2008a) are now recognized as valid species (
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<taxonomicName id="4C0D4D43FFC4FFD6ACA538B2FD9DFB33" authorityName=", Voss and Giarla" authorityYear="2020" box="[495,630,1217,1241]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="12" phylum="Chordata" rank="species" species="isthmica">
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<emphasis id="B979EAD2FFC4FFD6ACA538B2FD9DFB33" box="[495,630,1217,1241]" italics="true" pageId="12" pageNumber="12">M. isthmica</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C0D4D43FFC4FFD6AD263893FF0CFB10" authorityName="Thomas" authorityYear="1899" box="[108,231,1251,1274]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="12" phylum="Chordata" rank="species" species="simonsi">
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<emphasis id="B979EAD2FFC4FFD6AD263893FF0CFB10" box="[108,231,1251,1274]" italics="true" pageId="12" pageNumber="12">M. simonsi</emphasis>
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</taxonomicName>
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) or have been relegated to the synonymies of other species. Even in its currently restricted sense (
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<bibRefCitation id="EF9C4B31FFC4FFD6AC783954FDEAFAD6" author="Rossi, R. V. & R. S. Voss & D. P. Lunde" box="[306,513,1316,1340]" pageId="12" pageNumber="12" pagination="1 - 83" refId="ref50085" refString="Rossi, R. V., R. S. Voss, and D. P. Lunde. 2010. A revision of the didelphid marsupial genus Marmosa. Part 1. The species in Tate's " Mexicana " and " Mitis " sections and other closely related forms. Bulletin of the American Museum of Natural History 334: 1 - 83." type="journal article" year="2010">Rossi et al., 2010</bibRefCitation>
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), however,
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<taxonomicName id="4C0D4D43FFC4FFD6AD263936FF16FAB7" authorityName="Bangs" authorityYear="1898" box="[108,253,1349,1373]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="12" phylum="Chordata" rank="species" species="robinsoni">
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<emphasis id="B979EAD2FFC4FFD6AD263936FF16FAB7" box="[108,253,1349,1373]" italics="true" pageId="12" pageNumber="12">M. robinsoni</emphasis>
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</taxonomicName>
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is a geographically and ecologically widespread species that includes numerous nominal taxa as subjective synonyms. Analyses of DNA sequence data (
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<bibRefCitation id="EF9C4B31FFC4FFD6ACBA39D8FF30FA08" author="Gutierrez, E. E." pageId="12" pageNumber="12" pagination="1175 - 1188" refId="ref47358" refString="Gutierrez, E. E., et al. [five additional coauthors]. 2014 a. Phylogeography of Marmosa robinsoni: insights into the biogeography of dry forests in northern South America. Journal of Mammalogy 95: 1175 - 1188." type="journal article" year="2014">Gutiérrez et al., 2014a</bibRefCitation>
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) have shown that geographic populations of this species sort out into two strongly supported phylogroups: an eastern clade for which the oldest available trinomen would be
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<taxonomicName id="4C0D4D43FFC4FFD6AD263A3EFE82F98C" authorityName="Bangs" authorityYear="1898" box="[108,361,1614,1638]" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="12" phylum="Chordata" rank="subSpecies" species="robinsoni" subSpecies="robinsoni">
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<emphasis id="B979EAD2FFC4FFD6AD263A3EFE82F98C" box="[108,361,1614,1638]" italics="true" pageId="12" pageNumber="12">M. robinsoni robinsoni</emphasis>
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</taxonomicName>
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, and a western clade for which the oldest trinomen would be
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<taxonomicName id="4C0D4D43FFC4FFD6AF753A00FF48F942" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageId="12" pageNumber="12" phylum="Chordata" rank="subSpecies" subSpecies="mitis">
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<emphasis id="B979EAD2FFC4FFD6AF753A00FF48F942" italics="true" pageId="12" pageNumber="12">M. r. mitis</emphasis>
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</taxonomicName>
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. Unfortunately, these putative subspecies appear to be phenotypically indistinguishable, such that specimens cannot be assigned to one or the other without DNA sequence data.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |