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<document ID-DOI="10.5252/geodiversitas2019v41a15" ID-GBIF-Dataset="1e89d723-7f54-4d6b-85dd-6bcd573eca32" ID-ISSN="1638-9395" ID-Zenodo-Dep="3694209" ID-ZooBank="urn:lsid:zoobank.org:pub:9DD3CC29-3AEA-44B8-8E8F-6AD882DF5B1C" checkinTime="1583172268754" checkinUser="valdenar" docAuthor="Bonis, Louis de, Gardin, Axelle &amp; Blondel, Cécile" docDate="2019" docId="03A487991A64FF9BFBAEF8BE6C95FC1F" docLanguage="en" docName="Geodiversitas.41.15.601-621.pdf.imf" docOrigin="Geodiversitas 41 (15)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Amphicynodon sp. 2 Filhol 1881" docType="treatment" docVersion="9" lastPageNumber="604" masterDocId="FF9DFFE11A60FF9EFFADFF8D6E62FF8E" masterDocTitle="Carnivora from the early Oligocene of the Phosphorites du Quercy in southwestern France" masterLastPageNumber="621" masterPageNumber="601" pageNumber="603" updateTime="1675702587573" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo>
<mods:title>Carnivora from the early Oligocene of the Phosphorites du Quercy in southwestern France</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bonis, Louis de</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">louis.debonis@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gardin, Axelle</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">axelle.gardin@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Blondel, Cécile</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">cecile.blondel@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
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<mods:titleInfo>
<mods:title>Geodiversitas</mods:title>
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<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-09-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>41</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>15</mods:number>
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<mods:start>601</mods:start>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/geodiversitas2019v41a15</mods:identifier>
<mods:identifier type="GBIF-Dataset">1e89d723-7f54-4d6b-85dd-6bcd573eca32</mods:identifier>
<mods:identifier type="ISSN">1638-9395</mods:identifier>
<mods:identifier type="Zenodo-Dep">3694209</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:9DD3CC29-3AEA-44B8-8E8F-6AD882DF5B1C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3703501" ID-GBIF-Taxon="162367541" ID-Zenodo-Dep="3703501" LSID="urn:lsid:plazi:treatment:03A487991A64FF9BFBAEF8BE6C95FC1F" httpUri="http://treatment.plazi.org/id/03A487991A64FF9BFBAEF8BE6C95FC1F" lastPageId="5" lastPageNumber="604" pageId="4" pageNumber="603">
<subSubSection box="[1027,1240,1843,1870]" pageId="4" pageNumber="603" type="nomenclature">
<paragraph blockId="4.[1027,1240,1843,1902]" box="[1027,1240,1843,1870]" pageId="4" pageNumber="603">
<heading box="[1027,1240,1843,1870]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="603" reason="2">
<taxonomicName authorityName="Filhol" authorityYear="1881" box="[1027,1240,1843,1870]" class="Mammalia" family="Ursidae" genus="Amphicynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="604" phylum="Chordata" rank="species" species="sp. 2">
<emphasis box="[1027,1182,1843,1869]" italics="true" pageId="4" pageNumber="603">Amphicynodon</emphasis>
sp. 2
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1083,1186,1875,1902]" pageId="4" pageNumber="603" type="description">
<paragraph blockId="4.[1027,1240,1843,1902]" box="[1083,1186,1875,1902]" pageId="4" pageNumber="603">
(
<figureCitation box="[1092,1176,1875,1902]" captionStart="FIG" captionStartId="6.[132,143,1964,1982]" captionTargetBox="[212,1402,209,1931]" captionTargetId="figure@6.[755,1301,780,1349]" captionTargetPageId="6" captionText="FIG. 1. — A, Amphicynodon sp. 1: left hemi-mandible UP VAL2, in buccal (A1), lingual (A2), and occlusal (A3) views; B, Amphicynodon typicus: left mandible UP VAL1, in buccal (B1), lingual (B2), and occlusal (B3) views; C, Amphicynodon sp. 3: right M1, UM VD49, in occlusal view; D, Amphicynodon sp. 2: right M1, UM VBO492, in occlusal view. Scale bars: A, B, 10 mm; C, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3694211" httpUri="https://zenodo.org/record/3694211/files/figure.png" pageId="4" pageNumber="603">Fig. 1D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="604" pageId="4" pageNumber="603" type="description">
<paragraph blockId="4.[813,1456,1939,2030]" box="[813,957,1939,1965]" pageId="4" pageNumber="603">DESCRIPTION</paragraph>
<paragraph blockId="4.[813,1456,1939,2030]" pageId="4" pageNumber="603">
A right M1 (UM VBO492) from Valbro (
<bibRefCitation author="Peigne" box="[1260,1448,1971,1997]" etAl="et al." firstAuthor="Peigne" pageId="4" pageNumber="603" pagination="1 - 45" refId="ref18618" refString="PEIGNE S., VIANEY- LIAUD M., PELISSIE T. &amp; SIGE B. 2014. -- Valbro: un nouveau site a vertebres de l'Oligocene inferieur (MP 22) de France (Quercy). I - Contexte geologique; Mammalia: Rodentia, Hyaenodontida, Carnivora. Annales de Paleontologie 100: 1 - 45. https: // doi. org / 10.1016 / j. annpal. 2013.11.002" type="journal article" year="2014">
Peigné
<emphasis box="[1336,1387,1971,1997]" italics="true" pageId="4" pageNumber="603">et al.</emphasis>
2014
</bibRefCitation>
: fig. 22b) has an overall pattern indicating
<taxonomicName authorityName="Filhol" authorityYear="1881" box="[1296,1456,2003,2029]" class="Mammalia" family="Ursidae" genus="Amphicynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis box="[1296,1456,2003,2029]" italics="true" pageId="4" pageNumber="603">Amphicynodon</emphasis>
</taxonomicName>
</paragraph>
<caption ID-Table-UUID="DF7266071A65FF9BFE35FF526AFFFF7C" box="[408,1181,222,242]" httpUri="http://table.plazi.org/id/DF7266071A65FF9BFE35FF526AFFFF7C" pageId="5" pageNumber="604" startId="5.[408,419,223,241]" targetBox="[134,1442,300,469]" targetIsTable="true" targetPageId="5">
<paragraph blockId="5.[408,1181,222,242]" box="[408,1181,222,242]" pageId="5" pageNumber="604">
TABLE 1. — Measurements (in mm) of teeth of
<taxonomicName authority="Filhol, 1881" authorityName="Filhol" authorityYear="1881" box="[789,1017,223,241]" class="Mammalia" family="Ursidae" genus="Amphicynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis box="[789,915,223,241]" italics="true" pageId="5" pageNumber="604">Amphicynodon</emphasis>
Filhol, 1881
</taxonomicName>
, from the alveoli.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="604">
<table box="[134,1442,300,469]" gridcols="16" gridrows="4" pageId="5" pageNumber="604">
<tr box="[134,1442,300,320]" gridrow="0" pageId="5" pageNumber="604">
<th box="[134,287,300,320]" gridcol="0" gridrow="0" pageId="5" pageNumber="604">
<taxonomicName authorityName="Filhol" authorityYear="1881" box="[134,287,300,320]" class="Mammalia" family="Ursidae" genus="Amphicynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="604" phylum="Chordata" rank="genus">
<emphasis bold="true" box="[134,287,300,320]" italics="true" pageId="5" pageNumber="604">Amphicynodon</emphasis>
</taxonomicName>
</th>
<th box="[344,392,300,320]" gridcol="1" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[348,385,300,320]" pageId="5" pageNumber="604">p2L</emphasis>
</th>
<th box="[411,472,300,320]" gridcol="2" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[420,462,300,320]" pageId="5" pageNumber="604">p2w</emphasis>
</th>
<th box="[486,547,300,320]" gridcol="3" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[498,535,300,320]" pageId="5" pageNumber="604">p3L</emphasis>
</th>
<th box="[571,614,300,320]" gridcol="4" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[571,613,300,320]" pageId="5" pageNumber="604">p3w</emphasis>
</th>
<th box="[648,690,300,320]" gridcol="5" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[648,685,300,320]" pageId="5" pageNumber="604">p4L</emphasis>
</th>
<th box="[721,765,300,320]" gridcol="6" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[721,763,300,320]" pageId="5" pageNumber="604">p4w</emphasis>
</th>
<th box="[792,840,300,320]" gridcol="7" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[795,839,300,320]" pageId="5" pageNumber="604">m1L</emphasis>
</th>
<th box="[862,922,300,320]" gridcol="8" gridrow="0" pageId="5" pageNumber="604">m1Ltr</th>
<th box="[935,999,300,320]" gridcol="9" gridrow="0" pageId="5" pageNumber="604">m1wtr</th>
<th box="[1012,1074,300,320]" gridcol="10" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[1012,1074,300,320]" pageId="5" pageNumber="604">m1wtl</emphasis>
</th>
<th box="[1096,1140,300,320]" gridcol="11" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[1096,1139,300,320]" pageId="5" pageNumber="604">m2L</emphasis>
</th>
<th box="[1161,1225,300,320]" gridcol="12" gridrow="0" pageId="5" pageNumber="604">m2wtr</th>
<th box="[1237,1299,300,320]" gridcol="13" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[1237,1299,300,320]" pageId="5" pageNumber="604">m2wtl</emphasis>
</th>
<th box="[1321,1366,300,320]" gridcol="14" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[1321,1365,300,320]" pageId="5" pageNumber="604">m3L</emphasis>
</th>
<th box="[1394,1442,300,320]" gridcol="15" gridrow="0" pageId="5" pageNumber="604">
<emphasis bold="true" box="[1394,1442,300,320]" pageId="5" pageNumber="604">m3w</emphasis>
</th>
</tr>
<tr box="[134,1442,335,379]" gridrow="1" pageId="5" pageNumber="604">
<th box="[134,287,335,379]" gridcol="0" gridrow="1" pageId="5" pageNumber="604">UP VAL1 UP VAL2</th>
<td box="[344,392,335,379]" gridcol="1" gridrow="1" pageId="5" pageNumber="604"> 4.86</td>
<td box="[411,472,335,379]" gridcol="2" gridrow="1" pageId="5" pageNumber="604"> 2.3</td>
<td box="[486,547,335,379]" gridcol="3" gridrow="1" pageId="5" pageNumber="604">6.68 4.61</td>
<td box="[571,614,335,379]" gridcol="4" gridrow="1" pageId="5" pageNumber="604">3.33 2.33</td>
<td box="[648,690,335,379]" gridcol="5" gridrow="1" pageId="5" pageNumber="604">6.98 7.09</td>
<td box="[721,765,335,379]" gridcol="6" gridrow="1" pageId="5" pageNumber="604">3.33 2.82</td>
<td box="[792,840,335,379]" gridcol="7" gridrow="1" pageId="5" pageNumber="604">(8.1) 7.99</td>
<td box="[862,922,335,379]" gridcol="8" gridrow="1" pageId="5" pageNumber="604"> 5.56</td>
<td box="[935,999,335,379]" gridcol="9" gridrow="1" pageId="5" pageNumber="604"> 4.17</td>
<td box="[1012,1074,335,379]" gridcol="10" gridrow="1" pageId="5" pageNumber="604"> 3.83</td>
<td box="[1096,1140,335,379]" gridcol="11" gridrow="1" pageId="5" pageNumber="604">6.23 4.27</td>
<td box="[1161,1225,335,379]" gridcol="12" gridrow="1" pageId="5" pageNumber="604">6.23 3.47</td>
<td box="[1237,1299,335,379]" gridcol="13" gridrow="1" pageId="5" pageNumber="604">4.1 2.89</td>
<td box="[1321,1366,335,379]" gridcol="14" gridrow="1" pageId="5" pageNumber="604">3.59 2.43</td>
<td box="[1394,1442,335,379]" gridcol="15" gridrow="1" pageId="5" pageNumber="604"> 2.01</td>
</tr>
<tr box="[134,1442,395,415]" gridrow="2" pageId="5" pageNumber="604" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td box="[344,392,395,415]" gridcol="1" gridrow="2" pageId="5" pageNumber="604">M1L</td>
<td box="[411,472,395,415]" gridcol="2" gridrow="2" pageId="5" pageNumber="604">M1w1</td>
<td box="[486,547,395,415]" gridcol="3" gridrow="2" pageId="5" pageNumber="604">
<emphasis bold="true" box="[486,547,395,415]" pageId="5" pageNumber="604">M1w2</emphasis>
</td>
</tr>
<tr box="[134,1442,424,469]" gridrow="3" pageId="5" pageNumber="604" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th box="[134,287,424,469]" gridcol="0" gridrow="3" pageId="5" pageNumber="604">UM VBO492 UM VD49</th>
<td box="[344,392,424,469]" gridcol="1" gridrow="3" pageId="5" pageNumber="604">6.5 6.9</td>
<td box="[411,472,424,469]" gridcol="2" gridrow="3" pageId="5" pageNumber="604">9.2 10.3</td>
<td box="[486,547,424,469]" gridcol="3" gridrow="3" pageId="5" pageNumber="604">
<tableNote box="[497,527,425,469]" pageId="5" pageNumber="604" targetBox="[134,1442,300,379]" targetPageId="5">8.8 8.3</tableNote>
</td>
</tr>
</table>
</paragraph>
<paragraph blockId="5.[132,776,536,913]" pageId="5" pageNumber="604">
morphology. Thus UM VBO492, shares with the M1 of
<emphasis box="[132,310,568,594]" italics="true" pageId="5" pageNumber="604">A. leptorhynchus</emphasis>
(
<bibRefCitation author="Filhol" box="[326,468,568,595]" firstAuthor="Filhol" pageId="5" pageNumber="604" pagination="1 - 338" refId="ref17470" refString="FILHOL H. 1876 - 1877. - Recherches sur les phosphorites du Quercy. Etudes des fossiles qu'on y rencontre et specialement des mammiferes. Bibliotheque de l'Ecole des Hautes Etudes, Section des Sciences naturelles. Premiere partie, 15 (4): 1 - 220. Deuxieme partie 16 (1): 1 - 338." type="journal article" year="1876">Filhol, 1876</bibRefCitation>
) found in the locality Itardies (
<bibRefCitation author="Cirot &amp; Bonis" box="[193,421,600,626]" firstAuthor="Cirot" pageId="5" pageNumber="604" pagination="103 - 130" refId="ref17043" refString="CIROT E. &amp; BONIS L. DE 1992. - Revision du genre Amphicynodon, carnivore de l'Oligocene. Palaeontographica 220 (4 - 6): 103 - 130." type="journal article" year="1992">Cirot &amp; Bonis 1992</bibRefCitation>
), a similar aspect of the trigon, the presence of a metaconule and a large lingual cingulum (
<figureCitation box="[141,231,664,690]" captionStart="FIG" captionStartId="6.[132,143,1964,1982]" captionTargetBox="[212,1402,209,1931]" captionTargetId="figure@6.[755,1301,780,1349]" captionTargetPageId="6" captionText="FIG. 1. — A, Amphicynodon sp. 1: left hemi-mandible UP VAL2, in buccal (A1), lingual (A2), and occlusal (A3) views; B, Amphicynodon typicus: left mandible UP VAL1, in buccal (B1), lingual (B2), and occlusal (B3) views; C, Amphicynodon sp. 3: right M1, UM VD49, in occlusal view; D, Amphicynodon sp. 2: right M1, UM VBO492, in occlusal view. Scale bars: A, B, 10 mm; C, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3694211" httpUri="https://zenodo.org/record/3694211/files/figure.png" pageId="5" pageNumber="604">Fig. 1D</figureCitation>
). On the other hand, it is narrower distally than mesially, indicating a reduction of the distal part of the jaw. This character is present in
<emphasis box="[429,550,727,754]" italics="true" pageId="5" pageNumber="604">A. velaunus</emphasis>
and
<emphasis box="[604,723,727,754]" italics="true" pageId="5" pageNumber="604">A. teilhardi</emphasis>
, but the outline of the tooth is different, with a greater length of the buccal border in those two species. We do not know if in our material there is a lower dentition corresponding to UM VBO492 and thus we cannot assign a species name to this M1, although
<taxonomicName baseAuthorityName="Schlosser" baseAuthorityYear="1888" box="[289,519,887,913]" class="Mammalia" family="Ursidae" genus="Amphicynodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="5" pageNumber="604" phylum="Chordata" rank="species" species="typicus">
<emphasis box="[289,519,887,913]" italics="true" pageId="5" pageNumber="604">Amphicynodon typicus</emphasis>
</taxonomicName>
could be a possibility.
</paragraph>
</subSubSection>
</treatment>
</document>