treatments-xml/data/03/92/77/0392774B78431075B6DD55D0FB98F90C.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.3791602" ID-GBIF-Dataset="2bb0a745-07bd-4df6-99e7-4d8a6b242aa5" ID-GBIF-Taxon="163977044" ID-ISSN="13132970" ID-Zenodo-Dep="3791602" ID-ZooBank="DB5973A9-8CF6-400B-87C4-7A4521BD3117" checkinTime="1587971617180" checkinUser="plazi" docAuthor="Dunlop, Jason &amp; Mitov, Plamen" docDate="2009" docId="0392774B78431075B6DD55D0FB98F90C" docLanguage="en" docName="ZK_article_2024.pdf" docOrigin="ZooKeys 16" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@18c8ecf5" docStyleName="zookeys.2008.journal_article" docTitle="Leiobunum longipes Menge 1854" docType="treatment" docVersion="4" lastPageNumber="369" masterDocId="FFAB0F3378551063B6515743FFCCFF8B" masterDocTitle="Fossil harvestmen (Arachnida, Opiliones) from Bitterfeld amber" masterLastPageNumber="375" masterPageNumber="347" pageId="22" pageNumber="369" updateTime="1643568438625" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Fossil harvestmen (Arachnida, Opiliones) from Bitterfeld amber</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dunlop, Jason</mods:namePart>
<mods:affiliation>Museum für Naturkunde, Berlin, Berlin, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mitov, Plamen</mods:namePart>
<mods:affiliation>University of Sofia, Sofia ,, Bulgaria</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-07-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>16</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>347</mods:start>
<mods:end>375</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.16.224</mods:identifier>
<mods:identifier type="GBIF-Dataset">2bb0a745-07bd-4df6-99e7-4d8a6b242aa5</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576490</mods:identifier>
<mods:identifier type="ZooBank">DB5973A9-8CF6-400B-87C4-7A4521BD3117</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791602" ID-GBIF-Taxon="163977044" ID-Zenodo-Dep="3791602" LSID="urn:lsid:plazi:treatment:0392774B78431075B6DD55D0FB98F90C" httpUri="http://treatment.plazi.org/id/0392774B78431075B6DD55D0FB98F90C" lastPageNumber="369" pageId="22" pageNumber="369">
<subSubSection box="[140,543,659,686]" pageId="22" pageNumber="369" type="nomenclature">
<paragraph blockId="22.[140,543,659,721]" box="[140,543,659,686]" pageId="22" pageNumber="369">
<heading box="[140,543,659,686]" fontSize="11" level="2" pageId="22" pageNumber="369" reason="3">
<taxonomicName authority="Menge, 1854" authorityName="Menge" authorityYear="1854" box="[140,543,659,686]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="22" pageNumber="369" phylum="Arthropoda" rank="species" species="longipes">
<emphasis bold="true" box="[140,543,659,686]" pageId="22" pageNumber="369">
<emphasis bold="true" box="[140,379,659,686]" italics="true" pageId="22" pageNumber="369">Leiobunum longipes</emphasis>
Menge, 1854
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,265,695,721]" pageId="22" pageNumber="369" type="description">
<paragraph blockId="22.[140,543,659,721]" box="[140,265,695,721]" pageId="22" pageNumber="369">
<figureCitation box="[140,224,695,721]" captionStart="Figures 27-30. 27" captionStartId="23.[139,221,1558,1580]" captionTargetBox="[140,1108,165,1538]" captionTargetId="figure@23.[136,1109,165,1539]" captionTargetPageId="23" captionText="Figures 27-30. 27 Fossil?Lacinius erinaceus Staręga, 1966, MB.A. 1661, from Bitterfeld amber (Palaeogene: Oligocene) 28-29 Recent male Lacinius erinaceus Staręga, 1966 (Holotype) in dorsal and lateral view for comparison, revealing an almost identical morphology to the amber fossil 30 Leiobunum longipes Menge, 1854 from Bitterfeld amber, MB.A. 1662. Scale bars equal 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3814848" httpUri="https://zenodo.org/record/3814848/files/figure.png" pageId="22" pageNumber="369">Figs 30</figureCitation>
,
<figureCitation box="[236,265,695,721]" captionStart="Figures 31-33" captionStartId="24.[140,222,1558,1580]" captionTargetBox="[185,1080,177,1537]" captionTargetId="figure@24.[139,1108,165,1541]" captionTargetPageId="24" captionText="Figures 31-33. Camera lucida drawings of the specimens shown in 27 and 30. 31?Lacinius erinaceus Staręga, 1966, MB.A. 1661 32 the same detail of ocularium and its spination 33 Leiobunum longipes Menge, 1854, MB.A. 1662. Both from Bitterfeld amber (Palaeogene: Oligocene). Scale bars equal 1.0 mm (31, 33)." figureDoi="http://doi.org/10.5281/zenodo.3814850" httpUri="https://zenodo.org/record/3814850/files/figure.png" pageId="22" pageNumber="369">33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[140,661,764,792]" pageId="22" pageNumber="369" type="reference_group">
<paragraph blockId="22.[140,1108,764,1672]" box="[140,661,764,792]" pageId="22" pageNumber="369">
<emphasis bold="true" box="[140,272,764,790]" pageId="22" pageNumber="369">
<emphasis box="[140,267,764,790]" inLineHeading="true" pageId="22" pageNumber="369" reason="1">Synonymy</emphasis>
.
</emphasis>
See
<bibRefCitation author="Dunlop" box="[324,492,765,791]" firstAuthor="Dunlop" pageId="22" pageNumber="369" pagination="167 - 182" refId="ref10654" refString="Dunlop JA (2006) Baltic amber harvestman types (Arachnida: Opiliones: Eupnoi and Dyspnoi). Fossil Record 9: 167 - 182." type="journal article" year="2006">Dunlop (2006</bibRefCitation>
, pp. 172-173).
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="369" type="materials_examined">
<paragraph blockId="22.[140,1108,764,1672]" pageId="22" pageNumber="369">
<materialsCitation ID-GBIF-Occurrence="2610756302" country="Poland" county="Eocene" location="Probably" municipality="Palaeogene" pageId="22" pageNumber="369" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[185,306,800,826]" pageId="22" pageNumber="369">
<emphasis box="[185,301,800,826]" inLineHeading="true" pageId="22" pageNumber="369" reason="1">
<typeStatus box="[185,301,800,826]" pageId="22" pageNumber="369">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<location LSID="urn:lsid:plazi:treatment:0392774B78431075B6DD55D0FB98F90C:8EE4908678431075B7695463FE50FCB1" box="[312,412,800,826]" country="Poland" county="Eocene" municipality="Palaeogene" name="Probably" pageId="22" pageNumber="369">Probably</location>
originally in
<location LSID="urn:lsid:plazi:treatment:0392774B78431075B6DD55D0FB98F90C:8EE4908678431075B4635463FD48FCB1" box="[562,644,800,826]" country="Poland" county="Eocene" municipality="Palaeogene" name="Danzig" pageId="22" pageNumber="369">Danzig</location>
(=
<location LSID="urn:lsid:plazi:treatment:0392774B78431075B6DD55D0FB98F90C:8EE4908678431075B4F35463FD32FCB1" box="[674,766,800,826]" country="Poland" county="Eocene" municipality="Palaeogene" name="Gdansk" pageId="22" pageNumber="369">Gdańsk</location>
,
<collectingCountry box="[776,859,800,827]" name="Poland" pageId="22" pageNumber="369">Poland</collectingCountry>
), current whereabouts unknown (see also
<bibRefCitation author="Dunlop" box="[364,527,835,862]" firstAuthor="Dunlop" pageId="22" pageNumber="369" pagination="167 - 182" refId="ref10654" refString="Dunlop JA (2006) Baltic amber harvestman types (Arachnida: Opiliones: Eupnoi and Dyspnoi). Fossil Record 9: 167 - 182." type="journal article" year="2006">Dunlop 2006</bibRefCitation>
), from
<location LSID="urn:lsid:plazi:treatment:0392774B78431075B6DD55D0FB98F90C:8EE4908678431075B4395400FD65FCD6" box="[616,681,835,861]" country="Poland" county="Eocene" municipality="Palaeogene" name="Baltic" pageId="22" pageNumber="369">Baltic</location>
amber (
<collectingMunicipality box="[779,908,835,861]" pageId="22" pageNumber="369">Palaeogene</collectingMunicipality>
,
<collectingCounty box="[922,1008,836,862]" pageId="22" pageNumber="369">Eocene</collectingCounty>
); precise locality unclear.
</materialsCitation>
</paragraph>
<paragraph blockId="22.[140,1108,764,1672]" pageId="22" pageNumber="369">
<emphasis bold="true" box="[185,426,905,931]" pageId="22" pageNumber="369">
<emphasis box="[185,313,905,931]" inLineHeading="true" pageId="22" pageNumber="369" reason="1">Additional</emphasis>
material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2610756304" collectionCode="MB" country="Germany" county="Bitterfeld" location="Goitsche" municipality="Mine" pageId="22" pageNumber="369" specimenCount="1" stateProvince="Sachsen-Anhalt">
<collectionCode box="[432,480,906,932]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="22" pageNumber="369">MB</collectionCode>
.A. 1662 (series 15/10, provisionally labelled “Gyantinae cf.
<taxonomicName authorityName="Martens" authorityYear="1969" box="[174,275,941,967]" class="Arachnida" family="Phalangiidae" genus="Amilenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="22" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis box="[174,275,941,967]" italics="true" pageId="22" pageNumber="369">Amilenus</emphasis>
</taxonomicName>
” but not in fact referable to this genus; see above), probably from the site of the
<location LSID="urn:lsid:plazi:treatment:0392774B78431075B6DD55D0FB98F90C:8EE4908678431075B6825493FEFBFC61" box="[211,311,976,1002]" country="Germany" county="Bitterfeld" municipality="Mine" name="Goitsche" pageId="22" pageNumber="369" stateProvince="Sachsen-Anhalt">Goitsche</location>
open-cast
<collectingMunicipality box="[432,493,976,1002]" pageId="22" pageNumber="369">Mine</collectingMunicipality>
near
<collectingCounty box="[554,664,976,1002]" pageId="22" pageNumber="369">Bitterfeld</collectingCounty>
,
<collectingRegion box="[675,854,976,1002]" country="Germany" name="Sachsen-Anhalt" pageId="22" pageNumber="369">Sachsen-Anhalt</collectingRegion>
,
<collectingCountry box="[864,973,976,1002]" name="Germany" pageId="22" pageNumber="369">Germany</collectingCountry>
</materialsCitation>
; Palaeogene (Oligocene: Chattian).
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="369" type="description">
<paragraph blockId="22.[140,1108,764,1672]" pageId="22" pageNumber="369">
<emphasis bold="true" box="[185,335,1046,1072]" pageId="22" pageNumber="369">
<emphasis box="[185,330,1046,1072]" inLineHeading="true" pageId="22" pageNumber="369" reason="1">Description</emphasis>
.
</emphasis>
MB.A. 1656 (
<figureCitation box="[517,608,1047,1073]" captionStart="Figures 27-30. 27" captionStartId="23.[139,221,1558,1580]" captionTargetBox="[140,1108,165,1538]" captionTargetId="figure@23.[136,1109,165,1539]" captionTargetPageId="23" captionText="Figures 27-30. 27 Fossil?Lacinius erinaceus Staręga, 1966, MB.A. 1661, from Bitterfeld amber (Palaeogene: Oligocene) 28-29 Recent male Lacinius erinaceus Staręga, 1966 (Holotype) in dorsal and lateral view for comparison, revealing an almost identical morphology to the amber fossil 30 Leiobunum longipes Menge, 1854 from Bitterfeld amber, MB.A. 1662. Scale bars equal 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3814848" httpUri="https://zenodo.org/record/3814848/files/figure.png" pageId="22" pageNumber="369">Figs 30</figureCitation>
,
<figureCitation box="[624,654,1047,1073]" captionStart="Figures 31-33" captionStartId="24.[140,222,1558,1580]" captionTargetBox="[185,1080,177,1537]" captionTargetId="figure@24.[139,1108,165,1541]" captionTargetPageId="24" captionText="Figures 31-33. Camera lucida drawings of the specimens shown in 27 and 30. 31?Lacinius erinaceus Staręga, 1966, MB.A. 1661 32 the same detail of ocularium and its spination 33 Leiobunum longipes Menge, 1854, MB.A. 1662. Both from Bitterfeld amber (Palaeogene: Oligocene). Scale bars equal 1.0 mm (31, 33)." figureDoi="http://doi.org/10.5281/zenodo.3814850" httpUri="https://zenodo.org/record/3814850/files/figure.png" pageId="22" pageNumber="369">33</figureCitation>
) is an almost complete specimen of a fairly large harvestman in dorsal view. Body length:
<quantity box="[742,837,1082,1108]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="22" pageNumber="369" unit="mm" value="2.1">2.1 mm</quantity>
; width of prosoma and opisthosoma
<quantity box="[294,393,1117,1144]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="22" pageNumber="369" unit="mm" value="1.3">1.3 mm</quantity>
. Carapace procurved anteriorly, with clear division into promeso- and metapeltidium, and centrally located ocular tubercle. Length of ocularium
<quantity box="[140,263,1187,1214]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" pageId="22" pageNumber="369" unit="mm" value="0.225">0.225 mm</quantity>
, width
<quantity box="[349,472,1187,1214]" metricMagnitude="-4" metricUnit="m" metricValue="2.75" pageId="22" pageNumber="369" unit="mm" value="0.275">0.275 mm</quantity>
, distance from ocular tubercle to front of prosoma
<quantity metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="22" pageNumber="369" unit="mm" value="0.25">0.25 mm</quantity>
. Opisthosoma covered by scutum parvum bearing in places slightly granular ornament; posterior region and any segmentation here obscure. Body covered with small black dots and a subcuticular, silvery pigment. Chelicerae equivocal. Pedipalps short and pediform, but details lacking. Legs relatively complete, at least proximally, elongate and slender. Leg 1 femur length
<quantity box="[562,670,1364,1390]" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="22" pageNumber="369" unit="mm" value="1.83">1.83 mm</quantity>
, patella, 0.24. Leg 2 only known from incomplete femora. Leg 3 complete with podomere lengths (in mm) of femur, 1.79; patella, 0.45; tibia, 1.52; metatarsus, 1.86, tarsus, 3.20. Tarsus divided into 24 individual elements, becoming shorter distally, and ending in a single claw (apotele). Leg 4 only known from incomplete femora.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="369" type="discussion">
<paragraph blockId="22.[140,1108,764,1672]" pageId="22" pageNumber="369">
<emphasis bold="true" box="[185,296,1539,1565]" pageId="22" pageNumber="369">
<emphasis box="[185,291,1539,1565]" inLineHeading="true" pageId="22" pageNumber="369" reason="1">Remarks</emphasis>
.
</emphasis>
<taxonomicName authority="Menge, 1854" authorityName="Menge" authorityYear="1854" box="[306,675,1539,1566]" class="Arachnida" family="Phalangiidae" genus="Leiobunum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="22" pageNumber="369" phylum="Arthropoda" rank="species" species="longipes">
<emphasis box="[306,518,1539,1565]" italics="true" pageId="22" pageNumber="369">Leiobunum longipes</emphasis>
Menge, 1854
</taxonomicName>
is present in both Baltic and Bitterfeld amber. Apparently quite common in Baltic amber (
<bibRefCitation author="Starega" box="[698,845,1575,1601]" firstAuthor="Starega" pageId="22" pageNumber="369" pagination="601 - 604" refId="ref11794" refString="Starega W (2002) Baltic amber harvestmen (Opiliones) from Polish collections. Annales Zoologici 52: 601 - 604." type="journal article" year="2002">Staręga 2002</bibRefCitation>
), this fossil is fairly large compared to the other Bitterfeld finds. Nevertheless, it is probably still only a juvenile, although we see nothing which differentiates it from the species known from Baltic amber.
</paragraph>
</subSubSection>
</treatment>
</document>