644 lines
54 KiB
XML
644 lines
54 KiB
XML
<document ID-DOI="10.5281/ZENODO.30012" ID-GBIF-Dataset="9c0666f8-85bf-45bd-b77a-fec54d7e88cc" ID-GBIF-Taxon="117164520" checkinTime="1456617533890" checkinUser="donat" docAuthor="Peter J. Schwendinger & Ondřej Košulič" docDate="2015" docId="03917400BD50D73DD2072F11FA2DF874" docLanguage="en" docName="423_436_Schwendiger.pdf.gs.imf" docOrigin="Revue suisse de Zoologie 122 (2)" docStyle="DocumentStyle{}" docTitle="Perania annam Schwendinger & Košulič, 2015, sp. nov." docType="treatment" docVersion="5" lastPageId="4" lastPageNumber="427" masterDocId="FFA80C78BD52D739D3452D60FFACFF9C" masterDocTitle="Two new species of armoured spiders from Vietnam and Cambodia (Araneae: Tetrablemmidae: Pacullinae)" masterLastPageNumber="436" masterPageNumber="423" pageId="2" pageNumber="425" updateTime="1643505865877" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of armoured spiders from Vietnam and Cambodia (Araneae: Tetrablemmidae: Pacullinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Peter J. Schwendinger</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ondřej Košulič</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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||
<mods:titleInfo>
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||
<mods:title>Revue suisse de Zoologie</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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||
<mods:date>2015</mods:date>
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||
<mods:detail type="volume">
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||
<mods:number>122</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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||
<mods:number>2</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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||
<mods:start>423</mods:start>
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||
<mods:end>436</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/ZENODO.30012</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="117164520" LSID="urn:lsid:plazi:treatment:03917400BD50D73DD2072F11FA2DF874" httpUri="http://treatment.plazi.org/id/03917400BD50D73DD2072F11FA2DF874" lastPageId="4" lastPageNumber="427" pageId="2" pageNumber="425">
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<subSubSection box="[322,585,625,650]" pageId="2" pageNumber="425" type="nomenclature">
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||
<paragraph box="[322,585,625,650]" pageId="2" pageNumber="425">
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||
<emphasis bold="true" box="[322,585,625,650]" pageId="2" pageNumber="425">
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||
<heading bold="true" box="[322,585,625,650]" fontSize="10" level="2" pageId="2" pageNumber="425" reason="6">
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||
<emphasis bold="true" box="[322,493,625,648]" italics="true" pageId="2" pageNumber="425">
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<taxonomicName box="[322,493,625,648]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="425" phylum="Arthropoda" rank="species" species="annam" status="sp. nov.">Perania annam</taxonomicName>
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</emphasis>
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<taxonomicNameLabel box="[500,585,626,650]" pageId="2" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
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||
</heading>
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||
</emphasis>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[387,520,657,682]" pageId="2" pageNumber="425" type="description">
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||
<paragraph box="[387,520,657,682]" pageId="2" pageNumber="425">
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||
<figureCitation box="[387,477,657,682]" captionStart-0="Fig. 2" captionStart-1="Fig. 3" captionStartId-0="2.[151,186,1998,2019]" captionStartId-1="3.[151,186,1326,1347]" captionTargetBox-0="[177,1419,1104,1949]" captionTargetBox-1="[190,1394,260,1265]" captionTargetId-0="figure@2.[151,1436,1083,1973]" captionTargetId-1="figure@3.[148,1427,238,1303]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 2. Perania annam sp. nov., male holotype (A-B, H), male paratypes (C-D), female “ allotype ” (E-G). (A) Carapace, dorsal view. (B) Same, lateral view. (C-E) Anterior portion of carapace, dorsal view. (F) Same, lateral view. (G-H) Dorsal scutum of opisthosoma, dorsal view. Scale lines 1.0 mm (A-B; C-D; E-F; G-H)." captionText-1="Fig. 3. Perania annam sp. nov., male paratype (A), male holotype (B-E), female allotype (F-H). (A-B) Distal portion of left palp, distal view. (C) Same, prolateral view. (D) Left cymbium and palpal organ, dorsal view. (E) Apex of embolus, retrolateral view. (F) Vulva, dorsal view. (G) Same, ventral view. (H) Same, anterior view. Abbreviations: ac = anterior collar; ep = embolic part; ga = genital atrium; pe = posterior extension of anterior collar; s = spermatheca with gland pores; sl = subterminal lamella. Scale lines 0.5 mm (F-H), 1.0 mm (A-E)." pageId="2" pageNumber="425">
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Figs
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<number box="[440,455,658,682]" pageId="2" pageNumber="425" value="2.0">2</number>
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-3
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</figureCitation>
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||
,
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||
<figureCitation box="[489,501,658,682]" captionStart="Fig. 8" captionStartId="12.[155,190,1828,1849]" captionTargetBox="[190,1406,269,1765]" captionTargetId="figure@12.[181,1415,257,1776]" captionTargetPageId="12" captionText="Fig. 8. Setae on legs of Tetrablemmidae. (A) Lamania kraui Shear, 1978, two short, sigmoid, serrate setae (SSSS) on tarsus I of subadult male from Gua Musang. (B) Perania annam sp. nov., scattered SSSS (not forming scopula) on tarsus I of male paratype. (C) Lamania kraui, scattered SSSS (indicated by arrows) on tarsus I of subadult male from Gua Musang. (D) Lamania kraui, scattered SSSS (indicated by arrows) on tarsus II of male from Gua Musang. (E) Lamania kraui, SSSS forming distinct ventral scopula on tarsus I and metatarsus I of male from Gua Musang. (F) Paculla cameronensis Shear, 1978, light scopula on tarsus I of male from Maxwell Hill. (G) Sabahya kinabaluana Deeleman-Reinhold, 1980, light scopula on tarsus I and metatarsus I of male from type locality. (H) Ablemma unicornis Burger, 2008, light scopula in distal half of tarsus I of male paratype. (I) Borneomma roberti Deeleman-Reinhold, 1980, distinct scopula in distal half of tarsus I of male from type locality. (J) Lamania kraui, pinnate seta on metatarsus III of female from Gua Musang." pageId="2" pageNumber="425">8</figureCitation>
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B
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="2" pageNumber="425" type="materials_examined">
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<paragraph pageId="2" pageNumber="425">
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<materialsCitation ID-GBIF-Occurrence="1249949206" collectionCode="MHNG" country="Vietnam" latitude="11.95" location="Dalat" longitude="108.4333" pageId="2" pageNumber="425" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong Province" typeStatus="holotype">
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<emphasis bold="true" box="[151,267,705,729]" pageId="2" pageNumber="425">
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<typeStatus box="[151,262,705,729]" pageId="2" pageNumber="425">Holotype</typeStatus>
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:
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</emphasis>
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||
<collectionCode box="[293,379,705,729]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="2" pageNumber="425">MHNG</collectionCode>
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||
;
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<specimenCount box="[409,465,704,728]" pageId="2" pageNumber="425" type="male">male</specimenCount>
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;
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<collectingCountry box="[495,591,704,728]" name="Vietnam" pageId="2" pageNumber="425" value="Vietnam">Vietnam</collectingCountry>
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,
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<collectingRegion pageId="2" pageNumber="425">
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Lam
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<heading box="[696,756,705,729]" centered="true" fontSize="10" level="4" pageId="2" pageNumber="425" reason="4">Dong</heading>
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Province
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</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:03917400BD50D73DD2072F11FA2DF874:8EE793CDBD50D73BD2462F80FEECFD64" box="[259,320,736,760]" country="Vietnam" name="Dalat" pageId="2" pageNumber="425" stateProvince="Lam Dong Province">Dalat</location>
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; collector unknown, don. J. Král
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||
</materialsCitation>
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||
<materialsCitation ID-GBIF-Occurrence="1249949205" box="[151,661,783,808]" collectionCode="MHNG" country="Vietnam" latitude="11.95" longitude="108.4333" pageId="2" pageNumber="425" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
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||
<emphasis bold="true" box="[151,276,784,808]" pageId="2" pageNumber="425">
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||
<typeStatus box="[151,271,784,808]" pageId="2" pageNumber="425">Paratypes</typeStatus>
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||
<paragraph box="[672,676,736,760]" pageId="2" pageNumber="425">.</paragraph>
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||
<paragraph pageId="2" pageNumber="425">
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||
:
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||
<specimenCount box="[290,373,783,808]" pageId="2" pageNumber="425" type="male">
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||
<number box="[290,304,784,808]" pageId="2" pageNumber="425" value="1.0">1</number>
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male
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||
</specimenCount>
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||
,
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||
<specimenCount box="[392,493,783,808]" pageId="2" pageNumber="425" type="female">
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<number box="[392,405,784,808]" pageId="2" pageNumber="425" value="1.0">1</number>
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||
female
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||
</specimenCount>
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||
(
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||
<collectionCode box="[515,600,783,807]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="2" pageNumber="425">MHNG</collectionCode>
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||
) and
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<materialsCitation ID-GBIF-Occurrence="1249949208" collectionCode="NMP" country="Vietnam" latitude="11.95" longitude="108.4333" pageId="2" pageNumber="425" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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||
<specimenCount box="[676,756,783,808]" pageId="2" pageNumber="425" type="male">
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||
<number box="[676,689,784,808]" pageId="2" pageNumber="425" value="1.0">1</number>
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male
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||
</specimenCount>
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(
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<collectionCode box="[159,222,815,839]" collectionName="Czech Republic, Prague, National Museum (Natural History)" pageId="2" pageNumber="425">NMP</collectionCode>
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); same data as for holotype.
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</materialsCitation>
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</paragraph>
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</emphasis>
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection pageId="2" pageNumber="425" type="etymology">
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<paragraph pageId="2" pageNumber="425">
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<emphasis bold="true" box="[151,284,864,887]" pageId="2" pageNumber="425">Etymology:</emphasis>
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“
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<taxonomicName box="[305,386,862,886]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="annam" status="sp. nov.">Annam</taxonomicName>
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” is the former name of southern Vietnam. Noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="425" type="diagnosis">
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<paragraph pageId="2" pageNumber="425">
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<emphasis bold="true" box="[151,270,942,966]" pageId="2" pageNumber="425">Diagnosis:</emphasis>
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The new species differs from all congeners by the presence of conically elevated hair bases on tibia II of males and on metartarsus II of females. Males share with those of
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<taxonomicName authority="Schwendinger, 2013" authorityName="Schwendinger" authorityYear="2013" box="[1063,1412,258,282]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="425" phylum="Arthropoda" rank="species" species="egregia">
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<emphasis box="[1063,1176,259,282]" italics="true" pageId="2" pageNumber="425">P. egregia</emphasis>
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Schwendinger,
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<number box="[1359,1412,259,282]" pageId="2" pageNumber="425" value="2013.0">2013</number>
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||
</taxonomicName>
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||
a very short cymbium (apex reduced) and a palpal organ with a thick embolus without marked transition between bulbus and base of embolus;
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||
<emphasis box="[1163,1266,355,378]" italics="true" pageId="2" pageNumber="425">
|
||
<taxonomicName box="[1163,1266,355,378]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="425" phylum="Arthropoda" rank="species" species="annam" status="sp. nov.">P. annam</taxonomicName>
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</emphasis>
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||
<taxonomicNameLabel box="[1277,1364,355,378]" pageId="2" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
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||
males are distinguished from
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||
<emphasis box="[1086,1195,387,410]" italics="true" pageId="2" pageNumber="425">
|
||
<taxonomicName box="[1086,1195,387,410]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="425" phylum="Arthropoda" rank="species" species="egregia">P. egregia</taxonomicName>
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</emphasis>
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||
males by the apex of the clypeal process being spade-shaped (with broadly rounded distal margin) rather than anvil-shaped (with slightly rounded or straight distal margin), by having a slightly more pronounced cymbial apex, a subterminal lamella which is pointed instead of rounded, and a longer embolic part (
|
||
<figureCitation box="[1056,1121,578,602]" captionStart="Fig. 3" captionStartId="3.[151,186,1326,1347]" captionTargetBox="[190,1394,260,1265]" captionTargetId="figure@3.[148,1427,238,1303]" captionTargetPageId="3" captionText="Fig. 3. Perania annam sp. nov., male paratype (A), male holotype (B-E), female allotype (F-H). (A-B) Distal portion of left palp, distal view. (C) Same, prolateral view. (D) Left cymbium and palpal organ, dorsal view. (E) Apex of embolus, retrolateral view. (F) Vulva, dorsal view. (G) Same, ventral view. (H) Same, anterior view. Abbreviations: ac = anterior collar; ep = embolic part; ga = genital atrium; pe = posterior extension of anterior collar; s = spermatheca with gland pores; sl = subterminal lamella. Scale lines 0.5 mm (F-H), 1.0 mm (A-E)." pageId="2" pageNumber="425">
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||
Figs
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||
<number box="[1111,1121,579,602]" pageId="2" pageNumber="425" value="3.0">3</number>
|
||
</figureCitation>
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||
A-E cf. Schwendinger,
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||
<number box="[1376,1433,579,602]" pageId="2" pageNumber="425" value="2013.0">2013</number>
|
||
: figs
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||
<number box="[880,932,610,633]" pageId="2" pageNumber="425" value="9.5" valueMax="10.0" valueMin="9.0">9-10</number>
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||
). The female of the new species differs from females of
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||
<emphasis box="[952,1060,643,666]" italics="true" pageId="2" pageNumber="425">
|
||
<taxonomicName box="[952,1060,643,666]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="425" phylum="Arthropoda" rank="species" species="egregia">P. egregia</taxonomicName>
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||
</emphasis>
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||
by retaining a small preanal plate; its spermathecae have inclined instead of horizontal posterior margins; the central sclerite (pe = posterior extension of anterior collar in
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<figureCitation box="[1156,1218,738,761]" captionStart="Fig. 3" captionStartId="3.[151,186,1326,1347]" captionTargetBox="[190,1394,260,1265]" captionTargetId="figure@3.[148,1427,238,1303]" captionTargetPageId="3" captionText="Fig. 3. Perania annam sp. nov., male paratype (A), male holotype (B-E), female allotype (F-H). (A-B) Distal portion of left palp, distal view. (C) Same, prolateral view. (D) Left cymbium and palpal organ, dorsal view. (E) Apex of embolus, retrolateral view. (F) Vulva, dorsal view. (G) Same, ventral view. (H) Same, anterior view. Abbreviations: ac = anterior collar; ep = embolic part; ga = genital atrium; pe = posterior extension of anterior collar; s = spermatheca with gland pores; sl = subterminal lamella. Scale lines 0.5 mm (F-H), 1.0 mm (A-E)." pageId="2" pageNumber="425">
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||
Fig.
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||
<number box="[1206,1218,738,761]" pageId="2" pageNumber="425" value="3.0">3</number>
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||
</figureCitation>
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||
<collectionCode box="[1218,1234,738,761]" pageId="2" pageNumber="425">F</collectionCode>
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||
) in the dorsal wall of the vulva is much larger and thicker, with a widely rounded posterior margin (straight in
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||
<emphasis box="[1254,1365,803,826]" italics="true" pageId="2" pageNumber="425">
|
||
<taxonomicName box="[1254,1365,803,826]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="425" phylum="Arthropoda" rank="species" species="egregia">P. egregia</taxonomicName>
|
||
</emphasis>
|
||
) (
|
||
<figureCitation captionStart="Fig. 3" captionStartId="3.[151,186,1326,1347]" captionTargetBox="[190,1394,260,1265]" captionTargetId="figure@3.[148,1427,238,1303]" captionTargetPageId="3" captionText="Fig. 3. Perania annam sp. nov., male paratype (A), male holotype (B-E), female allotype (F-H). (A-B) Distal portion of left palp, distal view. (C) Same, prolateral view. (D) Left cymbium and palpal organ, dorsal view. (E) Apex of embolus, retrolateral view. (F) Vulva, dorsal view. (G) Same, ventral view. (H) Same, anterior view. Abbreviations: ac = anterior collar; ep = embolic part; ga = genital atrium; pe = posterior extension of anterior collar; s = spermatheca with gland pores; sl = subterminal lamella. Scale lines 0.5 mm (F-H), 1.0 mm (A-E)." pageId="2" pageNumber="425">
|
||
Fig.
|
||
<number box="[831,842,835,858]" pageId="2" pageNumber="425" value="3.0">3</number>
|
||
</figureCitation>
|
||
F-H cf. Schwendinger,
|
||
<number box="[1093,1151,835,858]" pageId="2" pageNumber="425" value="2013.0">2013</number>
|
||
: fig.
|
||
<number box="[1202,1228,834,857]" pageId="2" pageNumber="425" value="10.0">10</number>
|
||
A-E).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="4" lastPageNumber="427" pageId="2" pageNumber="425" type="description">
|
||
<paragraph lastPageId="3" lastPageNumber="426" pageId="2" pageNumber="425">
|
||
<emphasis bold="true" box="[831,1190,882,906]" pageId="2" pageNumber="425">
|
||
Description of male (
|
||
<typeStatus box="[1073,1177,882,906]" pageId="2" pageNumber="425">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Colour (in alcohol) of sclerotised parts mostly dark brown, except for lighter patellae and tibiae I-II, patella to metatarsus III and patella to tarsus IV; membranous parts of opisthosoma very light brown. Clypeus with long, distally widened and flattened median process projecting foreward and slightly upward, occupying
|
||
<number box="[474,501,1564,1588]" pageId="3" pageNumber="426" value="34.0">34</number>
|
||
% of carapace length, its distal margin broadly arched (
|
||
<figureCitation box="[543,614,1596,1619]" captionStart="Fig. 2" captionStartId="2.[151,186,1998,2019]" captionTargetBox="[177,1419,1104,1949]" captionTargetId="figure@2.[151,1436,1083,1973]" captionTargetPageId="2" captionText="Fig. 2. Perania annam sp. nov., male holotype (A-B, H), male paratypes (C-D), female “ allotype ” (E-G). (A) Carapace, dorsal view. (B) Same, lateral view. (C-E) Anterior portion of carapace, dorsal view. (F) Same, lateral view. (G-H) Dorsal scutum of opisthosoma, dorsal view. Scale lines 1.0 mm (A-B; C-D; E-F; G-H)." pageId="3" pageNumber="426">
|
||
Fig.
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||
<number box="[601,614,1596,1619]" pageId="3" pageNumber="426" value="2.0">2</number>
|
||
</figureCitation>
|
||
A-B). Fovea indistinct and narrow, including two shallow pits (
|
||
<figureCitation box="[159,225,1660,1683]" captionStart="Fig. 2" captionStartId="2.[151,186,1998,2019]" captionTargetBox="[177,1419,1104,1949]" captionTargetId="figure@2.[151,1436,1083,1973]" captionTargetPageId="2" captionText="Fig. 2. Perania annam sp. nov., male holotype (A-B, H), male paratypes (C-D), female “ allotype ” (E-G). (A) Carapace, dorsal view. (B) Same, lateral view. (C-E) Anterior portion of carapace, dorsal view. (F) Same, lateral view. (G-H) Dorsal scutum of opisthosoma, dorsal view. Scale lines 1.0 mm (A-B; C-D; E-F; G-H)." pageId="3" pageNumber="426">
|
||
Fig.
|
||
<number box="[211,225,1660,1683]" pageId="3" pageNumber="426" value="2.0">2</number>
|
||
</figureCitation>
|
||
<collectionCode box="[225,247,1660,1683]" pageId="3" pageNumber="426">A</collectionCode>
|
||
). Total length
|
||
<number box="[407,470,1660,1684]" pageId="3" pageNumber="426" value="15.88">15.88</number>
|
||
. Carapace
|
||
<number box="[591,637,1660,1684]" pageId="3" pageNumber="426" value="8.73">8.73</number>
|
||
long,
|
||
<number box="[709,756,1660,1684]" pageId="3" pageNumber="426" value="3.09">3.09</number>
|
||
wide, its cephalic portion only slightly domed (
|
||
<figureCitation box="[659,724,1692,1715]" captionStart="Fig. 2" captionStartId="2.[151,186,1998,2019]" captionTargetBox="[177,1419,1104,1949]" captionTargetId="figure@2.[151,1436,1083,1973]" captionTargetPageId="2" captionText="Fig. 2. Perania annam sp. nov., male holotype (A-B, H), male paratypes (C-D), female “ allotype ” (E-G). (A) Carapace, dorsal view. (B) Same, lateral view. (C-E) Anterior portion of carapace, dorsal view. (F) Same, lateral view. (G-H) Dorsal scutum of opisthosoma, dorsal view. Scale lines 1.0 mm (A-B; C-D; E-F; G-H)." pageId="3" pageNumber="426">
|
||
Fig.
|
||
<number box="[709,724,1692,1715]" pageId="3" pageNumber="426" value="2.0">2</number>
|
||
</figureCitation>
|
||
<collectionCode box="[724,743,1692,1715]" pageId="3" pageNumber="426">B</collectionCode>
|
||
). Eye sizes and interdistances: PME
|
||
<number box="[537,583,1724,1748]" pageId="3" pageNumber="426" value="0.28">0.28</number>
|
||
long, separated by
|
||
<number box="[192,243,1756,1780]" pageId="3" pageNumber="426" value="0.24">0.24</number>
|
||
; ALE
|
||
<number box="[326,373,1756,1780]" pageId="3" pageNumber="426" value="0.28">0.28</number>
|
||
long; ALE-PLE
|
||
<number box="[578,629,1756,1780]" pageId="3" pageNumber="426" value="0.15">0.15</number>
|
||
; PLE
|
||
<number box="[709,756,1756,1780]" pageId="3" pageNumber="426" value="0.37">0.37</number>
|
||
long, separated by
|
||
<number box="[363,413,1788,1812]" pageId="3" pageNumber="426" value="1.42">1.42</number>
|
||
. Labium
|
||
<number box="[520,566,1788,1812]" pageId="3" pageNumber="426" value="0.98">0.98</number>
|
||
long,
|
||
<number box="[641,687,1788,1812]" pageId="3" pageNumber="426" value="1.27">1.27</number>
|
||
wide. Sternum
|
||
<number box="[251,298,1820,1844]" pageId="3" pageNumber="426" value="3.53">3.53</number>
|
||
long,
|
||
<number box="[371,418,1820,1844]" pageId="3" pageNumber="426" value="2.25">2.25</number>
|
||
wide; sternal apophysis anvilshaped. Chelicerae without modifications; two small teeth distal to median lamina.
|
||
</paragraph>
|
||
<caption pageId="2" pageNumber="425" targetBox="[177,1419,1104,1949]" targetPageId="2">
|
||
<paragraph pageId="2" pageNumber="425">
|
||
Fig. 2.
|
||
<emphasis box="[227,376,1999,2020]" italics="true" pageId="2" pageNumber="425">
|
||
<taxonomicName box="[227,376,1999,2020]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="425" phylum="Arthropoda" rank="species" species="annam" status="sp. nov.">Perania annam</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[382,460,1999,2020]" pageId="2" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male holotype (A-B, H), male paratypes (C-D), female “allotype” (E-G). (A) Carapace, dorsal view. (B) Same, lateral view. (C-E) Anterior portion of carapace, dorsal view. (F) Same, lateral view. (G-H) Dorsal scutum of opisthosoma, dorsal view. Scale lines 1.0 mm (A-B; C-D; E-F; G-H).
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="3" pageNumber="426" targetBox="[190,1394,260,1265]" targetPageId="3">
|
||
<paragraph pageId="3" pageNumber="426">
|
||
Fig. 3.
|
||
<emphasis box="[227,374,1327,1348]" italics="true" pageId="3" pageNumber="426">
|
||
<taxonomicName box="[227,374,1327,1348]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="426" phylum="Arthropoda" rank="species" species="annam" status="sp. nov.">Perania annam</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[379,455,1327,1348]" pageId="3" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male paratype (A), male holotype (B-E), female allotype (F-H). (A-B) Distal portion of left palp, distal view. (C) Same, prolateral view. (D) Left cymbium and palpal organ, dorsal view. (E) Apex of embolus, retrolateral view. (F) Vulva, dorsal view. (G) Same, ventral view. (H) Same, anterior view. Abbreviations: ac = anterior collar; ep = embolic part; ga = genital atrium; pe = posterior extension of anterior collar; s = spermatheca with gland pores; sl = subterminal lamella. Scale lines 0.5 mm (F-H), 1.0 mm (A-E).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="3" pageNumber="426">
|
||
Palp
|
||
<number box="[224,270,1916,1940]" pageId="3" pageNumber="426" value="5.42">5.42</number>
|
||
long (
|
||
<number box="[376,422,1916,1939]" pageId="3" pageNumber="426" value="1.67">1.67</number>
|
||
+
|
||
<number box="[438,484,1916,1939]" pageId="3" pageNumber="426" value="0.71">0.71</number>
|
||
+
|
||
<number box="[500,549,1916,1939]" pageId="3" pageNumber="426" value="2.16">2.16</number>
|
||
+
|
||
<number box="[565,613,1916,1939]" pageId="3" pageNumber="426" value="0.88">0.88</number>
|
||
). Cymbium (
|
||
<figureCitation box="[159,224,1948,1971]" captionStart="Fig. 3" captionStartId="3.[151,186,1326,1347]" captionTargetBox="[190,1394,260,1265]" captionTargetId="figure@3.[148,1427,238,1303]" captionTargetPageId="3" captionText="Fig. 3. Perania annam sp. nov., male paratype (A), male holotype (B-E), female allotype (F-H). (A-B) Distal portion of left palp, distal view. (C) Same, prolateral view. (D) Left cymbium and palpal organ, dorsal view. (E) Apex of embolus, retrolateral view. (F) Vulva, dorsal view. (G) Same, ventral view. (H) Same, anterior view. Abbreviations: ac = anterior collar; ep = embolic part; ga = genital atrium; pe = posterior extension of anterior collar; s = spermatheca with gland pores; sl = subterminal lamella. Scale lines 0.5 mm (F-H), 1.0 mm (A-E)." pageId="3" pageNumber="426">
|
||
Fig.
|
||
<number box="[212,224,1948,1971]" pageId="3" pageNumber="426" value="3.0">3</number>
|
||
</figureCitation>
|
||
<collectionCode box="[224,245,1948,1971]" pageId="3" pageNumber="426">D</collectionCode>
|
||
) short, with indistinct retrodorsal lobe (apex). Palpal organ (
|
||
<figureCitation box="[304,367,1980,2003]" captionStart="Fig. 3" captionStartId="3.[151,186,1326,1347]" captionTargetBox="[190,1394,260,1265]" captionTargetId="figure@3.[148,1427,238,1303]" captionTargetPageId="3" captionText="Fig. 3. Perania annam sp. nov., male paratype (A), male holotype (B-E), female allotype (F-H). (A-B) Distal portion of left palp, distal view. (C) Same, prolateral view. (D) Left cymbium and palpal organ, dorsal view. (E) Apex of embolus, retrolateral view. (F) Vulva, dorsal view. (G) Same, ventral view. (H) Same, anterior view. Abbreviations: ac = anterior collar; ep = embolic part; ga = genital atrium; pe = posterior extension of anterior collar; s = spermatheca with gland pores; sl = subterminal lamella. Scale lines 0.5 mm (F-H), 1.0 mm (A-E)." pageId="3" pageNumber="426">
|
||
Fig.
|
||
<number box="[356,367,1980,2003]" pageId="3" pageNumber="426" value="3.0">3</number>
|
||
</figureCitation>
|
||
B-E) strongly modified: bulbus and base of embolus of same thickness, without constriction between them; apex of embolus compressed and deeply split into a triangular, acutely pointed subterminal lamella and a longer and basally wider embolic part with a small, narrowly rounded tip.
|
||
</paragraph>
|
||
<paragraph pageId="3" pageNumber="426">
|
||
Legs
|
||
<number box="[891,948,1628,1652]" pageId="3" pageNumber="426" value="3421.0">3421</number>
|
||
. About
|
||
<number box="[1033,1059,1628,1652]" pageId="3" pageNumber="426" value="50.0">50</number>
|
||
dark conical setal bases of various sizes prolaterally to ventrally on metatarsus I, about
|
||
<number box="[1410,1436,1660,1684]" pageId="3" pageNumber="426" value="30.0">30</number>
|
||
shorter ones proventrally on tibia I; about
|
||
<number box="[1312,1339,1692,1716]" pageId="3" pageNumber="426" value="25.0">25</number>
|
||
ones on metatarsus II and six on tibia II. All leg femora ventrally rugose (with wart-like hair bases); palpal femur essentially smooth. Paired leg claws with a row of
|
||
<number box="[1277,1339,1788,1812]" pageId="3" pageNumber="426" value="12.0" valueMax="13.0" valueMin="11.0">11-13</number>
|
||
teeth on anteriorlegs,
|
||
<number box="[970,1005,1820,1844]" pageId="3" pageNumber="426" value="8.5" valueMax="9.0" valueMin="8.0">8-9</number>
|
||
onposteriors; unpairedlegclawswithone denticle. Leg I
|
||
<number box="[996,1056,1852,1876]" pageId="3" pageNumber="426" value="22.54">22.54</number>
|
||
long (
|
||
<number box="[1126,1174,1852,1875]" pageId="3" pageNumber="426" value="6.96">6.96</number>
|
||
+
|
||
<number box="[1190,1238,1852,1875]" pageId="3" pageNumber="426" value="2.45">2.45</number>
|
||
+
|
||
<number box="[1253,1301,1852,1875]" pageId="3" pageNumber="426" value="6.76">6.76</number>
|
||
+
|
||
<number box="[1316,1363,1852,1875]" pageId="3" pageNumber="426" value="4.61">4.61</number>
|
||
+
|
||
<number box="[1378,1424,1852,1875]" pageId="3" pageNumber="426" value="1.76">1.76</number>
|
||
), leg II
|
||
<number box="[905,965,1884,1908]" pageId="3" pageNumber="426" value="17.9">17.90</number>
|
||
long (
|
||
<number box="[1044,1091,1884,1907]" pageId="3" pageNumber="426" value="5.25">5.25</number>
|
||
+
|
||
<number box="[1106,1153,1884,1907]" pageId="3" pageNumber="426" value="1.96">1.96</number>
|
||
+
|
||
<number box="[1168,1216,1884,1907]" pageId="3" pageNumber="426" value="5.1">5.10</number>
|
||
+
|
||
<number box="[1231,1280,1884,1907]" pageId="3" pageNumber="426" value="4.02">4.02</number>
|
||
+
|
||
<number box="[1296,1342,1884,1907]" pageId="3" pageNumber="426" value="1.57">1.57</number>
|
||
), leg III
|
||
<number box="[832,892,1916,1940]" pageId="3" pageNumber="426" value="13.43">13.43</number>
|
||
long (
|
||
<number box="[970,1020,1916,1939]" pageId="3" pageNumber="426" value="4.02">4.02</number>
|
||
+
|
||
<number box="[1035,1081,1916,1939]" pageId="3" pageNumber="426" value="1.47">1.47</number>
|
||
+
|
||
<number box="[1096,1143,1916,1939]" pageId="3" pageNumber="426" value="3.43">3.43</number>
|
||
+
|
||
<number box="[1158,1207,1916,1939]" pageId="3" pageNumber="426" value="3.24">3.24</number>
|
||
+
|
||
<number box="[1222,1268,1916,1939]" pageId="3" pageNumber="426" value="1.27">1.27</number>
|
||
), leg IV
|
||
<number box="[1376,1436,1916,1940]" pageId="3" pageNumber="426" value="16.62">16.62</number>
|
||
long (5.00+
|
||
<number box="[957,1004,1948,1971]" pageId="3" pageNumber="426" value="1.57">1.57</number>
|
||
+
|
||
<number box="[1019,1066,1948,1971]" pageId="3" pageNumber="426" value="4.51">4.51</number>
|
||
+
|
||
<number box="[1081,1131,1948,1971]" pageId="3" pageNumber="426" value="4.22">4.22</number>
|
||
+
|
||
<number box="[1146,1192,1948,1971]" pageId="3" pageNumber="426" value="1.32">1.32</number>
|
||
).
|
||
</paragraph>
|
||
<paragraph lastPageId="4" lastPageNumber="427" pageId="3" pageNumber="426">
|
||
Opisthosoma
|
||
<number box="[991,1037,1980,2004]" pageId="3" pageNumber="426" value="7.16">7.16</number>
|
||
long,
|
||
<number box="[1128,1175,1980,2004]" pageId="3" pageNumber="426" value="3.92">3.92</number>
|
||
wide. Dorsal scutum (
|
||
<figureCitation box="[840,907,2012,2035]" captionStart="Fig. 2" captionStartId="2.[151,186,1998,2019]" captionTargetBox="[177,1419,1104,1949]" captionTargetId="figure@2.[151,1436,1083,1973]" captionTargetPageId="2" captionText="Fig. 2. Perania annam sp. nov., male holotype (A-B, H), male paratypes (C-D), female “ allotype ” (E-G). (A) Carapace, dorsal view. (B) Same, lateral view. (C-E) Anterior portion of carapace, dorsal view. (F) Same, lateral view. (G-H) Dorsal scutum of opisthosoma, dorsal view. Scale lines 1.0 mm (A-B; C-D; E-F; G-H)." pageId="3" pageNumber="426">
|
||
Fig.
|
||
<number box="[894,907,2012,2035]" pageId="3" pageNumber="426" value="2.0">2</number>
|
||
</figureCitation>
|
||
<collectionCode box="[907,928,2012,2035]" pageId="3" pageNumber="426">H</collectionCode>
|
||
) exceptionally narrow,
|
||
<number box="[1195,1242,2012,2036]" pageId="3" pageNumber="426" value="5.88">5.88</number>
|
||
long,
|
||
<number box="[1319,1366,2012,2036]" pageId="3" pageNumber="426" value="2.99">2.99</number>
|
||
wide, somewhat oval, quite smooth, its surface with only few indistinct elevations. Most lateral microplates flat, only a posterolateral one on each side of second band from above slightly elevated. Most microplates in bands on posterior side of opisthosoma closely packed, those in uppermost band largely fused into strap-like plate (
|
||
<figureCitation box="[159,223,418,441]" captionStart="Fig. 2" captionStartId="2.[151,186,1998,2019]" captionTargetBox="[177,1419,1104,1949]" captionTargetId="figure@2.[151,1436,1083,1973]" captionTargetPageId="2" captionText="Fig. 2. Perania annam sp. nov., male holotype (A-B, H), male paratypes (C-D), female “ allotype ” (E-G). (A) Carapace, dorsal view. (B) Same, lateral view. (C-E) Anterior portion of carapace, dorsal view. (F) Same, lateral view. (G-H) Dorsal scutum of opisthosoma, dorsal view. Scale lines 1.0 mm (A-B; C-D; E-F; G-H)." pageId="4" pageNumber="427">
|
||
Fig.
|
||
<number box="[209,223,418,441]" pageId="4" pageNumber="427" value="2.0">2</number>
|
||
</figureCitation>
|
||
<collectionCode box="[223,245,418,441]" pageId="4" pageNumber="427">H</collectionCode>
|
||
). Pulmonary plate
|
||
<number box="[448,495,419,442]" pageId="4" pageNumber="427" value="4.02">4.02</number>
|
||
long,
|
||
<number box="[563,610,419,442]" pageId="4" pageNumber="427" value="3.33">3.33</number>
|
||
wide; genital region flat. Anterior pair of ventrolateral plates largely free, only their posterior tips fused with margin of pulmonary plate. Postgenital plate
|
||
<number box="[523,570,515,538]" pageId="4" pageNumber="427" value="0.29">0.29</number>
|
||
long, completely fused to posterior margin of pulmonary plate, both linked by a sclerotised suture. Preanal plate well developed,
|
||
<number box="[151,198,611,634]" pageId="4" pageNumber="427" value="0.78">0.78</number>
|
||
long,
|
||
<number box="[265,312,611,634]" pageId="4" pageNumber="427" value="1.27">1.27</number>
|
||
wide. Anal plate
|
||
<number box="[499,546,611,634]" pageId="4" pageNumber="427" value="1.23">1.23</number>
|
||
long,
|
||
<number box="[613,660,611,634]" pageId="4" pageNumber="427" value="1.62">1.62</number>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph pageId="4" pageNumber="427">
|
||
<emphasis bold="true" box="[151,422,658,682]" pageId="4" pageNumber="427">Description of female:</emphasis>
|
||
As in male, except for the following. Total length
|
||
<number box="[436,500,690,714]" pageId="4" pageNumber="427" value="12.94">12.94</number>
|
||
. Carapace
|
||
<number box="[637,685,690,714]" pageId="4" pageNumber="427" value="5.29">5.29</number>
|
||
long,
|
||
<number box="[151,198,722,746]" pageId="4" pageNumber="427" value="3.04">3.04</number>
|
||
wide; cephalic portion more strongly domed than in male, with a short but distinct conical hump on clypeus occupying about
|
||
<number box="[439,453,786,810]" pageId="4" pageNumber="427" value="4.0">4</number>
|
||
% of carapace length (
|
||
<figureCitation captionStart="Fig. 2" captionStartId="2.[151,186,1998,2019]" captionTargetBox="[177,1419,1104,1949]" captionTargetId="figure@2.[151,1436,1083,1973]" captionTargetPageId="2" captionText="Fig. 2. Perania annam sp. nov., male holotype (A-B, H), male paratypes (C-D), female “ allotype ” (E-G). (A) Carapace, dorsal view. (B) Same, lateral view. (C-E) Anterior portion of carapace, dorsal view. (F) Same, lateral view. (G-H) Dorsal scutum of opisthosoma, dorsal view. Scale lines 1.0 mm (A-B; C-D; E-F; G-H)." pageId="4" pageNumber="427">
|
||
Fig.
|
||
<number box="[151,165,817,840]" pageId="4" pageNumber="427" value="2.0">2</number>
|
||
</figureCitation>
|
||
E-F). PME
|
||
<number box="[305,352,818,842]" pageId="4" pageNumber="427" value="0.34">0.34</number>
|
||
long,
|
||
<number box="[436,483,818,842]" pageId="4" pageNumber="427" value="0.21">0.21</number>
|
||
apart; ALE
|
||
<number box="[639,686,818,842]" pageId="4" pageNumber="427" value="0.3">0.30</number>
|
||
long; ALE-PLE
|
||
<number box="[277,328,850,874]" pageId="4" pageNumber="427" value="0.1">0.10</number>
|
||
; PLE
|
||
<number box="[413,461,850,874]" pageId="4" pageNumber="427" value="0.3">0.30</number>
|
||
long,
|
||
<number box="[549,596,850,874]" pageId="4" pageNumber="427" value="1.57">1.57</number>
|
||
apart. Fovea indistinct, with only one shallow pit. Labium
|
||
<number box="[647,694,882,906]" pageId="4" pageNumber="427" value="0.98">0.98</number>
|
||
long,
|
||
<number box="[151,198,914,938]" pageId="4" pageNumber="427" value="1.76">1.76</number>
|
||
wide. Sternum
|
||
<number box="[388,435,914,938]" pageId="4" pageNumber="427" value="3.04">3.04</number>
|
||
long,
|
||
<number box="[516,562,914,938]" pageId="4" pageNumber="427" value="2.11">2.11</number>
|
||
wide. Palp
|
||
<number box="[709,756,914,938]" pageId="4" pageNumber="427" value="4.26">4.26</number>
|
||
long (
|
||
<number box="[224,271,945,968]" pageId="4" pageNumber="427" value="1.27">1.27</number>
|
||
+
|
||
<number box="[287,337,945,968]" pageId="4" pageNumber="427" value="0.59">0.59</number>
|
||
+
|
||
<number box="[352,402,945,968]" pageId="4" pageNumber="427" value="0.98">0.98</number>
|
||
+
|
||
<number box="[418,465,945,968]" pageId="4" pageNumber="427" value="1.42">1.42</number>
|
||
). Legs
|
||
<number box="[562,619,946,970]" pageId="4" pageNumber="427" value="3421.0">3421</number>
|
||
; metatarsus I with about
|
||
<number box="[304,331,978,1002]" pageId="4" pageNumber="427" value="25.0">25</number>
|
||
conical setal bases of different sizes prolaterally and proventrally, tibia I with seven quite small ones prolaterally, metatarsus II with six very small ones prolaterally and proventrally; all conical setal bases weaker than those in males. Paired leg claws with a row of
|
||
<number box="[385,434,1138,1162]" pageId="4" pageNumber="427" value="11.5" valueMax="14.0" valueMin="9.0">9-14</number>
|
||
teeth. Leg I about
|
||
<number box="[652,698,1138,1162]" pageId="4" pageNumber="427" value="17.5">17.5</number>
|
||
long [
|
||
<number box="[157,206,1169,1192]" pageId="4" pageNumber="427" value="5.2">5.20</number>
|
||
+
|
||
<number box="[221,267,1169,1192]" pageId="4" pageNumber="427" value="1.82">1.82</number>
|
||
+
|
||
<number box="[282,330,1169,1192]" pageId="4" pageNumber="427" value="5.1">5.10</number>
|
||
+
|
||
<number box="[346,393,1169,1192]" pageId="4" pageNumber="427" value="3.73">3.73</number>
|
||
+
|
||
<number box="[408,439,1169,1192]" pageId="4" pageNumber="427" value="1.7">1.7</number>
|
||
?(distal half missing)], leg II
|
||
<number box="[151,212,1202,1226]" pageId="4" pageNumber="427" value="14.8">14.80</number>
|
||
long (
|
||
<number box="[290,337,1201,1224]" pageId="4" pageNumber="427" value="4.31">4.31</number>
|
||
+
|
||
<number box="[353,399,1201,1224]" pageId="4" pageNumber="427" value="1.62">1.62</number>
|
||
+
|
||
<number box="[415,464,1201,1224]" pageId="4" pageNumber="427" value="4.12">4.12</number>
|
||
+
|
||
<number box="[480,528,1201,1224]" pageId="4" pageNumber="427" value="3.28">3.28</number>
|
||
+
|
||
<number box="[544,590,1201,1224]" pageId="4" pageNumber="427" value="1.47">1.47</number>
|
||
), leg III
|
||
<number box="[696,756,1202,1226]" pageId="4" pageNumber="427" value="11.32">11.32</number>
|
||
long (
|
||
<number box="[220,269,1233,1256]" pageId="4" pageNumber="427" value="3.28">3.28</number>
|
||
+
|
||
<number box="[284,331,1233,1256]" pageId="4" pageNumber="427" value="1.27">1.27</number>
|
||
+
|
||
<number box="[346,396,1233,1256]" pageId="4" pageNumber="427" value="2.84">2.84</number>
|
||
+
|
||
<number box="[411,460,1233,1256]" pageId="4" pageNumber="427" value="2.75">2.75</number>
|
||
+
|
||
<number box="[475,521,1233,1256]" pageId="4" pageNumber="427" value="1.18">1.18</number>
|
||
), leg IV
|
||
<number box="[634,695,1234,1258]" pageId="4" pageNumber="427" value="14.07">14.07</number>
|
||
long (
|
||
<number box="[159,208,1265,1288]" pageId="4" pageNumber="427" value="4.07">4.07</number>
|
||
+
|
||
<number box="[224,270,1265,1288]" pageId="4" pageNumber="427" value="1.37">1.37</number>
|
||
+
|
||
<number box="[285,334,1265,1288]" pageId="4" pageNumber="427" value="3.82">3.82</number>
|
||
+
|
||
<number box="[349,397,1265,1288]" pageId="4" pageNumber="427" value="3.58">3.58</number>
|
||
+
|
||
<number box="[413,458,1265,1288]" pageId="4" pageNumber="427" value="1.23">1.23</number>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="4" pageNumber="427">
|
||
Opisthosoma
|
||
<number box="[301,348,1298,1322]" pageId="4" pageNumber="427" value="7.65">7.65</number>
|
||
long,
|
||
<number box="[421,468,1298,1322]" pageId="4" pageNumber="427" value="4.41">4.41</number>
|
||
wide; several microplates in posterior half larger than others, slightly elevated and conical; microplates on posterior side of opisthosoma not forming strap-like horizontal plates. Dorsal scutum (
|
||
<figureCitation box="[159,231,1425,1448]" captionStart="Fig. 2" captionStartId="2.[151,186,1998,2019]" captionTargetBox="[177,1419,1104,1949]" captionTargetId="figure@2.[151,1436,1083,1973]" captionTargetPageId="2" captionText="Fig. 2. Perania annam sp. nov., male holotype (A-B, H), male paratypes (C-D), female “ allotype ” (E-G). (A) Carapace, dorsal view. (B) Same, lateral view. (C-E) Anterior portion of carapace, dorsal view. (F) Same, lateral view. (G-H) Dorsal scutum of opisthosoma, dorsal view. Scale lines 1.0 mm (A-B; C-D; E-F; G-H)." pageId="4" pageNumber="427">
|
||
Fig.
|
||
<number box="[218,231,1425,1448]" pageId="4" pageNumber="427" value="2.0">2</number>
|
||
</figureCitation>
|
||
<collectionCode box="[231,251,1425,1448]" pageId="4" pageNumber="427">G</collectionCode>
|
||
)
|
||
<number box="[275,321,1426,1450]" pageId="4" pageNumber="427" value="4.51">4.51</number>
|
||
long,
|
||
<number box="[406,453,1426,1450]" pageId="4" pageNumber="427" value="2.55">2.55</number>
|
||
wide, its anterior margin relatively wider and its surface with more pronounced elevations than in males. Anterior opisthosomal plate completely fragmented into microplates (nine of them slightly larger than others). Pulmonary plate
|
||
<number box="[646,692,1554,1578]" pageId="4" pageNumber="427" value="3.24">3.24</number>
|
||
long,
|
||
<number box="[151,198,1586,1610]" pageId="4" pageNumber="427" value="3.04">3.04</number>
|
||
wide. Preanal plate small, spindle-shaped,
|
||
<number box="[709,756,1586,1610]" pageId="4" pageNumber="427" value="0.15">0.15</number>
|
||
long,
|
||
<number box="[212,259,1618,1642]" pageId="4" pageNumber="427" value="0.69">0.69</number>
|
||
wide. Anal plate
|
||
<number box="[446,493,1618,1642]" pageId="4" pageNumber="427" value="1.23">1.23</number>
|
||
long,
|
||
<number box="[560,607,1618,1642]" pageId="4" pageNumber="427" value="1.57">1.57</number>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph pageId="4" pageNumber="427">
|
||
Vulva (
|
||
<figureCitation box="[230,291,1649,1672]" captionStart="Fig. 3" captionStartId="3.[151,186,1326,1347]" captionTargetBox="[190,1394,260,1265]" captionTargetId="figure@3.[148,1427,238,1303]" captionTargetPageId="3" captionText="Fig. 3. Perania annam sp. nov., male paratype (A), male holotype (B-E), female allotype (F-H). (A-B) Distal portion of left palp, distal view. (C) Same, prolateral view. (D) Left cymbium and palpal organ, dorsal view. (E) Apex of embolus, retrolateral view. (F) Vulva, dorsal view. (G) Same, ventral view. (H) Same, anterior view. Abbreviations: ac = anterior collar; ep = embolic part; ga = genital atrium; pe = posterior extension of anterior collar; s = spermatheca with gland pores; sl = subterminal lamella. Scale lines 0.5 mm (F-H), 1.0 mm (A-E)." pageId="4" pageNumber="427">
|
||
Fig.
|
||
<number box="[280,291,1649,1672]" pageId="4" pageNumber="427" value="3.0">3</number>
|
||
</figureCitation>
|
||
F-H) with very long anterior collar, median zone of its dorsal and ventral walls distinctly lighter than lateral zones, posteriorly extended into a large and thick sclerite; transition between anterior collar and its posterior extensions weakly sclerotised and weakly pigmented, posterior margin of extension in dorsal wall widely rounded. Spermathecae and ventral wall of genital atrium lightly and uniformly pigmented, with a transparent zone between them. Spermathecae elliptical, with a distinct and wide median invagination in common anterior margin and with inclined posterior margins; anterolateral sides with relatively small porepatches on flat surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="427" type="discussion">
|
||
<paragraph pageId="4" pageNumber="427">
|
||
<emphasis bold="true" box="[831,948,259,282]" pageId="4" pageNumber="427">Variation:</emphasis>
|
||
Carapace length in three males (one female) ranges
|
||
<number box="[922,1026,291,314]" pageId="4" pageNumber="427" value="7.65" valueMax="8.73" valueMin="6.57">6.57-8.73</number>
|
||
(
|
||
<number box="[1053,1105,290,313]" pageId="4" pageNumber="427" value="5.29">5.29</number>
|
||
), carapace width
|
||
<number box="[1332,1436,291,314]" pageId="4" pageNumber="427" value="2.92" valueMax="3.09" valueMin="2.75">2.75-3.09</number>
|
||
(
|
||
<number box="[840,891,322,345]" pageId="4" pageNumber="427" value="3.04">3.04</number>
|
||
). In two males (including the
|
||
<typeStatus box="[1233,1328,322,345]" pageId="4" pageNumber="427">holotype</typeStatus>
|
||
) the cleft between the subterminal lamella and the embolic part of the palpal organ is acute at its base (
|
||
<figureCitation box="[1300,1369,386,409]" captionStart="Fig. 3" captionStartId="3.[151,186,1326,1347]" captionTargetBox="[190,1394,260,1265]" captionTargetId="figure@3.[148,1427,238,1303]" captionTargetPageId="3" captionText="Fig. 3. Perania annam sp. nov., male paratype (A), male holotype (B-E), female allotype (F-H). (A-B) Distal portion of left palp, distal view. (C) Same, prolateral view. (D) Left cymbium and palpal organ, dorsal view. (E) Apex of embolus, retrolateral view. (F) Vulva, dorsal view. (G) Same, ventral view. (H) Same, anterior view. Abbreviations: ac = anterior collar; ep = embolic part; ga = genital atrium; pe = posterior extension of anterior collar; s = spermatheca with gland pores; sl = subterminal lamella. Scale lines 0.5 mm (F-H), 1.0 mm (A-E)." pageId="4" pageNumber="427">
|
||
Fig.
|
||
<number box="[1356,1369,386,409]" pageId="4" pageNumber="427" value="3.0">3</number>
|
||
</figureCitation>
|
||
<collectionCode box="[1369,1389,386,409]" pageId="4" pageNumber="427">B</collectionCode>
|
||
), in the third male it is narrowly rounded (
|
||
<figureCitation box="[1279,1347,418,441]" captionStart="Fig. 3" captionStartId="3.[151,186,1326,1347]" captionTargetBox="[190,1394,260,1265]" captionTargetId="figure@3.[148,1427,238,1303]" captionTargetPageId="3" captionText="Fig. 3. Perania annam sp. nov., male paratype (A), male holotype (B-E), female allotype (F-H). (A-B) Distal portion of left palp, distal view. (C) Same, prolateral view. (D) Left cymbium and palpal organ, dorsal view. (E) Apex of embolus, retrolateral view. (F) Vulva, dorsal view. (G) Same, ventral view. (H) Same, anterior view. Abbreviations: ac = anterior collar; ep = embolic part; ga = genital atrium; pe = posterior extension of anterior collar; s = spermatheca with gland pores; sl = subterminal lamella. Scale lines 0.5 mm (F-H), 1.0 mm (A-E)." pageId="4" pageNumber="427">
|
||
Fig.
|
||
<number box="[1334,1347,418,441]" pageId="4" pageNumber="427" value="3.0">3</number>
|
||
</figureCitation>
|
||
<collectionCode box="[1347,1370,418,441]" pageId="4" pageNumber="427">A</collectionCode>
|
||
). The number of conical hair bases on tibia II of males ranges
|
||
<number box="[831,870,483,506]" pageId="4" pageNumber="427" value="4.5" valueMax="6.0" valueMin="3.0">3-6</number>
|
||
. Variation in the shape of clypeus modifications of males see
|
||
<figureCitation box="[980,1046,514,538]" captionStart="Fig. 2" captionStartId="2.[151,186,1998,2019]" captionTargetBox="[177,1419,1104,1949]" captionTargetId="figure@2.[151,1436,1083,1973]" captionTargetPageId="2" captionText="Fig. 2. Perania annam sp. nov., male holotype (A-B, H), male paratypes (C-D), female “ allotype ” (E-G). (A) Carapace, dorsal view. (B) Same, lateral view. (C-E) Anterior portion of carapace, dorsal view. (F) Same, lateral view. (G-H) Dorsal scutum of opisthosoma, dorsal view. Scale lines 1.0 mm (A-B; C-D; E-F; G-H)." pageId="4" pageNumber="427">
|
||
Fig.
|
||
<number box="[1032,1046,515,538]" pageId="4" pageNumber="427" value="2.0">2</number>
|
||
</figureCitation>
|
||
A, C-D. All three males have most microplates in the uppermost band on the posterior side of the opisthosoma fused; one of them (a
|
||
<typeStatus box="[1285,1378,578,601]" pageId="4" pageNumber="427">paratype</typeStatus>
|
||
) also has fused microplates in the second band from above. In all males the postgenital plate is relatively long (in comparison with males of other
|
||
<emphasis box="[1225,1313,675,698]" italics="true" pageId="4" pageNumber="427">
|
||
<taxonomicName box="[1225,1313,675,698]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="427" phylum="Arthropoda" rank="genus">Perania</taxonomicName>
|
||
</emphasis>
|
||
spp.) and completely fused to the pulmonary plate.
|
||
</paragraph>
|
||
<paragraph pageId="4" pageNumber="427">
|
||
<emphasis bold="true" box="[831,945,754,778]" pageId="4" pageNumber="427">Remarks:</emphasis>
|
||
When we received the specimens, they all had their opisthosomata detached, widely cut open and the gonads removed for an examination of the chromosomes. This may have caused some minor deformation of the dorsal plates which may make them look slightly narrower than on intact specimens. The
|
||
<typeStatus box="[884,978,945,968]" pageId="4" pageNumber="427">holotype</typeStatus>
|
||
has all legs of the left side missing, the allotype all legs on the right side. The only available female (the allotype) has its only remaining tarsus I incomplete. Since its tarsi II-IV are each one millimetre shorter than those of the male
|
||
<typeStatus box="[1163,1259,1073,1096]" pageId="4" pageNumber="427">holotype</typeStatus>
|
||
, we assume that this is also the case in tarsus I. This puts its length at
|
||
<quantity box="[831,915,1138,1162]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="427" unit="mm" value="1.7">1.7 mm</quantity>
|
||
, and the entire length of leg I at
|
||
<quantity box="[1261,1359,1138,1162]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="4" pageNumber="427" unit="mm" value="17.5">17.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="4" pageNumber="427">
|
||
<emphasis bold="true" box="[831,998,1185,1209]" pageId="4" pageNumber="427">Relationships:</emphasis>
|
||
Males of
|
||
<emphasis box="[1147,1261,1185,1209]" italics="true" pageId="4" pageNumber="427">
|
||
<taxonomicName box="[1147,1261,1185,1209]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="annam" status="sp. nov.">P. annam</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[1280,1378,1185,1209]" pageId="4" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authority="Schwendinger, 2013" authorityName="Schwendinger" authorityYear="2013" box="[831,1172,1216,1241]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="utara">
|
||
<emphasis box="[831,926,1217,1241]" italics="true" pageId="4" pageNumber="427">P. utara</emphasis>
|
||
Schwendinger,
|
||
<number box="[1117,1172,1217,1241]" pageId="4" pageNumber="427" value="2013.0">2013</number>
|
||
</taxonomicName>
|
||
(from Sumatra) share the conspicuous presence of conical setal bases on metatarsus II (not present in males of other congeners), but this is certainly a homoplasy. The strongly apomorphic palps of
|
||
<emphasis box="[1069,1171,1345,1369]" italics="true" pageId="4" pageNumber="427">
|
||
<taxonomicName box="[1069,1171,1345,1369]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="annam" status="sp. nov.">P. annam</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[1181,1268,1345,1369]" pageId="4" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<emphasis box="[1326,1436,1345,1369]" italics="true" pageId="4" pageNumber="427">
|
||
<taxonomicName box="[1326,1436,1345,1369]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="egregia">P. egregia</taxonomicName>
|
||
</emphasis>
|
||
are phylogenetically much more informative than non-genital characters and clearly show that these two species are very closely related. This is also reflected in the female copulatory organs: the dorsal and ventral wall of their anterior collar each has a posterior extension that ends in a more or less pronounced sclerite, the ventral one partly covering the entrance to the spermathecae. Such structures were not observed in other congeners.
|
||
<emphasis box="[1076,1262,1633,1657]" italics="true" pageId="4" pageNumber="427">
|
||
<taxonomicName box="[1076,1262,1633,1657]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="egregia">Perania egregia</taxonomicName>
|
||
</emphasis>
|
||
occurs at the northeastern periphery of the previously know range of the genus and thus is – together with
|
||
<taxonomicName authority="Schwendinger, 2013" authorityName="Schwendinger" authorityYear="2013" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="quadrifurcata">
|
||
<emphasis box="[1258,1436,1696,1721]" italics="true" pageId="4" pageNumber="427">P. quadrifurcata</emphasis>
|
||
Schwendinger,
|
||
<number box="[1007,1063,1729,1753]" pageId="4" pageNumber="427" value="2013.0">2013</number>
|
||
</taxonomicName>
|
||
, found a bit further south (see Schwendinger,
|
||
<number box="[997,1054,1761,1785]" pageId="4" pageNumber="427" value="2013.0">2013</number>
|
||
: fig.
|
||
<number box="[1107,1118,1760,1784]" pageId="4" pageNumber="427" value="1.0">1</number>
|
||
) – the geographically closest conspecific species of
|
||
<emphasis box="[1071,1169,1793,1817]" italics="true" pageId="4" pageNumber="427">
|
||
<taxonomicName box="[1071,1169,1793,1817]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="annam" status="sp. nov.">P. annam</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[1176,1259,1793,1817]" pageId="4" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="427" type="distribution">
|
||
<paragraph pageId="4" pageNumber="427">
|
||
<emphasis bold="true" box="[831,981,1840,1864]" pageId="4" pageNumber="427">Distribution:</emphasis>
|
||
This species is known only from Dalat (= Da Lat; exact locality unknown; approximately
|
||
<geoCoordinate box="[831,924,1904,1928]" direction="north" orientation="latitude" pageId="4" pageNumber="427" precision="925" value="11.95">
|
||
<number box="[831,856,1904,1928]" pageId="4" pageNumber="427" value="11.0">11</number>
|
||
°
|
||
<number box="[866,896,1904,1928]" pageId="4" pageNumber="427" value="57.0">57</number>
|
||
’
|
||
<collectionCode box="[901,924,1904,1928]" pageId="4" pageNumber="427">N</collectionCode>
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[937,1042,1903,1927]" direction="east" orientation="longitude" pageId="4" pageNumber="427" precision="925" value="108.433334">
|
||
<number box="[937,980,1903,1927]" pageId="4" pageNumber="427" value="108.0">108</number>
|
||
°
|
||
<number box="[989,1019,1903,1927]" pageId="4" pageNumber="427" value="26.0">26</number>
|
||
’
|
||
<collectionCode box="[1024,1042,1903,1927]" pageId="4" pageNumber="427">E</collectionCode>
|
||
</geoCoordinate>
|
||
), situated in the southern part of the Central Highlands of Vietnam, at about
|
||
<quantity box="[1266,1348,1936,1960]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="427" unit="m" value="1500.0">1500 m</quantity>
|
||
altitude (
|
||
<figureCitation box="[840,906,1967,1992]" captionStart="Fig. 1" captionStartId="1.[151,186,1679,1700]" captionTargetBox="[151,1436,257,1618]" captionTargetId="figure@1.[151,1436,257,1618]" captionTargetPageId="1" captionText="Fig. 1. Map of Southeast Asia showing localities of Perania annam sp. nov. (inverted triangle), Lamania bokor sp. nov. (triangle) and L. kraui Shear, 1978 (circles; empty circle indicates the type locality), and the currently known biogeographic range of the genera Perania (hatched areas) and Lamania (grey areas)." pageId="4" pageNumber="427">
|
||
Fig.
|
||
<number box="[893,906,1968,1992]" pageId="4" pageNumber="427" value="1.0">1</number>
|
||
</figureCitation>
|
||
, inverted triangle). Further
|
||
<emphasis box="[1216,1302,1968,1992]" italics="true" pageId="4" pageNumber="427">
|
||
<taxonomicName box="[1216,1302,1968,1992]" class="Arachnida" family="Tetrablemmidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="427" phylum="Arthropoda" rank="genus">Perania</taxonomicName>
|
||
</emphasis>
|
||
species can be expected to occur in Vietnam, Cambodia and Laos.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |