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<document id="4BF30AA318C9FE133B9C87B04640B6DB" ID-DOI="10.5070/P9401462457" ID-ISSN="0031-0298" ID-Zenodo-Dep="10913330" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1712167746961" checkinUser="felipe" docAuthor="Wheeler, Elisabeth A., Manchester, Steven R. &amp; Baas, Pieter" docDate="2023" docId="038AF505A33196315575FD9DFEC59FCD" docLanguage="en" docName="PaleoBios.40.14.1-55.pdf" docOrigin="PaleoBios 40 (14)" docSource="http://dx.doi.org/10.5070/p9401462457" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.3:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="3" docTitle="Styphnolobium SCHOTT 1830" docType="treatment" docVersion="4" lastPageNumber="15" masterDocId="FFB38D7DA33F9621547BFFC0FFD49B4F" masterDocTitle="A late Eocene wood assemblage from the Crooked River Basin, Oregon, USA" masterLastPageNumber="55" masterPageNumber="1" pageNumber="13" updateTime="1728048930804" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="369F2D0001A2EB2F486EFB4BF47116AC">A late Eocene wood assemblage from the Crooked River Basin, Oregon, USA</mods:title>
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<treatment id="038AF505A33196315575FD9DFEC59FCD" LSID="urn:lsid:plazi:treatment:038AF505A33196315575FD9DFEC59FCD" httpUri="http://treatment.plazi.org/id/038AF505A33196315575FD9DFEC59FCD" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13">
<subSubSection id="C3391798A331962F5575FD9DFDB199EF" pageId="14" pageNumber="13" type="nomenclature">
<paragraph id="8B9C4413A331962F5575FD9DFDB199EF" blockId="14.[270,642,533,712]" pageId="14" pageNumber="13">
<heading id="D0D4F37FA331962F5575FD9DFD569936" allCaps="true" box="[270,642,605,633]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="13" reason="2">
<taxonomicName id="4C233F90A331962F5575FD9DFD569936" ID-CoL="9CM5J" authority="SCHOTT, 1830" authorityName="SCHOTT" authorityYear="1830" box="[270,642,605,633]" class="Magnoliopsida" family="Fabaceae" genus="Styphnolobium" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="genus">
<emphasis id="B9579801A331962F5575FD9DFE309936" box="[270,484,605,633]" italics="true" pageId="14" pageNumber="13">STYPHNOLOBIUM</emphasis>
SCHOTT, 1830
</taxonomicName>
</heading>
CF.
<emphasis id="B9579801A331962F552EFD44FDE899EF" box="[341,572,644,672]" italics="true" pageId="14" pageNumber="13">STYPHONOLOBIUM</emphasis>
<collectionCode id="ED32DCD6A331962F5639FD43FDB599EF" box="[578,609,643,672]" country="Brazil" name="Instituto de Botânica" pageId="14" pageNumber="13" type="Herbarium">SP</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3391798A331962F55EDFD6FFDEC9EFB" pageId="14" pageNumber="13" type="description">
<paragraph id="8B9C4413A331962F55EDFD6FFE2E9987" blockId="14.[270,642,533,712]" box="[406,506,687,712]" pageId="14" pageNumber="13">
<figureCitation id="13185896A331962F55EDFD6FFE2E9987" box="[406,506,687,712]" captionStart="Figure 7" captionStartId="15.[185,260,1923,1947]" captionTargetBox="[151,1475,195,1887]" captionTargetId="figure-22@15.[151,1475,195,1887]" captionTargetPageId="15" captionText="Figure 7. Caption on pg, 15 Figure 7. Fabaceae. cf. Styphonolobium sp., UF 278-84867. A. Ring-porous wood, with distinct earlywood zone 23 vessels deep, confluent-banded axial parenchyma, TS. B. Pith and first formed secondary xylem, first formed growth ring not ring-porous, pith with thin-walled parenchyma cells, TS. C. Earlywood vessels mostly solitary, one radial pair; latewood vessels in small multiples, confluent parenchyma, TS. D. Crowded alternate pits, TLS. E. Vessel-ray parenchyma pits similar to intervessel pits, RLS. F. Multiseriate rays; axial parenchyma strands of 24 cells, TLS. G. Multiseriate rays; non-septate fibers, TLS. H. Simple perforation plate in narrow vessel element; heterocellular rays with procumbent and square cells, RLS. I. Pith (at right) with thin-walled parenchyma cells, upright cells common in rays nearest to pith (at left), RLS. Scale bars=500 µm in A; 200 µm in B, I; 100 µm in C, F; 50 µm in G, H; 20 µm in D, E." figureDoi="http://doi.org/10.5281/zenodo.10913344" httpUri="https://zenodo.org/record/10913344/files/figure.png" pageId="14" pageNumber="13">FIG. 7AI</figureCitation>
</paragraph>
<paragraph id="8B9C4413A331962F54EBFD1EFE3E9F16" blockId="14.[112,801,734,1947]" pageId="14" pageNumber="13">
<emphasis id="B9579801A331962F54EBFD1EFEC899B7" bold="true" box="[144,284,734,760]" pageId="14" pageNumber="13">Descriptio</emphasis>
n
<emphasis id="B9579801A331962F5556FD1EFE9E99B7" bold="true" box="[301,330,734,760]" pageId="14" pageNumber="13"></emphasis>
Growth rings distinct. Wood ring-porous. Earlywood vessels mostly solitary, rarely in radial multiples of two (
<figureCitation id="13185896A331962F5539FCE4FE6E9870" box="[322,442,804,831]" captionStart="Figure 7" captionStartId="15.[185,260,1923,1947]" captionTargetBox="[151,1475,195,1887]" captionTargetId="figure-22@15.[151,1475,195,1887]" captionTargetPageId="15" captionText="Figure 7. Caption on pg, 15 Figure 7. Fabaceae. cf. Styphonolobium sp., UF 278-84867. A. Ring-porous wood, with distinct earlywood zone 23 vessels deep, confluent-banded axial parenchyma, TS. B. Pith and first formed secondary xylem, first formed growth ring not ring-porous, pith with thin-walled parenchyma cells, TS. C. Earlywood vessels mostly solitary, one radial pair; latewood vessels in small multiples, confluent parenchyma, TS. D. Crowded alternate pits, TLS. E. Vessel-ray parenchyma pits similar to intervessel pits, RLS. F. Multiseriate rays; axial parenchyma strands of 24 cells, TLS. G. Multiseriate rays; non-septate fibers, TLS. H. Simple perforation plate in narrow vessel element; heterocellular rays with procumbent and square cells, RLS. I. Pith (at right) with thin-walled parenchyma cells, upright cells common in rays nearest to pith (at left), RLS. Scale bars=500 µm in A; 200 µm in B, I; 100 µm in C, F; 50 µm in G, H; 20 µm in D, E." figureDoi="http://doi.org/10.5281/zenodo.10913344" httpUri="https://zenodo.org/record/10913344/files/figure.png" pageId="14" pageNumber="13">Fig. 7AC</figureCitation>
), average tangential diameter 99 (
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=21) µm, range 56134 µm, earlywood zone 23 vessels deep with an abrupt transition to latewood with vessels solitary and in short radial multiples (
<figureCitation id="13185896A331962F56C8FC4EFCC698E7" box="[691,786,910,936]" captionStart="Figure 7" captionStartId="15.[185,260,1923,1947]" captionTargetBox="[151,1475,195,1887]" captionTargetId="figure-22@15.[151,1475,195,1887]" captionTargetPageId="15" captionText="Figure 7. Caption on pg, 15 Figure 7. Fabaceae. cf. Styphonolobium sp., UF 278-84867. A. Ring-porous wood, with distinct earlywood zone 23 vessels deep, confluent-banded axial parenchyma, TS. B. Pith and first formed secondary xylem, first formed growth ring not ring-porous, pith with thin-walled parenchyma cells, TS. C. Earlywood vessels mostly solitary, one radial pair; latewood vessels in small multiples, confluent parenchyma, TS. D. Crowded alternate pits, TLS. E. Vessel-ray parenchyma pits similar to intervessel pits, RLS. F. Multiseriate rays; axial parenchyma strands of 24 cells, TLS. G. Multiseriate rays; non-septate fibers, TLS. H. Simple perforation plate in narrow vessel element; heterocellular rays with procumbent and square cells, RLS. I. Pith (at right) with thin-walled parenchyma cells, upright cells common in rays nearest to pith (at left), RLS. Scale bars=500 µm in A; 200 µm in B, I; 100 µm in C, F; 50 µm in G, H; 20 µm in D, E." figureDoi="http://doi.org/10.5281/zenodo.10913344" httpUri="https://zenodo.org/record/10913344/files/figure.png" pageId="14" pageNumber="13">Fig. 7A</figureCitation>
); perforation plates simple (
<figureCitation id="13185896A331962F55C8FC71FDF99883" box="[435,557,945,972]" captionStart="Figure 7" captionStartId="15.[185,260,1923,1947]" captionTargetBox="[151,1475,195,1887]" captionTargetId="figure-22@15.[151,1475,195,1887]" captionTargetPageId="15" captionText="Figure 7. Caption on pg, 15 Figure 7. Fabaceae. cf. Styphonolobium sp., UF 278-84867. A. Ring-porous wood, with distinct earlywood zone 23 vessels deep, confluent-banded axial parenchyma, TS. B. Pith and first formed secondary xylem, first formed growth ring not ring-porous, pith with thin-walled parenchyma cells, TS. C. Earlywood vessels mostly solitary, one radial pair; latewood vessels in small multiples, confluent parenchyma, TS. D. Crowded alternate pits, TLS. E. Vessel-ray parenchyma pits similar to intervessel pits, RLS. F. Multiseriate rays; axial parenchyma strands of 24 cells, TLS. G. Multiseriate rays; non-septate fibers, TLS. H. Simple perforation plate in narrow vessel element; heterocellular rays with procumbent and square cells, RLS. I. Pith (at right) with thin-walled parenchyma cells, upright cells common in rays nearest to pith (at left), RLS. Scale bars=500 µm in A; 200 µm in B, I; 100 µm in C, F; 50 µm in G, H; 20 µm in D, E." figureDoi="http://doi.org/10.5281/zenodo.10913344" httpUri="https://zenodo.org/record/10913344/files/figure.png" pageId="14" pageNumber="13">Fig. 7D, H</figureCitation>
); intervessel pitting crowded alternate, polygonal in outline (
<figureCitation id="13185896A331962F5612FC14FD1298A0" box="[617,710,980,1007]" captionStart="Figure 7" captionStartId="15.[185,260,1923,1947]" captionTargetBox="[151,1475,195,1887]" captionTargetId="figure-22@15.[151,1475,195,1887]" captionTargetPageId="15" captionText="Figure 7. Caption on pg, 15 Figure 7. Fabaceae. cf. Styphonolobium sp., UF 278-84867. A. Ring-porous wood, with distinct earlywood zone 23 vessels deep, confluent-banded axial parenchyma, TS. B. Pith and first formed secondary xylem, first formed growth ring not ring-porous, pith with thin-walled parenchyma cells, TS. C. Earlywood vessels mostly solitary, one radial pair; latewood vessels in small multiples, confluent parenchyma, TS. D. Crowded alternate pits, TLS. E. Vessel-ray parenchyma pits similar to intervessel pits, RLS. F. Multiseriate rays; axial parenchyma strands of 24 cells, TLS. G. Multiseriate rays; non-septate fibers, TLS. H. Simple perforation plate in narrow vessel element; heterocellular rays with procumbent and square cells, RLS. I. Pith (at right) with thin-walled parenchyma cells, upright cells common in rays nearest to pith (at left), RLS. Scale bars=500 µm in A; 200 µm in B, I; 100 µm in C, F; 50 µm in G, H; 20 µm in D, E." figureDoi="http://doi.org/10.5281/zenodo.10913344" httpUri="https://zenodo.org/record/10913344/files/figure.png" pageId="14" pageNumber="13">Fig. 7D</figureCitation>
), 68.5 µm in horizontal diameter; vessel-ray parenchyma pits similar to intervessel pits (
<figureCitation id="13185896A331962F55D7FBDBFDD79F7A" box="[428,515,1051,1077]" captionStart="Figure 7" captionStartId="15.[185,260,1923,1947]" captionTargetBox="[151,1475,195,1887]" captionTargetId="figure-22@15.[151,1475,195,1887]" captionTargetPageId="15" captionText="Figure 7. Caption on pg, 15 Figure 7. Fabaceae. cf. Styphonolobium sp., UF 278-84867. A. Ring-porous wood, with distinct earlywood zone 23 vessels deep, confluent-banded axial parenchyma, TS. B. Pith and first formed secondary xylem, first formed growth ring not ring-porous, pith with thin-walled parenchyma cells, TS. C. Earlywood vessels mostly solitary, one radial pair; latewood vessels in small multiples, confluent parenchyma, TS. D. Crowded alternate pits, TLS. E. Vessel-ray parenchyma pits similar to intervessel pits, RLS. F. Multiseriate rays; axial parenchyma strands of 24 cells, TLS. G. Multiseriate rays; non-septate fibers, TLS. H. Simple perforation plate in narrow vessel element; heterocellular rays with procumbent and square cells, RLS. I. Pith (at right) with thin-walled parenchyma cells, upright cells common in rays nearest to pith (at left), RLS. Scale bars=500 µm in A; 200 µm in B, I; 100 µm in C, F; 50 µm in G, H; 20 µm in D, E." figureDoi="http://doi.org/10.5281/zenodo.10913344" httpUri="https://zenodo.org/record/10913344/files/figure.png" pageId="14" pageNumber="13">Fig. 7E</figureCitation>
); vessel elements short, average 213 µm (
<collectionCode id="ED32DCD6A331962F553DFBFEFEBA9F16" box="[326,366,1086,1113]" country="USA" lsid="urn:lsid:biocol.org:col:15659" name="San Diego Natural History Museum" pageId="14" pageNumber="13" type="Herbarium">SD</collectionCode>
=53, n=7).
</paragraph>
<paragraph id="8B9C4413A331962F54EBFBA1FD269F34" blockId="14.[112,801,734,1947]" box="[144,754,1121,1148]" pageId="14" pageNumber="13">
Fibers non-septate, pits not observed (
<figureCitation id="13185896A331962F560BFBA1FD329F34" box="[624,742,1121,1148]" captionStart="Figure 7" captionStartId="15.[185,260,1923,1947]" captionTargetBox="[151,1475,195,1887]" captionTargetId="figure-22@15.[151,1475,195,1887]" captionTargetPageId="15" captionText="Figure 7. Caption on pg, 15 Figure 7. Fabaceae. cf. Styphonolobium sp., UF 278-84867. A. Ring-porous wood, with distinct earlywood zone 23 vessels deep, confluent-banded axial parenchyma, TS. B. Pith and first formed secondary xylem, first formed growth ring not ring-porous, pith with thin-walled parenchyma cells, TS. C. Earlywood vessels mostly solitary, one radial pair; latewood vessels in small multiples, confluent parenchyma, TS. D. Crowded alternate pits, TLS. E. Vessel-ray parenchyma pits similar to intervessel pits, RLS. F. Multiseriate rays; axial parenchyma strands of 24 cells, TLS. G. Multiseriate rays; non-septate fibers, TLS. H. Simple perforation plate in narrow vessel element; heterocellular rays with procumbent and square cells, RLS. I. Pith (at right) with thin-walled parenchyma cells, upright cells common in rays nearest to pith (at left), RLS. Scale bars=500 µm in A; 200 µm in B, I; 100 µm in C, F; 50 µm in G, H; 20 µm in D, E." figureDoi="http://doi.org/10.5281/zenodo.10913344" httpUri="https://zenodo.org/record/10913344/files/figure.png" pageId="14" pageNumber="13">Fig. 7F, G</figureCitation>
).
</paragraph>
<paragraph id="8B9C4413A331962F54EBFB44FD179F8D" blockId="14.[112,801,734,1947]" pageId="14" pageNumber="13">
Axial parenchyma aliform-confluent-banded (
<figureCitation id="13185896A331962F56B3FB44FF519F8D" captionStart="Figure 7" captionStartId="15.[185,260,1923,1947]" captionTargetBox="[151,1475,195,1887]" captionTargetId="figure-22@15.[151,1475,195,1887]" captionTargetPageId="15" captionText="Figure 7. Caption on pg, 15 Figure 7. Fabaceae. cf. Styphonolobium sp., UF 278-84867. A. Ring-porous wood, with distinct earlywood zone 23 vessels deep, confluent-banded axial parenchyma, TS. B. Pith and first formed secondary xylem, first formed growth ring not ring-porous, pith with thin-walled parenchyma cells, TS. C. Earlywood vessels mostly solitary, one radial pair; latewood vessels in small multiples, confluent parenchyma, TS. D. Crowded alternate pits, TLS. E. Vessel-ray parenchyma pits similar to intervessel pits, RLS. F. Multiseriate rays; axial parenchyma strands of 24 cells, TLS. G. Multiseriate rays; non-septate fibers, TLS. H. Simple perforation plate in narrow vessel element; heterocellular rays with procumbent and square cells, RLS. I. Pith (at right) with thin-walled parenchyma cells, upright cells common in rays nearest to pith (at left), RLS. Scale bars=500 µm in A; 200 µm in B, I; 100 µm in C, F; 50 µm in G, H; 20 µm in D, E." figureDoi="http://doi.org/10.5281/zenodo.10913344" httpUri="https://zenodo.org/record/10913344/files/figure.png" pageId="14" pageNumber="13">Fig 7A, C</figureCitation>
); strands of 24 cells; non-storied. (
<figureCitation id="13185896A331962F563BFB68FD629F8D" box="[576,694,1192,1218]" captionStart="Figure 7" captionStartId="15.[185,260,1923,1947]" captionTargetBox="[151,1475,195,1887]" captionTargetId="figure-22@15.[151,1475,195,1887]" captionTargetPageId="15" captionText="Figure 7. Caption on pg, 15 Figure 7. Fabaceae. cf. Styphonolobium sp., UF 278-84867. A. Ring-porous wood, with distinct earlywood zone 23 vessels deep, confluent-banded axial parenchyma, TS. B. Pith and first formed secondary xylem, first formed growth ring not ring-porous, pith with thin-walled parenchyma cells, TS. C. Earlywood vessels mostly solitary, one radial pair; latewood vessels in small multiples, confluent parenchyma, TS. D. Crowded alternate pits, TLS. E. Vessel-ray parenchyma pits similar to intervessel pits, RLS. F. Multiseriate rays; axial parenchyma strands of 24 cells, TLS. G. Multiseriate rays; non-septate fibers, TLS. H. Simple perforation plate in narrow vessel element; heterocellular rays with procumbent and square cells, RLS. I. Pith (at right) with thin-walled parenchyma cells, upright cells common in rays nearest to pith (at left), RLS. Scale bars=500 µm in A; 200 µm in B, I; 100 µm in C, F; 50 µm in G, H; 20 µm in D, E." figureDoi="http://doi.org/10.5281/zenodo.10913344" httpUri="https://zenodo.org/record/10913344/files/figure.png" pageId="14" pageNumber="13">Fig. 7F, G</figureCitation>
).
</paragraph>
<paragraph id="8B9C4413A331962F54EBFB0BFDB69E03" blockId="14.[112,801,734,1947]" pageId="14" pageNumber="13">
Rays 14 (-5)-seriate, heterocellular (
<figureCitation id="13185896A331962F560AFB0AFD249FAA" box="[625,752,1226,1253]" captionStart="Figure 7" captionStartId="15.[185,260,1923,1947]" captionTargetBox="[151,1475,195,1887]" captionTargetId="figure-22@15.[151,1475,195,1887]" captionTargetPageId="15" captionText="Figure 7. Caption on pg, 15 Figure 7. Fabaceae. cf. Styphonolobium sp., UF 278-84867. A. Ring-porous wood, with distinct earlywood zone 23 vessels deep, confluent-banded axial parenchyma, TS. B. Pith and first formed secondary xylem, first formed growth ring not ring-porous, pith with thin-walled parenchyma cells, TS. C. Earlywood vessels mostly solitary, one radial pair; latewood vessels in small multiples, confluent parenchyma, TS. D. Crowded alternate pits, TLS. E. Vessel-ray parenchyma pits similar to intervessel pits, RLS. F. Multiseriate rays; axial parenchyma strands of 24 cells, TLS. G. Multiseriate rays; non-septate fibers, TLS. H. Simple perforation plate in narrow vessel element; heterocellular rays with procumbent and square cells, RLS. I. Pith (at right) with thin-walled parenchyma cells, upright cells common in rays nearest to pith (at left), RLS. Scale bars=500 µm in A; 200 µm in B, I; 100 µm in C, F; 50 µm in G, H; 20 µm in D, E." figureDoi="http://doi.org/10.5281/zenodo.10913344" httpUri="https://zenodo.org/record/10913344/files/figure.png" pageId="14" pageNumber="13">Fig. 7FH</figureCitation>
), in tangential view cells appear isodiametric and angular in outline, ray heights average 195 µm (
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=66), range 104313 µm.; 58 per mm; non-storied.
</paragraph>
<paragraph id="8B9C4413A331962F54EBFA95FE959EDD" blockId="14.[112,801,734,1947]" pageId="14" pageNumber="13">
<emphasis id="B9579801A331962F54EBFA95FEE39E20" bold="true" box="[144,311,1365,1391]" pageId="14" pageNumber="13">Specimen—</emphasis>
<collectionCode id="ED32DCD6A331962F5543FA96FE889E3F" box="[312,348,1366,1392]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="14" pageNumber="13" type="Museum">UF</collectionCode>
278-84867, estimated maximum diameter
<quantity id="4CDBE9F6A331962F5493FAB7FEE99EDD" box="[232,317,1399,1426]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="13" unit="cm" value="2.5">2.5 cm</quantity>
.
</paragraph>
<paragraph id="8B9C4413A331962F54EBFA5AFDEC9EFB" blockId="14.[112,801,734,1947]" box="[144,568,1434,1460]" pageId="14" pageNumber="13">
<emphasis id="B9579801A331962F54EBFA5AFE939EFB" bold="true" box="[144,327,1434,1460]" pageId="14" pageNumber="13">Occurrence—</emphasis>
Dietz Hill (
<collectionCode id="ED32DCD6A331962F55B1FA5AFE249EFB" box="[458,496,1434,1460]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="14" pageNumber="13" type="Museum">UF</collectionCode>
278).
</paragraph>
</subSubSection>
<subSubSection id="C3391798A331963154EBFA7DFEC59FCD" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13" type="discussion">
<paragraph id="8B9C4413A331962F54EBFA7DFD8F9C1B" blockId="14.[112,801,734,1947]" pageId="14" pageNumber="13">
<emphasis id="B9579801A331962F54EBFA7DFEEB9E98" bold="true" box="[144,319,1469,1495]" pageId="14" pageNumber="13">Comments—</emphasis>
Rays near the pith had more upright cells than did the rays farther away. We are not sure what the ray cellular composition of an older stem would be, although the tangential section taken at the periphery suggests they will become homocellular and/or heterocellular with a single marginal row of square cells. This sample contained insect frass and invading roots and had a total of 16 growth rings. Also, in such a narrow stem, vessel diameters probably are narrower than would characterize mature wood of this wood
<typeStatus id="5498FAB1A331962F560FF936FD7A9C5F" box="[628,686,1782,1808]" pageId="14" pageNumber="13">type</typeStatus>
. We did not observe helical thickenings in this sample, which is unusual for ring-porous legume woods.
</paragraph>
<paragraph id="8B9C4413A331962F54EBF89DFAA59835" blockId="14.[112,801,734,1947]" lastBlockId="14.[832,1521,195,1946]" pageId="14" pageNumber="13">
<emphasis id="B9579801A331962F54EBF89DFDE59C38" bold="true" box="[144,561,1885,1911]" pageId="14" pageNumber="13">Comparisons to extant woods—</emphasis>
One of our searches used 1p 3p (ring-porous wood), 7a 8a 9a 10a 11a (vessels randomly distributed, solitary and in short radial multiples), 13p 14a (perforation plates exclusively simple), 22p 26p (medium-sized alternate intervessel pitting), 30p (vessel-ray parenchyma pitting similar to intervessel pitting), 61p 66p (non-septate fibers with simple pits), 83p 85p (axial parenchyma confluent and in bands), 98p 105a (maximum ray width 410-seriate, and not composed exclusively of square/upright cells). This particular search only returned
<emphasis id="B9579801A331962F5174FE1CFCBA9955" italics="true" pageId="14" pageNumber="13">
<taxonomicName id="4C233F90A331962F5174FE1CFA1C9AB9" authorityName="SCHOTT" authorityYear="1830" box="[1295,1480,476,502]" class="Magnoliopsida" family="Fabaceae" genus="Styphnolobium" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="genus">Styphnolobium</taxonomicName>
af- fine
</emphasis>
(Torr. and
<collectionCode id="ED32DCD6A331962F578FFDC0FBDD9955" box="[1012,1033,512,538]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="14" pageNumber="13" type="Herbarium">A</collectionCode>
. Gray) Walp. (1842) (synonym
<taxonomicName id="4C233F90A331962F51F1FDC0FB6F9972" authority="Torr. and A. Gray, 1840" authorityName="Torr. and A. Gray" authorityYear="1840" class="Magnoliopsida" family="Fabaceae" genus="Sophora" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis id="B9579801A331962F51F1FDC0FC5F9972" italics="true" pageId="14" pageNumber="13">Sophora affinis</emphasis>
Torr. and A. Gray, 1840
</taxonomicName>
). The other searches we did, which were variations on the search given above, returned more legumes, as well as
<taxonomicName id="4C233F90A331962F517CFDA8FBCA99E8" authority="Siebold and Zucc. (1835)" authorityName="Siebold and Zucc." authorityYear="1835" class="Magnoliopsida" family="Paulowniaceae" genus="Paulownia" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="genus">
<emphasis id="B9579801A331962F517CFDA8FA5D99CB" box="[1287,1417,616,644]" italics="true" pageId="14" pageNumber="13">Paulownia</emphasis>
Siebold and Zucc. (1835)
</taxonomicName>
and some
<taxonomicName id="4C233F90A331962F50CAFD4CFA3F99E8" authority="Mart. (1820)" authorityName="Mart." authorityYear="1820" box="[1201,1515,652,679]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="family">Lamiaceae Mart. (1820)</taxonomicName>
, but invariably included
<taxonomicName id="4C233F90A331962F500CFD70FA249985" authority="Schott (1830)" authorityName="SCHOTT" authorityYear="1830" box="[1143,1520,688,714]" class="Magnoliopsida" family="Fabaceae" genus="Styphnolobium" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="genus">
<emphasis id="B9579801A331962F500CFD70FAE79985" box="[1143,1331,688,714]" italics="true" pageId="14" pageNumber="13">Styphnolobium</emphasis>
Schott (1830)
</taxonomicName>
in the results. This fossil has features in common with the
<taxonomicName id="4C233F90A331962F5715FD37FC00985C" box="[878,980,759,787]" class="Magnoliopsida" family="Fabaceae" genus="Sophora" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="genus">
<emphasis id="B9579801A331962F5715FD37FC00985C" box="[878,980,759,787]" italics="true" pageId="14" pageNumber="13">Sophora</emphasis>
</taxonomicName>
<collectionCode id="ED32DCD6A331962F57A2FD36FC3F985F" box="[985,1003,758,784]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="14" pageNumber="13" type="Herbarium">A</collectionCode>
group of
<bibRefCitation id="EFB239E2A331962F5024FD36FAFC985F" author="Fujii, T. &amp; P. Baas &amp; P. Gasson &amp; J. W. A. Ridder-Numan" box="[1119,1320,758,785]" pageId="14" pageNumber="13" refId="ref34938" refString="Fujii, T., P. Baas, P. Gasson and J. W. A. Ridder-Numan. 1994. Wood anatomy of the Sophora group (Leguminosae). Pp 205 - 249 in I. K. Ferguson and S. C. Tucker (eds.). Advances in Legume Systematics. 6. Structural Botany, Royal Botanic Gardens, Kew." type="journal volume" year="1994">Fujii et al. (1994</bibRefCitation>
,
<tableCitation id="C6A171A8A331962F514FFD36FA5F985F" box="[1332,1419,758,784]" captionStart="Table 1" captionStartId="10.[113,177,194,218]" captionTargetBox="[126,1490,424,1246]" captionTargetId="graphics-14@10.[112,1520,457,1205]" captionText="Table 1. Comparison of fossil Magnoliaceae woods in chronological order. K=Cretaceous, (C)=Campanian, (M)=Maastrichtian, Eo=Eocene, Olig=Oligocene; Mio=Miocene, e=early, m=middle, up=upper; IVP=intervessel pitting, S=scalariform, O=opposite; Bars/PP=number of bars per scalariform perforation plate; Si=simple perforation plates present; HT=helical thickenings in vessel elements; O/M= oil/mucilage cells; Y=present; N=absent; *=small axis, ()=not as common. References. 1=Cevallos-Ferriz and Stockey (1990); 2=Huang et al. (2020; 3=Huard (1967); 4=Iamandei et al. (2011); 5=Iamandei et al. (2020); 6=Page (1970); 7=Prakash et al. (1971); 8=Sakala et al. (2010); 9=Srivastava and Suzuki (2001); 10=Suzuki (1976); 11=Takahashi and Suzuki (2003); 12=van der Burgh (1973); 13=Wheeler et al. (1977); 14=Wheeler and Manchester (2002)." httpUri="http://table.plazi.org/id/DF5C149BA335962B540AFF02FB4C9A35" pageId="14" pageNumber="13" tableUuid="DF5C149BA335962B540AFF02FB4C9A35">Table 1</tableCitation>
), which includes
<emphasis id="B9579801A331962F57CBFCDAFBC3987C" box="[944,1047,793,820]" italics="true" pageId="14" pageNumber="13">
<taxonomicName id="4C233F90A331962F57CBFCDAFBC7987C" box="[944,1043,793,820]" class="Magnoliopsida" family="Fabaceae" genus="Sophora" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="species" species="affine">S. affine</taxonomicName>
.
</emphasis>
Gasson's 1994 review of the anatomy of the
<taxonomicName id="4C233F90A331962F57EAFCFCFBC29819" box="[913,1046,828,854]" pageId="14" pageNumber="51" rank="tribe" tribe="Sophoreae">Sophoreae</taxonomicName>
Spreng and DC. (1825) also indicates this fossil has a similarity with
<emphasis id="B9579801A331962F50B9FCA0FAA59835" box="[1218,1393,864,890]" italics="true" pageId="14" pageNumber="13">Stypholobium.</emphasis>
</paragraph>
<paragraph id="8B9C4413A331962F571BFC43FAC39DA5" blockId="14.[832,1521,195,1946]" pageId="14" pageNumber="13">
The
<bibRefCitation id="EFB239E2A331962F57E1FC43FB9C98D1" author="LPWG &amp; Legume Phylogeny Working Group" box="[922,1096,899,926]" pageId="14" pageNumber="13" pagination="44 - 77" refId="ref36907" refString="LPWG (Legume Phylogeny Working Group). 2017. A new subfamily classification of the Leguminosae based on a taxonomically comprehensive phylogeny. Taxon 66 (1): 44 - 77. [https: // www. jstor. org / stable / 90010911]." type="journal article" year="2017">LPWG (2017)</bibRefCitation>
proposed a new classification of the
<taxonomicName id="4C233F90A331962F5716FC66FC34988F" authorityName="LINDLEY" authorityYear="1836" box="[877,992,934,960]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
replacing the three traditional subfamilies: Caesalpinioideae DC. (1825), Mimosoideae DC. (1825), and Papilionoideae DC. (1825), which were based on floral characters, with six subfamilies: Caesalpinoideae, Cercidoideae
<bibRefCitation id="EFB239E2A331962F578BFBF3FB749F01" author="LPWG &amp; Legume Phylogeny Working Group" box="[1008,1184,1075,1102]" pageId="14" pageNumber="13" pagination="44 - 77" refId="ref36907" refString="LPWG (Legume Phylogeny Working Group). 2017. A new subfamily classification of the Leguminosae based on a taxonomically comprehensive phylogeny. Taxon 66 (1): 44 - 77. [https: // www. jstor. org / stable / 90010911]." type="journal article" year="2017">LPWG (2017)</bibRefCitation>
, Detarioideae Burmeister (1837), Dialioideae
<bibRefCitation id="EFB239E2A331962F503DFB96FB239F3E" author="LPWG &amp; Legume Phylogeny Working Group" box="[1094,1271,1110,1137]" pageId="14" pageNumber="13" pagination="44 - 77" refId="ref36907" refString="LPWG (Legume Phylogeny Working Group). 2017. A new subfamily classification of the Leguminosae based on a taxonomically comprehensive phylogeny. Taxon 66 (1): 44 - 77. [https: // www. jstor. org / stable / 90010911]." type="journal article" year="2017">LPWG (2017)</bibRefCitation>
, Duparquetioideae
<bibRefCitation id="EFB239E2A331962F573BFBB9FC3C9FDB" author="LPWG &amp; Legume Phylogeny Working Group" box="[832,1000,1145,1172]" pageId="14" pageNumber="13" pagination="44 - 77" refId="ref36907" refString="LPWG (Legume Phylogeny Working Group). 2017. A new subfamily classification of the Leguminosae based on a taxonomically comprehensive phylogeny. Taxon 66 (1): 44 - 77. [https: // www. jstor. org / stable / 90010911]." type="journal article" year="2017">LPWG (2017)</bibRefCitation>
, and Papilionoideae, as well as a mimosoid clade, based primarily on molecular work and supported by morphological data.
<taxonomicName id="4C233F90A331962F5025FB00FAC29F95" authorityName="SCHOTT" authorityYear="1830" box="[1118,1302,1216,1242]" class="Magnoliopsida" family="Fabaceae" genus="Styphnolobium" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="genus">
<emphasis id="B9579801A331962F5025FB00FAC29F95" box="[1118,1302,1216,1242]" italics="true" pageId="14" pageNumber="13">Styphnolobium</emphasis>
</taxonomicName>
is one of 503 genera in Papilionoideae and includes nine species (
<bibRefCitation id="EFB239E2A331962F51E4FB23FC559E6F" author="POWO" pageId="14" pageNumber="13" refId="ref38449" refString="POWO. 2023. Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet, https: // www. plantsoftheworldonline. org / Retrieved 2023." type="url" year="2023">POWO 2023</bibRefCitation>
). As best we can determine, there are wood anatomical descriptions for only two species:
<taxonomicName id="4C233F90A331962F5128FAEAFA249E0C" box="[1363,1520,1322,1348]" class="Magnoliopsida" family="Fabaceae" genus="Styphnolobium" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="species" species="japonicum">
<emphasis id="B9579801A331962F5128FAEAFA249E0C" box="[1363,1520,1322,1348]" italics="true" pageId="14" pageNumber="13">S. japonicum</emphasis>
</taxonomicName>
(
<collectionCode id="ED32DCD6A331962F5732FA8DFC8A9E28" box="[841,862,1357,1383]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="14" pageNumber="13" type="Herbarium">L</collectionCode>
.) Schott (1830) (native to central and southern.
<collectingCountry id="F3340483A331962F51E5FA8CFA3C9E28" box="[1438,1512,1356,1383]" name="China" pageId="14" pageNumber="13">China</collectingCountry>
) and
<taxonomicName id="4C233F90A331962F570CFAB0FC0D9EC5" box="[887,985,1392,1418]" class="Magnoliopsida" family="Fabaceae" genus="Sophora" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="species" species="affine">
<emphasis id="B9579801A331962F570CFAB0FC0D9EC5" box="[887,985,1392,1418]" italics="true" pageId="14" pageNumber="13">S. affine</emphasis>
</taxonomicName>
(native to
<collectingRegion id="49E78AF1A331962F5019FAB0FB039EC5" box="[1122,1239,1392,1418]" country="United States of America" name="Arkansas" pageId="14" pageNumber="13">Arkansas</collectingRegion>
,
<collectingRegion id="49E78AF1A331962F5098FAB0FA8A9EC5" box="[1251,1374,1392,1418]" country="United States of America" name="Louisiana" pageId="14" pageNumber="13">Louisiana</collectingRegion>
,
<collectingRegion id="49E78AF1A331962F5110FAB0FA389EC5" box="[1387,1516,1392,1418]" country="United States of America" name="Oklahoma" pageId="14" pageNumber="13">Oklahoma</collectingRegion>
, and
<collectingRegion id="49E78AF1A331962F5708FA54FC6C9EE1" box="[883,952,1428,1454]" country="United States of America" name="Texas" pageId="14" pageNumber="13">Texas</collectingRegion>
in the
<collectingCountry id="F3340483A331962F507DFA53FBE99EE2" box="[1030,1085,1427,1453]" name="United States of America" pageId="14" pageNumber="13">USA</collectingCountry>
) (
<bibRefCitation id="EFB239E2A331962F5028FA53FADF9EE2" author="Fujii, T. &amp; P. Baas &amp; P. Gasson &amp; J. W. A. Ridder-Numan" box="[1107,1291,1427,1454]" pageId="14" pageNumber="13" refId="ref34938" refString="Fujii, T., P. Baas, P. Gasson and J. W. A. Ridder-Numan. 1994. Wood anatomy of the Sophora group (Leguminosae). Pp 205 - 249 in I. K. Ferguson and S. C. Tucker (eds.). Advances in Legume Systematics. 6. Structural Botany, Royal Botanic Gardens, Kew." type="journal volume" year="1994">Fujii et al. 1994</bibRefCitation>
,
<bibRefCitation id="EFB239E2A331962F516DFA53FA679EE2" author="Gasson, P." box="[1302,1459,1427,1453]" pageId="14" pageNumber="13" refId="ref35008" refString="Gasson, P. 1994. Wood anatomy of the Tribe Sophoreae and related Caesalpinioidea and Papilionoideae. Pp. 165 - 203 in I. K. Ferguson and S. C. Tucker (eds.). Advances in Legume Systematics. 6. Structural Botany, Royal Botanic Gardens, Kew." type="book" year="1994">Gasson 1994</bibRefCitation>
,
<bibRefCitation id="EFB239E2A331962F51C5FA53FC109E9F" author="Itoh, T. B. &amp; Pan, P. &amp; Baas, J. &amp; Luo, D. &amp; Li, Y. &amp; Cui, F. &amp; Wang &amp; M. Mertz &amp; Y. Yasumoto" pageId="14" pageNumber="13" refId="ref36047" refString="Itoh, T. B. Pan, P. Baas, J. Luo, D. Li, Y. Cui, F. Wang. M. Mertz and Y. Yasumoto. 2022. Anatomical Atlas and Database of Chinese Woods. Kaiseisha Press. 2679 pp." type="book" year="2022">Itoh et al. 2022</bibRefCitation>
); both are ring-porous. The other species of
<taxonomicName id="4C233F90A331962F573BFA19FC2D9EBC" authorityName="SCHOTT" authorityYear="1830" box="[832,1017,1497,1523]" class="Magnoliopsida" family="Fabaceae" genus="Styphnolobium" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="genus">
<emphasis id="B9579801A331962F573BFA19FC2D9EBC" box="[832,1017,1497,1523]" italics="true" pageId="14" pageNumber="13">Styphnolobium</emphasis>
</taxonomicName>
are native to Central America, with one species extending into Columbia; some species grow in seasonally dry tropical biomes, some in wet tropics. The Chinese species belongs to Section
<taxonomicName id="4C233F90A331962F517BF983FC369DCF" authority="(Rudd) M. Sousa (1993)" authorityName="M. Sousa" authorityYear="1993" baseAuthorityName="Rudd" pageId="14" pageNumber="13" rank="section" section="Oresbios">
<emphasis id="B9579801A331962F517BF983FABC9D10" box="[1280,1384,1603,1631]" italics="true" pageId="14" pageNumber="13">Oresbios</emphasis>
(Rudd) M. Sousa (1993)
</taxonomicName>
; the American species to Section
<emphasis id="B9579801A331962F5114F9A7FC509DEC" italics="true" pageId="14" pageNumber="13">
<taxonomicName id="4C233F90A331962F5114F9A7FC549DEC" authorityName="SCHOTT" authorityYear="1830" class="Magnoliopsida" family="Fabaceae" genus="Styphnolobium" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="13" phylum="Tracheophyta" rank="genus">Styphnolobium</taxonomicName>
.
</emphasis>
Because of the relatively few samples available to us we are not sure whether there are wood anatomical differences between the two Sections.
</paragraph>
<paragraph id="8B9C4413A3319631571BF933FD449824" blockId="14.[832,1521,195,1946]" lastBlockId="16.[112,800,461,1154]" lastPageId="16" lastPageNumber="15" pageId="14" pageNumber="13">
<emphasis id="B9579801A331962F571BF933FB2B9C42" bold="true" box="[864,1279,1779,1805]" pageId="14" pageNumber="13">Comparisons to fossil woods—</emphasis>
The middle Eocene Clarno Nut Beds has
<specimenCount id="9D258F9AA331962F501BF8D6FB239C7E" box="[1120,1271,1814,1841]" count="3" pageId="14" pageNumber="13" type="generic" typeStatus="types">three types</specimenCount>
of legume woods; all differ from the Dietz Hill wood as they are diffuse-porous (
<bibRefCitation id="EFB239E2A331962F57D4F89CFAEB9C38" author="Wheeler, E. A. &amp; S. R. Manchester" box="[943,1343,1884,1911]" pageId="14" pageNumber="13" pagination="1 - 188" refId="ref41110" refString="Wheeler, E. A. and S. R. Manchester. 2002. Woods of the Eocene Nut Beds flora, Clarno Formation, Oregon, USA. IAWA Journal Supplement 3: 1 - 188." type="journal article" year="2002">
Wheeler and
<collectingRegion id="49E78AF1A331962F5025F89CFB279C39" box="[1118,1267,1884,1910]" country="United Kingdom" name="Manchester" pageId="14" pageNumber="13">Manchester</collectingRegion>
2002
</bibRefCitation>
).
<bibRefCitation id="EFB239E2A331962F5121F89CFBF39CD5" author="Muller-Stoll, W. R. &amp; E. Madel" pageId="14" pageNumber="13" pagination="95 - 174" refId="ref37836" refString="Muller-Stoll, W. R. and E. Madel. 1967. Die fossilen Leguminosen- Holzer. Eine Revision der mit Leguminosen verglichen fossilen Holzer und Beschreibungen alterer und neuer Arten. Palaeontographica 119 B: 95 - 174." type="journal article" year="1967">Müller-Stoll and Mädel (1967)</bibRefCitation>
reviewed the fossil wood record of the Leguminosae and at that time recognized only 19 fossil genera. Because it is difficult to distinguish some present-day legume genera from one another, most of their genera accommodate more than one present-day genus. Their key to the genera included only two ring-porous genera:
<taxonomicName id="4C233F90A32F96315576FDBEFCF499D7" authority="Muller-Stoll and Madel (1967)" authorityName="Muller-Stoll and Madel" authorityYear="1967" box="[269,800,637,666]" genus="Robinioxylon" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="16" pageNumber="15" rank="genus">
<emphasis id="B9579801A32F96315576FDBEFE7E99D5" box="[269,426,638,666]" italics="true" pageId="16" pageNumber="15">Robinioxylon</emphasis>
<bibRefCitation id="EFB239E2A32F963155D4FDBDFCF499D7" author="Muller-Stoll, W. R. &amp; E. Madel" box="[431,800,637,664]" pageId="16" pageNumber="15" pagination="95 - 174" refId="ref37836" refString="Muller-Stoll, W. R. and E. Madel. 1967. Die fossilen Leguminosen- Holzer. Eine Revision der mit Leguminosen verglichen fossilen Holzer und Beschreibungen alterer und neuer Arten. Palaeontographica 119 B: 95 - 174." type="journal article" year="1967">Müller-Stoll and Mädel (1967)</bibRefCitation>
</taxonomicName>
(differs in having latewood vessel clusters, storied axial parenchyma) and
<taxonomicName id="4C233F90A32F96315528FD04FF13984D" authority="Muller-Stoll and Madel (1967)" authorityName="Muller-Stoll and Madel" authorityYear="1967" class="Magnoliopsida" family="Fabaceae" genus="Gleditsioxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="16" pageNumber="15" phylum="Tracheophyta" rank="genus">
<emphasis id="B9579801A32F96315528FD04FE2999AF" box="[339,509,708,736]" italics="true" pageId="16" pageNumber="15">Gleditsioxylon</emphasis>
<bibRefCitation id="EFB239E2A32F9631567FFD04FF13984D" author="Muller-Stoll, W. R. &amp; E. Madel" pageId="16" pageNumber="15" pagination="95 - 174" refId="ref37836" refString="Muller-Stoll, W. R. and E. Madel. 1967. Die fossilen Leguminosen- Holzer. Eine Revision der mit Leguminosen verglichen fossilen Holzer und Beschreibungen alterer und neuer Arten. Palaeontographica 119 B: 95 - 174." type="journal article" year="1967">Müller-Stoll and Mädel (1967)</bibRefCitation>
</taxonomicName>
(rays often&gt;10-seriate). Given that present-day Leguminosae comprises over 750 genera (
<bibRefCitation id="EFB239E2A32F96315611FCCAFD2E986B" author="LPWG &amp; Legume Phylogeny Working Group" box="[618,762,778,804]" pageId="16" pageNumber="15" pagination="44 - 77" refId="ref36907" refString="LPWG (Legume Phylogeny Working Group). 2017. A new subfamily classification of the Leguminosae based on a taxonomically comprehensive phylogeny. Taxon 66 (1): 44 - 77. [https: // www. jstor. org / stable / 90010911]." type="journal article" year="2017">LPWG 2017</bibRefCitation>
),it is not surprising that there are fossil legume woods that do not fit the 19 genera recognized in 1967.
</paragraph>
<caption id="DF5C149BA330963154C2F843FEF69ADB" ID-DOI="http://doi.org/10.5281/zenodo.10913344" ID-Zenodo-Dep="10913344" httpUri="https://zenodo.org/record/10913344/files/figure.png" lastPageId="16" lastPageNumber="15" pageId="15" pageNumber="14" startId="15.[185,260,1923,1947]" targetBox="[151,1475,195,1887]" targetPageId="15" targetType="figure">
<paragraph id="8B9C4413A330962E54C2F843FE349CD4" blockId="15.[185,480,1923,1948]" box="[185,480,1923,1948]" pageId="15" pageNumber="14">
<emphasis id="B9579801A330962E54C2F843FECB9CD3" bold="true" box="[185,287,1923,1948]" pageId="15" pageNumber="14">Figure 7.</emphasis>
Caption on pg, 15
</paragraph>
<paragraph id="8B9C4413A32F9631540BFF01FEF69ADB" blockId="16.[112,1515,193,405]" pageId="16" pageNumber="15">
<emphasis id="B9579801A32F9631540BFF01FF029B95" bold="true" box="[112,214,193,218]" pageId="16" pageNumber="15">Figure 7.</emphasis>
<taxonomicName id="4C233F90A32F963154A0FF01FEEB9B96" authorityName="LINDLEY" authorityYear="1836" box="[219,319,193,217]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="16" pageNumber="15" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
. cf.
<emphasis id="B9579801A32F96315513FF01FDC09B96" box="[360,532,193,217]" italics="true" pageId="16" pageNumber="15">Styphonolobium</emphasis>
sp., UF 278-84867.
<emphasis id="B9579801A32F96315692FF01FD2E9B96" bold="true" box="[745,762,193,217]" pageId="16" pageNumber="15">A</emphasis>
. Ring-porous wood, with distinct earlywood zone 23 vessels deep, confluent-banded axial parenchyma, TS.
<emphasis id="B9579801A32F9631565EFF1CFDE19BBB" bold="true" box="[549,565,220,244]" pageId="16" pageNumber="15">B</emphasis>
. Pith and first formed secondary xylem, first formed growth ring not ring-porous, pith with thin-walled parenchyma cells, TS.
<emphasis id="B9579801A32F9631566EFF37FDF09A40" bold="true" box="[533,548,247,271]" pageId="16" pageNumber="15">C</emphasis>
. Earlywood vessels mostly solitary, one radial pair; latewood vessels in small multiples, confluent parenchyma, TS.
<emphasis id="B9579801A32F963155E8FED2FE719A65" bold="true" box="[403,421,274,298]" pageId="16" pageNumber="15">D</emphasis>
. Crowded alternate pits, TLS.
<emphasis id="B9579801A32F9631569EFED2FD209A65" bold="true" box="[741,756,274,298]" pageId="16" pageNumber="15">E</emphasis>
. Vessel-ray parenchyma pits similar to intervessel pits, RLS.
<emphasis id="B9579801A32F96315107FED2FA5E9A65" bold="true" box="[1404,1418,274,298]" pageId="16" pageNumber="15">F</emphasis>
. Multiseriate rays; axial parenchyma strands of 24 cells, TLS.
<emphasis id="B9579801A32F963156B7FEECFD089A0B" bold="true" box="[716,732,300,324]" pageId="16" pageNumber="15">G</emphasis>
. Multiseriate rays; non-septate fibers, TLS.
<emphasis id="B9579801A32F963150D6FEECFB6B9A0B" bold="true" box="[1197,1215,300,324]" pageId="16" pageNumber="15">H</emphasis>
. Simple perforation plate in narrow vessel element; heterocellular rays with procumbent and square cells, RLS.
<emphasis id="B9579801A32F96315794FE87FC2C9A10" bold="true" box="[1007,1016,327,351]" pageId="16" pageNumber="15">I</emphasis>
. Pith (at right) with thin-walled parenchyma cells, upright cells common in rays nearest to pith (at left), RLS. Scale bars=500 µm in A; 200 µm in B, I; 100 µm in C, F; 50 µm in G, H; 20 µm in D, E.
</paragraph>
</caption>
<paragraph id="8B9C4413A32F963154EBFCB4FEC59FCD" blockId="16.[112,800,461,1154]" pageId="16" pageNumber="15">
More recently, the genus
<taxonomicName id="4C233F90A32F963155B8FCB4FCF498C1" authority="Akkemik (2019)" authorityName="Akkemik" authorityYear="2019" box="[451,800,884,912]" genus="Cercioxylon" pageId="16" pageNumber="15" rank="genus">
<emphasis id="B9579801A32F963155B8FCB4FD9B98DF" box="[451,591,884,912]" italics="true" pageId="16" pageNumber="15">Cercioxylon</emphasis>
Akkemik (2019)
</taxonomicName>
was created for a Pliocene wood considered to resemble
<taxonomicName id="4C233F90A32F9631540BFC7AFF6C9899" box="[112,184,954,982]" class="Magnoliopsida" family="Fabaceae" genus="Cercis" kingdom="Plantae" order="Fabales" pageId="16" pageNumber="15" phylum="Tracheophyta" rank="genus">
<emphasis id="B9579801A32F9631540BFC7AFF6C9899" box="[112,184,954,982]" italics="true" pageId="16" pageNumber="15">Cercis</emphasis>
</taxonomicName>
<collectionCode id="ED32DCD6A32F963154BBFC7AFF06989B" box="[192,210,954,980]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="16" pageNumber="15" type="Herbarium">L</collectionCode>
. (1753) and described as having ring- to semi-ring-porous wood. It differs from this Dietz Hill wood because it does not have a well-defined narrow early- wood zone (Akkemik 2019, Plate
<collectionCode id="ED32DCD6A32F9631566EFBE3FDF39F72" box="[533,551,1059,1085]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="15" type="Museum">V</collectionCode>
) and it is described as having vessel clusters common and storied axial parenchyma.
</paragraph>
</subSubSection>
</treatment>
</document>