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<document id="4F21DE5273FA7BB396BEA2C1CA2951E5" ID-DOI="10.11646/phytotaxa.597.2.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="7929279" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1683889203312" checkinUser="plazi" docAuthor="Wan, Xia, Knapp, Ralf, Chen, De-Kui, He, Hai &amp; Zhang, Li-Bing" docDate="2023" docId="0384467FFFBDFF873FC307C3FDB52FC3" docLanguage="en" docName="phytotaxa.597.2.3.pdf" docOrigin="Phytotaxa 597 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.597.2.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.6:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="6" docTitle="Polystichum subobliquum Tagawa 1935" docType="treatment" docVersion="4" lastPageNumber="132" masterDocId="FFBD3E07FFB9FF833F4B026EFFD72907" masterDocTitle="Taxonomic notes on some East Asian pteridophytes" masterLastPageNumber="140" masterPageNumber="128" pageNumber="132" updateTime="1684155635051" updateUser="juliana" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="992A0B7BF72AD787CAF252927520EAEC">Taxonomic notes on some East Asian pteridophytes</mods:title>
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<mods:namePart id="2F23E9B4E501CC022E5A2172E876FEB7">Wan, Xia</mods:namePart>
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<mods:affiliation id="110ECDAC717107008925E85D48CA21FA">CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization, Chengdu Institute of Biology, Chinese Academy of Sciences, P. O. Box 416, Chengdu, Sichuan 610041, China; &amp; College of Life Sciences, Sichuan University, Chengdu 610065, China; &amp; University of Chinese Academy of Sciences, Beijing 100049, China; &amp; Missouri Botanical Garden, P. O. Box 299, St. Louis, Missouri 63166 - 0299, U. S. A. &amp; xwan @ mobot. org; https: // orcid. org / 0000 - 0001 - 8312 - 6917</mods:affiliation>
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<mods:namePart id="952B02C91AB717476DD71E8C30281105">Knapp, Ralf</mods:namePart>
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<mods:affiliation id="5DE8E6542AB54F38CEAC5DABA1588F87">Correspondent, The Muséum national dHistoire naturelle (MNHN, Paris, France), Steigestrasse 78, 69412 Eberbach, Germany; &amp; ralf. knapp @ gmail. org; https: // orcid. org / 0000 - 0002 - 0147 - 5687</mods:affiliation>
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<mods:namePart id="15F3DD6DE227BBEC55ABD41F14A11A4C">Chen, De-Kui</mods:namePart>
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<mods:affiliation id="64DC2B38A36C540B2A8D225F5D17E2A5">College of Life Sciences, Chongqing Normal University, Shapingba, Chongqing 401331, China; &amp; No. 1 Middle School of Tongren, Tong Ren, Guizhou 554300, China &amp; cjxby 739847 @ 163. com; https: // orcid. org / 0000 - 0002 - 1224 - 7372</mods:affiliation>
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<mods:namePart id="C69094E5DE0AF9FB3370FD12923618BD">He, Hai</mods:namePart>
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<mods:affiliation id="7FCF24723DF637AD75C18C31E5FB9125">College of Life Sciences, Chongqing Normal University, Shapingba, Chongqing 401331, China; &amp; hehai @ cqnu. edu. cn; https: // orcid. org / 0000 - 0001 - 9896 - 0971</mods:affiliation>
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<mods:namePart id="B1BA6055C51B334484506007033ADE24">Zhang, Li-Bing</mods:namePart>
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<mods:affiliation id="CF9810F1A636EB47FA445526E5E83DCC">CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization, Chengdu Institute of Biology, Chinese Academy of Sciences, P. O. Box 416, Chengdu, Sichuan 610041, China; &amp; Missouri Botanical Garden, P. O. Box 299, St. Louis, Missouri 63166 - 0299, U. S. A. &amp; Libing. Zhang @ mobot. org; https: // orcid. org / 0000 - 0002 - 4905 - 040 X</mods:affiliation>
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<mods:title id="6EA730D1BF2075132D9C95AAE685B2AC">Phytotaxa</mods:title>
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<mods:date id="7E4151E920848E308B9F9DB4791FCBDC">2023</mods:date>
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<subSubSection id="C337A4E2FFBDFF873FC307C3FCC72CCF" box="[136,784,1453,1481]" pageId="4" pageNumber="132" type="nomenclature">
<paragraph id="8B92F769FFBDFF873FC307C3FCC72CCF" blockId="4.[136,784,1453,1481]" box="[136,784,1453,1481]" pageId="4" pageNumber="132">
<heading id="D0DA4005FFBDFF873FC307C3FCC72CCF" box="[136,784,1453,1481]" fontSize="11" level="2" pageId="4" pageNumber="132" reason="3">
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<treatmentCitation id="0A8CD178FFBDFF873FC307C3FD772CCF" author="Tagawa, M." box="[136,672,1453,1481]" page=")" pageId="4" pageNumber="132" year="1935">
<taxonomicName id="4C2D8CEAFFBDFF873FC307C3FD772CCF" ID-CoL="786VJ" authority="Tagawa (1935: 138)" authorityName="Tagawa" authorityPageNumber="138" authorityYear="1935" box="[136,672,1453,1481]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="subobliquum">
<emphasis id="B9592B7BFFBDFF873FC307C3FE622CCF" bold="true" box="[136,437,1453,1480]" italics="true" pageId="4" pageNumber="132">Polystichum subobliquum</emphasis>
<bibRefCitation id="EFBC8A98FFBDFF873EF707C0FD772CCF" author="Tagawa, M." box="[444,672,1454,1481]" pageId="4" pageNumber="132" pagination="132 - 148" refId="ref5665" refString="Tagawa, M. (1935) Spicilegium Pteridographiae Asiae Orientalis 8. Acta Phytotaxonomica et Geobotanica 4: 132 - 148. https: // doi. org / 10.18942 / bunruichiri. KJ 00001078841" type="journal article" year="1935">Tagawa (1935: 138)</bibRefCitation>
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.
<figureCitation id="1316EBECFFBDFF873DE007C0FCDB2CCF" box="[683,780,1454,1480]" captionStart="FIGURE 6" captionStartId="8.[136,229,1935,1957]" captionTargetBox="[157,1426,199,1901]" captionTargetId="figure-17@8.[151,1436,190,1911]" captionTargetPageId="8" captionText="FIGURE 6. Polystichum subobliquum.—A. Habitat &amp; habit.—B. Lower portions of petioles showing scales.—C.Adaxial view of middle portion of lamina.—D, E. Abaxial views of portions of laminae (photo credit: Ralf Knapp)." figureDoi="http://doi.org/10.5281/zenodo.7929295" httpUri="https://zenodo.org/record/7929295/files/figure.png" pageId="4" pageNumber="132">Figure 6</figureCitation>
.
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</paragraph>
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<paragraph id="8B92F769FFBDFF873FC307BBFA9B2CED" blockId="4.[136,1356,1493,1515]" box="[136,1356,1493,1515]" pageId="4" pageNumber="132">
<materialsCitation id="3B45FD34FFBDFF873FC307BBFBA62CEC" box="[136,1137,1493,1515]" collectingDate="1934-10-04" collectionCode="M" country="Taiwan Island" elevation="2000" location="Taito" pageId="4" pageNumber="132" specimenCount="1" typeStatus="holotype">
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:—
<collectingCountry id="F33AB7F9FFBDFF873F9E07BBFEB62CEC" box="[213,353,1493,1515]" name="Taiwan Island" pageId="4" pageNumber="132">Taiwan Island</collectingCountry>
:
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(Taitung):
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, about
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,
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<collectingDate id="EFD72841FFBDFF873C9607BBFB872CED" box="[989,1104,1493,1514]" pageId="4" pageNumber="132" value="1934-10-04">4 Oct. 1934</collectingDate>
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,
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<emphasis id="B9592B7BFFBDFF873B1107B8FBA62CEC" box="[1114,1137,1494,1515]" italics="true" pageId="4" pageNumber="132">M</emphasis>
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<emphasis id="B9592B7BFFBDFF873B3A07B8FBA22CEC" box="[1137,1141,1494,1515]" italics="true" pageId="4" pageNumber="132">.</emphasis>
<materialsCitation id="3B45FD34FFBDFF873B3107B8FA9F2CED" box="[1146,1352,1493,1515]" collectionCode="KYO" country="Taiwan Island" location="Tagawa" pageId="4" pageNumber="132" specimenCount="1" typeStatus="holotype">
<emphasis id="B9592B7BFFBDFF873B3107B8FB2C2CED" box="[1146,1275,1493,1515]" italics="true" pageId="4" pageNumber="132">
<location id="8EF2A1B2FFBDFF873B3107B8FB122CEC" LSID="urn:lsid:plazi:treatment:0384467FFFBDFF873FC307C3FDB52FC3:8EF2A1B2FFBDFF873B3107B8FB122CEC" box="[1146,1221,1494,1515]" country="Taiwan Island" name="Tagawa" pageId="4" pageNumber="132">Tagawa</location>
1092
</emphasis>
(
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337A4E2FFBDFF873FC30474FDB52FC3" pageId="4" pageNumber="132" type="discussion">
<paragraph id="8B92F769FFBDFF873FC30474FDB52FC3" blockId="4.[136,1452,1562,1732]" pageId="4" pageNumber="132">
<taxonomicName id="4C2D8CEAFFBDFF873FC30474FE7B2F33" authorityName="Tagawa" authorityYear="1935" box="[136,428,1562,1588]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="subobliquum">
<emphasis id="B9592B7BFFBDFF873FC30474FE7B2F33" box="[136,428,1562,1588]" italics="true" pageId="4" pageNumber="132">Polystichum subobliquum</emphasis>
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is often treated as a synonym of
<taxonomicName id="4C2D8CEAFFBDFF873C6C0475FABD2F33" authority="(D. Don 1825: 3) T. Moore (1858: 87)" authorityName="T. Moore" authorityPageNumber="87" authorityYear="1858" baseAuthorityName="D. Don" baseAuthorityPageNumber="3" baseAuthorityYear="1825" box="[807,1386,1562,1589]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="obliquum">
<emphasis id="B9592B7BFFBDFF873C6C0475FC792F33" box="[807,942,1562,1588]" italics="true" pageId="4" pageNumber="132">P. obliquum</emphasis>
(D.
<bibRefCitation id="EFBC8A98FFBDFF873CA80474FBA62F33" author="Don, D." box="[995,1137,1562,1589]" pageId="4" pageNumber="132" refId="ref3357" refString="Don, D. (1825) Prodromus Florae Nepalensis. J. Gale, Londini, 256 pp." type="book" year="1825">Don 1825: 3</bibRefCitation>
) T.
<bibRefCitation id="EFBC8A98FFBDFF873BEA0474FABD2F33" author="Moore, T." box="[1185,1386,1562,1589]" pageId="4" pageNumber="132" refId="ref5189" refString="Moore, T. (1858) Index Filicum. W. Pamplin, London, 204 pp." type="book" year="1858">Moore (1858: 87)</bibRefCitation>
</taxonomicName>
(e.g.,
<bibRefCitation id="EFBC8A98FFBDFF873FC30450FEC42F5F" author="Knapp, R." box="[136,275,1598,1624]" pageId="4" pageNumber="132" refId="ref4087" refString="Knapp, R. (2011) Ferns and fern allies of Taiwan. Taipei: KBCC Press &amp; Yuan-Liou Publishing." type="book" year="2011">Knapp 2011</bibRefCitation>
,
<bibRefCitation id="EFBC8A98FFBDFF873E560450FD952F5F" author="Zhang, L. - B. &amp; Barrington, D. S." box="[285,578,1598,1624]" pageId="4" pageNumber="132" pagination="629 - 713" refId="ref7470" refString="Zhang, L. - B. &amp; Barrington, D. S. (2013) Polystichum. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. Y. (eds.) Flora of China, vol. 2 - 3 (Lycopodiaceae- Polypodiaceae). Science Press and St. Louis, Missouri Botanical Garden Press, Beijing, pp. 629 - 713." type="book chapter" year="2013">Zhang &amp; Barrington 2013</bibRefCitation>
,
<bibRefCitation id="EFBC8A98FFBDFF873D060450FD2C2F5F" author="Hsu, T. - C. &amp; Chen, C. - W. &amp; Knapp, R. &amp; Hung, H. - C." box="[589,763,1598,1624]" pageId="4" pageNumber="132" refId="ref4034" refString="Hsu, T. - C., Chen, C. - W., Knapp, R. &amp; Hung, H. - C. (2019) Illustrated Flora of Taiwan vol. 8: Ferns &amp; Lycophytes (I &amp; II). Maotouying Chuban, Taiwan." type="book" year="2019">
Hsu
<emphasis id="B9592B7BFFBDFF873DCB0451FD6F2F5F" box="[640,696,1598,1624]" italics="true" pageId="4" pageNumber="132">et al.</emphasis>
2019
</bibRefCitation>
). However,
<taxonomicName id="4C2D8CEAFFBDFF873CCB0451FBFB2F5F" authorityName="Tagawa" authorityYear="1935" box="[896,1068,1598,1624]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="subobliquum">
<emphasis id="B9592B7BFFBDFF873CCB0451FBFB2F5F" box="[896,1068,1598,1624]" italics="true" pageId="4" pageNumber="132">P. subobliquum</emphasis>
</taxonomicName>
(
<emphasis id="B9592B7BFFBDFF873B700451FADD2F5F" box="[1083,1290,1598,1624]" italics="true" pageId="4" pageNumber="132">Knapp 3625, 4178</emphasis>
, P!) has lower pinnae that are not reflexed and sori close to pinna margins, but
<taxonomicName id="4C2D8CEAFFBDFF873C10040DFC362F7B" authorityName="T. Moore" authorityYear="1858" baseAuthorityName="D. Don" baseAuthorityYear="1825" box="[859,993,1634,1660]" class="Polypodiopsida" family="Dryopteridaceae" genus="Polystichum" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="obliquum">
<emphasis id="B9592B7BFFBDFF873C10040DFC362F7B" box="[859,993,1634,1660]" italics="true" pageId="4" pageNumber="132">P. obliquum</emphasis>
</taxonomicName>
has lower pinnae often reflexed and sori closer to the midrib. Therefore, here we reinstate this species. It is endemic to
<collectingCountry id="F33AB7F9FFBDFF873CAC04E8FBEA2FA7" box="[999,1085,1670,1696]" name="Taiwan" pageId="4" pageNumber="132">Taiwan</collectingCountry>
. Our unpublished molecular data supports the separation of the two species.
</paragraph>
</subSubSection>
</treatment>
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