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<document id="823B1953D9BFB3376EBF71E204933181" ID-CLB-Dataset="26396" ID-DOI="10.11646/zootaxa.4621.1.1" ID-GBIF-Dataset="a381190d-7f50-4a66-bc33-3a0417c9f89f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3257228" ID-ZooBank="3D77F329-9BB8-4231-90E4-6A9E0250FCDF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1561539959806" checkinUser="plazi" docAuthor="Asenjo, Angélico, Irmler, Ulrich, Klimaszewski, Jan, Chandler, Donald S., Fierros-López, Hugo Eduardo &amp; Vieira, Juliana S." docDate="2019" docId="03978788B9BFFF08C1E36FAAACED985D" docLanguage="en" docName="zootaxa.4621.1.1.pdf" docOrigin="Zootaxa 4621 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Piestinae Erichson 1839" docType="treatment" docVersion="13" lastPageNumber="206" masterDocId="FFAEFFF0B974FFC5C1746833AE7B997A" masterDocTitle="Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution" masterLastPageNumber="406" masterPageNumber="1" pageNumber="204" updateTime="1698737271145" updateUser="ExternalLinkService">
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<mods:title id="99E814EACE91A762A9AD8258A8E67F99">Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution</mods:title>
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<mods:namePart id="9FD55D7EF5F91F61D514B0BCB2C7AE48">Chandler, Donald S.</mods:namePart>
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Subfamily
<taxonomicName id="4C3E4D1DB9BFFF0EC06B6FAAACF79EC9" ID-CoL="8VR5Z" authority="Erichson, 1839 a" authorityName="Erichson" authorityYear="1839" box="[287,652,1945,1971]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="203" pageNumber="204" phylum="Arthropoda" rank="subFamily" subFamily="Piestinae">
PIESTINAE
<bibRefCitation id="EFAF4B6FB9BFFF0EC0B06FAAACF79EC9" author="Erichson, G. F." box="[452,652,1945,1971]" pageId="203" pageNumber="204" refId="ref190452" refString="Erichson, G. F. (1839 a) Genera et species staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin. viii + 400 pp." type="book" year="1839">Erichson, 1839a</bibRefCitation>
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<emphasis id="B94AEA8CB9B8FF09C1E368ADAEAD99CD" bold="true" box="[151,214,158,183]" pageId="204" pageNumber="205">0709.</emphasis>
<taxonomicName id="4C3E4D1DB9B8FF09C1A968ADAC4299CD" authority="Erichson, 1839 a: 31" authorityName="Erichson" authorityPageNumber="31" authorityYear="1839" box="[221,569,158,183]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9B8FF09C1A968ADAF2A99CD" bold="true" box="[221,337,158,183]" italics="true" pageId="204" pageNumber="205">Hypotelus</emphasis>
<bibRefCitation id="EFAF4B6FB9B8FF09C02C68ADAC4299CD" author="Erichson, G. F." box="[344,569,158,183]" pageId="204" pageNumber="205" refId="ref190452" refString="Erichson, G. F. (1839 a) Genera et species staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin. viii + 400 pp." type="book" year="1839">Erichson, 1839a: 31</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13052A1BB9B8FF09C31968ADAC8F99CD" box="[621,756,158,183]" captionStart="FIGURES 76–87" captionStartId="327.[151,264,1860,1883]" captionTargetBox="[155,1435,193,1832]" captionTargetId="figure@327.[150,1436,193,1840]" captionTargetPageId="327" captionText="FIGURES 76–87. Habitus; Aleocharinae (76–80); Scaphidiinae (81–85); Piestinae (86–87); 76, after Kistner 2006; 77, after Borgmeier 1950; 78, after Pace 2007; 79, after Kistner 2006; 80, after Pace 2014c; 81, after Matthews 1888; 82, after Fierros- Lopéz 2005; 83, after Fierros-Lopéz 2005; 84, after Fierros-Lopéz 2006; 85, after Matthews 1888; 86, after Bortoluzzi et al. 2017; 87, Peru: department of Cuzco (CEMT); slightly modified. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3257246/files/figure.png" pageId="204" pageNumber="205">Figs. 86–87</figureCitation>
)
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<typeStatus id="5485883CB9B8FF09C1E368F5AEAA99A4" box="[151,209,198,222]" pageId="204" pageNumber="205">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9B8FF09C04368F6ACCC99A4" authority="Erichson, 1840" authorityName="Erichson" authorityYear="1840" box="[311,695,197,222]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B94AEA8CB9B8FF09C04368F6AC7C99A4" box="[311,519,197,222]" italics="true" pageId="204" pageNumber="205">Hypotelus pusillus</emphasis>
<bibRefCitation id="EFAF4B6FB9B8FF09C37968F6ACCC99A4" author="Erichson, G. F." box="[525,695,197,222]" pageId="204" pageNumber="205" pagination="1 - 5" refId="ref190520" refString="Erichson, G. F. (1840) Genera et species staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin. pp. 401 - 954, pls. 1 - 5. https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter" year="1840">Erichson, 1840</bibRefCitation>
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<treatmentCitationGroup id="AB2E11B0B9B8FF09C1BB68DEAA52987F" box="[207,1065,236,261]" pageId="204" pageNumber="205">
<taxonomicName id="4C3E4D1DB9B8FF09C1BB68DEAC2E987F" authority="Bernhauer, 1917 a: 45" authorityName="Bernhauer" authorityPageNumber="45" authorityYear="1917" box="[207,597,236,261]" class="Insecta" family="Staphylinidae" genus="Antropiestus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9B8FF09C1BB68DEAF26987E" box="[207,349,237,260]" italics="true" pageId="204" pageNumber="205">Antropiestus</emphasis>
<treatmentCitation id="0A9F108FB9B8FF09C01768DFAC2E987F" author="Bernhauer, M." box="[355,597,236,261]" page="45" pageId="204" pageNumber="205" year="1917">
<bibRefCitation id="EFAF4B6FB9B8FF09C01768DFAC2E987F" author="Bernhauer, M." box="[355,597,236,261]" pageId="204" pageNumber="205" pagination="45 - 48" refId="ref171806" refString="Bernhauer, M. (1917 a) Neue Arten der Gattungen Piestus, Leptochirus und Conosoma aus Sudamerika. (20. Beitrag.). Neue Beitrage zur systematischen Insektenkunde, 1 (6), 45 - 48." type="journal article" year="1917">Bernhauer, 1917a: 45</bibRefCitation>
</treatmentCitation>
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[synonymized by
<treatmentCitation id="0A9F108FB9B8FF09C25168DEAA58987F" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." box="[805,1059,236,261]" page="550" pageId="204" pageNumber="205" year="2012">
<bibRefCitation id="EFAF4B6FB9B8FF09C25168DEAA58987F" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." box="[805,1059,236,261]" pageId="204" pageNumber="205" pagination="490 - 585" refId="ref182077" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25 (6), 490 - 585. [Note: nominal year of publication 2011] https: // doi. org / 10.1071 / is 10016" type="journal article" year="2012">
Caron
<emphasis id="B94AEA8CB9B8FF09C20468DEADD3987F" box="[880,936,236,261]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2012: 550
</bibRefCitation>
</treatmentCitation>
]
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</paragraph>
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<paragraph id="8B81369EB9B8FF09C0736927AD629851" blockId="204.[151,1122,158,378]" box="[263,793,275,300]" pageId="204" pageNumber="205">
<typeStatus id="5485883CB9B8FF09C0736927AF3A9856" box="[263,321,276,300]" pageId="204" pageNumber="205">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9B8FF09C0D36927AD629851" authority="Bernhauer, 1917" authorityName="Bernhauer" authorityYear="1917" box="[423,793,275,300]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B94AEA8CB9B8FF09C0D36927AC239856" box="[423,600,275,300]" italics="true" pageId="204" pageNumber="205">Piestus andinus</emphasis>
Bernhauer, 1917
</taxonomicName>
</paragraph>
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<subSubSection id="C3246515B9B8FF09C1BB6908AA199829" box="[207,1122,314,339]" pageId="204" pageNumber="205" type="reference_group">
<paragraph id="8B81369EB9B8FF09C1BB6908AA199829" blockId="204.[151,1122,158,378]" box="[207,1122,314,339]" pageId="204" pageNumber="205">
<treatmentCitationGroup id="AB2E11B0B9B8FF09C1BB6908AA199829" box="[207,1122,314,339]" pageId="204" pageNumber="205">
<taxonomicName id="4C3E4D1DB9B8FF09C1BB6908ACF69829" authority="Scheerpeltz, 1952 a: 295" authorityName="Scheerpeltz" authorityPageNumber="295" authorityYear="1952" box="[207,653,314,339]" class="Insecta" family="Staphylinidae" genus="Eccoptopiestus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9B8FF09C1BB6908AF029828" box="[207,377,315,338]" italics="true" pageId="204" pageNumber="205">Eccoptopiestus</emphasis>
<treatmentCitation id="0A9F108FB9B8FF09C00B6909ACF69829" author="Scheerpeltz, O." box="[383,653,314,339]" page="295" pageId="204" pageNumber="205" year="1952">
<bibRefCitation id="EFAF4B6FB9B8FF09C00B6909ACF69829" author="Scheerpeltz, O." box="[383,653,314,339]" pageId="204" pageNumber="205" pagination="281 - 305" refId="ref234063" refString="Scheerpeltz, O. (1952 a) Revision der Gattung Piestus Gravh. (Coleoptera Staphyinidae). Revista Chilena de Entomologia, 2, 281 - 305." type="journal article" year="1952">Scheerpeltz, 1952a: 295</bibRefCitation>
</treatmentCitation>
</taxonomicName>
[synonymized by
<treatmentCitation id="0A9F108FB9B8FF09C2296909AA209829" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." box="[861,1115,314,339]" page="550" pageId="204" pageNumber="205" year="2012">
<bibRefCitation id="EFAF4B6FB9B8FF09C2296909AA209829" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." box="[861,1115,314,339]" pageId="204" pageNumber="205" pagination="490 - 585" refId="ref182077" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25 (6), 490 - 585. [Note: nominal year of publication 2011] https: // doi. org / 10.1071 / is 10016" type="journal article" year="2012">
Caron
<emphasis id="B94AEA8CB9B8FF09C2DC6908AD9A9829" box="[936,993,314,339]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2012: 550
</bibRefCitation>
</treatmentCitation>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9B8FF09C0736951ACB99AE6" pageId="204" pageNumber="205" type="materials_examined">
<paragraph id="8B81369EB9B8FF09C0736951ACA29803" blockId="204.[151,1122,158,378]" box="[263,729,353,378]" pageId="204" pageNumber="205">
<typeStatus id="5485883CB9B8FF09C0736951AF3A9800" box="[263,321,354,378]" pageId="204" pageNumber="205">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9B8FF09C0D36951ACA29803" authority="Solsky, 1872" authorityName="Solsky" authorityYear="1872" box="[423,729,353,378]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="B94AEA8CB9B8FF09C0D36951AC3A9800" box="[423,577,353,378]" italics="true" pageId="204" pageNumber="205">Piestus laevis</emphasis>
Solsky, 1872
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB9B8FF09C1E3699CAD6498B2" blockId="204.[151,1436,431,806]" box="[151,799,431,456]" pageId="204" pageNumber="205">13 species are included in this genus from Latin America.</paragraph>
<paragraph id="8B81369EB9B8FF09C1E369E5AFA89B47" blockId="204.[151,1436,431,806]" pageId="204" pageNumber="205">
Characters/Description:
<bibRefCitation id="EFAF4B6FB9B8FF09C0DC69E5ACF89895" author="Erichson, G. F." box="[424,643,470,495]" pageId="204" pageNumber="205" pagination="1 - 5" refId="ref190520" refString="Erichson, G. F. (1840) Genera et species staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin. pp. 401 - 954, pls. 1 - 5. https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter" year="1840">Erichson 1840: 840</bibRefCitation>
(characters);
<bibRefCitation id="EFAF4B6FB9B8FF09C26969E5AA6E9895" author="Blackwelder, R. E." box="[797,1045,470,495]" pageId="204" pageNumber="205" refId="ref175651" refString="Blackwelder, R. E. (1943) Monograph of the West-Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, viii + 658 pp. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book" year="1943">Blackwelder 1943: 41</bibRefCitation>
(characters in key and characters);
<bibRefCitation id="EFAF4B6FB9B8FF09C1BB69CEAC279B6C" author="Navarrete-Heredia, J. L. &amp; Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." box="[207,604,509,534]" pageId="204" pageNumber="205" refId="ref220178" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Universidad de Guadalajara y CONABIO, Mexico. xii + 401 pp." type="book" year="2002">
Navarrete-Heredia
<emphasis id="B94AEA8CB9B8FF09C0DC69CDAF999B6C" box="[424,482,509,534]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2002: 207
</bibRefCitation>
(characters in key);
<bibRefCitation id="EFAF4B6FB9B8FF09C23369CDAA3C9B6C" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." box="[839,1095,509,534]" pageId="204" pageNumber="205" pagination="490 - 585" refId="ref182077" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25 (6), 490 - 585. [Note: nominal year of publication 2011] https: // doi. org / 10.1071 / is 10016" type="journal article" year="2012">
Caron
<emphasis id="B94AEA8CB9B8FF09C2E069CDADB69B6C" box="[916,973,509,534]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2012: 550
</bibRefCitation>
(characters);
<bibRefCitation id="EFAF4B6FB9B8FF09C59069CEAF3B9B47" author="Bortoluzzi, S. &amp; Caron, E. &amp; Ribeiro-Costa, C. S." pageId="204" pageNumber="205" pagination="451 - 487" refId="ref177089" refString="Bortoluzzi, S., Caron, E. &amp; Ribeiro-Costa, C. S. (2017) Revision and phylogeny of Hypotelus Erichson: a Neotropical genus of minute rove beetles (Coleoptera, Staphylinidae, Piestinae). Zootaxa, 4273 (4), 451 - 487. https: // doi. org / 10.11646 / zootaxa. 4273.4.1" type="journal article" year="2017">
Bortoluzzi
<emphasis id="B94AEA8CB9B8FF09C41769CDABE79B6C" box="[1379,1436,509,534]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2017: 456
</bibRefCitation>
(characters).
</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36A7FAD789B19" blockId="204.[151,1436,431,806]" box="[151,771,587,612]" pageId="204" pageNumber="205">
Key:
<bibRefCitation id="EFAF4B6FB9B8FF09C1A06A78AC7A9B1E" author="Bortoluzzi, S. &amp; Caron, E. &amp; Ribeiro-Costa, C. S." box="[212,513,587,612]" pageId="204" pageNumber="205" pagination="451 - 487" refId="ref177089" refString="Bortoluzzi, S., Caron, E. &amp; Ribeiro-Costa, C. S. (2017) Revision and phylogeny of Hypotelus Erichson: a Neotropical genus of minute rove beetles (Coleoptera, Staphylinidae, Piestinae). Zootaxa, 4273 (4), 451 - 487. https: // doi. org / 10.11646 / zootaxa. 4273.4.1" type="journal article" year="2017">
Bortoluzzi
<emphasis id="B94AEA8CB9B8FF09C0256A7FAFF19B1E" box="[337,394,587,612]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2017: 458
</bibRefCitation>
(species of the genus).
</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36A41AF5E9BF0" blockId="204.[151,1436,431,806]" box="[151,293,626,651]" pageId="204" pageNumber="205">Distribution.</paragraph>
<paragraph id="8B81369EB9B8FF09C1DE6AA9AA009BC8" blockId="204.[151,1436,431,806]" box="[170,1147,665,690]" pageId="204" pageNumber="205">
West Indies:
<collectingCountry id="F329760EB9B8FF09C0496AAAAF009BC8" box="[317,379,665,690]" name="Cuba" pageId="204" pageNumber="205">Cuba</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C0F26AAAAF989BC8" box="[390,483,665,690]" name="Jamaica" pageId="204" pageNumber="205">Jamaica</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C09B6AAAACA19BC8" box="[495,730,665,690]" name="Dominican Republic" pageId="204" pageNumber="205">Dominican Republic</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C3916AAAAA729BC8" box="[741,1033,665,690]" name="Saint Vincent and the Grenadines" pageId="204" pageNumber="205">St. Vincent &amp; Grenadines</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C5616AAAAA039BC8" box="[1045,1144,665,690]" name="Grenada" pageId="204" pageNumber="205">Grenada</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9B8FF09C1DE6AF3AD3D9BA2" blockId="204.[151,1436,431,806]" box="[170,838,704,729]" pageId="204" pageNumber="205">
Central America:
<collectingCountry id="F329760EB9B8FF09C0066AF3AFB19BA3" box="[370,458,704,729]" name="Mexico" pageId="204" pageNumber="205">Mexico</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C0A26AF3AC299BA3" box="[470,594,704,729]" name="Guatemala" pageId="204" pageNumber="205">Guatemala</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C3296AF3ACA19BA3" box="[605,730,704,729]" name="Costa Rica" pageId="204" pageNumber="205">Costa Rica</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C3916AF2AD3A9BA2" box="[741,833,705,728]" name="Panama" pageId="204" pageNumber="205">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9B8FF09C1DE6AD4AA149A7A" blockId="204.[151,1436,431,806]" box="[170,1135,743,768]" pageId="204" pageNumber="205">
South America:
<collectingCountry id="F329760EB9B8FF09C0156AD4AFA99A7A" box="[353,466,743,768]" name="Colombia" pageId="204" pageNumber="205">Colombia</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C0AA6AD4AC2C9A7A" box="[478,599,743,768]" name="Venezuela" pageId="204" pageNumber="205">Venezuela</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C3166AD4AD709A7A" box="[610,779,743,768]" name="French Guiana" pageId="204" pageNumber="205">French Guiana</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C2626AD4AD0D9A7A" box="[790,886,743,768]" name="Ecuador" pageId="204" pageNumber="205">Ecuador</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C2F56ADBADCC9B85" box="[897,951,744,767]" name="Peru" pageId="204" pageNumber="205">Peru</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C2B66AD4AA6C9A7A" box="[962,1047,743,768]" name="Bolivia" pageId="204" pageNumber="205">Bolivia</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C5576AD4AA109A7A" box="[1059,1131,743,768]" name="Brazil" pageId="204" pageNumber="205">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9B8FF09C1DE6B3EAFC29A5C" blockId="204.[151,1436,431,806]" box="[170,441,781,806]" pageId="204" pageNumber="205">Other regions: Nearctic.</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36B6FAD799A0E" blockId="204.[151,1436,860,1781]" box="[151,770,860,885]" pageId="204" pageNumber="205">
<emphasis id="B94AEA8CB9B8FF09C1E36B6FAEAD9A0F" bold="true" box="[151,214,860,885]" pageId="204" pageNumber="205">0710.</emphasis>
<taxonomicName id="4C3E4D1DB9B8FF09C1A96B6FAC449A0E" authority="Gravenhorst, 1806: 223" authorityName="Gravenhorst" authorityPageNumber="223" authorityYear="1806" box="[221,575,860,885]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9B8FF09C1A96B6FAF4B9A0F" bold="true" box="[221,304,860,885]" italics="true" pageId="204" pageNumber="205">Piestus</emphasis>
<bibRefCitation id="EFAF4B6FB9B8FF09C0426B6FAC449A0E" author="Gravenhorst, J. L. C." box="[310,575,860,885]" pageId="204" pageNumber="205" refId="ref197065" refString="Gravenhorst, J. L. C. (1806) Monographia Coleopterorum Micropterorum. H. Dieterich, Gottingae [Gottingen]. xvi + 248 pp., 1 pl." type="book" year="1806">Gravenhorst, 1806: 223</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13052A1BB9B8FF09C3076B6FAC819A0E" box="[627,762,860,885]" captionStart="FIGURES 88–99" captionStartId="328.[151,264,1862,1885]" captionTargetBox="[156,1436,198,1832]" captionTargetId="figure@328.[151,1436,193,1840]" captionTargetPageId="328" captionText="FIGURES 88–99. Habitus; Piestinae (88–91); Osoriinae (92–99); 88, after Pérez et al. 2017; 89, Peru: department of Madre de Dios (MUSM); 90, Peru: department of Cuzco (MUSM); 91, after Brunke et al. 2011; 92, after Sharp 1887a; 93, after Sharp 1887a; 94, after Irmler 1991; 95, after Irmler 2001b; 96, after Bierig 1931; 97, after Sharp 1887a; 98, after Caron et al. 2012; 99, after Bruch 1942; slightly modified. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3257248/files/figure.png" pageId="204" pageNumber="205">Figs. 88–90</figureCitation>
)
</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36BB7ACB99AE6" blockId="204.[151,1436,860,1781]" box="[151,706,899,924]" pageId="204" pageNumber="205">
<typeStatus id="5485883CB9B8FF09C1E36BB7AEAA9AE6" box="[151,209,900,924]" pageId="204" pageNumber="205">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9B8FF09C0436BB7ACB99AE6" authority="Gravenhorst, 1806" authorityName="Gravenhorst" authorityYear="1806" box="[311,706,899,924]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="B94AEA8CB9B8FF09C0436BB7AF909AE6" box="[311,491,899,924]" italics="true" pageId="204" pageNumber="205">Piestus sulcatus</emphasis>
<bibRefCitation id="EFAF4B6FB9B8FF09C0866BB0ACB99AE6" author="Gravenhorst, J. L. C." box="[498,706,899,924]" pageId="204" pageNumber="205" refId="ref197065" refString="Gravenhorst, J. L. C. (1806) Monographia Coleopterorum Micropterorum. H. Dieterich, Gottingae [Gottingen]. xvi + 248 pp., 1 pl." type="book" year="1806">Gravenhorst, 1806</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9B8FF09C1BB6B99AA379AB9" box="[207,1100,938,963]" pageId="204" pageNumber="205" type="reference_group">
<paragraph id="8B81369EB9B8FF09C1BB6B99AA379AB9" blockId="204.[151,1436,860,1781]" box="[207,1100,938,963]" pageId="204" pageNumber="205">
<treatmentCitationGroup id="AB2E11B0B9B8FF09C1BB6B99AA379AB9" box="[207,1100,938,963]" pageId="204" pageNumber="205">
<taxonomicName id="4C3E4D1DB9B8FF09C1BB6B99AC039AB8" authority="Scheerpeltz, 1952 a: 294" authorityName="Scheerpeltz" authorityPageNumber="294" authorityYear="1952" box="[207,632,938,963]" class="Insecta" family="Staphylinidae" genus="Elytropiestus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9B8FF09C1BB6B99AF189AB9" box="[207,355,938,963]" italics="true" pageId="204" pageNumber="205">Elytropiestus</emphasis>
<treatmentCitation id="0A9F108FB9B8FF09C01E6B99AC039AB8" author="Scheerpeltz, O." box="[362,632,938,963]" page="294" pageId="204" pageNumber="205" year="1952">
<bibRefCitation id="EFAF4B6FB9B8FF09C01E6B99AC039AB8" author="Scheerpeltz, O." box="[362,632,938,963]" pageId="204" pageNumber="205" pagination="281 - 305" refId="ref234063" refString="Scheerpeltz, O. (1952 a) Revision der Gattung Piestus Gravh. (Coleoptera Staphyinidae). Revista Chilena de Entomologia, 2, 281 - 305." type="journal article" year="1952">Scheerpeltz, 1952a: 294</bibRefCitation>
</treatmentCitation>
</taxonomicName>
[synonymized by
<treatmentCitation id="0A9F108FB9B8FF09C2336B99AA3E9AB9" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." box="[839,1093,938,963]" page="500" pageId="204" pageNumber="205" year="2012">
<bibRefCitation id="EFAF4B6FB9B8FF09C2336B99AA3E9AB9" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." box="[839,1093,938,963]" pageId="204" pageNumber="205" pagination="490 - 585" refId="ref182077" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25 (6), 490 - 585. [Note: nominal year of publication 2011] https: // doi. org / 10.1071 / is 10016" type="journal article" year="2012">
Caron
<emphasis id="B94AEA8CB9B8FF09C2E66B98ADB09AB9" box="[914,971,938,963]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2012: 500
</bibRefCitation>
</treatmentCitation>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9B8FF09C0736BE1AD4C9A93" box="[263,823,977,1002]" pageId="204" pageNumber="205" type="materials_examined">
<paragraph id="8B81369EB9B8FF09C0736BE1AD4C9A93" blockId="204.[151,1436,860,1781]" box="[263,823,977,1002]" pageId="204" pageNumber="205">
<typeStatus id="5485883CB9B8FF09C0736BE1AF3A9A90" box="[263,321,978,1002]" pageId="204" pageNumber="205">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9B8FF09C0D36BE1AD4C9A93" authority="Bernhauer, 1917" authorityName="Bernhauer" authorityYear="1917" box="[423,823,977,1002]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="B94AEA8CB9B8FF09C0D36BE1AC0E9A90" box="[423,629,977,1002]" italics="true" pageId="204" pageNumber="205">Piestus paradoxus</emphasis>
Bernhauer, 1917
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9B8FF09C1BB6BCBAA2E9D6B" box="[207,1109,1016,1041]" pageId="204" pageNumber="205" type="reference_group">
<paragraph id="8B81369EB9B8FF09C1BB6BCBAA2E9D6B" blockId="204.[151,1436,860,1781]" box="[207,1109,1016,1041]" pageId="204" pageNumber="205">
<treatmentCitationGroup id="AB2E11B0B9B8FF09C1BB6BCBAA2E9D6B" box="[207,1109,1016,1041]" pageId="204" pageNumber="205">
<taxonomicName id="4C3E4D1DB9B8FF09C1BB6BCBACFA9D6A" authority="Scheerpeltz, 1952 a: 291" authorityName="Scheerpeltz" authorityPageNumber="291" authorityYear="1952" box="[207,641,1016,1041]" class="Insecta" family="Staphylinidae" genus="Trachypiestus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9B8FF09C1BB6BCBAF179D6B" box="[207,364,1016,1041]" italics="true" pageId="204" pageNumber="205">Trachypiestus</emphasis>
<treatmentCitation id="0A9F108FB9B8FF09C0076BCBACFA9D6A" author="Scheerpeltz, O." box="[371,641,1016,1041]" page="291" pageId="204" pageNumber="205" year="1952">
<bibRefCitation id="EFAF4B6FB9B8FF09C0076BCBACFA9D6A" author="Scheerpeltz, O." box="[371,641,1016,1041]" pageId="204" pageNumber="205" pagination="281 - 305" refId="ref234063" refString="Scheerpeltz, O. (1952 a) Revision der Gattung Piestus Gravh. (Coleoptera Staphyinidae). Revista Chilena de Entomologia, 2, 281 - 305." type="journal article" year="1952">Scheerpeltz, 1952a: 291</bibRefCitation>
</treatmentCitation>
</taxonomicName>
[synonymized by
<treatmentCitation id="0A9F108FB9B8FF09C2246BCBAA359D6B" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." box="[848,1102,1016,1041]" page="500" pageId="204" pageNumber="205" year="2012">
<bibRefCitation id="EFAF4B6FB9B8FF09C2246BCBAA359D6B" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." box="[848,1102,1016,1041]" pageId="204" pageNumber="205" pagination="490 - 585" refId="ref182077" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25 (6), 490 - 585. [Note: nominal year of publication 2011] https: // doi. org / 10.1071 / is 10016" type="journal article" year="2012">
Caron
<emphasis id="B94AEA8CB9B8FF09C2E86BCAADAF9D6B" box="[924,980,1016,1041]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2012: 500
</bibRefCitation>
</treatmentCitation>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9B8FF09C0736C13AD7F9D4D" box="[263,772,1054,1080]" pageId="204" pageNumber="205" type="materials_examined">
<paragraph id="8B81369EB9B8FF09C0736C13AD7F9D4D" blockId="204.[151,1436,860,1781]" box="[263,772,1054,1080]" pageId="204" pageNumber="205">
<typeStatus id="5485883CB9B8FF09C0736C13AF3A9D42" box="[263,321,1056,1080]" pageId="204" pageNumber="205">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9B8FF09C0D36C13AD7F9D4D" authority="Fauvel, 1864" authorityName="Fauvel" authorityYear="1864" box="[423,772,1054,1079]" class="Insecta" family="Staphylinidae" genus="Piestus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="angularis">
<emphasis id="B94AEA8CB9B8FF09C0D36C13AC109D4D" box="[423,619,1054,1079]" italics="true" pageId="204" pageNumber="205">Piestus angularis</emphasis>
<bibRefCitation id="EFAF4B6FB9B8FF09C3056C2DAD7F9D4D" author="Fauvel, A." box="[625,772,1054,1079]" pageId="204" pageNumber="205" pagination="3 - 62" refId="ref191389" refString="Fauvel, A. (1864) Etudes sur les Staphylinides de l'Amerique centrale, principalement du Mexique. Notices Entomologiques, 2, 3 - 62, 66, pl. 1." type="journal article" year="1864">Fauvel, 1864</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9B8FF09C1BB6C76AABF9D24" box="[207,1220,1093,1118]" pageId="204" pageNumber="205" type="reference_group">
<paragraph id="8B81369EB9B8FF09C1BB6C76AABF9D24" blockId="204.[151,1436,860,1781]" box="[207,1220,1093,1118]" pageId="204" pageNumber="205">
<treatmentCitationGroup id="AB2E11B0B9B8FF09C1BB6C76AABF9D24" box="[207,1220,1093,1118]" pageId="204" pageNumber="205">
<taxonomicName id="4C3E4D1DB9B8FF09C1BB6C76AD679D24" ID-CoL="63W2B" authority="Gray in Griffith &amp; Pidgeon, 1831: 306" authorityName="Gray in Griffith &amp; Pidgeon" authorityPageNumber="306" authorityYear="1831" box="[207,796,1093,1118]" class="Magnoliopsida" family="Asteraceae" genus="Trichocoryne" kingdom="Plantae" order="Asterales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AEA8CB9B8FF09C1BB6C76AF1D9D24" box="[207,358,1093,1118]" italics="true" pageId="204" pageNumber="205">Trichocoryne</emphasis>
Gray
<emphasis id="B94AEA8CB9B8FF09C0D86C74AFB99D24" box="[428,450,1095,1118]" italics="true" pageId="204" pageNumber="205">in</emphasis>
<treatmentCitation id="0A9F108FB9B8FF09C0BC6C76AD679D24" author="Griffith, E. &amp; Pidgeon, E." box="[456,796,1093,1118]" page="306" pageId="204" pageNumber="205" year="1831">
<bibRefCitation id="EFAF4B6FB9B8FF09C0BC6C76AD679D24" author="Griffith, E. &amp; Pidgeon, E." box="[456,796,1093,1118]" pageId="204" pageNumber="205" pagination="1832" refId="ref197260" refString="Griffith, E. &amp; Pidgeon, E. (1831) The Class Insecta arranged by the Baron Cuvier, with supplementary additions to each order. Volume 1. In: Cuvier, G. (Ed.), The animal kingdom arranged in conformity with its organization. Vol. 14. Whittaker, Treacher and Co., London. viii + 570. [Note: nominal year of publication 1832]" type="journal article" year="1831">Griffith &amp; Pidgeon, 1831: 306</bibRefCitation>
</treatmentCitation>
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[synonymized by
<treatmentCitation id="0A9F108FB9B8FF09C2986C74AAC69D24" author="Laporte, F. L." box="[1004,1213,1093,1118]" page="126" pageId="204" pageNumber="205" year="1835">
<bibRefCitation id="EFAF4B6FB9B8FF09C2986C74AAC69D24" author="Laporte, F. L." box="[1004,1213,1093,1118]" pageId="204" pageNumber="205" pagination="95 - 159" refId="ref211693" refString="Laporte, F. L. (1835) Etudes Entomologiques, ou description d'insectes nouveaux, et observations sur la synonymie. Premiere Partie, Carnassiers. Mequignon-Marvis Pere et Fils, Paris, pp. 95 - 159, 4 pls." type="book chapter" year="1835">Laporte 1835: 126</bibRefCitation>
</treatmentCitation>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9B8FF09C0736C5DAD2F9DFF" box="[263,852,1132,1158]" pageId="204" pageNumber="205" type="materials_examined">
<paragraph id="8B81369EB9B8FF09C0736C5DAD2F9DFF" blockId="204.[151,1436,860,1781]" box="[263,852,1132,1158]" pageId="204" pageNumber="205">
<typeStatus id="5485883CB9B8FF09C0736C5DAF3A9DFC" box="[263,321,1134,1158]" pageId="204" pageNumber="205">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9B8FF09C0D36C5FAD2F9DFF" authority="Dalman, 1821" authorityName="Dalman" authorityYear="1821" box="[423,852,1132,1157]" class="Insecta" family="Staphylinidae" genus="Zirophorus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="penicillatus">
<emphasis id="B94AEA8CB9B8FF09C0D36C5FACD49DFF" box="[423,687,1132,1157]" italics="true" pageId="204" pageNumber="205">Zirophorus penicillatus</emphasis>
<bibRefCitation id="EFAF4B6FB9B8FF09C3C26C5FAD2F9DFF" author="Dalman, J. W." box="[694,852,1132,1157]" pageId="204" pageNumber="205" pagination="372 - 387" refId="ref189029" refString="Dalman, J. W. (1821) Nya genera och species af insecter. Kongelige Vetenskaps Academiens Handlingar. 1821, 372 - 387, pl. VII." type="journal article" year="1821">Dalman, 1821</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9B8FF09C1BB6CA7ADA89DD7" box="[207,979,1172,1197]" pageId="204" pageNumber="205" type="reference_group">
<paragraph id="8B81369EB9B8FF09C1BB6CA7ADA89DD7" blockId="204.[151,1436,860,1781]" box="[207,979,1172,1197]" pageId="204" pageNumber="205">
<treatmentCitationGroup id="AB2E11B0B9B8FF09C1BB6CA7ADA89DD7" box="[207,979,1172,1197]" pageId="204" pageNumber="205">
<taxonomicName id="4C3E4D1DB9B8FF09C1BB6CA7AC509DD6" ID-CoL="8XQTD" authority="Dalman, 1821: 372" authorityName="Dalman" authorityPageNumber="372" authorityYear="1821" box="[207,555,1172,1197]" class="Insecta" family="Staphylinidae" genus="Zirophorus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9B8FF09C1BB6CA7AF369DD7" box="[207,333,1172,1197]" italics="true" pageId="204" pageNumber="205">Zirophorus</emphasis>
<treatmentCitation id="0A9F108FB9B8FF09C0206CA7AC509DD6" author="Dalman, J. W." box="[340,555,1172,1197]" page="372" pageId="204" pageNumber="205" year="1821">
<bibRefCitation id="EFAF4B6FB9B8FF09C0206CA7AC509DD6" author="Dalman, J. W." box="[340,555,1172,1197]" pageId="204" pageNumber="205" pagination="372 - 387" refId="ref189029" refString="Dalman, J. W. (1821) Nya genera och species af insecter. Kongelige Vetenskaps Academiens Handlingar. 1821, 372 - 387, pl. VII." type="journal article" year="1821">Dalman, 1821: 372</bibRefCitation>
</treatmentCitation>
</taxonomicName>
[synonymized by
<treatmentCitation id="0A9F108FB9B8FF09C38F6CA6ADB79DD7" author="Laporte, F. L." box="[763,972,1172,1197]" page="126" pageId="204" pageNumber="205" year="1835">
<bibRefCitation id="EFAF4B6FB9B8FF09C38F6CA6ADB79DD7" author="Laporte, F. L." box="[763,972,1172,1197]" pageId="204" pageNumber="205" pagination="95 - 159" refId="ref211693" refString="Laporte, F. L. (1835) Etudes Entomologiques, ou description d'insectes nouveaux, et observations sur la synonymie. Premiere Partie, Carnassiers. Mequignon-Marvis Pere et Fils, Paris, pp. 95 - 159, 4 pls." type="book chapter" year="1835">Laporte 1835: 126</bibRefCitation>
</treatmentCitation>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9B8FF08C0736C8FAC3A987F" lastPageId="205" lastPageNumber="206" pageId="204" pageNumber="205" type="materials_examined">
<paragraph id="8B81369EB9B8FF09C0736C8FAD2E9DA9" blockId="204.[151,1436,860,1781]" box="[263,853,1211,1236]" pageId="204" pageNumber="205">
<typeStatus id="5485883CB9B8FF09C0736C8FAF3A9DAE" box="[263,321,1212,1236]" pageId="204" pageNumber="205">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9B8FF09C0D36C88AD2E9DA9" authority="Dalman, 1821" authorityName="Dalman" authorityYear="1821" box="[423,853,1211,1236]" class="Insecta" family="Staphylinidae" genus="Zirophorus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="fronticornis">
<emphasis id="B94AEA8CB9B8FF09C0D36C88ACCA9DA9" box="[423,689,1211,1236]" italics="true" pageId="204" pageNumber="205">Zirophorus fronticornis</emphasis>
<bibRefCitation id="EFAF4B6FB9B8FF09C3C36C88AD2E9DA9" author="Dalman, J. W." box="[695,853,1211,1236]" pageId="204" pageNumber="205" pagination="372 - 387" refId="ref189029" refString="Dalman, J. W. (1821) Nya genera och species af insecter. Kongelige Vetenskaps Academiens Handlingar. 1821, 372 - 387, pl. VII." type="journal article" year="1821">Dalman, 1821</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36CD1AD649D81" blockId="204.[151,1436,860,1781]" box="[151,799,1250,1275]" pageId="204" pageNumber="205">47 species are included in this genus from Latin America.</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36D3AAA879C32" blockId="204.[151,1436,860,1781]" pageId="204" pageNumber="205">
Characters/Description:
<bibRefCitation id="EFAF4B6FB9B8FF09C0DD6D3AACD99C5B" author="Blackwelder, R. E." box="[425,674,1289,1314]" pageId="204" pageNumber="205" refId="ref175651" refString="Blackwelder, R. E. (1943) Monograph of the West-Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, viii + 658 pp. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book" year="1943">Blackwelder 1943: 41</bibRefCitation>
(characters in key), 43 (characters);
<bibRefCitation id="EFAF4B6FB9B8FF09C5376D3AAE829C32" author="Navarrete-Heredia, J. L. &amp; Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." pageId="204" pageNumber="205" refId="ref220178" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Universidad de Guadalajara y CONABIO, Mexico. xii + 401 pp." type="book" year="2002">
Navarrete-Heredia
<emphasis id="B94AEA8CB9B8FF09C46F6D39AB2F9C58" box="[1307,1364,1289,1314]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2002: 207
</bibRefCitation>
(characters in key);
<bibRefCitation id="EFAF4B6FB9B8FF09C0946D03ACA09C32" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." box="[480,731,1327,1352]" pageId="204" pageNumber="205" pagination="490 - 585" refId="ref182077" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25 (6), 490 - 585. [Note: nominal year of publication 2011] https: // doi. org / 10.1071 / is 10016" type="journal article" year="2012">
Caron
<emphasis id="B94AEA8CB9B8FF09C35F6D02AC189C32" box="[555,611,1327,1352]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2012: 500
</bibRefCitation>
(characters);
<bibRefCitation id="EFAF4B6FB9B8FF09C2006D03AA129C32" author="Perez, L. &amp; Rodriguez, M. &amp; Asenjo, A." box="[884,1129,1327,1352]" pageId="204" pageNumber="205" pagination="417 - 426" refId="ref226664" refString="Perez, L., Rodriguez, M. &amp; Asenjo, A. (2017) A new Peruvian species and new records of the genus Piestus Gravenhorst (Coleoptera: Staphylinidae). Zootaxa, 4303 (3), 417 - 426. https: // doi. org / 10.11646 / zootaxa. 4303.3.7" type="journal article" year="2017">
Pérez
<emphasis id="B94AEA8CB9B8FF09C2CE6D02AD899C32" box="[954,1010,1327,1352]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2017: 422
</bibRefCitation>
(characters).
</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36D6BAF129C9E" blockId="204.[151,1436,860,1781]" pageId="204" pageNumber="205">
Key:
<bibRefCitation id="EFAF4B6FB9B8FF09C1A06D65AFB69C15" author="Blackwelder, R. E." box="[212,461,1366,1391]" pageId="204" pageNumber="205" refId="ref175651" refString="Blackwelder, R. E. (1943) Monograph of the West-Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, viii + 658 pp. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book" year="1943">Blackwelder 1943: 44</bibRefCitation>
(species of the West Indies);
<bibRefCitation id="EFAF4B6FB9B8FF09C2686D65AD809C15" author="Scheerpeltz, O." box="[796,1019,1366,1391]" pageId="204" pageNumber="205" pagination="1 - 16" refId="ref234012" refString="Scheerpeltz, O. (1951) Los Staphylinidae (Coleoptera) encontrados en Bolivia por el Sr. Rodolfo Zischka, primera parte. (cuarta contribucion de los Staphylinidae Neotropicales). Folia Universitaria [Universidad de Cochabamba], 5 (5), 1 - 16." type="journal article" year="1951">Scheerpeltz 1951: 5</bibRefCitation>
(species of
<taxonomicName id="4C3E4D1DB9B8FF09C5F76D65AB7A9C15" authorityName="Dalman" authorityYear="1821" box="[1155,1281,1366,1391]" class="Insecta" family="Staphylinidae" genus="Zirophorus" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9B8FF09C5F76D65AB7A9C15" box="[1155,1281,1366,1391]" italics="true" pageId="204" pageNumber="205">Zirophorus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFAF4B6FB9B8FF09C46D6D65AF359CEC" author="Scheerpeltz, O." pageId="204" pageNumber="205" pagination="281 - 305" refId="ref234063" refString="Scheerpeltz, O. (1952 a) Revision der Gattung Piestus Gravh. (Coleoptera Staphyinidae). Revista Chilena de Entomologia, 2, 281 - 305." type="journal article" year="1952">Scheerpeltz 1952a: 282</bibRefCitation>
(subgenera and species);
<bibRefCitation id="EFAF4B6FB9B8FF09C3016D4CADDE9CEC" author="Newton, A. F. &amp; Peck, S. B." box="[629,933,1405,1430]" pageId="204" pageNumber="205" pagination="131 - 158" refId="ref220547" refString="Newton, A. F. &amp; Peck, S. B. (2006) Family Staphylinidae, the rove beetles. In: Peck, S. B. (Ed.), The beetles of the Galapagos Islands, Ecuador: evolution, ecology, and diversity (Insecta: Coleoptera). NRC Research Press, Ottawa, pp. 131 - 158." type="book chapter" year="2006">Newton &amp; Peck 2006: 141</bibRefCitation>
(species of the
<collectingRegion id="49FAF87CB9B8FF09C52F6D4EAAAA9CEC" box="[1115,1233,1405,1430]" country="Ecuador" name="Galapagos" pageId="204" pageNumber="205">Galapagos</collectingRegion>
Isl.);
<bibRefCitation id="EFAF4B6FB9B8FF09C4626D4DAF3B9CC7" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." pageId="204" pageNumber="205" pagination="490 - 585" refId="ref182077" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2012) Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 25 (6), 490 - 585. [Note: nominal year of publication 2011] https: // doi. org / 10.1071 / is 10016" type="journal article" year="2012">
Caron
<emphasis id="B94AEA8CB9B8FF09C4176D4CABE79CEC" box="[1379,1436,1405,1430]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2012: 504
</bibRefCitation>
(species of the genus);
<bibRefCitation id="EFAF4B6FB9B8FF09C3336D96AD3A9CC7" author="Caron, E. &amp; Ribeiro-Costa, C. S. &amp; Newton, A. F." box="[583,833,1444,1469]" pageId="204" pageNumber="205" pagination="129" refId="ref182150" refString="Caron, E., Ribeiro-Costa, C. S. &amp; Newton, A. F. (2013) Corrigendum to: Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera: Staphylinidae: Piestinae). Invertebrate Systematics, 27 (1), 129. https: // doi. org / 10.1071 / IS 10016 _ CO" type="journal article" year="2013">
Caron
<emphasis id="B94AEA8CB9B8FF09C3E66D95ACB19CC7" box="[658,714,1444,1469]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2013: 129
</bibRefCitation>
(species of the genus);
<bibRefCitation id="EFAF4B6FB9B8FF09C53C6D96AB409CC7" author="Perez, L. &amp; Rodriguez, M. &amp; Asenjo, A." box="[1096,1339,1444,1469]" pageId="204" pageNumber="205" pagination="417 - 426" refId="ref226664" refString="Perez, L., Rodriguez, M. &amp; Asenjo, A. (2017) A new Peruvian species and new records of the genus Piestus Gravenhorst (Coleoptera: Staphylinidae). Zootaxa, 4303 (3), 417 - 426. https: // doi. org / 10.11646 / zootaxa. 4303.3.7" type="journal article" year="2017">
Pérez
<emphasis id="B94AEA8CB9B8FF09C5F86D95AABF9CC7" box="[1164,1220,1444,1469]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2017: 422
</bibRefCitation>
(species of the genus).
</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36DC0AF5E9F71" blockId="204.[151,1436,860,1781]" box="[151,293,1523,1548]" pageId="204" pageNumber="205">Distribution.</paragraph>
<paragraph id="8B81369EB9B8FF09C1DE6E29AA4B9F23" blockId="204.[151,1436,860,1781]" pageId="204" pageNumber="205">
West Indies:
<collectingCountry id="F329760EB9B8FF09C0496E2AAF009F48" box="[317,379,1561,1586]" name="Cuba" pageId="204" pageNumber="205">Cuba</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C0F26E2AAF989F48" box="[390,483,1561,1586]" name="Jamaica" pageId="204" pageNumber="205">Jamaica</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C09A6E2AAC529F48" box="[494,553,1561,1586]" name="Haiti" pageId="204" pageNumber="205">Haiti</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C3416E2AAD5B9F48" box="[565,800,1561,1586]" name="Dominican Republic" pageId="204" pageNumber="205">Dominican Republic</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C25F6E29ADC89F48" box="[811,947,1561,1586]" name="Puerto Rico" pageId="204" pageNumber="205">Puerto Rico</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C2CA6E2AAAFD9F48" box="[958,1158,1561,1586]" name="Saint Barthelemy" pageId="204" pageNumber="205">Saint-Barthelemy</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C5E66E2AAB679F48" box="[1170,1308,1561,1586]" name="Guadeloupe" pageId="204" pageNumber="205">Guadeloupe</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C4536E2AABE39F48" box="[1319,1432,1561,1586]" name="Dominica" pageId="204" pageNumber="205">Dominica</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C1BB6E73AF359F23" box="[207,334,1600,1625]" name="Martinique" pageId="204" pageNumber="205">Martinique</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C02E6E72AFB99F23" box="[346,450,1600,1625]" name="Saint Lucia" pageId="204" pageNumber="205">St. Lucia</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C0B96E72AC8A9F23" box="[461,753,1600,1625]" name="Saint Vincent and the Grenadines" pageId="204" pageNumber="205">St. Vincent &amp; Grenadines</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C3886E73AD259F23" box="[764,862,1600,1625]" name="Grenada" pageId="204" pageNumber="205">Grenada</collectingCountry>
, Trinidad-Tobago.
</paragraph>
<paragraph id="8B81369EB9B8FF09C1DE6E54AB529F05" blockId="204.[151,1436,860,1781]" box="[170,1321,1639,1664]" pageId="204" pageNumber="205">
Central America:
<collectingCountry id="F329760EB9B8FF09C0066E54AFB19FFA" box="[370,458,1639,1664]" name="Mexico" pageId="204" pageNumber="205">Mexico</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C0A26E54AC299FFA" box="[470,594,1639,1664]" name="Guatemala" pageId="204" pageNumber="205">Guatemala</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C3296E54ACDC9FFA" box="[605,679,1639,1664]" name="Belize" pageId="204" pageNumber="205">Belize</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C3C76E54AD589FFA" box="[691,803,1639,1664]" name="Honduras" pageId="204" pageNumber="205">Honduras</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C25A6E54ADCF9FFA" box="[814,948,1639,1664]" name="El Salvador" pageId="204" pageNumber="205">El Salvador</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C2CB6E54AA4E9FFA" box="[959,1077,1639,1664]" name="Nicaragua" pageId="204" pageNumber="205">Nicaragua</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C5346E5BAAC69FFA" box="[1088,1213,1639,1664]" name="Costa Rica" pageId="204" pageNumber="205">Costa Rica</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C5BC6E5BAB5F9F05" box="[1224,1316,1640,1663]" name="Panama" pageId="204" pageNumber="205">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9B8FF09C1DE6EBDAC9B9FB4" blockId="204.[151,1436,860,1781]" pageId="204" pageNumber="205">
South America:
<collectingCountry id="F329760EB9B8FF09C0056EBDAF999FDD" box="[369,482,1678,1703]" name="Colombia" pageId="204" pageNumber="205">Colombia</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C0816EBDAC159FDD" box="[501,622,1678,1703]" name="Venezuela" pageId="204" pageNumber="205">Venezuela</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C3F56EBCACA09FDD" box="[641,731,1679,1703]" name="Guyana" pageId="204" pageNumber="205">Guyana</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C3996EBDAD209FDD" box="[749,859,1678,1703]" name="Suriname" pageId="204" pageNumber="205">Suriname</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C21A6EBDAA649FDD" box="[878,1055,1678,1703]" name="French Guiana" pageId="204" pageNumber="205">French Guiana</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C5456EBDAAEA9FDD" box="[1073,1169,1678,1703]" name="Ecuador" pageId="204" pageNumber="205">Ecuador</collectingCountry>
,
<collectingRegion id="49FAF87CB9B8FF09C5D06EBDAB619FDD" box="[1188,1306,1678,1703]" country="Ecuador" name="Galapagos" pageId="204" pageNumber="205">Galapagos</collectingRegion>
Isl.,
<collectingCountry id="F329760EB9B8FF09C4166EA3ABE39FDD" box="[1378,1432,1680,1703]" name="Peru" pageId="204" pageNumber="205">Peru</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C1BB6E86AF5F9FB4" box="[207,292,1717,1742]" name="Bolivia" pageId="204" pageNumber="205">Bolivia</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C0446E86AF0C9FB4" box="[304,375,1717,1742]" name="Brazil" pageId="204" pageNumber="205">Brazil</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C0F66E84AF969FB5" box="[386,493,1719,1743]" name="Paraguay" pageId="204" pageNumber="205">Paraguay</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C08D6E84AC259FB5" box="[505,606,1719,1743]" name="Uruguay" pageId="204" pageNumber="205">Uruguay</collectingCountry>
,
<collectingCountry id="F329760EB9B8FF09C31C6E86ACA79FB4" box="[616,732,1717,1742]" name="Argentina" pageId="204" pageNumber="205">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9B8FF09C1DE6EEFAFC29F8F" blockId="204.[151,1436,860,1781]" box="[170,441,1756,1781]" pageId="204" pageNumber="205">Other regions: Nearctic.</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36F19AD5F9E39" blockId="204.[151,835,1834,1898]" box="[151,804,1834,1859]" pageId="204" pageNumber="205">
<emphasis id="B94AEA8CB9B8FF09C1E36F19AEAD9E39" bold="true" box="[151,214,1834,1859]" pageId="204" pageNumber="205">0711.</emphasis>
<taxonomicName id="4C3E4D1DB9B8FF09C1A96F19AC199E39" authority="Kirby &amp; Spence, 1815" authorityName="Kirby &amp; Spence" authorityYear="1815" box="[221,610,1834,1859]" class="Insecta" family="Staphylinidae" genus="Siagonium" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9B8FF09C1A96F19AF209E39" bold="true" box="[221,347,1834,1859]" italics="true" pageId="204" pageNumber="205">Siagonium</emphasis>
<bibRefCitation id="EFAF4B6FB9B8FF09C0166F19AC199E39" author="Kirby, W. &amp; Spence, W." box="[354,610,1834,1859]" pageId="204" pageNumber="205" pagination="1 - 512" refId="ref208368" refString="Kirby, W. &amp; Spence, W. (1815) An introduction to entomology: or elements of the natural history of insects: with plates. Vol. 1. Longman, Hurst, Rees, Orme, and Brown, London, xxiii + 1 + 1 - 512, 3 pls." type="book chapter" year="1815">Kirby &amp; Spence, 1815</bibRefCitation>
</taxonomicName>
: pl.1 (
<figureCitation id="13052A1BB9B8FF09C3BF6F19AD669E39" box="[715,797,1834,1859]" captionStart="FIGURES 88–99" captionStartId="328.[151,264,1862,1885]" captionTargetBox="[156,1436,198,1832]" captionTargetId="figure@328.[151,1436,193,1840]" captionTargetPageId="328" captionText="FIGURES 88–99. Habitus; Piestinae (88–91); Osoriinae (92–99); 88, after Pérez et al. 2017; 89, Peru: department of Madre de Dios (MUSM); 90, Peru: department of Cuzco (MUSM); 91, after Brunke et al. 2011; 92, after Sharp 1887a; 93, after Sharp 1887a; 94, after Irmler 1991; 95, after Irmler 2001b; 96, after Bierig 1931; 97, after Sharp 1887a; 98, after Caron et al. 2012; 99, after Bruch 1942; slightly modified. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3257248/files/figure.png" pageId="204" pageNumber="205">Fig. 91</figureCitation>
)
</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36F61AD389E10" blockId="204.[151,835,1834,1898]" box="[151,835,1873,1898]" pageId="204" pageNumber="205">
<typeStatus id="5485883CB9B8FF09C1E36F61AEAA9E10" box="[151,209,1874,1898]" pageId="204" pageNumber="205">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9B8FF09C0436F61AD389E10" authority="Kirby &amp; Spence, 1815" authorityName="Kirby &amp; Spence" authorityYear="1815" box="[311,835,1873,1898]" class="Insecta" family="Staphylinidae" genus="Siagonium" kingdom="Animalia" order="Coleoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="quadricorne">
<emphasis id="B94AEA8CB9B8FF09C0436F61AC3A9E10" box="[311,577,1873,1898]" italics="true" pageId="204" pageNumber="205">Siagonium quadricorne</emphasis>
<bibRefCitation id="EFAF4B6FB9B8FF09C3336F62AD389E10" author="Kirby, W. &amp; Spence, W." box="[583,835,1873,1898]" pageId="204" pageNumber="205" pagination="1 - 512" refId="ref208368" refString="Kirby, W. &amp; Spence, W. (1815) An introduction to entomology: or elements of the natural history of insects: with plates. Vol. 1. Longman, Hurst, Rees, Orme, and Brown, London, xxiii + 1 + 1 - 512, 3 pls." type="book chapter" year="1815">Kirby &amp; Spence, 1815</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36F93AD629EC2" blockId="204.[151,1038,1951,2015]" box="[151,793,1951,1976]" pageId="204" pageNumber="205">1 species is included in this genus from Central America.</paragraph>
<paragraph id="8B81369EB9B8FF09C1E36FF5AA759EA5" blockId="204.[151,1038,1951,2015]" box="[151,1038,1990,2015]" pageId="204" pageNumber="205">
Characters/Description:
<bibRefCitation id="EFAF4B6FB9B8FF09C0DC6FF5AD549EA5" author="Navarrete-Heredia, J. L. &amp; Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." box="[424,815,1990,2015]" pageId="204" pageNumber="205" refId="ref220178" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Universidad de Guadalajara y CONABIO, Mexico. xii + 401 pp." type="book" year="2002">
Navarrete-Heredia
<emphasis id="B94AEA8CB9B8FF09C30B6FFBACCC9EA5" box="[639,695,1990,2015]" italics="true" pageId="204" pageNumber="205">et al.</emphasis>
2002: 207
</bibRefCitation>
(characters in key).
</paragraph>
<paragraph id="8B81369EB9B9FF08C1E368ADAF5E99CC" blockId="205.[151,662,158,295]" box="[151,293,158,183]" pageId="205" pageNumber="206">Distribution.</paragraph>
<paragraph id="8B81369EB9B9FF08C1DE68F6AFAB99A4" blockId="205.[151,662,158,295]" box="[170,464,197,222]" pageId="205" pageNumber="206">
Central America:
<collectingCountry id="F329760EB9B9FF08C00668F6AFB099A4" box="[370,459,197,222]" name="Mexico" pageId="205" pageNumber="206">Mexico</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9B9FF08C1DE68DFAC3A987F" blockId="205.[151,662,158,295]" box="[170,577,236,261]" pageId="205" pageNumber="206">Other regions: Nearctic, Palaearctic.</paragraph>
</subSubSection>
<subSubSection id="C3246515B9B9FF08C1E3693EACED985D" box="[151,662,269,295]" pageId="205" pageNumber="206" type="reference_group">
<paragraph id="8B81369EB9B9FF08C1E3693EACED985D" blockId="205.[151,662,158,295]" box="[151,662,269,295]" pageId="205" pageNumber="206">
<emphasis id="B94AEA8CB9B9FF08C1E3693EACED985D" bold="true" box="[151,662,269,295]" pageId="205" pageNumber="206">
Subfamily
<taxonomicName id="4C3E4D1DB9B9FF08C06B693EACED985D" ID-CoL="8VR5S" authority="Erichson, 1839 a" authorityName="Erichson" authorityYear="1839" box="[287,662,269,295]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="205" pageNumber="206" phylum="Arthropoda" rank="subFamily" subFamily="Osoriinae">
OSORIINAE
<bibRefCitation id="EFAF4B6FB9B9FF08C0BA693EACED985D" author="Erichson, G. F." box="[462,662,269,295]" pageId="205" pageNumber="206" refId="ref190452" refString="Erichson, G. F. (1839 a) Genera et species staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin. viii + 400 pp." type="book" year="1839">Erichson, 1839a</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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