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<document id="EA2188C4DF70005581F8E2515AE23763" ID-CLB-Dataset="29175" ID-DOI="10.11646/zootaxa.4453.1.1" ID-GBIF-Dataset="348e1a52-72a5-4809-b533-a158fc7a298a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1454920" ID-ZooBank="866690A9-0462-4892-AE29-9AAC623F87B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539178978648" checkinUser="plazi" docAuthor="Paweł Jałoszyński" docDate="2018" docId="2161879CFFF28A2EFF7A358C633CDF8F" docLanguage="en" docName="zootaxa.4453.1.1.pdf.imf" docOrigin="Zootaxa 4453 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Leptochromus Motschulsky" docType="treatment" docVersion="8" lastPageNumber="62" masterDocId="DD58FFE4FFC68A13FFED3418614DD933" masterDocTitle="World genera of Mastigitae: review of morphological structures and new ecological data (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="119" masterPageNumber="1" pageNumber="53" updateTime="1698572836684" updateUser="ExternalLinkService">
<mods:mods id="448EB4BD4661A275E406D5268A4CB6B2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F48DB699BB2CB3DC857D4C36692697AA">World genera of Mastigitae: review of morphological structures and new ecological data (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:name id="ACC1FBE71E975A7092F59C69CDE80F33" type="personal">
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<mods:namePart id="AFFEACF858FAB441A1A20AFC66C45A05">Paweł Jałoszyński</mods:namePart>
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<mods:title id="F87D6C35F0381585E013838F48EDDDCA">Zootaxa</mods:title>
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<mods:part id="CCEC93167D2A8BA900460F5FD834EB79">
<mods:date id="A99391480EB88CBC8C63346F9B292105">2018</mods:date>
<mods:detail id="57770D01E088F397CF1807B32684EB58" type="pubDate">
<mods:number id="B9C3BC99B897794C502C474738CD1647">2018-07-30</mods:number>
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<mods:number id="E5780C4FC0BB91ECA29209BE4602AE37">4453</mods:number>
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<mods:classification id="DEA761D8C95494B63DFE298CC91521BB">journal article</mods:classification>
<mods:identifier id="8BB9FCDBA7625F8B09A5CEA9675EB616" type="CLB-Dataset">29175</mods:identifier>
<mods:identifier id="050C6B64BE1287EC756251367F9814DE" type="DOI">10.11646/zootaxa.4453.1.1</mods:identifier>
<mods:identifier id="E4B9F1C42E304D6AAD63815821830006" type="GBIF-Dataset">348e1a52-72a5-4809-b533-a158fc7a298a</mods:identifier>
<mods:identifier id="DD17F788DE65FB3BF5C3FF90B6D6F79A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="2161879CFFF28A2EFF7A358C633CDF8F" ID-DOI="http://doi.org/10.5281/zenodo.5976946" ID-GBIF-Taxon="148669751" ID-Zenodo-Dep="5976946" LSID="urn:lsid:plazi:treatment:2161879CFFF28A2EFF7A358C633CDF8F" httpUri="http://treatment.plazi.org/id/2161879CFFF28A2EFF7A358C633CDF8F" lastPageId="61" lastPageNumber="62" pageId="52" pageNumber="53">
<subSubSection id="E1D26501FFF28A27FF7A358C60A1D89D" box="[151,492,404,430]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="A977368AFFF28A27FF7A358C60A1D89D" blockId="52.[151,492,404,430]" box="[151,492,404,430]" pageId="52" pageNumber="53">
<heading id="F23F81E6FFF28A27FF7A358C60A1D89D" bold="true" box="[151,492,404,430]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
<taxonomicName id="6EC84D09FFF28A27FF7A358C60A1D89D" ID-CoL="8XM9V" authority="Motschulsky" authorityName="Motschulsky" box="[151,492,404,430]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFF28A27FF7A358C60A1D89D" bold="true" box="[151,492,404,430]" pageId="52" pageNumber="53">
<emphasis id="9BBCEA98FFF28A27FF7A358C6008D89D" bold="true" box="[151,325,404,430]" italics="true" pageId="52" pageNumber="53">Leptochromus</emphasis>
Motschulsky
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E1D26501FFF28A27FF7A35C2637BDB00" pageId="52" pageNumber="53" type="reference_group">
<paragraph id="A977368AFFF28A27FF7A35C2637BDB00" blockId="52.[151,1435,474,563]" pageId="52" pageNumber="53">
<taxonomicName id="6EC84D09FFF28A27FF7A35C26372D8C0" ID-CoL="8XM9V" authority="Motschulsky, 1855: 12" authorityName="Motschulsky" authorityPageNumber="12" authorityYear="1855" box="[151,575,474,499]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFF28A27FF7A35C2607BD8C0" box="[151,310,474,499]" italics="true" pageId="52" pageNumber="53">Leptochromus</emphasis>
Motschulsky, 1855: 12
</taxonomicName>
.
<typeStatus id="76738828FFF28A27FDA735C363CFD8C0" box="[586,642,475,499]" pageId="52" pageNumber="53">Type</typeStatus>
species:
<taxonomicName id="6EC84D09FFF28A27FD0435C26593D8C0" ID-CoL="3TM7D" authority="Motschulsky, 1855" authorityName="Motschulsky" authorityYear="1855" box="[745,1246,474,499]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="fulvescens">
<emphasis id="9BBCEA98FFF28A27FD0435C2654FD8C0" box="[745,1026,474,499]" italics="true" pageId="52" pageNumber="53">Leptochromus fulvescens</emphasis>
Motschulsky, 1855
</taxonomicName>
(monotypy).
<taxonomicName id="6EC84D09FFF28A27FF7A361860D9DB27" ID-CoL="8XLQ8" authority="Sharp, 1887: 70" authorityName="Sharp" authorityPageNumber="70" authorityYear="1887" box="[151,404,510,533]" class="Insecta" family="Staphylinidae" genus="Hecotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFF28A27FF7A361861A7DB26" box="[151,234,512,533]" italics="true" pageId="52" pageNumber="53">Hecotus</emphasis>
Sharp, 1887: 70
</taxonomicName>
.
<typeStatus id="76738828FFF28A27FE7035E76082DB27" box="[413,463,511,532]" pageId="52" pageNumber="53">Type</typeStatus>
species:
<taxonomicName id="6EC84D09FFF28A27FDC13618623CDB27" ID-CoL="3JYTJ" authority="SHARP, 1887" authorityName="SHARP" authorityYear="1887" box="[556,881,510,533]" class="Insecta" family="Staphylinidae" genus="Hecotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="107" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="9BBCEA98FFF28A27FDC1361863BCDB26" box="[556,753,512,533]" italics="true" pageId="52" pageNumber="53">Hecotus mexicanus</emphasis>
Sharp, 1887
</taxonomicName>
(des. Franz in Newton &amp; Franz (1998)). Synonymized with
<taxonomicName id="6EC84D09FFF28A27FF17360560C7DB00" ID-CoL="8XM9V" box="[250,394,541,563]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFF28A27FF17360560C7DB00" box="[250,394,541,563]" italics="true" pageId="52" pageNumber="53">Leptochromus</emphasis>
</taxonomicName>
by
<bibRefCitation id="CD594B7BFFF28A27FE5D3605637EDB00" box="[432,563,541,563]" pageId="52" pageNumber="53" refId="ref52042" refString="CSIkI, E. (1919) SCYdMAENIdAE. In: SCHENkLING, S. &amp; JUNk, W. (EdS.), Coleopterorum Catalogus. Pars 70. W. JUNk, BERLIN, PP. 106." type="journal article">Csiki (1919)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E1D26501FFF28A2EFF7A364760D9DDF6" lastPageId="61" lastPageNumber="62" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="A977368AFFF28A27FF7A36476344DBD7" blockId="52.[151,1437,607,2037]" pageId="52" pageNumber="53">
<emphasis id="9BBCEA98FFF28A27FF7A3647605CDB4B" bold="true" box="[151,273,607,632]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
<taxonomicName id="6EC84D09FFF28A27FEF4364760F5DB4B" box="[281,440,607,632]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFF28A27FEF4364760F5DB4B" box="[281,440,607,632]" italics="true" pageId="52" pageNumber="53">Leptochromus</emphasis>
</taxonomicName>
differs from all remaining Leptochromini in unique apomorphies: maxillary palpomere II with a long and slender cuticular projection with bifurcate apex and a pair of thick apical bristles (
<figureCitation id="31F32A0FFFF28A27FB10369C6412DBAF" box="[1277,1375,644,669]" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="52" pageNumber="53">Fig. 121</figureCitation>
); and maxillary palpomere IV strongly elongate, nearly as long as palpomere III, only slightly broadened in subapical region and with subconical apex.
</paragraph>
<paragraph id="A977368AFFF28A27FF2A36F76438DA7C" blockId="52.[151,1437,607,2037]" pageId="52" pageNumber="53">
<emphasis id="9BBCEA98FFF28A27FF2A36F760AFDA3B" bold="true" box="[199,482,751,776]" pageId="52" pageNumber="53">Characteristics. Adults.</emphasis>
Body (
<figureCitation id="31F32A0FFFF28A27FDDA36F763D6DA3B" box="[567,667,751,776]" captionStart="FIGURES 107–108" captionStartId="53.[151,264,1815,1838]" captionTargetBox="[169,1424,206,1772]" captionTargetId="figure@53.[151,1436,193,1794]" captionTargetPageId="53" captionText="FIGURES 107–108. Leptochromini, examples of extant species; dorsal (107, 109) and dorsolateral (108) habitus. Leptochromus laselva (107–108); Leptochromus agilis (109)." httpUri="https://zenodo.org/record/1454978/files/figure.png" pageId="52" pageNumber="53">Figs 107</figureCitation>
–109) moderately large, 2.70̄
<quantity id="6E309B6FFFF28A27FC0236E86515DA34" box="[1007,1112,752,776]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="52" pageNumber="53" unit="mm" value="4.2">4.20 mm</quantity>
in length, from yellowish to dark brown, strongly convex or slightly flattened, dorsally densely setose, setae long and suberect to erect, unmodified except for several long and thick bristles on protrochanters, postgenae, and maxillary palpomere II.
</paragraph>
<paragraph id="A977368AFFF28A27FF2A37446283DC7B" blockId="52.[151,1437,607,2037]" pageId="52" pageNumber="53">
Head capsule (
<figureCitation id="31F32A0FFFF28A27FE8337446341DA46" box="[366,524,860,885]" captionStart="FIGURES 114–118" captionStartId="55.[151,264,1704,1727]" captionTargetBox="[163,1407,193,1665]" captionTargetId="figure@55.[163,1407,193,1665]" captionTargetPageId="55" captionText="FIGURES 114–118. Leptochromini, morphological structures, Leptochromus agilis (114) and Leptochromus laselva (115– 118). Head in dorsal (114), dorsolateral (115) and anteroventral (116) views; thorax and abdomen in ventral view (117); right procoxal, protrochanter and base of profemur in ventral view (118). Abbreviations: fem1, profemur; mxp2–4, maxillary palpomere II–IV; pd, pedicel; pgp, postgenal (= subocular) process; sc, scape; tr1, protrochanter." httpUri="https://zenodo.org/record/1454984/files/figure.png" pageId="52" pageNumber="53">Figs 114–116</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FDF537446336DA47" box="[536,635,860,884]" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="52" pageNumber="53">119–120</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FD65374463F9DA47" box="[648,692,860,884]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="52" pageNumber="53">124</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FD2C37446268DA47" box="[705,805,860,884]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="52" pageNumber="53">127–129</figureCitation>
) divided into large and exposed anterior part and much smaller, subcylindrical 'neck' region retracted into prothorax and demarcated by distinct occipital constriction; 'neck' region about as broad as half width of head. Anterior part of head strongly flattened, subequal in width with prothorax, strongly transverse, broadest at eyes, slightly in front of middle. Composite eyes dorsolateral, near anterior margin of head, moderately large, composed of numerous small ommatidia, strongly convex. Vertex strongly transverse and posteriorly impressed at middle, convex at sides, with posterior margin concave. Tempora long and rounded. Frons between antennal insertions with straight margin, not forming a demarcated 'platform', anteriorly demarcated by a deep and complete frontoclypeal groove. Clypeus very short and broad, with rounded or straight sides. Antennal insertions located anterodorsally, very broadly separated. Gular plate (
<figureCitation id="31F32A0FFFF28A27FB5F3064645EDDA7" box="[1202,1299,1148,1173]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="52" pageNumber="53">Fig. 129</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FAF030656474DDA7" box="[1309,1337,1149,1172]" italics="true" pageId="52" pageNumber="53">gp</emphasis>
) lacking sutures, nearly smooth; posterior tentorial pits (
<figureCitation id="31F32A0FFFF28A27FD2A3087627DDD8B" box="[711,816,1183,1208]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="52" pageNumber="53">Figs 124</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FCAC30B86223DD8B" box="[833,878,1184,1208]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="52" pageNumber="53">129</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FC9230B962EEDD8B" box="[895,931,1185,1208]" italics="true" pageId="52" pageNumber="53">ptp</emphasis>
) arcuate, in front of transverse impression demarcating 'neck' region ventrally; hypostomal ridges (
<figureCitation id="31F32A0FFFF28A27FCF430DC62CDDDEF" box="[793,896,1220,1245]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="52" pageNumber="53">Figs 124</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FC6230DC62F1DDEF" box="[911,956,1220,1244]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="52" pageNumber="53">129</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FC2430DC62AFDDEE" box="[969,994,1220,1245]" italics="true" pageId="52" pageNumber="53">hr</emphasis>
) nearly straight or arcuate, posteriorly reaching middle between anterior submental margin and posterior tentorial pits. Postgenae (
<figureCitation id="31F32A0FFFF28A27FB5430FF641ADC33" box="[1209,1367,1255,1280]" captionStart="FIGURES 114–118" captionStartId="55.[151,264,1704,1727]" captionTargetBox="[163,1407,193,1665]" captionTargetId="figure@55.[163,1407,193,1665]" captionTargetPageId="55" captionText="FIGURES 114–118. Leptochromini, morphological structures, Leptochromus agilis (114) and Leptochromus laselva (115– 118). Head in dorsal (114), dorsolateral (115) and anteroventral (116) views; thorax and abdomen in ventral view (117); right procoxal, protrochanter and base of profemur in ventral view (118). Abbreviations: fem1, profemur; mxp2–4, maxillary palpomere II–IV; pd, pedicel; pgp, postgenal (= subocular) process; sc, scape; tr1, protrochanter." httpUri="https://zenodo.org/record/1454984/files/figure.png" pageId="52" pageNumber="53">Figs 115–116</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FA8830F0618EDC17" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="52" pageNumber="53">119– 120</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FF3D311461B1DC17" box="[208,252,1292,1316]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="52" pageNumber="53">124</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FEE731146020DC17" box="[266,365,1292,1316]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="52" pageNumber="53">127–129</figureCitation>
) in most species with a tubercle or projection (postgenal or subocular process) with 2–3 thick apical bristles. Head with or without punctures, densely setose (
<figureCitation id="31F32A0FFFF28A27FC8C3137628FDC7B" box="[865,962,1327,1352]" captionStart="FIGURES 114–118" captionStartId="55.[151,264,1704,1727]" captionTargetBox="[163,1407,193,1665]" captionTargetId="figure@55.[163,1407,193,1665]" captionTargetPageId="55" captionText="FIGURES 114–118. Leptochromini, morphological structures, Leptochromus agilis (114) and Leptochromus laselva (115– 118). Head in dorsal (114), dorsolateral (115) and anteroventral (116) views; thorax and abdomen in ventral view (117); right procoxal, protrochanter and base of profemur in ventral view (118). Abbreviations: fem1, profemur; mxp2–4, maxillary palpomere II–IV; pd, pedicel; pgp, postgenal (= subocular) process; sc, scape; tr1, protrochanter." httpUri="https://zenodo.org/record/1454984/files/figure.png" pageId="52" pageNumber="53">Fig. 115</figureCitation>
).
</paragraph>
<paragraph id="A977368AFFF28A27FF2A314C64C4DF13" blockId="52.[151,1437,607,2037]" pageId="52" pageNumber="53">
Antennae (
<figureCitation id="31F32A0FFFF28A27FEA9314C60E4DC5F" box="[324,425,1364,1389]" captionStart="FIGURES 107–108" captionStartId="53.[151,264,1815,1838]" captionTargetBox="[169,1424,206,1772]" captionTargetId="figure@53.[151,1436,193,1794]" captionTargetPageId="53" captionText="FIGURES 107–108. Leptochromini, examples of extant species; dorsal (107, 109) and dorsolateral (108) habitus. Leptochromus laselva (107–108); Leptochromus agilis (109)." httpUri="https://zenodo.org/record/1454978/files/figure.png" pageId="52" pageNumber="53">Figs 107</figureCitation>
–109, 114–116, 127) shorter or slightly longer than body, slender; scape (
<figureCitation id="31F32A0FFFF28A27FB17314C64D5DC5E" box="[1274,1432,1364,1389]" captionStart="FIGURES 114–118" captionStartId="55.[151,264,1704,1727]" captionTargetBox="[163,1407,193,1665]" captionTargetId="figure@55.[163,1407,193,1665]" captionTargetPageId="55" captionText="FIGURES 114–118. Leptochromini, morphological structures, Leptochromus agilis (114) and Leptochromus laselva (115– 118). Head in dorsal (114), dorsolateral (115) and anteroventral (116) views; thorax and abdomen in ventral view (117); right procoxal, protrochanter and base of profemur in ventral view (118). Abbreviations: fem1, profemur; mxp2–4, maxillary palpomere II–IV; pd, pedicel; pgp, postgenal (= subocular) process; sc, scape; tr1, protrochanter." httpUri="https://zenodo.org/record/1454984/files/figure.png" pageId="52" pageNumber="53">Figs 114–116</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FF7A31606189DCA3" box="[151,196,1400,1424]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="52" pageNumber="53">127</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FF3C316161A5DCA3" box="[209,232,1401,1424]" italics="true" pageId="52" pageNumber="53">sc</emphasis>
) 6 to more than 10 times as long as broad, usually slightly broadening distad, longer than head, with very deep lateroventral emargination; pedicel (
<figureCitation id="31F32A0FFFF28A27FD8031846247DC86" box="[621,778,1436,1461]" captionStart="FIGURES 114–118" captionStartId="55.[151,264,1704,1727]" captionTargetBox="[163,1407,193,1665]" captionTargetId="figure@55.[163,1407,193,1665]" captionTargetPageId="55" captionText="FIGURES 114–118. Leptochromini, morphological structures, Leptochromus agilis (114) and Leptochromus laselva (115– 118). Head in dorsal (114), dorsolateral (115) and anteroventral (116) views; thorax and abdomen in ventral view (117); right procoxal, protrochanter and base of profemur in ventral view (118). Abbreviations: fem1, profemur; mxp2–4, maxillary palpomere II–IV; pd, pedicel; pgp, postgenal (= subocular) process; sc, scape; tr1, protrochanter." httpUri="https://zenodo.org/record/1454984/files/figure.png" pageId="52" pageNumber="53">Figs 114, 115</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FCFA31846209DC87" box="[791,836,1436,1460]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="52" pageNumber="53">127</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FCBD31846221DC86" box="[848,876,1436,1461]" italics="true" pageId="52" pageNumber="53">pd</emphasis>
) subequal in length to antennomere III or shorter and broadening from narrow base to apex; antennomeres III–XI elongate (usually strongly, rarely weakly) and weakly thickening distad, basal stalks not exposed, basal rings absent or indistinct; antennomere XI elongate and slightly asymmetrical. Antennomeres covered with variously dense, long setae; surface of antennomeres smooth.
</paragraph>
<paragraph id="A977368AFFF28A25FF2A32346276DE97" blockId="52.[151,1437,607,2037]" lastBlockId="54.[151,1437,1643,1956]" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53">
Mouthparts. Labrum strongly transverse, with lateral margins divergent anterad, nearly straight or weakly rounded, and with anterior margin concave, with a pair of broad and short submedian teeth separated at middle by a relatively shallow emargination, and with a transverse dorsoanterior row of long setae. Mandibles symmetrical, subtriangular and robust, each with two subapical teeth, at least one of them above the coronal plane of mandible, setose prostheca present and long. Maxilla (
<figureCitation id="31F32A0FFFF28A27FD7832A463B0DFE7" box="[661,765,1724,1749]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="52" pageNumber="53">Figs 124</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FCE132A4627ADFE7" box="[780,823,1724,1748]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="52" pageNumber="53">129</figureCitation>
) with large but relatively short cardo (
<figureCitation id="31F32A0FFFF28A27FB1032A4642CDFE7" box="[1277,1377,1724,1749]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="52" pageNumber="53">Fig. 124</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FA9D32A464C7DFE6" box="[1392,1418,1724,1749]" italics="true" pageId="52" pageNumber="53">cd</emphasis>
); basistipes (
<figureCitation id="31F32A0FFFF28A27FEFB32C76036DFCB" box="[278,379,1759,1784]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="52" pageNumber="53">Fig. 124</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FE6632C760E1DFCB" box="[395,428,1759,1784]" italics="true" pageId="52" pageNumber="53">bst</emphasis>
) subtriangular and elongate; mediostipes (
<figureCitation id="31F32A0FFFF28A27FC7732C762B2DFCB" box="[922,1023,1759,1784]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="52" pageNumber="53">Fig. 124</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FBE332F96578DFCB" box="[1038,1077,1761,1784]" italics="true" pageId="52" pageNumber="53">mst</emphasis>
) large and sharply demarcated from lacinia and galea, which are both elongate and curved mesally and each with dense row of distal setae; palpifer (
<figureCitation id="31F32A0FFFF28A27FEEC333F6028DE73" box="[257,357,1831,1856]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="52" pageNumber="53">Fig. 124</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FE99333060DADE73" box="[372,407,1832,1856]" italics="true" pageId="52" pageNumber="53">ppf</emphasis>
) broad and elongate; maxillary palp (
<figureCitation id="31F32A0FFFF28A27FCBE333F62B8DE73" box="[851,1013,1831,1856]" captionStart="FIGURES 114–118" captionStartId="55.[151,264,1704,1727]" captionTargetBox="[163,1407,193,1665]" captionTargetId="figure@55.[163,1407,193,1665]" captionTargetPageId="55" captionText="FIGURES 114–118. Leptochromini, morphological structures, Leptochromus agilis (114) and Leptochromus laselva (115– 118). Head in dorsal (114), dorsolateral (115) and anteroventral (116) views; thorax and abdomen in ventral view (117); right procoxal, protrochanter and base of profemur in ventral view (118). Abbreviations: fem1, profemur; mxp2–4, maxillary palpomere II–IV; pd, pedicel; pgp, postgenal (= subocular) process; sc, scape; tr1, protrochanter." httpUri="https://zenodo.org/record/1454984/files/figure.png" pageId="52" pageNumber="53">Figs 114, 116</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FBE93330657DDE73" box="[1028,1072,1832,1856]" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="52" pageNumber="53">121</figureCitation>
,
<figureCitation id="31F32A0FFFF28A27FBAD333065EBDE73" box="[1088,1190,1832,1856]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="52" pageNumber="53">127, 129</figureCitation>
) conspicuously long, much longer than head capsule, composed of minute palpomere I (
<figureCitation id="31F32A0FFFF28A27FC7B335462B4DE57" box="[918,1017,1868,1893]" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="52" pageNumber="53">Fig. 121</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FBEA33546509DE57" box="[1031,1092,1868,1892]" italics="true" pageId="52" pageNumber="53">mxp1</emphasis>
), extremely elongate, slender, only slightly broadening distad palpomere II (
<figureCitation id="31F32A0FFFF28A27FD4733776243DEBB" box="[682,782,1903,1928]" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="52" pageNumber="53">Fig. 121</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FCF033686217DEBB" box="[797,858,1904,1928]" italics="true" pageId="52" pageNumber="53">mxp2</emphasis>
), which is distinctly recurved and bearing a long, slender bifurcate cuticular process with two apical bristles, palpomere III (
<figureCitation id="31F32A0FFFF28A27FBEA338C6520DE9F" box="[1031,1133,1940,1965]" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="52" pageNumber="53">Fig. 121</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FB90338C65F4DE9F" box="[1149,1209,1940,1964]" italics="true" pageId="52" pageNumber="53">mxp3</emphasis>
) strongly elongate, slender, broadened only slightly in its distal third or fourth, with transverse distal margin, palpomere IV (
<figureCitation id="31F32A0FFFF28A27FADB33A064DADEE3" box="[1334,1431,1976,2001]" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="52" pageNumber="53">Fig. 121</figureCitation>
;
<emphasis id="9BBCEA98FFF28A27FF7A33C46199DEC7" box="[151,212,2012,2036]" italics="true" pageId="52" pageNumber="53">mxp4</emphasis>
) nearly rod-like, strongly elongate, slender, only slightly shorter than III, slightly broadened distad up to its distal third and then narrowed to subconical and blunt apex. Palpomeres IĪIV round or nearly round in crosssection, covered with moderately long setae. Labium (
<figureCitation id="31F32A0FFFF08A25FD123297622BDF9B" box="[767,870,1679,1704]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="54" pageNumber="55">Figs 124</figureCitation>
,
<figureCitation id="31F32A0FFFF08A25FC9E328862D3DF9B" box="[883,926,1680,1704]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="54" pageNumber="55">129</figureCitation>
) with broad and short submentum (
<figureCitation id="31F32A0FFFF08A25FAD8329764D5DF9B" box="[1333,1432,1679,1704]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="54" pageNumber="55">Fig. 124</figureCitation>
;
<emphasis id="9BBCEA98FFF08A25FF7A32AD6189DFFF" box="[151,196,1717,1740]" italics="true" pageId="54" pageNumber="55">smn</emphasis>
) posteriorly not demarcated from gular region, bearing a pair of long thin setae in its subanterior region; mentum (
<figureCitation id="31F32A0FFFF08A25FEEE32CF602BDFC3" box="[259,358,1751,1776]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="54" pageNumber="55">Fig. 124</figureCitation>
;
<emphasis id="9BBCEA98FFF08A25FE9E32C160D8DFC3" box="[371,405,1753,1776]" italics="true" pageId="54" pageNumber="55">mn</emphasis>
) subtrapezoidal and strongly transverse, with anterior margin weakly concave; prementum (
<figureCitation id="31F32A0FFFF08A25FF7232E36045DE27" box="[159,264,1787,1812]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="54" pageNumber="55">Fig. 124</figureCitation>
;
<emphasis id="9BBCEA98FFF08A25FEF632E56070DE27" box="[283,317,1789,1812]" italics="true" pageId="54" pageNumber="55">pm</emphasis>
) long, subtrapezoidal, broadest distally, lacking demarcated ligula, with one or two pairs of anterolateral setae, with broadly separated bases of labial palps; lateral hypopharyngeal lobes (
<figureCitation id="31F32A0FFFF08A25FB1033076429DE0B" box="[1277,1380,1823,1848]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="54" pageNumber="55">Fig. 124</figureCitation>
;
<emphasis id="9BBCEA98FFF08A25FA98330764DFDE0B" box="[1397,1426,1823,1848]" italics="true" pageId="54" pageNumber="55">lhl</emphasis>
) moderately large; labial palp (
<figureCitation id="31F32A0FFFF08A25FE1C335B6318DE6F" box="[497,597,1859,1884]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="54" pageNumber="55">Fig. 124</figureCitation>
;
<emphasis id="9BBCEA98FFF08A25FD8E335B6334DE6F" box="[611,633,1859,1884]" italics="true" pageId="54" pageNumber="55">lp</emphasis>
) composed of three palpomeres: palpomere I small, weakly elongate, broadening distad, palpomere II largest, strongly elongate and barrel-shaped, palpomere III narrow, long and pointed. Porous fields on mouthparts and clypeus not found.
</paragraph>
<caption id="FDB76602FFF38A26FF7A330F6249DE7F" httpUri="https://zenodo.org/record/1454978/files/figure.png" pageId="53" pageNumber="54" startId="53.[151,264,1815,1838]" targetBox="[169,1424,206,1772]" targetPageId="53">
<paragraph id="A977368AFFF38A26FF7A330F6249DE7F" blockId="53.[151,1438,1815,1868]" pageId="53" pageNumber="54">
<emphasis id="9BBCEA98FFF38A26FF7A330F603ADE1E" bold="true" box="[151,375,1815,1838]" pageId="53" pageNumber="54">FIGURES 107–108.</emphasis>
Leptochromini, examples of extant species; dorsal (107, 109) and dorsolateral (108) habitus.
<taxonomicName id="6EC84D09FFF38A26FF7A332E6038DE7F" box="[151,373,1846,1868]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="9BBCEA98FFF38A26FF7A332E6038DE7F" box="[151,373,1846,1868]" italics="true" pageId="53" pageNumber="54">Leptochromus laselva</emphasis>
</taxonomicName>
(107–108);
<taxonomicName id="6EC84D09FFF38A26FE1F332E638DDE7F" box="[498,704,1846,1868]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="9BBCEA98FFF38A26FE1F332E638DDE7F" box="[498,704,1846,1868]" italics="true" pageId="53" pageNumber="54">Leptochromus agilis</emphasis>
</taxonomicName>
(109).
</paragraph>
</caption>
<caption id="FDB76602FFF08A25FF7A373F61A7DAA9" httpUri="https://zenodo.org/record/1454980/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,264,807,830]" targetBox="[151,1436,193,786]" targetPageId="54">
<paragraph id="A977368AFFF08A25FF7A373F61A7DAA9" blockId="54.[151,1438,807,922]" pageId="54" pageNumber="55">
<emphasis id="9BBCEA98FFF08A25FF7A373F6021DA0E" bold="true" box="[151,364,807,830]" pageId="54" pageNumber="55">FIGURES 110–112.</emphasis>
Leptochromini, examples of extinct species; dorsal (110), lateral (111) and laterodorsal (112) habitus.
<taxonomicName id="6EC84D09FFF08A25FF7A375E60AFDA6F" authorityName="O'Keefe" authorityYear="2002" box="[151,482,838,860]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="palaeomexicanus">
<emphasis id="9BBCEA98FFF08A25FF7A375E60AFDA6F" box="[151,482,838,860]" italics="true" pageId="54" pageNumber="55">Leptochromus palaeomexicanus</emphasis>
</taxonomicName>
(110; Middle Miocene of Mexico; photo George Poinar);
<taxonomicName id="6EC84D09FFF08A25FBB4375E6410DA6F" authorityName="Jaloszynski &amp; Brunke W" authorityYear="2018" box="[1113,1373,838,860]" class="Insecta" family="Staphylinidae" genus="Euroleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="setifer">
<emphasis id="9BBCEA98FFF08A25FBB4375E6410DA6F" box="[1113,1373,838,860]" italics="true" pageId="54" pageNumber="55">Euroleptochromus setifer</emphasis>
</taxonomicName>
(111; Upper Eocene of Europe);
<taxonomicName id="6EC84D09FFF08A25FE43377D6243DA48" authorityName="JAlOSzYnSkI &amp; PERkOVSkY" authorityYear="2016" box="[430,782,869,891]" class="Insecta" family="Staphylinidae" genus="Rovnoleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="ableptonoides">
<emphasis id="9BBCEA98FFF08A25FE43377D6243DA48" box="[430,782,869,891]" italics="true" pageId="54" pageNumber="55">Rovnoleptochromus ableptonoides</emphasis>
</taxonomicName>
(112; Upper Eocene of Europe; after Jałoszyński &amp; Perkovsky (2016)).
</paragraph>
</caption>
<caption id="FDB76602FFF08A25FF7A321D62E2DF28" box="[151,943,1541,1563]" httpUri="https://zenodo.org/record/1454982/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,250,1541,1563]" targetBox="[151,1436,971,1519]" targetPageId="54">
<paragraph id="A977368AFFF08A25FF7A321D62E2DF28" blockId="54.[151,943,1541,1563]" box="[151,943,1541,1563]" pageId="54" pageNumber="55">
<emphasis id="9BBCEA98FFF08A25FF7A321D6066DF29" bold="true" box="[151,299,1541,1563]" pageId="54" pageNumber="55">FIGURE 113.</emphasis>
Leptochromini, distribution of extant species of
<taxonomicName id="6EC84D09FFF08A25FCF4321D62E4DF28" box="[793,937,1541,1563]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFF08A25FCF4321D62E4DF28" box="[793,937,1541,1563]" italics="true" pageId="54" pageNumber="55">Leptochromus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="FDB76602FFF18A24FF7A32B06524DE28" httpUri="https://zenodo.org/record/1454984/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,264,1704,1727]" targetBox="[163,1407,193,1665]" targetPageId="55">
<paragraph id="A977368AFFF18A24FF7A32B06524DE28" blockId="55.[151,1436,1704,1819]" pageId="55" pageNumber="56">
<emphasis id="9BBCEA98FFF18A24FF7A32B06020DF8D" bold="true" box="[151,365,1704,1727]" pageId="55" pageNumber="56">FIGURES 114–118.</emphasis>
Leptochromini, morphological structures,
<taxonomicName id="6EC84D09FFF18A24FCC032B162B2DF8C" box="[813,1023,1705,1727]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="9BBCEA98FFF18A24FCC032B162B2DF8C" box="[813,1023,1705,1727]" italics="true" pageId="55" pageNumber="56">Leptochromus agilis</emphasis>
</taxonomicName>
(114) and
<taxonomicName id="6EC84D09FFF18A24FB9A32B16415DF8C" box="[1143,1368,1705,1727]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="9BBCEA98FFF18A24FB9A32B16415DF8C" box="[1143,1368,1705,1727]" italics="true" pageId="55" pageNumber="56">Leptochromus laselva</emphasis>
</taxonomicName>
(115– 118). Head in dorsal (114), dorsolateral (115) and anteroventral (116) views; thorax and abdomen in ventral view (117); right procoxal, protrochanter and base of profemur in ventral view (118). Abbreviations: fem1, profemur; mxp2–4, maxillary palpomere II–IV; pd, pedicel; pgp, postgenal (= subocular) process; sc, scape; tr1, protrochanter.
</paragraph>
</caption>
<caption id="FDB76602FFFE8A2BFF7A33206386DE98" httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,264,1848,1871]" targetBox="[151,1436,193,1828]" targetPageId="56">
<paragraph id="A977368AFFFE8A2BFF7A33206386DE98" blockId="56.[151,1436,1848,1963]" pageId="56" pageNumber="57">
<emphasis id="9BBCEA98FFFE8A2BFF7A33206020DE7D" bold="true" box="[151,365,1848,1871]" pageId="56" pageNumber="57">FIGURES 119–123.</emphasis>
Leptochromini, morphological structures,
<taxonomicName id="6EC84D09FFFE8A2BFCC93321654EDE7C" box="[804,1027,1849,1871]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="9BBCEA98FFFE8A2BFCC93321654EDE7C" box="[804,1027,1849,1871]" italics="true" pageId="56" pageNumber="57">Leptochromus laselva</emphasis>
</taxonomicName>
. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum.
</paragraph>
</caption>
<caption id="FDB76602FFFF8A2AFF7A31FA65AFDF82" httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,264,1506,1529]" targetBox="[157,1436,197,1462]" targetPageId="57">
<paragraph id="A977368AFFFF8A2AFF7A31FA65AFDF82" blockId="57.[151,1437,1506,1713]" pageId="57" pageNumber="58">
<emphasis id="9BBCEA98FFFF8A2AFF7A31FA6021DCCB" bold="true" box="[151,364,1506,1529]" pageId="57" pageNumber="58">FIGURES 124–126.</emphasis>
Leptochromini, morphological structures,
<taxonomicName id="6EC84D09FFFF8A2AFCF131FB62B4DCCA" box="[796,1017,1507,1529]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="9BBCEA98FFFF8A2AFCF131FB62B4DCCA" box="[796,1017,1507,1529]" italics="true" pageId="57" pageNumber="58">Leptochromus laselva</emphasis>
</taxonomicName>
. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite.
</paragraph>
</caption>
<paragraph id="A977368AFFFF8A2AFF2A32C565BEDEC2" blockId="57.[151,1436,1757,2034]" pageId="57" pageNumber="58">
Prothorax (
<figureCitation id="31F32A0FFFFF8A2AFEA932C560ADDFC5" box="[324,480,1757,1782]" captionStart="FIGURES 107–108" captionStartId="53.[151,264,1815,1838]" captionTargetBox="[169,1424,206,1772]" captionTargetId="figure@53.[151,1436,193,1794]" captionTargetPageId="53" captionText="FIGURES 107–108. Leptochromini, examples of extant species; dorsal (107, 109) and dorsolateral (108) habitus. Leptochromus laselva (107–108); Leptochromus agilis (109)." httpUri="https://zenodo.org/record/1454978/files/figure.png" pageId="57" pageNumber="58">Figs 107–108</figureCitation>
,
<figureCitation id="31F32A0FFFFF8A2AFE0132C56355DFC5" box="[492,536,1757,1782]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="57" pageNumber="58">125</figureCitation>
,
<figureCitation id="31F32A0FFFFF8A2AFDC932C66302DFC5" box="[548,591,1758,1782]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="57" pageNumber="58">131</figureCitation>
) elongate, strongly convex, broadest near anterior third or at least distinctly in front of middle. Pronotum with anterior and posterior margins arcuate or nearly straight, sides rounded in anterior half and sinuate in posterior half; anterior and posterior corners obtuse-angled; pronotal base with a short posterior 'collar' (
<figureCitation id="31F32A0FFFFF8A2AFEB6335160F0DE51" box="[347,445,1865,1890]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="57" pageNumber="58">Fig. 131</figureCitation>
;
<emphasis id="9BBCEA98FFFF8A2AFE24335160B8DE51" box="[457,501,1865,1890]" italics="true" pageId="57" pageNumber="58">prcl</emphasis>
) demarcated by a dorsal transverse row of several variously distinct pits connected by a groove. Prosternum (
<figureCitation id="31F32A0FFFFF8A2AFE2C33756365DEB5" box="[449,552,1901,1926]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="57" pageNumber="58">Figs 125</figureCitation>
,
<figureCitation id="31F32A0FFFFF8A2AFDDB3376632CDEB5" box="[566,609,1902,1926]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="57" pageNumber="58">130</figureCitation>
) with basisternal part (
<figureCitation id="31F32A0FFFFF8A2AFC8633756282DEB5" box="[875,975,1901,1926]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="57" pageNumber="58">Fig. 125</figureCitation>
;
<emphasis id="9BBCEA98FFFF8A2AFC3133756545DEB5" box="[988,1032,1901,1926]" italics="true" pageId="57" pageNumber="58">bstr</emphasis>
) much longer than coxal part (
<figureCitation id="31F32A0FFFFF8A2AFA8333756189DE99" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="57" pageNumber="58">Fig. 125</figureCitation>
;
<emphasis id="9BBCEA98FFFF8A2AFF3F338B61B0DE99" box="[210,253,1939,1962]" italics="true" pageId="57" pageNumber="58">cxst</emphasis>
). Prosternum laterally completely fused with hypomera. Coxal region demarcated anteriorly by carina extending laterally up to apices of very weakly developed adcoxal hypomeral lobes. Procoxal cavities broadly open. Prosternal intercoxal process indistinct or absent. Ventral surface of prothorax densely setose.
</paragraph>
<caption id="FDB76602FFFC8A29FF7A332F606BDED4" httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,264,1847,1870]" targetBox="[171,1415,193,1830]" targetPageId="58">
<paragraph id="A977368AFFFC8A29FF7A332F606BDED4" blockId="58.[151,1437,1847,2023]" pageId="58" pageNumber="59">
<emphasis id="9BBCEA98FFFC8A29FF7A332F603CDE7E" bold="true" box="[151,369,1847,1870]" pageId="58" pageNumber="59">FIGURES 127–131.</emphasis>
Leptochromini, morphological structures,
<taxonomicName id="6EC84D09FFFC8A29FCDF3320655EDE7D" box="[818,1043,1848,1870]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="9BBCEA98FFFC8A29FCDF3320655EDE7D" box="[818,1043,1848,1870]" italics="true" pageId="58" pageNumber="59">Leptochromus laselva</emphasis>
</taxonomicName>
. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter.
</paragraph>
</caption>
<caption id="FDB76602FFFD8A28FF7A31936463DF34" httpUri="https://zenodo.org/record/1454992/files/figure.png" pageId="59" pageNumber="60" startId="59.[151,264,1419,1442]" targetBox="[151,1436,193,1398]" targetPageId="59">
<paragraph id="A977368AFFFD8A28FF7A31936463DF34" blockId="59.[151,1438,1419,1543]" pageId="59" pageNumber="60">
<emphasis id="9BBCEA98FFFD8A28FF7A3193603DDC92" bold="true" box="[151,368,1419,1442]" pageId="59" pageNumber="60">FIGURES 132–134.</emphasis>
Leptochromini, morphological structures,
<taxonomicName id="6EC84D09FFFD8A28FCDC3194655EDC91" box="[817,1043,1420,1442]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="9BBCEA98FFFD8A28FCDC3194655EDC91" box="[817,1043,1420,1442]" italics="true" pageId="59" pageNumber="60">Leptochromus laselva</emphasis>
</taxonomicName>
. Pterothorax and abdomen in ventral view (132); metaventrite in lateral view (133); last visible abdominal sternites of male in ventral view (134). Abbreviations: aest3, metanepisternum; cp, copulatory piece; cx3, metacoxa; epm3, metepimeron; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; par, paramere; si, setose impression; st–s8; sternite III–VIII; v3, metaventrite.
</paragraph>
</caption>
<paragraph id="A977368AFFFD8A28FF2A322F6333DEA7" blockId="59.[151,1437,1591,2012]" pageId="59" pageNumber="60">
Mesoventrite (
<figureCitation id="31F32A0FFFFD8A28FE84322F6080DF63" box="[361,461,1591,1616]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="59" pageNumber="60">Figs 126</figureCitation>
,
<figureCitation id="31F32A0FFFFD8A28FE3432206349DF63" box="[473,516,1592,1616]" captionStart="FIGURES 132–134" captionStartId="59.[151,264,1419,1442]" captionTargetBox="[151,1436,193,1398]" captionTargetId="figure@59.[151,1436,193,1398]" captionTargetPageId="59" captionText="FIGURES 132–134. Leptochromini, morphological structures, Leptochromus laselva. Pterothorax and abdomen in ventral view (132); metaventrite in lateral view (133); last visible abdominal sternites of male in ventral view (134). Abbreviations: aest3, metanepisternum; cp, copulatory piece; cx3, metacoxa; epm3, metepimeron; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; par, paramere; si, setose impression; st–s8; sternite III–VIII; v3, metaventrite." httpUri="https://zenodo.org/record/1454992/files/figure.png" pageId="59" pageNumber="60">132</figureCitation>
) subtrapezoidal, broadening posteriorly. Prepecti (
<figureCitation id="31F32A0FFFFD8A28FBD5322F65D4DF63" box="[1080,1177,1591,1616]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="59" pageNumber="60">Fig. 126</figureCitation>
;
<emphasis id="9BBCEA98FFFD8A28FB4932216585DF63" box="[1188,1224,1593,1616]" italics="true" pageId="59" pageNumber="60">pre</emphasis>
) long and together with anteromedian mesoventral area forming a massive 'collar', which bears an indistinct transverse groove just behind its anterior ridge, posterior margin of collar broadly subtriangular, only slightly projecting posterad at middle, tip of this median projection is broadly separated from a similar anteromedian projection of posterior margin of setose impression (
<figureCitation id="31F32A0FFFFD8A28FE0632DF6302DFD3" box="[491,591,1735,1760]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="59" pageNumber="60">Fig. 126</figureCitation>
;
<emphasis id="9BBCEA98FFFD8A28FDB332D1633CDFD3" box="[606,625,1737,1760]" italics="true" pageId="59" pageNumber="60">si</emphasis>
), margins of impression are obscured by setae (
<figureCitation id="31F32A0FFFFD8A28FB7332DF644EDFD3" box="[1182,1283,1735,1760]" captionStart="FIGURES 132–134" captionStartId="59.[151,264,1419,1442]" captionTargetBox="[151,1436,193,1398]" captionTargetId="figure@59.[151,1436,193,1398]" captionTargetPageId="59" captionText="FIGURES 132–134. Leptochromini, morphological structures, Leptochromus laselva. Pterothorax and abdomen in ventral view (132); metaventrite in lateral view (133); last visible abdominal sternites of male in ventral view (134). Abbreviations: aest3, metanepisternum; cp, copulatory piece; cx3, metacoxa; epm3, metepimeron; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; par, paramere; si, setose impression; st–s8; sternite III–VIII; v3, metaventrite." httpUri="https://zenodo.org/record/1454992/files/figure.png" pageId="59" pageNumber="60">Fig. 132</figureCitation>
). Region just behind collar strongly and abruptly constricted, ventrally forming setose impression (
<figureCitation id="31F32A0FFFFD8A28FBB532F365F3DE37" box="[1112,1214,1771,1796]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="59" pageNumber="60">Figs 126</figureCitation>
,
<figureCitation id="31F32A0FFFFD8A28FB2732F465BADE37" box="[1226,1271,1772,1796]" captionStart="FIGURES 132–134" captionStartId="59.[151,264,1419,1442]" captionTargetBox="[151,1436,193,1398]" captionTargetId="figure@59.[151,1436,193,1398]" captionTargetPageId="59" captionText="FIGURES 132–134. Leptochromini, morphological structures, Leptochromus laselva. Pterothorax and abdomen in ventral view (132); metaventrite in lateral view (133); last visible abdominal sternites of male in ventral view (134). Abbreviations: aest3, metanepisternum; cp, copulatory piece; cx3, metacoxa; epm3, metepimeron; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; par, paramere; si, setose impression; st–s8; sternite III–VIII; v3, metaventrite." httpUri="https://zenodo.org/record/1454992/files/figure.png" pageId="59" pageNumber="60">132</figureCitation>
;
<emphasis id="9BBCEA98FFFD8A28FAE932F5645ADE37" box="[1284,1303,1773,1796]" italics="true" pageId="59" pageNumber="60">si</emphasis>
) filled with dense, unmodified setae. Mesoventral intercoxal process (
<figureCitation id="31F32A0FFFFD8A28FCCE331762C5DE1B" box="[803,904,1807,1832]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="59" pageNumber="60">Figs 126</figureCitation>
,
<figureCitation id="31F32A0FFFFD8A28FC783308628FDE1B" box="[917,962,1808,1832]" captionStart="FIGURES 132–134" captionStartId="59.[151,264,1419,1442]" captionTargetBox="[151,1436,193,1398]" captionTargetId="figure@59.[151,1436,193,1398]" captionTargetPageId="59" captionText="FIGURES 132–134. Leptochromini, morphological structures, Leptochromus laselva. Pterothorax and abdomen in ventral view (132); metaventrite in lateral view (133); last visible abdominal sternites of male in ventral view (134). Abbreviations: aest3, metanepisternum; cp, copulatory piece; cx3, metacoxa; epm3, metepimeron; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; par, paramere; si, setose impression; st–s8; sternite III–VIII; v3, metaventrite." httpUri="https://zenodo.org/record/1454992/files/figure.png" pageId="59" pageNumber="60">132</figureCitation>
;
<emphasis id="9BBCEA98FFFD8A28FC233309654ADE1B" box="[974,1031,1809,1832]" italics="true" pageId="59" pageNumber="60">msvp</emphasis>
) long, narrow and weakly elevated, nearly parallel-sided or indistinctly broadened near middle of mesocoxal cavities, fully separating mesocoxae, posteriorly with indistinct posterior tip, which is fused with metaventrite. Mesanepisterna large and subtriangular; mesepimera partly exposed in ventral view.
</paragraph>
<paragraph id="A977368AFFFD8A28FF2A33B865B6DEEF" blockId="59.[151,1437,1591,2012]" pageId="59" pageNumber="60">
Mesonotum with subtriangular, strongly elongate mesoscutellum (
<figureCitation id="31F32A0FFFFD8A28FC2433876563DE8B" box="[969,1070,1951,1976]" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="59" pageNumber="60">Fig. 122</figureCitation>
;
<emphasis id="9BBCEA98FFFD8A28FBD333876521DE8B" box="[1086,1132,1951,1976]" italics="true" pageId="59" pageNumber="60">scl2</emphasis>
) with pointed or rounded apex, in intact specimens only its very tip visible between elytral bases; scutoscutellar suture absent.
</paragraph>
<caption id="FDB76602FFFA8A2FFF7A33346364DE51" httpUri="https://zenodo.org/record/1454994/files/figure.png" pageId="60" pageNumber="61" startId="60.[151,264,1836,1859]" targetBox="[151,1436,193,1815]" targetPageId="60">
<paragraph id="A977368AFFFA8A2FFF7A33346364DE51" blockId="60.[151,1435,1836,1890]" pageId="60" pageNumber="61">
<emphasis id="9BBCEA98FFFA8A2FFF7A33346023DE71" bold="true" box="[151,366,1836,1859]" pageId="60" pageNumber="61">FIGURES 135–137.</emphasis>
Leptochromini, habitats. La Selva Biological Station env., Costa Rica, habitat of
<taxonomicName id="6EC84D09FFFA8A2FFB5B333564DBDE70" box="[1206,1430,1837,1859]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="9BBCEA98FFFA8A2FFB5B333564DBDE70" box="[1206,1430,1837,1859]" italics="true" pageId="60" pageNumber="61">Leptochromus laselva</emphasis>
</taxonomicName>
. Photo Rich Leschen &amp; Tara Cornelisse.
</paragraph>
</caption>
<paragraph id="A977368AFFFB8A2EFF2A348063F5D983" blockId="61.[151,1437,151,1725]" box="[199,696,151,176]" pageId="61" pageNumber="62">Metanotum and hind wings fully developed.</paragraph>
<paragraph id="A977368AFFFB8A2EFF2A34A46024D8FC" blockId="61.[151,1437,151,1725]" pageId="61" pageNumber="62">
Metaventrite (
<figureCitation id="31F32A0FFFFB8A2EFE8534A4609CD9E7" box="[360,465,188,213]" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="61" pageNumber="62">Figs 123</figureCitation>
,
<figureCitation id="31F32A0FFFFB8A2EFE0D34A46341D9E6" box="[480,524,188,213]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="61" pageNumber="62">126</figureCitation>
,
<figureCitation id="31F32A0FFFFB8A2EFDF634A4630BD9E7" box="[539,582,188,212]" captionStart="FIGURES 132–134" captionStartId="59.[151,264,1419,1442]" captionTargetBox="[151,1436,193,1398]" captionTargetId="figure@59.[151,1436,193,1398]" captionTargetPageId="59" captionText="FIGURES 132–134. Leptochromini, morphological structures, Leptochromus laselva. Pterothorax and abdomen in ventral view (132); metaventrite in lateral view (133); last visible abdominal sternites of male in ventral view (134). Abbreviations: aest3, metanepisternum; cp, copulatory piece; cx3, metacoxa; epm3, metepimeron; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; par, paramere; si, setose impression; st–s8; sternite III–VIII; v3, metaventrite." httpUri="https://zenodo.org/record/1454992/files/figure.png" pageId="61" pageNumber="62">132</figureCitation>
) short, subquadrate or subrectangular and slightly transverse, with lateral margins rounded; mesocoxal cavities not carinate; posterior margin of metaventrite deeply bisinuate laterally (in front of each metacoxa) and with a broad metaventral intercoxal process (
<figureCitation id="31F32A0FFFFB8A2EFC07351C6503D82E" box="[1002,1102,260,285]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="61" pageNumber="62">Fig. 126</figureCitation>
;
<emphasis id="9BBCEA98FFFB8A2EFBB1351D65DFD82F" box="[1116,1170,261,284]" italics="true" pageId="61" pageNumber="62">mtvp</emphasis>
) with variously deeply concave posterior margin and subtriangular posterolateral corners; metaventrite with lateral mesocoxal (
<figureCitation id="31F32A0FFFFB8A2EFADC353F64DAD873" box="[1329,1431,295,320]" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="61" pageNumber="62">Figs 123</figureCitation>
,
<figureCitation id="31F32A0FFFFB8A2EFF7A35546189D856" box="[151,196,332,357]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="61" pageNumber="62">126</figureCitation>
;
<emphasis id="9BBCEA98FFFB8A2EFF3D3554604DD856" box="[208,256,332,357]" italics="true" pageId="61" pageNumber="62">lmcf</emphasis>
) and postmesocoxal (
<figureCitation id="31F32A0FFFFB8A2EFE1835546317D857" box="[501,602,332,357]" captionStart="FIGURES 119–123" captionStartId="56.[151,264,1848,1871]" captionTargetBox="[151,1436,193,1828]" captionTargetId="figure@56.[151,1436,193,1828]" captionTargetPageId="56" captionText="FIGURES 119–123. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsal (119) and ventral (120) views; left maxillary palp in ventral view (121); mesonotum in dorsal view (122); pterothorax and abdominal base min ventral view (123). Abbreviations: pgp, postgenal (= subocular) process; lmcf, lateral mesocoxal fovea; mxp1–4, maxillary palpomere I–IV; pmcf, postmesocoxal fovea; scl2, mesoscutellum." httpUri="https://zenodo.org/record/1454986/files/figure.png" pageId="61" pageNumber="62">Figs 123</figureCitation>
,
<figureCitation id="31F32A0FFFFB8A2EFD8A355463D9D856" box="[615,660,332,357]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="61" pageNumber="62">126</figureCitation>
;
<emphasis id="9BBCEA98FFFB8A2EFD7235546398D857" box="[671,725,332,356]" italics="true" pageId="61" pageNumber="62">pmcf</emphasis>
) foveae. Posterior metaventral margin lacking adcoxal carinae. Metanepisterna (
<figureCitation id="31F32A0FFFFB8A2EFEB5357760F2D8BB" box="[344,447,367,392]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="61" pageNumber="62">Fig. 126</figureCitation>
;
<emphasis id="9BBCEA98FFFB8A2EFE3D35686347D8BB" box="[464,522,368,392]" italics="true" pageId="61" pageNumber="62">aest3</emphasis>
) broad and partly visible in ventral view, broadened posteriorly; metepimera broader than metanepisterna, with indistinctly demarcated inner and outer component, posteriorly extending far behind metacoxae.
</paragraph>
<paragraph id="A977368AFFFB8A2EFF2A35C462F7DB2B" blockId="61.[151,1437,151,1725]" pageId="61" pageNumber="62">
Metendosternite (metafurca) with stem much broader than long and broadly separated divergent lateral furcal arms (
<figureCitation id="31F32A0FFFFB8A2EFF3035E76073DB2B" box="[221,318,511,536]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="61" pageNumber="62">Fig. 126</figureCitation>
;
<emphasis id="9BBCEA98FFFB8A2EFEA735E76031DB2B" box="[330,380,511,536]" italics="true" pageId="61" pageNumber="62">lmfa</emphasis>
), additionally with median longitudinal projection.
</paragraph>
<paragraph id="A977368AFFFB8A2EFF2A363D65ABDA0B" blockId="61.[151,1437,151,1725]" pageId="61" pageNumber="62">
Legs (
<figureCitation id="31F32A0FFFFB8A2EFEF8363C60CCDB0F" box="[277,385,548,573]" captionStart="FIGURES 107–108" captionStartId="53.[151,264,1815,1838]" captionTargetBox="[169,1424,206,1772]" captionTargetId="figure@53.[151,1436,193,1794]" captionTargetPageId="53" captionText="FIGURES 107–108. Leptochromini, examples of extant species; dorsal (107, 109) and dorsolateral (108) habitus. Leptochromus laselva (107–108); Leptochromus agilis (109)." httpUri="https://zenodo.org/record/1454978/files/figure.png" pageId="61" pageNumber="62">Figs 107</figureCitation>
–109, 117–118, 126, 130, 132) conspicuously long, slender. Pro- and mesocoxa short subconical, metacoxa with nearly hemispherical basal part and subconical distal part. Mesocoxa lacking coxal bristles, but usually with a row or patch of thin long setae. All trochanters short and subtriangular; protrochanters (
<figureCitation id="31F32A0FFFFB8A2EFF7236976048DB9B" box="[159,261,655,680]" captionStart="FIGURES 114–118" captionStartId="55.[151,264,1704,1727]" captionTargetBox="[163,1407,193,1665]" captionTargetId="figure@55.[163,1407,193,1665]" captionTargetPageId="55" captionText="FIGURES 114–118. Leptochromini, morphological structures, Leptochromus agilis (114) and Leptochromus laselva (115– 118). Head in dorsal (114), dorsolateral (115) and anteroventral (116) views; thorax and abdomen in ventral view (117); right procoxal, protrochanter and base of profemur in ventral view (118). Abbreviations: fem1, profemur; mxp2–4, maxillary palpomere II–IV; pd, pedicel; pgp, postgenal (= subocular) process; sc, scape; tr1, protrochanter." httpUri="https://zenodo.org/record/1454984/files/figure.png" pageId="61" pageNumber="62">Figs 118</figureCitation>
,
<figureCitation id="31F32A0FFFFB8A2EFEF93697600DDB9B" box="[276,320,655,680]" captionStart="FIGURES 124–126" captionStartId="57.[151,264,1506,1529]" captionTargetBox="[157,1436,197,1462]" captionTargetId="figure@57.[151,1436,193,1485]" captionTargetPageId="57" captionText="FIGURES 124–126. Leptochromini, morphological structures, Leptochromus laselva. Head in ventral view (124); prothorax in ventral view (125); pterothorax and abdomen in ventral view (126). Abbreviations: aed, aedeagus; aest3, metanepisternum; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; cx1, procoxa; cxst, coxal part of prosternum; hr, hypostomal ridge; lhl, lateral hypopharyngeal lobe; lmcf, lateral mesocoxal fovea; lmfa, lateral metafurcal arm; lp, labial palp; mn, mentum; msf, mesofurca; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pgp, postgenal (=subocular) process; pm, prementum; pmcf, postmesocoxal fovea; ppf, palpifer; pre, prepectus; ptp, posterior tentorial pit; si, setose impression; smn, submentum; st3–8, abdominal sternite III–VIII; tr1, protrochanter; v3, metaventrite." httpUri="https://zenodo.org/record/1454988/files/figure.png" pageId="61" pageNumber="62">125</figureCitation>
,
<figureCitation id="31F32A0FFFFB8A2EFEA236886037DB9B" box="[335,378,656,680]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="61" pageNumber="62">130</figureCitation>
) in both sexes with a row of several thick bristles (in one species reduced to 1–2). Femora weakly clavate, profemur in both sexes with a ventral row of several conspicuously thick bristles, usually with papillate insertions (
<figureCitation id="31F32A0FFFFB8A2EFE9736CF6093DBC3" box="[378,478,727,752]" captionStart="FIGURES 114–118" captionStartId="55.[151,264,1704,1727]" captionTargetBox="[163,1407,193,1665]" captionTargetId="figure@55.[163,1407,193,1665]" captionTargetPageId="55" captionText="FIGURES 114–118. Leptochromini, morphological structures, Leptochromus agilis (114) and Leptochromus laselva (115– 118). Head in dorsal (114), dorsolateral (115) and anteroventral (116) views; thorax and abdomen in ventral view (117); right procoxal, protrochanter and base of profemur in ventral view (118). Abbreviations: fem1, profemur; mxp2–4, maxillary palpomere II–IV; pd, pedicel; pgp, postgenal (= subocular) process; sc, scape; tr1, protrochanter." httpUri="https://zenodo.org/record/1454984/files/figure.png" pageId="61" pageNumber="62">Figs 118</figureCitation>
,
<figureCitation id="31F32A0FFFFB8A2EFE0736C06355DBC3" box="[490,536,728,752]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="61" pageNumber="62">130</figureCitation>
). Tibiae slender, protibiae in both sexes curved, only rarely nearly straight. Tarsi long and slender, nearly subcylindrical, tarsomeres I–V subequal or reducing in length, tarsomere V strongly elongate, with curved and slender claws lacking elongate costae; empodial region was not studied.
</paragraph>
<paragraph id="A977368AFFFB8A2EFF2A375C6458DA97" blockId="61.[151,1437,151,1725]" pageId="61" pageNumber="62">
Elytra (
<figureCitation id="31F32A0FFFFB8A2EFEF6375C60F4DA6F" box="[283,441,836,861]" captionStart="FIGURES 107–108" captionStartId="53.[151,264,1815,1838]" captionTargetBox="[169,1424,206,1772]" captionTargetId="figure@53.[151,1436,193,1794]" captionTargetPageId="53" captionText="FIGURES 107–108. Leptochromini, examples of extant species; dorsal (107, 109) and dorsolateral (108) habitus. Leptochromus laselva (107–108); Leptochromus agilis (109)." httpUri="https://zenodo.org/record/1454978/files/figure.png" pageId="61" pageNumber="62">Figs 107–108</figureCitation>
,
<figureCitation id="31F32A0FFFFB8A2EFE2B375C60BCDA6F" box="[454,497,836,860]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="61" pageNumber="62">131</figureCitation>
) oval, strongly convex, with prominent humeral calli (
<figureCitation id="31F32A0FFFFB8A2EFBB3375C658CDA6F" box="[1118,1217,836,861]" captionStart="FIGURES 127–131" captionStartId="58.[151,264,1847,1870]" captionTargetBox="[171,1415,193,1830]" captionTargetId="figure@58.[171,1415,193,1830]" captionTargetPageId="58" captionText="FIGURES 127–131. Leptochromini, morphological structures, Leptochromus laselva. Head in dorsolateral (127–128) and ventral (129) views; prothorax in ventral view (130); prothorax and elytral base in lateral view (131). Abbreviations: bstr, basisternal part of prosternum; cx1, procoxa; fm1, profemur; gp, gular plate; hcl, humeral callus; hr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp1–4, maxillary palpomere I–IV; pd, pedicel; pgp, postgenal (= subocular) process; pm, prementum; prcl, procoxal collar; ptp, posterior tentorial pit; sc, scape; scl2, mesoscutellum; smn, submentum; tr1, protrochanter." httpUri="https://zenodo.org/record/1454990/files/figure.png" pageId="61" pageNumber="62">Fig. 131</figureCitation>
;
<emphasis id="9BBCEA98FFFB8A2EFB20375C65A2DA6E" box="[1229,1263,836,861]" italics="true" pageId="61" pageNumber="62">hcl</emphasis>
), lacking basal impressions, with rounded apices; elytral disc with distinct large and deep punctures arranged in six nearly complete and typically regular longitudinal rows. Elytra variously densely setose, setae long and erect.
</paragraph>
<paragraph id="A977368AFFFB8A2EFF2A37B763A1DADF" blockId="61.[151,1437,151,1725]" pageId="61" pageNumber="62">
Abdomen (
<figureCitation id="31F32A0FFFFB8A2EFEA837B760E9DAFB" box="[325,420,943,968]" captionStart="FIGURES 132–134" captionStartId="59.[151,264,1419,1442]" captionTargetBox="[151,1436,193,1398]" captionTargetId="figure@59.[151,1436,193,1398]" captionTargetPageId="59" captionText="FIGURES 132–134. Leptochromini, morphological structures, Leptochromus laselva. Pterothorax and abdomen in ventral view (132); metaventrite in lateral view (133); last visible abdominal sternites of male in ventral view (134). Abbreviations: aest3, metanepisternum; cp, copulatory piece; cx3, metacoxa; epm3, metepimeron; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; par, paramere; si, setose impression; st–s8; sternite III–VIII; v3, metaventrite." httpUri="https://zenodo.org/record/1454992/files/figure.png" pageId="61" pageNumber="62">Fig. 132</figureCitation>
) with sternite III not fused with metaventrite, about as long as sternites IV and V together; sternite VIII in male distinctly emarginate (
<figureCitation id="31F32A0FFFFB8A2EFD9137CC6393DADF" box="[636,734,980,1005]" captionStart="FIGURES 132–134" captionStartId="59.[151,264,1419,1442]" captionTargetBox="[151,1436,193,1398]" captionTargetId="figure@59.[151,1436,193,1398]" captionTargetPageId="59" captionText="FIGURES 132–134. Leptochromini, morphological structures, Leptochromus laselva. Pterothorax and abdomen in ventral view (132); metaventrite in lateral view (133); last visible abdominal sternites of male in ventral view (134). Abbreviations: aest3, metanepisternum; cp, copulatory piece; cx3, metacoxa; epm3, metepimeron; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; par, paramere; si, setose impression; st–s8; sternite III–VIII; v3, metaventrite." httpUri="https://zenodo.org/record/1454992/files/figure.png" pageId="61" pageNumber="62">Fig. 134</figureCitation>
).
</paragraph>
<paragraph id="A977368AFFFB8A2EFF2A37EF64DCDD4F" blockId="61.[151,1437,151,1725]" pageId="61" pageNumber="62">
Aedeagus (illustrated in Jałoszyński
<emphasis id="9BBCEA98FFFB8A2EFD8537E163EEDD23" box="[616,675,1015,1040]" italics="true" pageId="61" pageNumber="62">et al.</emphasis>
(2018), Lord
<emphasis id="9BBCEA98FFFB8A2EFCA937E16232DD23" box="[836,895,1015,1040]" italics="true" pageId="61" pageNumber="62">et al.</emphasis>
(2014) and O’Keefe (2002)) strongly elongate, with symmetrical median lobe, parameres heavily sclerotized, symmetrical, with arrowhead-shaped or spatulate and pointed apices; flagellum simple, not coiled. Ejaculatory duct with elongate and narrow sperm pump lacking funnel-like structures. Aedeagus in repose positioned symmetrically inside abdomen, with basal orifice facing up.
</paragraph>
<paragraph id="A977368AFFFB8A2EFF2A309F64DDDD93" blockId="61.[151,1437,151,1725]" box="[199,1424,1159,1185]" pageId="61" pageNumber="62">Spermatheca (illustrated by O’Keefe (2002)) globular, nearly spherical, with relatively large accessory gland.</paragraph>
<paragraph id="A977368AFFFB8A2EFF2A30B460D9DDF6" blockId="61.[151,1437,151,1725]" box="[199,404,1196,1221]" pageId="61" pageNumber="62">
<emphasis id="9BBCEA98FFFB8A2EFF2A30B4605ADDF6" bold="true" box="[199,279,1196,1221]" pageId="61" pageNumber="62">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="E1D26501FFFB8A2EFF2A30D764DADF46" pageId="61" pageNumber="62" type="description">
<paragraph id="A977368AFFFB8A2EFF2A30D764D0DC3E" blockId="61.[151,1437,151,1725]" pageId="61" pageNumber="62">
<emphasis id="9BBCEA98FFFB8A2EFF2A30D7637FDDDB" bold="true" box="[199,562,1231,1256]" pageId="61" pageNumber="62">Composition and distribution.</emphasis>
<taxonomicName id="6EC84D09FFFB8A2EFDD730D76394DDDB" box="[570,729,1231,1256]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFFB8A2EFDD730D76394DDDB" box="[570,729,1231,1256]" italics="true" pageId="61" pageNumber="62">Leptochromus</emphasis>
</taxonomicName>
comprises five extant species distributed in
<collectingCountry id="D1DF761AFFFB8A2EFB3E30D76457DDDB" box="[1235,1306,1231,1256]" name="Brazil" pageId="61" pageNumber="62">Brazil</collectingCountry>
,
<collectingCountry id="D1DF761AFFFB8A2EFACB30D764DADDDB" box="[1318,1431,1231,1256]" name="Colombia" pageId="61" pageNumber="62">Colombia</collectingCountry>
,
<collectingCountry id="D1DF761AFFFB8A2EFF7A30EC605DDC3E" box="[151,272,1268,1293]" name="Costa Rica" pageId="61" pageNumber="62">
<collectingRegion id="6B0CF868FFFB8A2EFF7A30EC619ADC3F" box="[151,215,1268,1292]" country="New Zealand" name="West Coast" pageId="61" pageNumber="62">Costa</collectingRegion>
Rica
</collectingCountry>
and
<collectingRegion id="6B0CF868FFFB8A2EFEAA30ED60EDDC3F" box="[327,416,1269,1292]" country="Panama" name="Panama" pageId="61" pageNumber="62">
<collectingCountry id="D1DF761AFFFB8A2EFEAA30ED60EDDC3F" box="[327,416,1269,1292]" name="Panama" pageId="61" pageNumber="62">Panama</collectingCountry>
</collectingRegion>
(
<figureCitation id="31F32A0FFFFB8A2EFE4230EC6342DC3F" box="[431,527,1268,1293]" captionStart="FIGURE 113" captionStartId="54.[151,250,1541,1563]" captionTargetBox="[151,1436,971,1519]" captionTargetId="figure@54.[151,1436,971,1519]" captionTargetPageId="54" captionText="FIGURE 113. Leptochromini, distribution of extant species of Leptochromus." httpUri="https://zenodo.org/record/1454982/files/figure.png" pageId="61" pageNumber="62">Fig. 113</figureCitation>
), and one extinct species (
<figureCitation id="31F32A0FFFFB8A2EFCD930EC62DEDC3F" box="[820,915,1268,1293]" captionStart="FIGURES 110–112" captionStartId="54.[151,264,807,830]" captionTargetBox="[151,1436,193,786]" captionTargetId="figure@54.[151,1436,193,786]" captionTargetPageId="54" captionText="FIGURES 110–112. Leptochromini, examples of extinct species; dorsal (110), lateral (111) and laterodorsal (112) habitus. Leptochromus palaeomexicanus (110; Middle Miocene of Mexico; photo George Poinar); Euroleptochromus setifer (111; Upper Eocene of Europe); Rovnoleptochromus ableptonoides (112; Upper Eocene of Europe; after Jałoszyński &amp; Perkovsky (2016))." httpUri="https://zenodo.org/record/1454980/files/figure.png" pageId="61" pageNumber="62">Fig. 110</figureCitation>
) found in the Middle Miocene
<collectingRegion id="6B0CF868FFFB8A2EFB1D30EC6407DC3E" box="[1264,1354,1268,1293]" country="Mexico" name="Chiapas" pageId="61" pageNumber="62">Chiapas</collectingRegion>
amber.
</paragraph>
<paragraph id="A977368AFFFB8A2EFF2A310F60EEDF1F" blockId="61.[151,1437,151,1725]" pageId="61" pageNumber="62">
<emphasis id="9BBCEA98FFFB8A2EFF2A310F60CADC03" bold="true" box="[199,391,1303,1328]" pageId="61" pageNumber="62">Natural history.</emphasis>
Little is known about natural history of
<taxonomicName id="6EC84D09FFFB8A2EFCB6310F62B7DC03" box="[859,1018,1303,1328]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFFB8A2EFCB6310F62B7DC03" box="[859,1018,1303,1328]" italics="true" pageId="61" pageNumber="62">Leptochromus</emphasis>
</taxonomicName>
, except collecting circumstances for some species. Adults were collected from moist leaf litter, moss and lichens in subtropical forests (
<figureCitation id="31F32A0FFFFB8A2EFB1D312464DDDC66" box="[1264,1424,1340,1365]" captionStart="FIGURES 135–137" captionStartId="60.[151,264,1836,1859]" captionTargetBox="[151,1436,193,1815]" captionTargetId="figure@60.[151,1436,193,1815]" captionTargetPageId="60" captionText="FIGURES 135–137. Leptochromini, habitats. La Selva Biological Station env., Costa Rica, habitat of Leptochromus laselva. Photo Rich Leschen &amp; Tara Cornelisse." httpUri="https://zenodo.org/record/1454994/files/figure.png" pageId="61" pageNumber="62">Figs 135–137</figureCitation>
). Numerous individuals of
<taxonomicName id="6EC84D09FFFB8A2EFE2C314763F6DC4B" box="[449,699,1375,1400]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="9BBCEA98FFFB8A2EFE2C314763F6DC4B" box="[449,699,1375,1400]" italics="true" pageId="61" pageNumber="62">Leptochromus laselva</emphasis>
</taxonomicName>
were collected &quot;lifting dead palm fronds lying on the floor of secondary-growth forest (...) and beating them over a white sheet during late afternoon and early evening hours&quot; (Lord
<emphasis id="9BBCEA98FFFB8A2EFF3331B1605ADCF3" box="[222,279,1447,1472]" italics="true" pageId="61" pageNumber="62">et al.</emphasis>
2014). The latter authors also noted that although adults of
<taxonomicName id="6EC84D09FFFB8A2EFC5331B16561DCF3" box="[958,1068,1447,1472]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="9BBCEA98FFFB8A2EFC5331B16561DCF3" box="[958,1068,1447,1472]" italics="true" pageId="61" pageNumber="62">L. laselva</emphasis>
</taxonomicName>
are fully winged, no individuals were collected at light (including UV) for two weeks at the collecting site.
<taxonomicName id="6EC84D09FFFB8A2EFC3231D465B5DCD6" authorityName="Motschulsky" authorityYear="1855" box="[991,1272,1484,1509]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="fulvescens">
<emphasis id="9BBCEA98FFFB8A2EFC3231D465B5DCD6" box="[991,1272,1484,1509]" italics="true" pageId="61" pageNumber="62">Leptochromus fulvescens</emphasis>
</taxonomicName>
and
<taxonomicName id="6EC84D09FFFB8A2EFAC231D564D1DCD6" box="[1327,1436,1484,1509]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="9BBCEA98FFFB8A2EFAC231D564D1DCD6" box="[1327,1436,1484,1509]" italics="true" pageId="61" pageNumber="62">L. laselva</emphasis>
</taxonomicName>
were also collected by canopy fogging, demonstrating that members of this genus are active not only on the forest floor (Lord
<emphasis id="9BBCEA98FFFB8A2EFEF6320D601EDF1E" box="[283,339,1556,1581]" italics="true" pageId="61" pageNumber="62">et al.</emphasis>
2014).
</paragraph>
<paragraph id="A977368AFFFB8A2EFF2A322F64DADF46" blockId="61.[151,1437,151,1725]" pageId="61" pageNumber="62">
The unusual modifications of the maxillary palpomere II and postgenae may be used by beetles as a sensory and/or prey-capture apparatus, but hunting and feeding-related behavior of
<taxonomicName id="6EC84D09FFFB8A2EFC0F324465CCDF46" box="[994,1153,1628,1653]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFFB8A2EFC0F324465CCDF46" box="[994,1153,1628,1653]" italics="true" pageId="61" pageNumber="62">Leptochromus</emphasis>
</taxonomicName>
have not been observed.
</paragraph>
</subSubSection>
<subSubSection id="E1D26501FFFB8A2EFF2A3267633CDF8F" pageId="61" pageNumber="62" type="discussion">
<paragraph id="A977368AFFFB8A2EFF2A3267633CDF8F" blockId="61.[151,1437,151,1725]" pageId="61" pageNumber="62">
<emphasis id="9BBCEA98FFFB8A2EFF2A32676076DFAB" bold="true" box="[199,315,1663,1688]" pageId="61" pageNumber="62">Remarks.</emphasis>
All nominal extant and extinct species of
<taxonomicName id="6EC84D09FFFB8A2EFCF2326762F3DFAB" box="[799,958,1663,1688]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFFB8A2EFCF2326762F3DFAB" box="[799,958,1663,1688]" italics="true" pageId="61" pageNumber="62">Leptochromus</emphasis>
</taxonomicName>
were adequately described or redescribed by O’Keefe (2002) and Lord
<emphasis id="9BBCEA98FFFB8A2EFE0D32BD6355DF8E" box="[480,536,1700,1725]" italics="true" pageId="61" pageNumber="62">et al.</emphasis>
(2014).
</paragraph>
</subSubSection>
</treatment>
</document>
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