plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/21/61/87/2161879CFFE38A34FF7A36466567DFE8.xml
maintenance 29c63c90d1 added 21
2024-06-21 12:31:17 +02:00

403 lines
111 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="C6C2029970D00C258A63D819541A2EE7" ID-CLB-Dataset="29175" ID-DOI="10.11646/zootaxa.4453.1.1" ID-GBIF-Dataset="348e1a52-72a5-4809-b533-a158fc7a298a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1454920" ID-ZooBank="866690A9-0462-4892-AE29-9AAC623F87B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539178978648" checkinUser="plazi" docAuthor="Paweł Jałoszyński" docDate="2018" docId="2161879CFFE38A34FF7A36466567DFE8" docLanguage="en" docName="zootaxa.4453.1.1.pdf.imf" docOrigin="Zootaxa 4453 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Clidicini Casey" docType="treatment" docVersion="8" lastPageNumber="40" masterDocId="DD58FFE4FFC68A13FFED3418614DD933" masterDocTitle="World genera of Mastigitae: review of morphological structures and new ecological data (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="119" masterPageNumber="1" pageNumber="38" updateTime="1698572836684" updateUser="ExternalLinkService">
<mods:mods id="32E6FBB93DCF4AAF6103EB5793C259B0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D24B20393872D937B497D65B94E74397">
<mods:title id="37CBD3487D6EC81FF52C6B716753C1B1">World genera of Mastigitae: review of morphological structures and new ecological data (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="C9527DB6BCB23112A35128A2D57E6C70" type="personal">
<mods:role id="1CA3659DD76BBD926899AAA3AA5B4426">
<mods:roleTerm id="032F04F451B1DE06250784EB6DF753FB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="407D271DDA6D9B3CF974E73148EBAA66">Paweł Jałoszyński</mods:namePart>
</mods:name>
<mods:typeOfResource id="29B637CF4126788E8E217ED3E8210E7A">text</mods:typeOfResource>
<mods:relatedItem id="D8485809AEB0B11BB41990E94B66B19C" type="host">
<mods:titleInfo id="2CD5FFF0DB0136CB071E600E62B0BD70">
<mods:title id="9AC42EA547943E0E652D8F0D3F3C060F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="982A6B963B29CD80E7EA580C66C258E3">
<mods:date id="98F426CB58C5BCF88B711DE6887ED8A5">2018</mods:date>
<mods:detail id="AF7F582F3146CB96D56713BC6CC09769" type="pubDate">
<mods:number id="47FB97DF1BEDF8E49E4574FD794127B3">2018-07-30</mods:number>
</mods:detail>
<mods:detail id="F74F4BE9F7A7191F962F0E718D842DF7" type="volume">
<mods:number id="016CF2293406B12B3DE1A6F14BD8140D">4453</mods:number>
</mods:detail>
<mods:detail id="0E28FF13A63182E0D37F2CF11CE6E072" type="issue">
<mods:number id="7F1844943B34D69E22426BF85CDCDB5B">1</mods:number>
</mods:detail>
<mods:extent id="FC1456738E727B5F3B6673A62A8A1C05" unit="page">
<mods:start id="1744EBCAC83CAAB9DDCAC7FE421450B0">1</mods:start>
<mods:end id="5CECCF79EDB67A78B91BE8EF2610143F">119</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BE5A8BBFF88D49D5C891BD6E9B25EC68">journal article</mods:classification>
<mods:identifier id="0AA09F50A23EA46AB888F49923919BED" type="CLB-Dataset">29175</mods:identifier>
<mods:identifier id="130BB494CBD8695FA9757D9DB91DCC47" type="DOI">10.11646/zootaxa.4453.1.1</mods:identifier>
<mods:identifier id="2C7605E1D085A17AAA14CD7C7D7232C0" type="GBIF-Dataset">348e1a52-72a5-4809-b533-a158fc7a298a</mods:identifier>
<mods:identifier id="F9EBD3EADE435314F1CCC7D379CE3CE1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D03A95ADA598CF0B3B93C63CD08B00B3" type="Zenodo-Dep">1454920</mods:identifier>
<mods:identifier id="8D74538F4E2B6642A0DF6802F400A53D" type="ZooBank">866690A9-0462-4892-AE29-9AAC623F87B3</mods:identifier>
</mods:mods>
<treatment id="2161879CFFE38A34FF7A36466567DFE8" ID-DOI="http://doi.org/10.5281/zenodo.5976936" ID-GBIF-Taxon="148669764" ID-Zenodo-Dep="5976936" LSID="urn:lsid:plazi:treatment:2161879CFFE38A34FF7A36466567DFE8" httpUri="http://treatment.plazi.org/id/2161879CFFE38A34FF7A36466567DFE8" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
<subSubSection id="E1D26501FFE38A36FF7A364660EDDB4B" box="[151,416,606,632]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="A977368AFFE38A36FF7A364660EDDB4B" blockId="37.[151,416,606,632]" box="[151,416,606,632]" pageId="37" pageNumber="38">
<heading id="F23F81E6FFE38A36FF7A364660EDDB4B" bold="true" box="[151,416,606,632]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
<emphasis id="9BBCEA98FFE38A36FF7A364660EDDB4B" bold="true" box="[151,416,606,632]" pageId="37" pageNumber="38">
Tribe
<taxonomicName id="6EC84D09FFE38A36FF0F364660EDDB4B" ID-CoL="8XHM4" authority="Casey" authorityName="Casey" box="[226,416,606,632]" pageId="37" pageNumber="38" rank="tribe" tribe="Clidicini">Clidicini Casey</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E1D26501FFE38A36FF7A368664D1DBE0" pageId="37" pageNumber="38" type="reference_group">
<paragraph id="A977368AFFE38A36FF7A368662F7DB87" blockId="37.[151,1436,670,723]" box="[151,954,670,692]" pageId="37" pageNumber="38">
Clidicinae
<bibRefCitation id="CD594B7BFFE38A36FEE8368660C9DB87" box="[261,388,670,692]" pageId="37" pageNumber="38" refId="ref51905" refString="CASEY, T. L. (1897) COLEOPTEROLOGICAL NOTICES, VII. Annals of the New York Academy of Sciences, 9, 285 - 684." type="journal article">Casey, 1897</bibRefCitation>
: 541.
<typeStatus id="76738828FFE38A36FE2C368760BEDB87" box="[449,499,671,692]" pageId="37" pageNumber="38">Type</typeStatus>
genus:
<taxonomicName id="6EC84D09FFE38A36FDAF368662FBDB87" ID-CoL="8X77G" authority="LAPORTE dE CASTELNAU, 1832" authorityName="Laporte de Castelnau" authorityYear="1832" box="[578,950,670,692]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFE38A36FDAF368663DBDB87" box="[578,662,670,692]" italics="true" pageId="37" pageNumber="38">Clidicus</emphasis>
Laporte de Castelnau, 1832
</taxonomicName>
.
</paragraph>
<paragraph id="A977368AFFE38A36FF7A36A564D1DBE0" blockId="37.[151,1436,670,723]" box="[151,1436,701,723]" pageId="37" pageNumber="38">
Clidicini
<bibRefCitation id="CD594B7BFFE38A36FF1B36A560E0DBE0" box="[246,429,701,723]" pageId="37" pageNumber="38" refId="ref52555" refString="GANGLBAUER, L. (1899) Die Kafer von Mitteleuropa. Die Kafer des osterreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzosischen und italienischen Alpengebietes. Band 3. Staphylinoidea. II Theil. Scydmaenidae, Silphidae, Clambidae, Leptinidae, Platypsyl. CARL GEROLd'S SOHN, WIEN, 1046 PP." type="book">Ganglbauer, 1899</bibRefCitation>
: 60.
<typeStatus id="76738828FFE38A36FE3036A66342DBE0" box="[477,527,702,723]" pageId="37" pageNumber="38">Type</typeStatus>
genus:
<taxonomicName id="6EC84D09FFE38A36FDB336A5629FDBE0" ID-CoL="8X77G" authority="LAPORTE dE CASTELNAU, 1832" authorityName="Laporte de Castelnau" authorityYear="1832" box="[606,978,701,723]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFE38A36FDB336A563FFDBE0" box="[606,690,701,723]" italics="true" pageId="37" pageNumber="38">Clidicus</emphasis>
Laporte de Castelnau, 1832
</taxonomicName>
. Tribe redefined by Jałoszyński
<emphasis id="9BBCEA98FFE38A36FAF436A66401DBE0" box="[1305,1356,701,723]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
(2018).
</paragraph>
</subSubSection>
<subSubSection id="E1D26501FFE38A36FF7A36E160DFDCDC" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="A977368AFFE38A36FF7A36E1632FDC73" blockId="37.[151,1437,761,2009]" pageId="37" pageNumber="38">
<emphasis id="9BBCEA98FFE38A36FF7A36E1605CDA21" bold="true" box="[151,273,761,786]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
Clidicini differ from all other Mastigitae in two unique apomorphies: some setae on frons and vertex conspicuously long and erect among much shorter and denser basic vestiture (
<figureCitation id="31F32A0FFFE38A36FBA9370465D0DA06" box="[1092,1181,796,821]" captionStart="FIGURES 84–85" captionStartId="41.[151,264,1421,1444]" captionTargetBox="[151,1436,265,1400]" captionTargetId="figure@41.[151,1436,265,1400]" captionTargetPageId="41" captionText="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." httpUri="https://zenodo.org/record/1454964/files/figure.png" pageId="37" pageNumber="38">Fig. 84</figureCitation>
, but in some species indistinct); one or two conspicuously long and erect ventral setae present in basal half of each profemur (reduced in some species); and a combination of synapomorphies that can be found in other tribes and genera, but not occurring all together: only scape enlarged and as long as head or (more frequently) longer (e.g.,
<figureCitation id="31F32A0FFFE38A36FB16379D641CDAAE" box="[1275,1361,901,926]" captionStart="FIGURES 84–85" captionStartId="41.[151,264,1421,1444]" captionTargetBox="[151,1436,265,1400]" captionTargetId="figure@41.[151,1436,265,1400]" captionTargetPageId="41" captionText="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." httpUri="https://zenodo.org/record/1454964/files/figure.png" pageId="37" pageNumber="38">Fig. 84</figureCitation>
); head capsule about as long as broad or slightly transverse (e.g.,
<figureCitation id="31F32A0FFFE38A36FCCE37BF623BDAF3" box="[803,886,935,960]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">Fig. 90</figureCitation>
); antennal insertions broadly separated (e.g.,
<figureCitation id="31F32A0FFFE38A36FA9D37BF61FADAD0" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">Fig. 90</figureCitation>
); occipital constriction about half as broad as width of head or narrower (e.g.,
<figureCitation id="31F32A0FFFE38A36FBCD37D3653EDAD0" box="[1056,1139,971,996]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">Fig. 90</figureCitation>
); eyes small, much shorter than tempora, anterior (e.g.,
<figureCitation id="31F32A0FFFE38A36FE1837F56302DD35" box="[501,591,1005,1030]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">Fig. 90</figureCitation>
); hypostomal ridges nearly reaching posterior tentorial pits; maxillary palpomere IV subtriangular (e.g.,
<figureCitation id="31F32A0FFFE38A36FDFA30086326DD1A" box="[535,619,1040,1065]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">Fig. 93</figureCitation>
); mandibles with at least three (usually more) subapical teeth of various lengths, some located above, and some beneath the coronal plane of mandible (
<figureCitation id="31F32A0FFFE38A36FBF3302B6539DD7F" box="[1054,1140,1075,1100]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">Fig. 92</figureCitation>
); pronotum with posterior collar demarcated by a transverse row of pits (e.g.,
<figureCitation id="31F32A0FFFE38A36FD0C304E6228DD5C" box="[737,869,1110,1135]" captionStart="FIGURES 78–83" captionStartId="40.[151,264,1712,1735]" captionTargetBox="[158,1426,201,1685]" captionTargetId="figure@40.[151,1436,193,1691]" captionTargetPageId="40" captionText="FIGURES 78–83. Clidicini, examples of extant genera; dorsal (78, 80, 82) and lateral (79, 81, 83) habitus. Clidicus gracillipes (78–79); Clidicus bellator (80–81); Clidicus formicarius (82–83)." httpUri="https://zenodo.org/record/1454962/files/figure.png" pageId="37" pageNumber="38">Figs 78–79</figureCitation>
); mesoventral intercoxal process narrow, carinate, with subparallel lateral margins (
<figureCitation id="31F32A0FFFE38A36FDEA30616311DDA1" box="[519,604,1145,1170]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="37" pageNumber="38">Fig. 97</figureCitation>
); metacoxal margin of metaventrite with an angulate adcoxal expansion at each side (
<figureCitation id="31F32A0FFFE38A36FEE330846029DD86" box="[270,356,1180,1205]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="37" pageNumber="38">Fig. 97</figureCitation>
); metaventrite lacking foveae; elytral rows of punctures deep, distinct and regular (
<figureCitation id="31F32A0FFFE38A36FAE1308464C2DD86" box="[1292,1423,1180,1205]" captionStart="FIGURES 78–83" captionStartId="40.[151,264,1712,1735]" captionTargetBox="[158,1426,201,1685]" captionTargetId="figure@40.[151,1436,193,1691]" captionTargetPageId="40" captionText="FIGURES 78–83. Clidicini, examples of extant genera; dorsal (78, 80, 82) and lateral (79, 81, 83) habitus. Clidicus gracillipes (78–79); Clidicus bellator (80–81); Clidicus formicarius (82–83)." httpUri="https://zenodo.org/record/1454962/files/figure.png" pageId="37" pageNumber="38">Figs 78–83</figureCitation>
); abdominal sternite VIII in males not emarginate; aedeagus heavily sclerotized and in repose positioned symmetrically inside abdomen; larva with subtrapezoidal head with short annular 'neck', each larval mandible with a preapical tooth, larval antennomere II undivided, larval antennomere III much longer than broad, sensory appendage spatulate, urogomphi present.
</paragraph>
<paragraph id="A977368AFFE38A36FF2A315360DFDCDC" blockId="37.[151,1437,761,2009]" pageId="37" pageNumber="38">
<emphasis id="9BBCEA98FFE38A36FF2A315360ACDC57" bold="true" box="[199,481,1355,1380]" pageId="37" pageNumber="38">Characteristics. Adults.</emphasis>
Body (
<figureCitation id="31F32A0FFFE38A36FDDE315363F9DC50" box="[563,692,1355,1380]" captionStart="FIGURES 78–83" captionStartId="40.[151,264,1712,1735]" captionTargetBox="[158,1426,201,1685]" captionTargetId="figure@40.[151,1436,193,1691]" captionTargetPageId="40" captionText="FIGURES 78–83. Clidicini, examples of extant genera; dorsal (78, 80, 82) and lateral (79, 81, 83) habitus. Clidicus gracillipes (78–79); Clidicus bellator (80–81); Clidicus formicarius (82–83)." httpUri="https://zenodo.org/record/1454962/files/figure.png" pageId="37" pageNumber="38">Figs 78–83</figureCitation>
,
<figureCitation id="31F32A0FFFE38A36FD523153624ADC57" box="[703,775,1355,1380]" captionStart="FIGURES 86–88" captionStartId="42.[151,264,1899,1922]" captionTargetBox="[151,1436,1201,1878]" captionTargetId="figure@42.[151,1436,1201,1878]" captionTargetPageId="42" captionText="FIGURES 86–88. Clidicini, examples of extinct species; lateral (86), ventral (87) and lateroventral (88) habitus. Palaeoleptochromus schaufussi (86; Upper Cretaceous of Canada; photo Brendon Strilisky); Clidicus archaicus (87; Upper Cretaceous of Myanmar); Clidicus sp. (88; Upper Eocene of Europe)." httpUri="https://zenodo.org/record/1454966/files/figure.png" pageId="37" pageNumber="38">86–88</figureCitation>
) small to large, in extant species 2.99̄8.5 mm in length, in nominal extinct species 2.23–7.33 mm, typically from light to dark, often reddish brown, in one species red with dark brown head; strongly convex, dorsally densely setose, setae typically conspicuously long and erect, unmodified except for variously developed longer and more erect bristles distributed on antennae, maxillary palps, head capsule and legs.
</paragraph>
</subSubSection>
<subSubSection id="E1D26501FFE38A34FF2A31E16203DC7D" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
<paragraph id="A977368AFFE38A36FF2A31E16352DEEB" blockId="37.[151,1437,761,2009]" pageId="37" pageNumber="38">
Head capsule (
<figureCitation id="31F32A0FFFE38A36FE6C31E16341DF21" box="[385,524,1529,1554]" captionStart="FIGURES 84–85" captionStartId="41.[151,264,1421,1444]" captionTargetBox="[151,1436,265,1400]" captionTargetId="figure@41.[151,1436,265,1400]" captionTargetPageId="41" captionText="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." httpUri="https://zenodo.org/record/1454964/files/figure.png" pageId="37" pageNumber="38">Figs 84–85</figureCitation>
,
<figureCitation id="31F32A0FFFE38A36FDCF31E26327DF21" box="[546,618,1530,1554]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">90–91</figureCitation>
) divided into large and exposed anterior part and much smaller, subcylindrical 'neck' region retracted into prothorax and demarcated by distinct occipital constriction; 'neck' region about as broad as half width of head. Anterior part of head strongly flattened, typically subequal in width with prothorax, weakly transverse or (rarely) about as long as broad, broadest behind middle or (rarely) at middle. Composite eyes dorsolateral, near anterior margin of head, small but composed of numerous small ommatidia, weakly convex. Vertex strongly transverse and posteriorly impressed at middle, convex at sides, with posterior margin concave; in one problematic extinct genus with a pair of long posterolateral setae. Tempora long and rounded. Frons between antennal insertions broadly subtrapezoidal, not forming a demarcated 'platform', anteriorly demarcated by a deep and complete frontoclypeal groove (
<figureCitation id="31F32A0FFFE38A36FCBC330962E4DE19" box="[849,937,1809,1834]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">Fig. 90</figureCitation>
;
<emphasis id="9BBCEA98FFE38A36FC57330A6291DE19" box="[954,988,1810,1834]" italics="true" pageId="37" pageNumber="38">fcg</emphasis>
). Clypeus very short and broad, with rounded or straight sides. Antennal insertions (
<figureCitation id="31F32A0FFFE38A36FD5D332C6248DE7E" box="[688,773,1844,1869]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">Fig. 90</figureCitation>
;
<emphasis id="9BBCEA98FFE38A36FCFE332D6264DE7F" box="[787,809,1845,1868]" italics="true" pageId="37" pageNumber="38">ai</emphasis>
) located anterodorsally, broadly separated. Gular plate (
<figureCitation id="31F32A0FFFE38A36FF72334F61B8DE5C" box="[159,245,1879,1904]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">Fig. 91</figureCitation>
;
<emphasis id="9BBCEA98FFE38A36FEEE33406052DE5C" box="[259,287,1880,1903]" italics="true" pageId="37" pageNumber="38">gp</emphasis>
) lacking sutures, nearly smooth; posterior tentorial pits (
<figureCitation id="31F32A0FFFE38A36FC42334F6549DE5C" box="[943,1028,1879,1904]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">Fig. 91</figureCitation>
;
<emphasis id="9BBCEA98FFE38A36FBFE3340657ADE5C" box="[1043,1079,1880,1903]" italics="true" pageId="37" pageNumber="38">ptp</emphasis>
) arcuate, in front of transverse impression demarcating 'neck' region ventrally; hypostomal ridges (
<figureCitation id="31F32A0FFFE38A36FC2B3362656CDEA1" box="[966,1057,1914,1939]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="37" pageNumber="38">Fig. 91</figureCitation>
;
<emphasis id="9BBCEA98FFE38A36FBD833626503DEA0" box="[1077,1102,1914,1939]" italics="true" pageId="37" pageNumber="38">hr</emphasis>
) nearly straight or arcuate, posteriorly nearly reaching posterior tentorial pits. Head with or without punctures, densely setose, often with conspicuously long setae (
<figureCitation id="31F32A0FFFE38A36FE5333D8635FDEEB" box="[446,530,1984,2009]" captionStart="FIGURES 84–85" captionStartId="41.[151,264,1421,1444]" captionTargetBox="[151,1436,265,1400]" captionTargetId="figure@41.[151,1436,265,1400]" captionTargetPageId="41" captionText="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." httpUri="https://zenodo.org/record/1454964/files/figure.png" pageId="37" pageNumber="38">Fig. 84</figureCitation>
).
</paragraph>
<paragraph id="A977368AFFE08A35FF2A348065F9D86C" blockId="38.[151,1437,151,2028]" pageId="38" pageNumber="39">
Antennae (
<figureCitation id="31F32A0FFFE08A35FEAF348F6088D983" box="[322,453,151,176]" captionStart-0="FIGURES 78–83" captionStart-1="FIGURES 84–85" captionStart-2="FIGURES 86–88" captionStartId-0="40.[151,264,1712,1735]" captionStartId-1="41.[151,264,1421,1444]" captionStartId-2="42.[151,264,1899,1922]" captionTargetBox-0="[158,1426,201,1685]" captionTargetBox-1="[151,1436,265,1400]" captionTargetBox-2="[151,1436,1201,1878]" captionTargetId-0="figure@40.[151,1436,193,1691]" captionTargetId-1="figure@41.[151,1436,265,1400]" captionTargetId-2="figure@42.[151,1436,1201,1878]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionTargetPageId-2="42" captionText-0="FIGURES 78–83. Clidicini, examples of extant genera; dorsal (78, 80, 82) and lateral (79, 81, 83) habitus. Clidicus gracillipes (78–79); Clidicus bellator (80–81); Clidicus formicarius (82–83)." captionText-1="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." captionText-2="FIGURES 86–88. Clidicini, examples of extinct species; lateral (86), ventral (87) and lateroventral (88) habitus. Palaeoleptochromus schaufussi (86; Upper Cretaceous of Canada; photo Brendon Strilisky); Clidicus archaicus (87; Upper Cretaceous of Myanmar); Clidicus sp. (88; Upper Eocene of Europe)." httpUri-0="https://zenodo.org/record/1454962/files/figure.png" httpUri-1="https://zenodo.org/record/1454964/files/figure.png" httpUri-2="https://zenodo.org/record/1454966/files/figure.png" pageId="38" pageNumber="39">Figs 78–88</figureCitation>
,
<figureCitation id="31F32A0FFFE08A35FE3C348F60A3D983" box="[465,494,151,176]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">95</figureCitation>
) shorter or about as long as body, slender; scape (
<figureCitation id="31F32A0FFFE08A35FBC5348F6530D983" box="[1064,1149,151,176]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 95</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FB67348165ECD983" box="[1162,1185,153,176]" italics="true" pageId="38" pageNumber="39">sc</emphasis>
) 4̄6 times as long as broad, usually slightly broadening distad, with very deep lateroventral emargination; pedicel (
<figureCitation id="31F32A0FFFE08A35FB4234A2644ED9E0" box="[1199,1283,186,211]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 95</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FAE334A26467D9E0" box="[1294,1322,186,211]" italics="true" pageId="38" pageNumber="39">pd</emphasis>
) typically shorter than, or less frequently about as long as antennomere III and broadening from narrow base to apex; antennomeres III–X elongate (usually strongly, less frequently weakly) and weakly thickening distad, basal stalks not exposed, basal rings absent or indistinct; antennomere XI elongate and slightly asymmetrical. Antennomeres covered with variously dense, long setae; surface of antennomeres in most specimens smooth.
</paragraph>
<paragraph id="A977368AFFE08A35FF2A35716417DDDF" blockId="38.[151,1437,151,2028]" pageId="38" pageNumber="39">
Mouthparts. Labrum (
<figureCitation id="31F32A0FFFE08A35FE533571635CD8B1" box="[446,529,361,386]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="38" pageNumber="39">Fig. 92</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FDF135716370D8B1" box="[540,573,361,386]" italics="true" pageId="38" pageNumber="39">lbr</emphasis>
) strongly transverse, variable in shape, with lateral margins parallel or (more frequently) divergent anterad, nearly straight or rounded, and with anterior margin straight, concave, emarginated or notched at middle, or with a row of variously shaped anterior teeth or denticles, with a transverse dorsoanterior row of long setae. Mandibles (
<figureCitation id="31F32A0FFFE08A35FE0735CA6324D8D8" box="[490,617,466,491]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="38" pageNumber="39">Figs 90–92</figureCitation>
) symmetrical, subtriangular and robust, with at least three (usually more) subapical teeth of various lengths, some located above, and some beneath the coronal plane of mandible, setose prostheca present and long. Maxilla (
<figureCitation id="31F32A0FFFE08A35FDDA360063C5DB03" box="[567,648,536,561]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
) with large but relatively short cardo (
<figureCitation id="31F32A0FFFE08A35FBDB360065C4DB03" box="[1078,1161,536,561]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FB78360065E2DB02" box="[1173,1199,536,561]" italics="true" pageId="38" pageNumber="39">cd</emphasis>
); basistipes (
<figureCitation id="31F32A0FFFE08A35FAA9360064D5DB03" box="[1348,1432,536,561]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FF7A362361F5DB67" box="[151,184,571,596]" italics="true" pageId="38" pageNumber="39">bst</emphasis>
) subtriangular and elongate; mediostipes (
<figureCitation id="31F32A0FFFE08A35FD78362363A5DB60" box="[661,744,571,596]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FD1936246256DB60" box="[756,795,572,595]" italics="true" pageId="38" pageNumber="39">mst</emphasis>
) large and sharply demarcated from lacinia (
<figureCitation id="31F32A0FFFE08A35FAFC36236428DB60" box="[1297,1381,571,596]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FA9D362364DFDB67" box="[1392,1426,571,596]" italics="true" pageId="38" pageNumber="39">lac</emphasis>
) and galea (
<figureCitation id="31F32A0FFFE08A35FEF836466026DB44" box="[277,363,606,631]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FE94364660D0DB44" box="[377,413,606,631]" italics="true" pageId="38" pageNumber="39">gal</emphasis>
), which are both elongate and curved mesally and each with a dense row of distal setae; palpifer (
<figureCitation id="31F32A0FFFE08A35FF1236996019DBAA" box="[255,340,641,666]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FE8C369960C8DBAA" box="[353,389,641,665]" italics="true" pageId="38" pageNumber="39">ppf</emphasis>
) broad and elongate; maxillary palp (
<figureCitation id="31F32A0FFFE08A35FCD9369962FADBA9" box="[820,951,641,666]" captionStart="FIGURES 84–85" captionStartId="41.[151,264,1421,1444]" captionTargetBox="[151,1436,265,1400]" captionTargetId="figure@41.[151,1436,265,1400]" captionTargetPageId="41" captionText="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." httpUri="https://zenodo.org/record/1454964/files/figure.png" pageId="38" pageNumber="39">Figs 84–85</figureCitation>
,
<figureCitation id="31F32A0FFFE08A35FC2936996541DBAA" box="[964,1036,641,665]" captionStart-0="FIGURES 90–93" captionStart-1="FIGURES 94–98" captionStartId-0="44.[151,264,1885,1908]" captionStartId-1="45.[151,264,1816,1839]" captionTargetBox-0="[151,1436,495,1865]" captionTargetBox-1="[151,1436,193,1796]" captionTargetId-0="figure@44.[151,1436,495,1865]" captionTargetId-1="figure@45.[151,1436,193,1796]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionText-0="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." captionText-1="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri-0="https://zenodo.org/record/1454970/files/figure.png" httpUri-1="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">93–94</figureCitation>
) composed of minute palpomere I, elongate and broadening distad palpomere II, palpomere III (
<figureCitation id="31F32A0FFFE08A35FCA836BC62D4DB8F" box="[837,921,676,701]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="38" pageNumber="39">Fig. 93</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FC4836BC62ACDB8F" box="[933,993,676,700]" italics="true" pageId="38" pageNumber="39">mxp3</emphasis>
) subtriangular, broadening distad, with oblique distal margin, palpomere IV (
<figureCitation id="31F32A0FFFE08A35FDAE36DE63D5DBEC" box="[579,664,710,735]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="38" pageNumber="39">Fig. 93</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FD4836DF63ACDBEC" box="[677,737,711,735]" italics="true" pageId="38" pageNumber="39">mxp4</emphasis>
) subtriangular, with pointed apex, its base typically narrower than apex of palpomere III or subequal in width; palpomeres III and IV often somewhat flattened. Maxillary palps can be much longer than head capsule (
<figureCitation id="31F32A0FFFE08A35FDB6371463E3DA16" box="[603,686,780,805]" captionStart="FIGURES 84–85" captionStartId="41.[151,264,1421,1444]" captionTargetBox="[151,1436,265,1400]" captionTargetId="figure@41.[151,1436,265,1400]" captionTargetPageId="41" captionText="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." httpUri="https://zenodo.org/record/1454964/files/figure.png" pageId="38" pageNumber="39">Fig. 84</figureCitation>
) or about as long as head (
<figureCitation id="31F32A0FFFE08A35FC0837146577DA16" box="[997,1082,780,805]" captionStart="FIGURES 84–85" captionStartId="41.[151,264,1421,1444]" captionTargetBox="[151,1436,265,1400]" captionTargetId="figure@41.[151,1436,265,1400]" captionTargetPageId="41" captionText="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." httpUri="https://zenodo.org/record/1454964/files/figure.png" pageId="38" pageNumber="39">Fig. 85</figureCitation>
), typically in strongly elongate species with very slender antennae and legs also palps are long and slender (
<figureCitation id="31F32A0FFFE08A35FC103737651CDA7B" box="[1021,1105,815,840]" captionStart="FIGURES 78–83" captionStartId="40.[151,264,1712,1735]" captionTargetBox="[158,1426,201,1685]" captionTargetId="figure@40.[151,1436,193,1691]" captionTargetPageId="40" captionText="FIGURES 78–83. Clidicini, examples of extant genera; dorsal (78, 80, 82) and lateral (79, 81, 83) habitus. Clidicus gracillipes (78–79); Clidicus bellator (80–81); Clidicus formicarius (82–83)." httpUri="https://zenodo.org/record/1454962/files/figure.png" pageId="38" pageNumber="39">Fig. 82</figureCitation>
), and in those with relatively stout body and short antennae and legs (
<figureCitation id="31F32A0FFFE08A35FDB3374A63FDDA58" box="[606,688,850,875]" captionStart="FIGURES 78–83" captionStartId="40.[151,264,1712,1735]" captionTargetBox="[158,1426,201,1685]" captionTargetId="figure@40.[151,1436,193,1691]" captionTargetPageId="40" captionText="FIGURES 78–83. Clidicini, examples of extant genera; dorsal (78, 80, 82) and lateral (79, 81, 83) habitus. Clidicus gracillipes (78–79); Clidicus bellator (80–81); Clidicus formicarius (82–83)." httpUri="https://zenodo.org/record/1454962/files/figure.png" pageId="38" pageNumber="39">Fig. 80</figureCitation>
) the palps are proportionally shorter and with broader palpomeres III and IV. All palpomeres densely covered with variously long setae, palpomere II and/or III with some long and more erect setae among suberect and shorter basic vestiture (well-visible in
<figureCitation id="31F32A0FFFE08A35FBEE3780651ADA82" box="[1027,1111,920,945]" captionStart="FIGURES 84–85" captionStartId="41.[151,264,1421,1444]" captionTargetBox="[151,1436,265,1400]" captionTargetId="figure@41.[151,1436,265,1400]" captionTargetPageId="41" captionText="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." httpUri="https://zenodo.org/record/1454964/files/figure.png" pageId="38" pageNumber="39">Fig. 84</figureCitation>
, in some species indistinct). Labium (
<figureCitation id="31F32A0FFFE08A35FF1037A36033DAE7" box="[253,382,955,980]" captionStart-0="FIGURES 90–93" captionStart-1="FIGURES 94–98" captionStartId-0="44.[151,264,1885,1908]" captionStartId-1="45.[151,264,1816,1839]" captionTargetBox-0="[151,1436,495,1865]" captionTargetBox-1="[151,1436,193,1796]" captionTargetId-0="figure@44.[151,1436,495,1865]" captionTargetId-1="figure@45.[151,1436,193,1796]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionText-0="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." captionText-1="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri-0="https://zenodo.org/record/1454970/files/figure.png" httpUri-1="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Figs 91–94</figureCitation>
) with broad submentum (
<figureCitation id="31F32A0FFFE08A35FD4D37A363B9DAE7" box="[672,756,955,980]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="38" pageNumber="39">Fig. 91</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FCED37A46260DAE0" box="[768,813,956,979]" italics="true" pageId="38" pageNumber="39">smn</emphasis>
) posteriorly not demarcated from gular region, bearing a pair of long thin setae near its anterior margin or in subanterior region; mentum (
<figureCitation id="31F32A0FFFE08A35FBDA37C665C7DAC4" box="[1079,1162,990,1015]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FB7B37C765F5DAC5" box="[1174,1208,991,1014]" italics="true" pageId="38" pageNumber="39">mn</emphasis>
) subtrapezoidal and strongly transverse, with anterior margin straight or broadly rounded; prementum (
<figureCitation id="31F32A0FFFE08A35FB4030196440DD29" box="[1197,1293,1025,1050]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FAC9301A640BDD2A" box="[1316,1350,1026,1049]" italics="true" pageId="38" pageNumber="39">pm</emphasis>
) long, subtrapezoidal, broadest distally, lacking demarcated ligula, with a pair of anterolateral setae, with broadly separated bases of labial palps; lateral hypopharyngeal lobes (
<figureCitation id="31F32A0FFFE08A35FCAA305F62D4DD6C" box="[839,921,1095,1120]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FC49305F628CDD53" box="[932,961,1095,1120]" italics="true" pageId="38" pageNumber="39">lhl</emphasis>
) moderately large; labial palp (
<figureCitation id="31F32A0FFFE08A35FAF2305F643FDD6C" box="[1311,1394,1095,1120]" captionStart="FIGURES 90–93" captionStartId="44.[151,264,1885,1908]" captionTargetBox="[151,1436,495,1865]" captionTargetId="figure@44.[151,1436,495,1865]" captionTargetPageId="44" captionText="FIGURES 90–93. Clidicini, morphological structures, Clidicus bellator. Head in dorsal (90) and ventral (91) views; mouthparts in dorsal view (92); left maxillary palpomeres III–IV in dorsal view (93). Abbreviations: ai, antennal insertion; bst, basistipes; cd, cardo; cl, clypeus; fcg, frontoclypeal groove; gal, galea; gp, gular plate; hr, hypostomal ridge; lac, lacinia; lbr, labrum; lhl, lateral hypopharyngeal lobe; lp, labial palp; mn, mentum; mst, mediostipes; mxp1–4, maxillary palpomere I–IV; pmn, prementum; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." httpUri="https://zenodo.org/record/1454970/files/figure.png" pageId="38" pageNumber="39">Fig. 91</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FA90305F64DEDD53" box="[1405,1427,1095,1120]" italics="true" pageId="38" pageNumber="39">lp</emphasis>
) composed of three palpomeres: palpomere I (
<figureCitation id="31F32A0FFFE08A35FD73307263BEDDB1" box="[670,755,1130,1155]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FCEC30726269DDB0" box="[769,804,1130,1155]" italics="true" pageId="38" pageNumber="39">lp1</emphasis>
) small, elongate, broadening distad, palpomere II (
<figureCitation id="31F32A0FFFE08A35FA82307261FBDD96" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FF29309561AADD95" box="[196,231,1165,1190]" italics="true" pageId="38" pageNumber="39">lp2</emphasis>
) largest, strongly elongate and only indistinctly broadening distad, only in one problematic extinct taxon narrow at base and strongly broadened in distal half, palpomere III (
<figureCitation id="31F32A0FFFE08A35FC7430B762A0DDFB" box="[921,1005,1199,1224]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 94</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FC1430B76550DDFB" box="[1017,1053,1199,1224]" italics="true" pageId="38" pageNumber="39">lp3</emphasis>
) narrow, long and pointed, in one problematic extinct taxon extremely slender, bristle-like. Porous fields on mouthparts and clypeus not found.
</paragraph>
<paragraph id="A977368AFFE08A35FF2A30ED63DFDF58" blockId="38.[151,1437,151,2028]" pageId="38" pageNumber="39">
Prothorax (
<figureCitation id="31F32A0FFFE08A35FEAB30ED6084DC3D" box="[326,457,1269,1294]" captionStart-0="FIGURES 78–83" captionStart-1="FIGURES 84–85" captionStartId-0="40.[151,264,1712,1735]" captionStartId-1="41.[151,264,1421,1444]" captionTargetBox-0="[158,1426,201,1685]" captionTargetBox-1="[151,1436,265,1400]" captionTargetId-0="figure@40.[151,1436,193,1691]" captionTargetId-1="figure@41.[151,1436,265,1400]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURES 78–83. Clidicini, examples of extant genera; dorsal (78, 80, 82) and lateral (79, 81, 83) habitus. Clidicus gracillipes (78–79); Clidicus bellator (80–81); Clidicus formicarius (82–83)." captionText-1="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." httpUri-0="https://zenodo.org/record/1454962/files/figure.png" httpUri-1="https://zenodo.org/record/1454964/files/figure.png" pageId="38" pageNumber="39">Figs 78–85</figureCitation>
,
<figureCitation id="31F32A0FFFE08A35FE3830ED60BFDC3D" box="[469,498,1269,1294]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">96</figureCitation>
) about as long as broad or elongate, strongly convex, broadest near anterior third. Pronotum with anterior and posterior margins arcuate or nearly straight, sides rounded in anterior half and usually sinuate in posterior half; anterior corners broadly rounded, posterior corners obtuse-angled; pronotal base with a short posterior 'collar' demarcated by a dorsal transverse row of several variously distinct pits, often connected by a groove. Prosternum (
<figureCitation id="31F32A0FFFE08A35FE683199609ADCA9" box="[389,471,1409,1434]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 96</figureCitation>
) with basisternal part (
<figureCitation id="31F32A0FFFE08A35FD303199627CDCA9" box="[733,817,1409,1434]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 96</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FCD331996224DCA9" box="[830,873,1409,1434]" italics="true" pageId="38" pageNumber="39">bstr</emphasis>
) subequal in length to coxal part, but proportions depend on the shape of pronotum, in species with short pronota the basisternal part tends to be indistinctly shorter than coxal part, in species with elongate pronota the basisternal part can be longer than coxal part. Prosternum laterally completely fused with hypomera. Coxal region demarcated anteriorly by carina extending laterally up to apices of very weakly developed adcoxal hypomeral lobes. Procoxal cavities broadly open. Prosternal intercoxal process indistinct, usually developed as a diffuse and weakly elevated ridge hidden between procoxae. Ventral surface of prothorax typically densely setose.
</paragraph>
<paragraph id="A977368AFFE08A35FF2A326E650EDE95" blockId="38.[151,1437,151,2028]" pageId="38" pageNumber="39">
Mesoventrite (
<figureCitation id="31F32A0FFFE08A35FE84326E60A7DFBC" box="[361,490,1654,1679]" captionStart="FIGURES 84–85" captionStartId="41.[151,264,1421,1444]" captionTargetBox="[151,1436,265,1400]" captionTargetId="figure@41.[151,1436,265,1400]" captionTargetPageId="41" captionText="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." httpUri="https://zenodo.org/record/1454964/files/figure.png" pageId="38" pageNumber="39">Figs 84–85</figureCitation>
,
<figureCitation id="31F32A0FFFE08A35FE18326F635FDFBC" box="[501,530,1655,1679]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">97</figureCitation>
) subtrapezoidal, broadening posteriorly. Prepecti (
<figureCitation id="31F32A0FFFE08A35FBAB326E65D4DFBC" box="[1094,1177,1654,1679]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FB49326F6585DFBD" box="[1188,1224,1655,1678]" italics="true" pageId="38" pageNumber="39">pre</emphasis>
) long and together with anteromedian mesoventral area forming a massive 'collar', which bears a transverse groove just behind its anterior ridge (
<figureCitation id="31F32A0FFFE08A35FED332A460DCDFE7" box="[318,401,1724,1749]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FE7032A560FBDFE7" box="[413,438,1725,1748]" italics="true" pageId="38" pageNumber="39">ar</emphasis>
), its posterior margin bisinuate with short subtriangular or subtrapezoidal posteromedian projection, which is broadly separated from a similar anteromedian projection of posterior margin of setose impression (
<figureCitation id="31F32A0FFFE08A35FECB33196033DE29" box="[294,382,1793,1818]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FE63331B60ECDE29" box="[398,417,1795,1818]" italics="true" pageId="38" pageNumber="39">si</emphasis>
). Region just behind collar strongly and abruptly constricted, ventrally forming setose impression (
<figureCitation id="31F32A0FFFE08A35FECF333C603ADE0E" box="[290,375,1828,1853]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FE69333E60DADE0E" box="[388,407,1830,1853]" italics="true" pageId="38" pageNumber="39">si</emphasis>
) filled with dense, unmodified setae. Mesoventral intercoxal process (
<figureCitation id="31F32A0FFFE08A35FB5B333C6446DE0E" box="[1206,1291,1828,1853]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FAF5333E641FDE0E" box="[1304,1362,1830,1853]" italics="true" pageId="38" pageNumber="39">msvp</emphasis>
) long, narrow and weakly convex, nearly parallel-sided or indistinctly broadened near middle of mesocoxal cavities, fully separating mesocoxae, posteriorly fused with metaventrite or with variously distinct posterior tip. Mesanepisterna large and subtriangular; mesepimera (
<figureCitation id="31F32A0FFFE08A35FDD3339563DCDE95" box="[574,657,1933,1958]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
;
<emphasis id="9BBCEA98FFE08A35FD7033966394DE95" box="[669,729,1934,1958]" italics="true" pageId="38" pageNumber="39">epm2</emphasis>
) partly exposed in ventral view.
</paragraph>
<paragraph id="A977368AFFE08A35FF2A33A9620ADEDF" blockId="38.[151,1437,151,2028]" pageId="38" pageNumber="39">Mesonotum with subtriangular, elongate mesoscutellum with rounded apex, in intact specimens only its very tip visible between elytral bases; scutoscutellar suture absent.</paragraph>
<paragraph id="A977368AFFE18A34FF2A34806011D9E7" blockId="39.[151,1437,151,1755]" pageId="39" pageNumber="40">Metanotum fully developed or somewhat reduced, with alacristae not or only slightly shortened; hind wings present or absent.</paragraph>
<paragraph id="A977368AFFE18A34FF2A34C6620FDB20" blockId="39.[151,1437,151,1755]" pageId="39" pageNumber="40">
Metaventrite (
<figureCitation id="31F32A0FFFE18A34FE8134C66089D9C4" box="[364,452,222,247]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="39" pageNumber="40">Fig. 97</figureCitation>
) short, subquadrate or subrectangular and slightly transverse, with lateral margins rounded; each mesocoxal cavity with posterior marginal carina, anterior margins not carinate; posterior margin of metaventrite deeply bisinuate laterally (in front of each metacoxa) and with a broad metaventral intercoxal process (
<figureCitation id="31F32A0FFFE18A34FF72355161B1D852" box="[159,252,329,354]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="39" pageNumber="40">Fig. 97</figureCitation>
;
<emphasis id="9BBCEA98FFE18A34FEFF35526005D852" box="[274,328,330,353]" italics="true" pageId="39" pageNumber="40">mtvp</emphasis>
) with variously deeply concave posterior margin and subtriangular posterolateral corners; metaventrite lacking foveae. External admetacoxal part of posterior metaventral margin with additional marginal thickening demarcated by a groove, forming adcoxal carina (
<figureCitation id="31F32A0FFFE18A34FCA6358862EDD89B" box="[843,928,400,425]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="39" pageNumber="40">Fig. 97</figureCitation>
;
<emphasis id="9BBCEA98FFE18A34FC4135896292D89B" box="[940,991,401,424]" italics="true" pageId="39" pageNumber="40">acxc</emphasis>
) at each side. Metanepisterna (
<figureCitation id="31F32A0FFFE18A34FAAE358864DAD89B" box="[1347,1431,400,425]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="39" pageNumber="40">Fig. 97</figureCitation>
;
<emphasis id="9BBCEA98FFE18A34FF7A35AC619FD8FF" box="[151,210,436,460]" italics="true" pageId="39" pageNumber="40">aest3</emphasis>
) broad and partly visible in ventral view, broadened posteriorly; metepimera broader than metanepisterna, with indistinctly demarcated inner and outer component, posteriorly extending far behind metacoxae. In one extinct species metaventrite with longitudinal median carina.
</paragraph>
<paragraph id="A977368AFFE18A34FF2A36056462DB05" blockId="39.[151,1437,151,1755]" box="[199,1327,541,566]" pageId="39" pageNumber="40">Metendosternite (metafurca) Y-shaped, with short but distinct stem and divergent lateral furcal arms.</paragraph>
<paragraph id="A977368AFFE18A34FF2A365B6347DA00" blockId="39.[151,1437,151,1755]" pageId="39" pageNumber="40">
Legs (
<figureCitation id="31F32A0FFFE18A34FEFE365A60D1DB69" box="[275,412,578,603]" captionStart-0="FIGURES 78–83" captionStart-1="FIGURES 84–85" captionStart-2="FIGURES 86–88" captionStartId-0="40.[151,264,1712,1735]" captionStartId-1="41.[151,264,1421,1444]" captionStartId-2="42.[151,264,1899,1922]" captionTargetBox-0="[158,1426,201,1685]" captionTargetBox-1="[151,1436,265,1400]" captionTargetBox-2="[151,1436,1201,1878]" captionTargetId-0="figure@40.[151,1436,193,1691]" captionTargetId-1="figure@41.[151,1436,265,1400]" captionTargetId-2="figure@42.[151,1436,1201,1878]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionTargetPageId-2="42" captionText-0="FIGURES 78–83. Clidicini, examples of extant genera; dorsal (78, 80, 82) and lateral (79, 81, 83) habitus. Clidicus gracillipes (78–79); Clidicus bellator (80–81); Clidicus formicarius (82–83)." captionText-1="FIGURES 84–85. Clidicini, examples of extant species; anterior portion of body, ventral. Clidicus formicarius (84); Clidicus abbotensis (85). Arrows showing outstanding setae on head, profemur and prothoracic hypomera, which are largely reduced in C. abbotensis." captionText-2="FIGURES 86–88. Clidicini, examples of extinct species; lateral (86), ventral (87) and lateroventral (88) habitus. Palaeoleptochromus schaufussi (86; Upper Cretaceous of Canada; photo Brendon Strilisky); Clidicus archaicus (87; Upper Cretaceous of Myanmar); Clidicus sp. (88; Upper Eocene of Europe)." httpUri-0="https://zenodo.org/record/1454962/files/figure.png" httpUri-1="https://zenodo.org/record/1454964/files/figure.png" httpUri-2="https://zenodo.org/record/1454966/files/figure.png" pageId="39" pageNumber="40">Figs 78–88</figureCitation>
,
<figureCitation id="31F32A0FFFE18A34FE42365A6081DB69" box="[431,460,578,602]" captionStart="FIGURES 94–98" captionStartId="45.[151,264,1816,1839]" captionTargetBox="[151,1436,193,1796]" captionTargetId="figure@45.[151,1436,193,1796]" captionTargetPageId="45" captionText="FIGURES 94–98. Clidicini, morphological structures, Clidicus bellator. Mouthparts in ventral view (94); left scape and pedicel in ventral view (95); prothorax in ventral view (96); pterothorax and first visible abdominal sternite in ventral view (97); aedeagus and sperm pump (98). Abbreviations: acxc, adcoxal carina; ade, adcoxal expansion; aed, aedeagus; aest3, metanepisternum; ar, anterior ridge; bst, basistipes; bstr, basisternal part of prosternum; cd, cardo; dej, ductus ejaculatorius; epm2, mesepimeron; gal, galea; lac, lacinia; lhl, lateral hypopharyngeal lobe; lp1–3, labial palpomere I–III; mn, mentum; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pd, pedicel; pm, prementum; ppf, palpifer; pre, prepectus; sc, scape; si, setose impression; spp, sperm pump; st3, abdominal sternite III; v3, metaventrite." httpUri="https://zenodo.org/record/1454972/files/figure.png" pageId="39" pageNumber="40">97</figureCitation>
) long and slender, often conspicuously so. Pro- and mesocoxa short subconical, metacoxa with nearly hemispherical basal part and subconical distal part. Mesocoxa lacking coxal bristles, but often with a comb of thin long setae. All trochanters short and subtriangular. Femora weakly clavate, profemur typically with at least one conspicuously long and erect ventral seta (rarely setae reduced). Tibiae robust, protibiae often curved in distal third or fourth. Tarsi long and slender, nearly subcylindrical, tarsomeres I–V subequal in length or reducing in length, tarsomere V strongly elongate, with curved and slender claws lacking elongate costae; empodial region was not studied.
</paragraph>
<paragraph id="A977368AFFE18A34FF2A37266535DAAD" blockId="39.[151,1437,151,1755]" pageId="39" pageNumber="40">
Elytra (
<figureCitation id="31F32A0FFFE18A34FEF0372660ECDA64" box="[285,417,830,855]" captionStart="FIGURES 78–83" captionStartId="40.[151,264,1712,1735]" captionTargetBox="[158,1426,201,1685]" captionTargetId="figure@40.[151,1436,193,1691]" captionTargetPageId="40" captionText="FIGURES 78–83. Clidicini, examples of extant genera; dorsal (78, 80, 82) and lateral (79, 81, 83) habitus. Clidicus gracillipes (78–79); Clidicus bellator (80–81); Clidicus formicarius (82–83)." httpUri="https://zenodo.org/record/1454962/files/figure.png" pageId="39" pageNumber="40">Figs 78–83</figureCitation>
) oval, strongly convex, with variously distinct humeral calli, lacking basal impressions, with rounded or pointed apices; elytral disc with distinct large and deep punctures arranged in nearly complete and regular longitudinal rows. Elytra variously densely setose, setae typically long and erect.
</paragraph>
<paragraph id="A977368AFFE18A34FF2A37B263C8DAD4" blockId="39.[151,1437,151,1755]" pageId="39" pageNumber="40">Abdomen with sternite III not fused with metaventrite, about as long as sternites IV and V together; sternite VIII in male with rounded posterior margin.</paragraph>
<paragraph id="A977368AFFE18A34FF2A37EA6308DD8D" blockId="39.[151,1437,151,1755]" pageId="39" pageNumber="40">
Aedeagus (illustrated in
<bibRefCitation id="CD594B7BFFE18A34FE0E37EA63EFDD39" box="[483,674,1010,1035]" pageId="39" pageNumber="40" refId="ref51580" refString="BESUCHET, C. (1971) LES Clidicus dE CEYLON (COL. SCYdMAENIdAE). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 43, 249 - 257." type="journal article">Besuchet (1971)</bibRefCitation>
, Jałoszyński (
<bibRefCitation id="CD594B7BFFE18A34FCA937EA62CDDD39" box="[836,896,1010,1034]" pageId="39" pageNumber="40" refId="ref52793" refString="JAlOSzYnSkI, P. (2009) Clidicus crocodylus N. SP. fROM MINdANAO, THE fIRST SPECIES Of THE MASTIGINAE fROM THE PHILIPPINES (COLEOPTERA: STAPHYLINIdAE: SCYdMAENINAE). Genus, Wroclaw, 20 (2), 203 - 207." type="journal article">2009</bibRefCitation>
, 2018), O’Keefe &amp; Monteith (2000), Orousset (2014), Zhou &amp; Li (2015)) strongly elongate, with symmetrical median lobe, parameres heavily sclerotized, symmetrical or one longer than the other; flagellum simple, not coiled, but often broadened and forming a sac-like structure. Ejaculatory duct with elongate and narrow sperm pump, typically with funnel-like structures at both ends, but in some species with one or both funnels absent. Aedeagus in repose positioned symmetrically inside abdomen, with basal orifice facing up.
</paragraph>
<paragraph id="A977368AFFE18A34FF2A30D262EADC34" blockId="39.[151,1437,151,1755]" pageId="39" pageNumber="40">Spermatheca (illustrated e.g., by Orousset (2014)) globular, variable in shape, in some species nearly spherical, in others elongate, thick-walled, with relatively large accessory gland.</paragraph>
<paragraph id="A977368AFFE18A34FF2A310A6203DC7D" blockId="39.[151,1437,151,1755]" pageId="39" pageNumber="40">
<emphasis id="9BBCEA98FFE18A34FF2A310A60A6DC18" bold="true" box="[199,491,1298,1323]" pageId="39" pageNumber="40">Characteristics. Larvae.</emphasis>
Only a larva of the Australian
<taxonomicName id="6EC84D09FFE18A34FC8D310A65A2DC19" authority="O'Keefe, 2000" authorityName="O'KEEfE" authorityYear="2000" box="[864,1263,1298,1323]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="abbotensis">
<emphasis id="9BBCEA98FFE18A34FC8D310A650DDC18" box="[864,1088,1298,1323]" italics="true" pageId="39" pageNumber="40">Clidicus abbotensis</emphasis>
O'Keefe, 2000
</taxonomicName>
was described (O’Keefe &amp; Monteith 2000); characteristics given at
<taxonomicName id="6EC84D09FFE18A34FD07312E620ADC7C" authorityName="Laporte de Castelnau" authorityYear="1832" box="[746,839,1334,1359]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFE18A34FD07312E620ADC7C" box="[746,839,1334,1359]" italics="true" pageId="39" pageNumber="40">Clidicus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1D26501FFE18A34FF2A31426567DFE8" pageId="39" pageNumber="40" type="discussion">
<paragraph id="A977368AFFE18A34FF2A31426378DF30" blockId="39.[151,1437,151,1755]" pageId="39" pageNumber="40">
<emphasis id="9BBCEA98FFE18A34FF2A31426372DC40" bold="true" box="[199,575,1370,1395]" pageId="39" pageNumber="40">Composition and distribution.</emphasis>
Clidicini, as currently defined, comprise 32 extant and extinct species classified in three genera. The extant species are distributed (
<figureCitation id="31F32A0FFFE18A34FCAC316662DEDCA4" box="[833,915,1406,1431]" captionStart="FIGURE 89" captionStartId="43.[151,250,1189,1211]" captionTargetBox="[151,1436,613,1168]" captionTargetId="figure@43.[151,1436,613,1168]" captionTargetPageId="43" captionText="FIGURE 89. Clidicini, distribution of extant species of Clidicus." httpUri="https://zenodo.org/record/1454968/files/figure.png" pageId="39" pageNumber="40">Fig. 89</figureCitation>
) in the Oriental region and northern
<collectingCountry id="D1DF761AFFE18A34FAC3316664D5DCA4" box="[1326,1432,1406,1431]" name="Australia" pageId="39" pageNumber="40">Australia</collectingCountry>
; definitive fossils come from Cenomanian of
<collectingCountry id="D1DF761AFFE18A34FD7C31BB63B3DC88" box="[657,766,1443,1467]" name="Myanmar" pageId="39" pageNumber="40">Myanmar</collectingCountry>
and Priabonian of central-eastern Europe. Extinct taxa that may belong to Clidicini are known from Upper Eocene Baltic amber, Lower Cretaceous Charentese French amber and Middle Miocene
<collectingRegion id="6B0CF868FFE18A34FE6A31F260ACDF30" box="[391,481,1514,1539]" country="Mexico" name="Chiapas" pageId="39" pageNumber="40">Chiapas</collectingRegion>
amber.
</paragraph>
<paragraph id="A977368AFFE18A34FF2A32166567DFE8" blockId="39.[151,1437,151,1755]" pageId="39" pageNumber="40">
<emphasis id="9BBCEA98FFE18A34FF2A32166076DF14" bold="true" box="[199,315,1550,1575]" pageId="39" pageNumber="40">Remarks.</emphasis>
Historically, Clidicini included
<taxonomicName id="6EC84D09FFE18A34FD5B3216625EDF14" authorityName="Laporte de Castelnau" authorityYear="1832" box="[694,787,1550,1575]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFE18A34FD5B3216625EDF14" box="[694,787,1550,1575]" italics="true" pageId="39" pageNumber="40">Clidicus</emphasis>
</taxonomicName>
,
<taxonomicName id="6EC84D09FFE18A34FCCB32166288DF14" box="[806,965,1550,1575]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFE18A34FCCB32166288DF14" box="[806,965,1550,1575]" italics="true" pageId="39" pageNumber="40">Leptochromus</emphasis>
</taxonomicName>
,
<taxonomicName id="6EC84D09FFE18A34FC353217657ADF15" box="[984,1079,1551,1574]" class="Insecta" family="Staphylinidae" genus="Papusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFE18A34FC353217657ADF15" box="[984,1079,1551,1574]" italics="true" pageId="39" pageNumber="40">Papusus</emphasis>
</taxonomicName>
, and later also several extinct genera. Jałoszyński
<emphasis id="9BBCEA98FFE18A34FE95322B60FDDF78" box="[376,432,1586,1611]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
(2018) redefined this tribe and restricted it only to
<taxonomicName id="6EC84D09FFE18A34FC1D322A6500DF78" authorityName="Laporte de Castelnau" authorityYear="1832" box="[1008,1101,1586,1611]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFE18A34FC1D322A6500DF78" box="[1008,1101,1586,1611]" italics="true" pageId="39" pageNumber="40">Clidicus</emphasis>
</taxonomicName>
, with the problematic Middle Miocene
<taxonomicName id="6EC84D09FFE18A34FEEF324E60A4DF5C" box="[258,489,1622,1647]" class="Insecta" family="Staphylinidae" genus="Palaeoleptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFE18A34FEEF324E60A4DF5C" box="[258,489,1622,1647]" italics="true" pageId="39" pageNumber="40">Palaeoleptochromus</emphasis>
</taxonomicName>
placed in this tribe tentatively and pending further study. The same applies to the Upper Eocene
<taxonomicName id="6EC84D09FFE18A34FEAE32636099DFA1" box="[323,468,1659,1682]" class="Insecta" family="Cerambycidae" genus="Aenictosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="9BBCEA98FFE18A34FEAE32636099DFA1" box="[323,468,1659,1682]" italics="true" pageId="39" pageNumber="40">Aenictosoma</emphasis>
</taxonomicName>
known from Baltic amber, which was originally described as possibly belonging to
<taxonomicName id="6EC84D09FFE18A34FF7A32866355DF84" authority="(Schaufuss 1891)" authorityName="Schaufuss" authorityYear="1891" baseAuthorityName="Schaufuss" baseAuthorityYear="1891" box="[151,536,1694,1719]" class="Insecta" family="Cerambycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="family">Cerambycidae (Schaufuss 1891)</taxonomicName>
, and later, solely on the basis of the original description, transferred to
<taxonomicName id="6EC84D09FFE18A34FF7A32DA6072DFE8" authorityName="Leach" authorityYear="1815" box="[151,319,1730,1755]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
: Clidicini (Vitali 2006). See Remarks for each of these taxa below.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink